971:
1329:(then known as Okinawa Ocean Expo Aquarium) in Japan. Since 1980, several have been kept at Okinawa, mostly obtained from incidental catches in coastal nets set by fishers (none after 2009), but two were strandings. Several of these were already weak from the capture/stranding and some were released, but initial captive survival rates were low. After the initial difficulties in maintaining the species had been resolved, some have survived long-term in captivity. The record for a whale shark in captivity is an individual that, as of 2021, has lived for more than 26 years in the Okinawa Churaumi Aquarium from Okinawa Ocean Expo Aquarium. Following Okinawa,
1291:
1403:, United States. This is unusual because of the comparatively long transport time and complex logistics required to bring the sharks to the aquarium, ranging between 28 and 36 hours. Georgia keeps two whale sharks: two males, Taroko and Yushan, who both arrived in 2007. Two earlier males at Georgia Aquarium, Ralph and Norton, both died in 2007. Trixie died in 2020. Alice died in 2021. Georgia's whale sharks were all imported from Taiwan and were taken from the commercial fishing quota for the species, usually used locally for food. Taiwan closed this fishery entirely in 2008.
736:
588:, as the human body temperature makes the pigment decay. This pigment becomes unstable in shallow water, where the temperature is higher and the full spectrum of light is present. To protect from this instability, the whale shark deactivates the pigment when in shallow water (as otherwise the pigment would hinder full color vision). In the colder environment at 2,000 meters below the surface where the shark dives, it is activated again. The mutations thus allow the shark to see well at both ends of its great vertical range. The eyes have also lost all
1026:. It also feeds on clouds of eggs during mass spawning of fish and corals. The many rows of vestigial teeth play no role in feeding. Feeding occurs either by ram filtration, in which the animal opens its mouth and swims forward, pushing water and food into the mouth, or by active suction feeding, in which the animal opens and closes its mouth, sucking in volumes of water that are then expelled through the gills. In both cases, the filter pads serve to separate food from water. These unique, black sieve-like structures are presumed to be modified
137:
116:
724:
767:, and can be found in both coastal and oceanic habitats. Tracking devices have shown that the whale shark displays dynamic patterns of habitat utilization, likely in response to availability of prey. Whale sharks observed off the northeast Yucatan Peninsula tend to engage in inshore surface swimming between sunrise and mid-afternoon, followed by regular vertical oscillations in oceanic waters during the afternoon and overnight. About 95% of the oscillating period was spent in
352:
1034:, in which the water travels nearly parallel to the filter pad surface, not perpendicularly through it, before passing to the outside, while denser food particles continue to the back of the throat. This is an extremely efficient filtration method that minimizes fouling of the filter pad surface. Whale sharks have been observed "coughing", presumably to clear a build-up of particles from the filter pads. Whale sharks migrate to feed and possibly to breed.
203:
1096:
1076:
712:
700:
5263:
161:
945:
1412:
658:(a Thai unit of length measuring between a person's outstretched arms). Smith noted that one wa could be interpreted as either 2 m (6.6 ft) or the approximate average of 1.7 to 1.8 m (5.6â5.9 ft), based on the local fishermen. Later sources have stated this whale shark as approximately 18 m (59 ft), but the accuracy of the estimate has been questioned.
1310:
5275:
918:, Philippines, and was released into the wild. Based on this discovery, some scientists no longer believe this area is just a feeding ground; this site may be a birthing ground, as well. Both young whale sharks and pregnant females have been seen in the waters of St Helena in the South Atlantic Ocean, where numerous whale sharks can be spotted during the summer.
748:
531:. Their skin is dark grey with a white belly marked with an arrangement of pale grey or white spots and stripes that is unique to each individual. The skin can be up to 15 cm (5.9 in) thick and is very hard and rough to the touch. The whale shark has three prominent ridges along its sides, which start above and behind the head and end at the
612:
14.5 m (48 ft) on average, based on more limited data. However, these are averages and do not represent the maximum possible sizes. Previous studies estimating the growth and longevity of whale sharks have produced estimates ranging from 14 to 21.9 meters (46 to 72 ft) in length. Limited evidence, mostly from males, suggests that
932:' photo identification activities in the first half of the year. There were a total 168 sightings â 64 of them âre-sightingsâ or reappearances of previously recorded whale sharks. WWF noted that âvery young whale shark juveniles" were identified among the 168 individuals spotted in the first half of 2019. Their presence suggests that the
960:
1268:, where one-third of all whale shark sightings in the northern part of the gulf have occurred in recent years. Sightings confirmed that the whale sharks were unable to avoid the oil slick, which was situated on the surface of the sea where the whale sharks feed for several hours at a time. No dead whale sharks were found.
1388:), with some maintaining whale sharks for years and others only for a very short period. The whale shark kept at Dubai's Atlantis, The Palm was rescued from shallow waters in 2008 with extensive abrasions to the fins and after rehabilitation it was released in 2010, having lived 19 months in captivity.
1305:
that keep it, but its large size means that a very large tank is required and it has specialized feeding requirements. Their large size and iconic status have also fueled an opposition to keeping the species in captivity, especially after the early death of some whale sharks in captivity and certain
518:
Whale sharks possess a broad, flattened head with a large mouth and two small eyes located at the front corners. Unlike many other sharks, whale shark mouths are located at the front of the head rather than on the underside of the head. A 12.1 m (39.7 ft) whale shark was reported to have a
470:
Whale sharks inhabit the open waters of all tropical oceans. They are rarely found in water below 21 °C (70 °F). Whale sharks' lifespans are estimated to be between 80 and 130 years, based on studies of their vertebral growth bands and the growth rates of free-swimming sharks. Whale sharks
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events in the 1950-60s, finding that growth bands are laid down annually. The study found an age of 50 years for a 10 m (33 ft) female and 35 years for a 9.9m male. Various studies looking at vertebrae growth bands and measuring whale sharks in the wild have estimated their lifespans from
1054:
Whale sharks are known to prey on a range of planktonic and small nektonic organisms that are spatiotemporally patchy. These include krill, crab larvae, jellyfish, sardines, anchovies, mackerels, small tunas, and squid. In ram filter feeding, the fish swims forward at constant speed with its mouth
620:
Large whale sharks are difficult to measure accurately, both on the land and in the water. When on land, the total length measurement can be affected by how the tail is positioned, either angled as it would be in life or stretched as far as possible. Historically, techniques such as comparisons to
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depths (<200 metres (660 ft)), but whale sharks also took regular deep dives (>500 metres (1,600 ft)), often descending in brief "stutter steps", perhaps for foraging. The deepest recorded dive was 1,928 metres (6,325 ft), making the whale shark the deepest diving fish to be
616:
occurs around 8 to 9 meters (26 to 30 ft) in length, with females possibly maturing at a similar size or larger. The maximum length of the species is uncertain due to a lack of detailed documentation of the largest reported individuals. Several whale sharks around 18 m (59 ft) in
869:
Coast. It was one of the largest gatherings of whale sharks recorded. Aggregations in that area are among the most reliable seasonal gatherings known for whale sharks, with large numbers occurring in most years between May and
September. Associated ecotourism has grown rapidly to unsustainable
611:
with regards to size, with females growing larger than males. A 2020 study looked at the growth of whale shark individuals over a 10-year period. It concluded that males on average reach 8 to 9 meters (26 to 30 ft) in length. The same study predicted females to reach a length of around
910:. The eggs remain in the body and the females give birth to live young which are 40 to 60 cm (16 to 24 in) long. Evidence indicates the pups are not all born at once, but rather the female retains sperm from one mating and produces a steady stream of pups over a prolonged period.
878:
Growth, longevity, and reproduction of the whale shark are poorly understood. There was uncertainty as to whether vertebrae growth bands are formed annually or biannually, which is important in determining the age, growth, and longevity of whale sharks. A 2020 study compared the ratio of
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The whale shark is an active feeder, targeting concentrations of plankton or fish. It is able to ram filter feed or can gulp in a stationary position. This is in contrast to the passive feeding basking shark, which does not pump water. Instead, it swims to force water across its gills.
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1413:
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Evidence suggests that males grow faster than females in the earlier stages of life but ultimately reach a smaller maximum size. Whale sharks exhibit late sexual maturity. One study looking at free-swimming whale sharks estimated the age at maturity in males at ~25 years.
665:
came across a whale shark in the southern
Pacific Ocean and rammed it. The shark became stuck on the prow of the ship, supposedly with 15 ft (4.6 m) on one side and 40 ft (12.2 m) on the other, suggesting a total length of about 55 ft (17 m).
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of whale sharks from between 1994 and 1996. Out of the 15 individuals tracked, two females were reported as measuring 15 m (49 ft) and 18 m (59 ft) respectively. A 20.75 m (68.1 ft) long whale shark was reported as being stranded along the
1806:
McClain CR, Balk MA, Benfield MC, Branch TA, Chen C, Cosgrove J, Dove ADM, Gaskins LC, Helm RR, Hochberg FG, Lee FB, Marshall A, McMurray SE, Schanche C, Stone SN, Thaler AD. 2015. "Sizing ocean giants: patterns of intraspecific size variation in marine megafauna".
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445:. The largest confirmed individual had a length of 18.8 m (61.7 ft). The whale shark holds many records for size in the animal kingdom, most notably being by far the most massive living non-cetacean animal. It is the sole member of the genus
787:, South Africa, and an Indo-Pacific subpopulation which holds 75% of the entire whale shark population. It usually roams between 30°N and 35°S where water temperatures are higher than 21 °C (70 °F) but have been spotted as far north as the
913:
On 7 March 2009, marine scientists in the
Philippines discovered what is believed to be the smallest living specimen of the whale shark. The young shark, measuring only 38 cm (15 in), was found with its tail tied to a stake at a beach in
584:) two amino acid substitutions make the pigment more sensitive to blue light instead, the light that dominates the deep ocean. One of these mutations also makes rhodopsin vulnerable to higher temperatures. In humans, a similar mutation leads to
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have photographed them swimming close to humans without any danger. Although whale sharks are docile fish, touching or riding the sharks is strictly forbidden and fineable in most countries, as it can cause serious harm to the animal.
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on the surface of their eyeballs that are structured differently from their body denticles. The dermal denticles, as well as the whale shark's ability to retract its eyes deep into their sockets, serve to protect the eyes from damage.
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filmed a whale shark feeding on a school of small fish. The same documentary showed footage of a whale shark timing its arrival to coincide with the mass spawning of fish shoals and feeding on the resultant clouds of eggs and sperm.
644:. Wright was informed of one whale shark that was measured as exceeding 45 ft (14 m). Wright claimed to have observed specimens over 50 ft (15 m) and was told of specimens upwards of 70 ft (21 m).
3174:
Borrell, AsunciĂłn; Aguilar, Alex; Gazo, Manel; Kumarran, R. P.; Cardona, Luis (2011). "Stable isotope profiles in whale shark (Rhincodon typus) suggest segregation and dissimilarities in the diet depending on sex and size".
4617:
Bennett; Kaiser; Selvan; Hueter; Tyminski; Lötter (2017). "Rescue, Rehabilitation and
Release of a Whale Shark, Rhincodon typus, in the Arabian Gulf". In Smith, Mark; Warmolts; Thoney; Hueter; Murray; Ezcurra (eds.).
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5419:
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Evidence suggests that whale sharks can recover from major injuries and may be able to regenerate small sections of their fins. Their spot markings have also been shown to reform over a previously wounded area.
3441:
de la Parra
Venegas, Rafael; Hueter, Robert; Cano, Jaime Gonzålez; Tyminski, John; Remolina, José Gregorio; Maslanka, Mike; Ormos, Andrea; Weigt, Lee; Carlson, Bruce; Dove, Alistair (29 April 2011).
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Matsumoto; Toda; Matsumoto; Ueda; Nakazato; Sato; Uchida (2017). "Notes on
Husbandry of Whale Sharks, Rhincodon typus, in Aquaria". In Smith, Mark; Warmolts; Thoney; Hueter; Murray; Ezcurra (eds.).
3960:
Yong, Mila Mi Hua; Leistenschneider, Clara; Miranda, Joni Anne; Paler, Maria
Kristina; Legaspi, Christine; Germanov, Elitza; Araujo, Gonzalo; Burkhardt-Holm, Patricia; Erni-Cassola, Gabriel (2021).
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Meekan, Mark G.; Taylor, Brett M.; Lester, Emily; Ferreira, Luciana C.; Sequeira, Ana M. M.; Dove, Alistair D. M.; Birt, Matthew J.; Aspinall, Alex; Brooks, Kim; Thums, Michele (2020).
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Eckert, Scott A.; Stewart, Brent S. (1 February 2001). "Telemetry and
Satellite Tracking of Whale Sharks, Rhincodon Typus, in the Sea of Cortez, Mexico, and the North Pacific Ocean".
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of 15 m (49.2 ft) and an estimated total length at 18.8 m (61.7 ft) was reported from the
Arabian Sea in 2001. In a 2015 study looking into the size of marine
1257:. In 1998, the Philippines banned all fishing, selling, importing, and exporting of whale sharks for commercial purposes, followed by India in May 2001 and Taiwan in May 2007.
4722:
2323:
Read, Timothy D.; Petit, Robert A.; Joseph, Sandeep J.; Alam, Md. Tauqeer; Weil, M. Ryan; Ahmad, Maida; Bhimani, Ravila; Vuong, Jocelyn S.; Haase, Chad P. (December 2017).
1459:. By law snorkelers must maintain a distance of 4 ft (1.2 m) from the sharks and there is a fine and possible prison sentence for anyone who touches the animals.
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fully open, straining prey particles from the water by forward propulsion. This is also called âpassive feedingâ, which usually occurs when prey is present at low density.
3371:
5173:
4603:
5779:
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3528:
Evidence that St. Helena island is an important multi-use habitat for whale sharks, Rhincodon typus , with the first description of putative mating in this species
1381:
1087:
Despite its size, the whale shark does not pose any danger to humans. Younger whale sharks are gentle and can play with divers. Underwater photographers such as
5562:
693:, Pakistan. The length of the specimen was said to be between 11 and 12 m (36 and 39 ft), with a weight of around 15,000 kg (33,000 lb).
4692:
2418:
5774:
5163:
3416:
735:
1926:"Comparing length-measurement methods and estimating growth parameters of free-swimming whale sharks (Rhincodon typus) near the South Ari Atoll, Maldives"
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3882:
5719:
5158:
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3505:
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3710:
1321:
The first attempt at keeping whale sharks in captivity was in 1934 when an individual was kept for about four months in a netted-off natural bay in
1244:
1240:
3909:
2629:
Wintner, Sabine P. (2000). "Preliminary Study of
Vertebral Growth Rings in the Whale Shark, Rhincodon typus, from the East Coast of South Africa".
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in 1919. The shark was too heavy to pull ashore, and no measurements were taken. Smith learned through independent sources that it was at least 10
5523:
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4103:
Establishing a quantifiable model of whale shark avoidance behaviours to anthropogenic impacts in tourism encounters to inform management actions
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2400:
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5598:
4169:
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2497:
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recorded. Whale sharks were also observed to remain continuously at depths of greater than 50 metres (160 ft) for three days or more.
4141:
2419:"Whale Sharks Can See in the Dark Due to Genetic Mutation; Light-Sensing Pigment in Retina Works in Dim Environment | Science Times"
621:
objects of known size and knotted ropes have been used for in-water measurements, but these techniques may be inaccurate. In 2011, laser
5002:
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mouth 1.55 m (5.1 ft) across. Whale shark mouths can contain over 300 rows of tiny teeth and 20 filter pads which it uses to
5458:
4185:
4627:
4575:
4433:
4279:
2945:
2093:
4726:
4085:
2256:
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5676:
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4035:
3283:
2188:
Tomita, Taketeru; Murakumo, Kiyomi; Komoto, Shinya; Dove, Alistair; Kino, Masakatsu; Miyamoto, Kei; Toda, Minoru (29 June 2020).
1456:
1353:
165:
4161:
Sharks of the World: An Annotated and Illustrated Catalogue of Shark Species Known to Date: Bullhead, Mackerel and Carpet Sharks
3655:
Schmidt, Jennifer V.; Chen, Chien-Chi; Sheikh, Saad I.; Meekan, Mark G.; Norman, Bradley M.; Joung, Shoou-Jeng (4 August 2010).
1282:
It was reported in 2014 that hundreds of whale sharks were illegally killed every year in China for their fins, skins, and oil.
906:
The capture of a ~10.6 m (35 ft) female in July 1996 that was pregnant with ~300 pups indicated that whale sharks are
498:, described it the following year. The name "whale shark" refers to the animal's appearance and large size; it is a fish, not a
5739:
4813:
4652:
5279:
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started keeping whale sharks and most of the basic research on the keeping of the species was made at these two institutions.
5749:
4566:
Schreiber, C; Coco, C (2017). "Husbandry of Whale Sharks". In Smith, Mark; Warmolts; Thoney; Hueter; Murray; Ezcurra (eds.).
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of the retina, is normally sensitive to green and used to see in dim light, but in the whale shark (and the bottom-dwelling
2773:
Rowat, D.; Brooks, K. S. (2012). "A review of the biology, fisheries and conservation of the whale shark Rhincodon typus".
5243:
4599:
4265:
Duffy, Clinton A. J.; Francis, Malcolm; Dunn, M. R.; Finucci, Brit; Ford, Richard; Hitchmough, Rod; Rolfe, Jeremy (2018).
3020:
Gudger, E. W. (1938). "Whale Sharks Rammed by Ocean Vessels: How These Sluggish Leviathans Aid in Their Own Destruction".
1978:
Brunnschweiler, J. M.; Baensch, H.; Pierce, S. J.; Sims, D. W. (3 February 2009). "Deep-diving behaviour of a whale shark
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1116:
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Stevens, J. D. (1 March 2007). "Whale shark (Rhincodon typus) biology and ecology: A review of the primary literature".
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via airplane in Australia in 2019, when a larger male unsuccessfully attempted to mate with a smaller, immature female.
3803:
5729:
5668:
5393:
5357:
2325:"Draft sequencing and assembly of the genome of the world's largest fish, the whale shark: Rhincodon typus Smith 1828"
1349:
1261:
1160:
975:
2681:"Annual Bands in Vertebrae Validated by Bomb Radiocarbon Assays Provide Estimates of Age and Growth of Whale Sharks"
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202:
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have very large mouths and are filter feeders, which is a feeding mode that occurs in only two other sharks, the
5248:
4116:
2522:
Yamaguchi, Kazuaki; Koyanagi, Mitsumasa; Sato, Keiichi; Terakita, Akihisa; Kuraku, Shigehiro (1 October 2021).
1924:
Perry, Cameron T.; Figueiredo, Joana; Vaudo, Jeremy J.; Hancock, James; Rees, Richard; Shivji, Mahmood (2018).
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723:
491:
341:
301:
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Yamaguchi, Kazuaki; Koyanagi, Mitsumasa; Sato, Keiichi; Terakita, Akihisa; Kuraku, Shigehiro (28 March 2023).
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losses, and vessel strikes, combined with its long lifespan and late maturation. In June 2018 the New Zealand
115:
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5769:
5744:
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5180:
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4958:
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1964:
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884:
2276:"Wound-healing capabilities of whale sharks (Rhincodon typus) and implications for conservation management"
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The species was distinguished in April 1828 after the harpooning of a 4.6 m (15 ft) specimen in
4017:
2881:"How large is the world's largest fish? Measuring whale sharks Rhincodon typus with laser photogrammetry"
5764:
5624:
5398:
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1031:
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On 7 February 2012, a large whale shark was found floating 150 kilometres (93 mi) off the coast of
324:
4678:
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3706:
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Ong, Joyce J. L.; Meekan, Mark G.; Hsu, Hua Hsun; Fanning, L. Paul; Campana, Steven E. (6 April 2020).
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3319:
1683:"Descriptions of New or imperfectly known Objects of the Animal Kingdom, found in the South of Africa"
1227:
There is currently no robust estimate of the global whale shark population. The species is considered
936:
may be a pupping ground for whale sharks, further increasing the ecological significance of the area.
5681:
5577:
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5344:
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3184:
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2834:"Size and maturity status of the whale shark (Rhincodon typus) at Ningaloo Reef in Western Australia"
2833:
2782:
2734:
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2445:
2434:"Whale shark rhodopsin adapted to deep-sea lifestyle by a substitution associated with human disease"
2201:
2039:
1991:
1254:
647:
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686:, McClain and colleagues considered this female as being the most reliable and accurately measured.
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Since the 1800s, there have been accounts of very large whale sharks. Some of these are as follows:
1180:
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839:
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150:
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4907:
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The whale shark is migratory and has two distinct subpopulations: an Atlantic subpopulation, from
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3607:
3208:
3107:
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1228:
699:
364:
197:
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4200:"Global collision-risk hotspots of marine traffic and the world's largest fish, the whale shark"
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1325:, Japan. The first attempt of keeping whale sharks in an aquarium was initiated in 1980 by the
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2007:
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608:
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Clingham, Elizabeth; Brown, Judith; Henry, Leeann; Beard, Annalea; Dove, Alistair D (2016).
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991:
915:
613:
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128:
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4857:
Colman, J.G. (December 1997). "A review of the biology and ecology of the whale shark".
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4215:
4061:"Whale Sharks â
Saint Helena Island Info â
About St Helena, in the South Atlantic Ocean"
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1995:
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of the Convention on International Trade in Endangered Species of Wild Fauna and Flora (
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resulted in 4,900,000 barrels (780,000 m) of oil flowing into an area south of the
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Rohner, C. A.; Richardson, A. J.; Marshall, A. D.; Weeks, S. J.; Pierce, S. J. (2011).
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The whale shark is also featured on the latest 2015â2017 edition of the Maldivian 1000
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classified the whale shark as "Migrant" with the qualifier "Secure Overseas" under the
1216:
1041:
A juvenile whale shark is estimated to eat 21 kg (46 pounds) of plankton per day.
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284:
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244:
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Pupping of whale sharks has not been observed, but mating has been witnessed twice in
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4268:
Conservation status of New Zealand chondrichthyans (chimaeras, sharks and rays), 2016
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2539:
2003:
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1062:. As such, the presence of microplastics in whale shark scat was recently confirmed.
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763:
The whale shark inhabits all tropical and warm-temperate seas. The fish is primarily
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5086:
4831:
3885:. Ichthyology at the Florida Museum of Natural History. 11 May 2017. Archived from
3537:
2922:
2818:
2274:
Womersley, Freya; Hancock, James; Perry, Cameron T.; Rowat, David (February 2021).
1778:
1539:
1508:
1322:
1152:
1148:
1132:
1108:
1088:
896:
827:
823:
819:
807:
803:
788:
784:
507:
436:
432:
274:
2969:
Smith, H. M. (13 November 1925). "A Whale Shark (Rhineodon) in the Gulf of Siam".
2857:
2746:
1838:"Age and growth of the whale shark (Rhincodon typus) in the north-western Pacific"
1440:, meaning "many stars", after the appearance of the markings on the shark's back.
5424:
4461:
4024:, "A 40-foot whale shark and a brave snorkeler swim off the South African coast."
3471:
3048:
2579:"Asymptotic Growth of Whale Sharks Suggests Sex-Specific Life-History Strategies"
2214:
1392:
in Singapore had planned on keeping whale sharks but scrapped this idea in 2009.
5655:
5549:
5497:
5329:
5081:
5035:
4164:. Vol. 2. Food & Agriculture Organization of the United Nations (FAO).
4060:
2990:
2030:
Colman, J. G. (1997). "A review of the biology and ecology of the whale shark".
1572:
1200:
1080:
944:
929:
835:
480:
124:
48:
4349:. Department of Sustainability, Environment, Water, Population and Communities.
4204:
Proceedings of the National Academy of Sciences of the United States of America
3982:
3834:
3150:"Stranding of a whale shark, Rhincodon typus (smith) at Pamban, Gulf of Mannar"
3149:
1309:
1279:) in 2003 to regulate the international trade of live specimens and its parts.
5119:
5101:
4637:
4585:
4443:
4289:
3546:
3296:
3196:
3095:
2650:
2595:
2578:
2531:
2498:"Mutation behind night blindness in humans helps whale sharks see in the dark"
2341:
2291:
1427:
1172:
1136:
1083:, whale sharks are fed shrimp to return every morning for tourists and divers.
1027:
933:
768:
641:
548:
536:
93:
58:
5463:
4974:
4932:
4893:
3561:"Attempted Whale Shark Mating Caught on Camera for the First Time in History"
3204:
3103:
2998:
2955:
2865:
2802:
2754:
2707:
2697:
2680:
2604:
2465:
2350:
2223:
2162:
2059:
1949:
1925:
1861:
1837:
1836:
Hsu, Hua Hsun; Joung, Shoou Jeng; Hueter, Robert E.; Liu, Kwang Ming (2014).
1336:
Since the mid-1990s, several other aquariums have kept the species in Japan (
5445:
5200:
4224:
3028:. New England Museum of Natural History: Boston Society of Natural History.
2457:
2103:
1712:
Penrith, M. J. (1972). "Earliest Description and Name for the Whale Shark".
1487:
1369:
1204:
1188:
1144:
1058:
Due their mode of feeding, whale sharks are susceptible to the ingestion of
1007:
880:
831:
683:
675:
655:
589:
573:
570:
The complete and annotated genome of the whale shark was published in 2017.
495:
487:
214:
98:
42:
5320:
4886:
4515:
4251:
4001:
3842:
3490:
3033:
3006:
2914:
2810:
2483:
2368:
2309:
2241:
2067:
2011:
1652:
899:. Mating in this species was filmed for the first time in whale sharks off
5590:
4723:"Transporting the World's Largest Fish: A Whale [Shark] of a Task"
3688:
17:
5471:
5385:
5314:
5091:
4947:
2938:
Six months at the Seychelles: letter to A. Searle Hart, LL. D., S.F.T.C.D
2144:"A note on the whale shark Rhincodon typus Smith, stranded off Mangalore"
2117:
1341:
1212:
1196:
999:
995:
952:
802:
Seasonal feeding aggregations occur at several coastal sites such as the
651:
604:
577:
544:
234:
88:
83:
68:
63:
53:
4233:
3733:"'Largest number in years': Over 100 new whale sharks spotted in Donsol"
3633:
2524:"Whale shark rhodopsin adapted to its vertically wide-ranging lifestyle"
527:
are located just behind the eyes. Whale sharks have five large pairs of
5689:
5489:
5476:
4086:"Government of Western Australia, Department of Fisheries. Whale Shark"
3603:
3320:"Whale sharks in Gulf of Mexico swim near the surface, take deep dives"
1733:
1504:
1497:
1400:
1373:
1236:
1120:
986:â one of only three known filter-feeding shark species (along with the
922:
866:
843:
764:
690:
442:
103:
78:
4327:
2143:
1816:
1395:
Outside Asia, the first and so far only place to keep whale sharks is
5502:
5370:
4304:"Memorandum of understanding on the conservation of migratory sharks"
3673:
3656:
3126:
2401:"Whale sharks see in dark due to mutation behind blindness in humans"
1475:
1422:
Snorkeling with whale shark near Isla Mujeres (Mexico) 30 August 2011
1377:
1365:
1168:
1015:
964:
851:
847:
780:
499:
224:
5291:
3808:
during surface ram filter feeding off the Yucatan Peninsula, Mexico"
3417:"'What a privilege': Swimming with endangered whale sharks in Qatar"
1941:
1853:
1725:
5450:
4274:. Wellington, New Zealand: Department of Conservation. p. 11.
3526:
3127:"Turtles and whale shark landed along ratnagiri coast, maharashtra"
1255:
Memorandum of Understanding on the Conservation of Migratory Sharks
144:
The size of various whale shark individuals with a human for scale
5650:
5637:
5011:
3582:, is a livebearer: 300 embryos found in one 'megamamma' supreme".
1469:
1410:
1308:
1289:
1276:
1184:
1094:
1074:
1019:
1003:
969:
959:
958:
943:
859:
855:
776:
592:
503:
264:
187:
181:
5437:
3709:. South Atlantic Media Services. 14 November 2013. Archived from
3391:"Whale sharks found off Gujarat coast no expats, they are Indian"
1682:
1306:
Chinese aquariums keeping the species in relatively small tanks.
607:
animal in the world. Evidence suggests that whale sharks exhibit
5603:
3804:"Feeding anatomy, filter-feeding rate, and diet of whale sharks
3261:
3148:
Venkatesan, V; Ramamurthy, N; Boominathan, N; Gandhi, A (2008).
2383:"How whale shark rhodopsin evolved to see, in the deep blue sea"
1250:
It is listed, along with six other species of sharks, under the
1192:
1023:
528:
494:, a military doctor associated with British troops stationed in
439:
5295:
4984:
4675:"Resorts World considering alternatives to whale shark exhibit"
625:
was proposed to improve the accuracy of in-water measurements.
4942:
3578:
Joung, Shoou-Jeng; et al. (July 1996). "The whale shark,
1779:"Contributions to the Natural History of South Africa, &c"
1601:
1599:
1597:
1595:
4922:
4142:"Friend of the Sea Sustainable Whale Shark Watching Standard"
4036:"Scuba diver fined for pulling whale shark tail in Thailand"
2142:
Kaikini, A. S.; Ramamohana Rao, V.; Dhulkhed, M. H. (1959).
1982:
during long-distance movement in the western Indian Ocean".
3689:"Tiny whale shark rescued â World news â World environment"
883:
isotopes found in growth bands of whale shark vertebrae to
4117:"Man filmed riding world's biggest shark in tourism stunt"
1472:(because the markings resemble patterns typically seen on
1103:
The shark is seen by divers in many places, including the
4980:
3346:"Whale Sharks Move in Mysterious Ways: Watch Them Online"
2169:. Food and Agriculture Organization of the United Nations
1965:"Rhincodon or Rhiniodon? A Whale Shark by Any Other Name"
4483:"Conservationists round on Chinese whale shark aquarium"
650:
described a huge animal caught in a bamboo fish trap in
506:. In addition, its filter feeding habits are similar to
4900:"Whale Sharks, Whale Shark Pictures, Whale Shark Facts"
3657:"Paternity analysis in a litter of whale shark embryos"
4516:"Whale Shark Rhincodon typus Smith, 1828 in Captivity"
3788:(2000) documentary by Jacinth O'Donnell; broadcast on
2840:. Whale Sharks: Science, Conservation and Management.
2729:. Whale Sharks: Science, Conservation and Management.
1473:
1463:
4600:"Dubai hotel releases whale shark back into the wild"
2832:
Norman, Bradley M.; Stevens, John D. (1 March 2007).
865:
In 2011, more than 400 whale sharks gathered off the
3228:"Experts to cut up 40.1-foot long whale shark today"
1886:
Colman, J. G. Froese, Ranier; Pauly, Daniel (eds.).
1455:. The whale shark is featured on the reverse of the
640:
obtained several small whale shark specimens in the
5304:
5216:
5139:
5059:
5018:
4954:
Whale Shark Fact Sheet, Fisheries Western Australia
4115:Reporter, Jess Thomson Science (30 November 2022).
4013:
4011:
925:last August 2019, whale sharks were sighted during
669:Scott A. Eckert & Brent S. Stewart reported on
4481:
4328:Whale Sharks Receive Protection in the Philippines
1681:
1384:), India (Thiruvananthapuram Aquarium) and Dubai (
502:, and (like all sharks) is not closely related to
4943:Foundation for the Protection of Marine Megafauna
4830:. Discovery.com. 5 September 2013. Archived from
3962:"Microplastics in fecal samples of whale sharks (
3447:, in Mexican Coastal Waters of the Caribbean Sea"
3050:The Jaws of Death: Shark as Predator, Man as Prey
2148:Central Marine Fisheries Research Unit, Mangalore
4814:"Rewards from Gurano Bintang Shark Conservation"
4716:
4714:
4561:
4559:
4557:
4419:
4417:
4415:
4413:
4411:
3781:
3779:
3507:Yucatan Whale Sharks Swimming in Troubled Waters
2257:"Whale Sharks Have Tiny Teeth on Their Eyeballs"
1745:
1743:
1491:
1481:
4788:"Swimming with whale sharks in the Philippines"
3443:"An Unprecedented Aggregation of Whale Sharks,
2438:Proceedings of the National Academy of Sciences
1207:. Juveniles can be found near the shore in the
539:set relatively far back on the body, a pair of
483:and small fishes and pose no threat to humans.
4677:. AsianOne Travel. 16 May 2009. Archived from
3912:. The Shark Research Institute. Archived from
3707:"St Helena whale sharks cause stir in Atlanta"
3378:. 30 August 2017 – via www.thehindu.com.
3318:Public Library of Science (18 November 2015).
2940:. British Library, Historical Print Editions.
1450:
1444:
1431:
1382:National Museum of Marine Biology and Aquarium
4996:
4923:Whale Shark Photograph-identification Library
4509:
4507:
4505:
4309:. Convention on migratory species. p. 10
3348:. National Geographic Society. Archived from
1675:
1673:
1631:10.2305/IUCN.UK.2016-1.RLTS.T19488A2365291.en
551:has a larger upper lobe than the lower lobe (
8:
4959:Albino whale shark photographed in Galapagos
4622:. Ohio Biological Survey. pp. 229â235.
4475:
4473:
4471:
3966:) and from surface water in the Philippines"
3252:Pierce, S.J.; Norman, Brad (18 March 2016).
543:, a pair of pelvic fins and a single medial
4904:Animals, Animal Pictures, Wild Animal Facts
4750:"Madagascar emerges as whale shark hotspot"
4396:Hilton/Greenpeace, Paul (5 February 2014).
4372:Whale Sharks Killed, Displaced by Gulf Oil?
4347:National Regulations on Whale Shark fishing
4018:Pictures of the Day: Tuesday, Aug. 04, 2009
1881:
1879:
1877:
1875:
1873:
1871:
5292:
5003:
4989:
4981:
4570:. Ohio Biological Survey. pp. 87â98.
4428:. Ohio Biological Survey. pp. 15â22.
1115:), the Philippines, the Maldives close to
678:coast in 1995. A female individual with a
350:
159:
135:
114:
31:
4460:. wsbtv. 30 November 2007. Archived from
4241:
4223:
4198:Womersley, Freya C.; et al. (2022).
4186:Snorkelling with whale sharks in Djibouti
4158:Compagno, Leonard J. V. (26 April 2002).
3991:
3981:
3672:
3536:
3480:
3470:
3295:
2904:
2706:
2696:
2594:
2473:
2358:
2340:
2299:
2231:
2213:
1629:
451:and the only extant member of the family
4034:Russell, Mark 'Crowley' (9 April 2024).
3908:Schmidt, Jennifer V. (4 December 2010).
1692:. Vol. 3, no. 145 – via
1245:New Zealand Threat Classification System
1030:. Food separation in whale sharks is by
558:Whale sharks have been found to possess
4938:Whale Sharks: Gentle Giants of the Seas
4928:Whale Shark And Oceanic Research Center
3854:
3852:
3247:
3245:
3243:
3241:
1760:The Bulletin of Zoological Nomenclature
1699:
1689:The South African Commercial Advertiser
1555:
974:Whale shark with open mouth feeding in
695:
5780:Taxa named by Andrew Smith (zoologist)
4776:. Oneocean.org. Retrieved 23 May 2013.
3802:Motta, Philip J.; et al. (2010).
2674:
2672:
2670:
2668:
2556:
2545:
1967:. ReefQuest Centre for Shark Research.
1301:The whale shark is popular in the few
1099:Swimming alongside an adult free diver
963:A whale shark in the Philippines with
4933:Maldives Whale Shark Research Program
4725:. Promega Connections. Archived from
4693:"Aquarium gains two new whale sharks"
2768:
2766:
2764:
2720:
2718:
2624:
2622:
2572:
2570:
2079:
2077:
2025:
2023:
2021:
1919:
1917:
1915:
1913:
1911:
1909:
636:In 1868, the Irish natural scientist
586:congenital stationary night blindness
7:
5695:36FC5495-4A9A-4CA0-88DC-6CE5B3561C88
5643:9A4F40FF-0697-93C4-39CD-E0527BB4DE82
5578:49350a19-d675-42ff-9984-9b0a304521aa
4620:The Elasmobranch Husbandry Manual II
4568:The Elasmobranch Husbandry Manual II
4490:from the original on 12 January 2022
4426:The Elasmobranch Husbandry Manual II
4398:"Hundreds of sharks killed in China"
4370:Handwerk, Brian (24 September 2010)
3389:Kaushik, Himanshu (30 August 2014).
3157:Marine Fisheries Information Service
3131:Marine Fisheries Information Service
3053:. HarperCollins Publishers Limited.
1831:
1829:
1827:
1825:
1802:
1800:
1798:
1796:
5274:
5072:Heterodontiformes (Bullhead sharks)
4906:. 10 September 2010. Archived from
4721:Sundquist, T. (18 September 2013).
4606:from the original on 7 August 2015.
4602:. Associated Press. 20 March 2010.
4458:"Whale Shark's Death Sparks Debate"
3258:IUCN Red List of Threatened Species
1756:A. Smith, 1829 (Pisces): Conserved"
1617:IUCN Red List of Threatened Species
795:north of Japan and as far south as
741:Close up showing eyeball denticles
603:The whale shark is the largest non-
576:, the light-sensing pigment in the
463:. Before 1984 it was classified as
5775:Extant Oligocene first appearances
4879:10.1111/j.1095-8649.1997.tb01138.x
4695:. CNN. 1 June 2007. Archived from
4361:. Taipei Times, 27 May 2007, p. 4.
3504:Dove, Alistair (27 January 2015),
2052:10.1111/j.1095-8649.1997.tb01138.x
1486:(roughly "star from the East") in
1107:in Honduras, Thailand, Indonesia (
479:. They feed almost exclusively on
25:
5067:Carcharhiniformes (Ground sharks)
4969:A whale shark recorded defecating
4359:COA bans fishing for whale sharks
4337:. hayop.0catch.com. 27 March 1998
3759:"Shark-cam captures ocean motion"
3415:Dekker, Stefanie (17 July 2020).
3344:Howard, Brian C. (28 June 2016).
2190:"Armored eyes of the whale shark"
1680:Smith, Andrew (5 November 1828).
1606:Pierce, S.J.; Norman, B. (2016).
1443:In the Philippines, it is called
1235:due to the impacts of fisheries,
5720:IUCN Red List endangered species
5273:
5262:
5261:
5087:Orectolobiformes (Carpet sharks)
4101:Whitehead, Darren Andrew (2014)
3226:Hasan, Saad (10 February 2012).
2906:10.1111/j.1095-8649.2010.02861.x
2795:10.1111/j.1095-8649.2012.03252.x
2004:10.1111/j.1095-8649.2008.02155.x
1752:"Opinion 1278. The Generic Name
1538:
1354:Yokohama Hakkeijima Sea Paradise
888:~80 years and up to ~130 years.
746:
734:
722:
710:
698:
455:, which belongs to the subclass
201:
4964:Photographs National Geographic
4786:Cannon, Marisa (21 July 2015).
4653:"No whale sharks at Sentosa IR"
3936:"Rhincodon typus (whale shark)"
3372:"Drive to conserve whale shark"
3282:Pain, Stephanie (31 May 2022).
3177:Environmental Biology of Fishes
3076:Environmental Biology of Fishes
2631:Environmental Biology of Fishes
2118:"Whale Sharks, Rhincodon typus"
1462:Whale sharks are also known as
1271:This species was also added to
5126:Xenacanthida (Xenacantiformes)
5092:Pristiophoriformes (Sawsharks)
4105:, University of Hertfordshire.
3970:Microplastics and Nanoplastics
3910:"Whale Sharks are BIG eaters!"
3538:10.7287/peerj.preprints.1885v1
1930:Marine and Freshwater Research
1842:Marine and Freshwater Research
1141:Greater St. Lucia Wetland Park
27:Largest living species of fish
1:
5244:Grey nurse shark conservation
5082:Lamniformes (Mackerel sharks)
4748:Briggs, Helen (17 May 2018).
4514:Mollet, H. (September 2012).
4480:Moore, M. (25 October 2010).
3735:. Rappler.com. 30 August 2019
3693:Associated Press via NBC News
2858:10.1016/j.fishres.2006.11.015
2747:10.1016/j.fishres.2006.11.008
1694:Center for Research Libraries
1568:in the Paleobiology Database"
5785:Symbols of Western Australia
5102:Squatiniformes (Angelsharks)
4948:Whale shark, Rhincodon typus
4184:Hawes, Craig (2 April 2013)
3630:"Frequently Asked Questions"
3472:10.1371/journal.pone.0018994
2936:Wright, E. Perceval (2011).
2215:10.1371/journal.pone.0235342
1139:in Mozambique, Sodwana Bay (
834:province in Mozambique, the
629:Reports of large individuals
5239:U.S. Shark Conservation Act
4894:FAO web page on Whale shark
4542:. Okinawa Churaumi Aquarium
3661:Endangered Species Research
3632:. Sharklady. Archived from
2991:10.1126/science.62.1611.438
2685:Frontiers in Marine Science
2583:Frontiers in Marine Science
2086:CITES identification manual
1474:
1464:
1372:Aquarium in Guangzhou Zoo,
1350:Oita Marine Palace Aquarium
1161:La Paz, Baja California Sur
976:La Paz, Baja California Sur
617:length have been reported.
5801:
4774:Ocean Ambassadors â Sharks
3983:10.1186/s43591-021-00017-9
3835:10.1016/j.zool.2009.12.001
1518:
1496:(literally "sir fish") in
1430:, whale sharks are called
1241:Department of Conservation
661:In 1934, a ship named the
5257:
5224:Sharks in popular culture
5041:Egg case (Chondrichthyes)
4374:National Geographic News.
3297:10.1146/knowable-052622-3
3197:10.1007/s10641-011-9879-y
3047:Maniguet, Xavier (1992).
2596:10.3389/fmars.2020.575683
2532:10.1101/2021.10.01.462724
2342:10.1186/s12864-017-3926-9
2088:. Environment Australia.
1696:Document Delivery System.
1533:List of threatened sharks
1519:For a topical guide, see
1507:banknote, along with the
1327:Okinawa Churaumi Aquarium
1315:Okinawa Churaumi Aquarium
1203:, and other parts of the
1012:Christmas Island red crab
370:
363:
358:
349:
330:
323:
198:Scientific classification
196:
179:
157:
148:
143:
134:
122:
113:
34:
5249:Save Our Seas Foundation
4651:Chua, G. (16 May 2009).
4333:16 February 2020 at the
2698:10.3389/fmars.2020.00188
1750:Melville, R. V. (1981).
1457:Philippine 100-peso bill
1262:Gulf of Mexico oil spill
1066:Relationship with humans
759:Distribution and habitat
409:MĂŒller & Henle, 1839
5755:Fish of Central America
5181:Shark attack prevention
4225:10.1073/pnas.2117440119
3096:10.1023/A:1007674716437
2885:Journal of Fish Biology
2775:Journal of Fish Biology
2651:10.1023/A:1026564707027
2458:10.1073/pnas.2220728120
2292:10.1093/conphys/coaa120
2280:Conservation Physiology
2032:Journal of Fish Biology
1984:Journal of Fish Biology
1451:
1445:
1362:Chimelong Ocean Kingdom
1266:Mississippi River Delta
1143:) in South Africa, the
1119:(South Ari Atoll), the
874:Growth and reproduction
842:in the Seychelles, the
5740:Fish described in 1828
4065:sainthelenaisland.info
3022:New England Naturalist
2555:Cite journal requires
1783:The Zoological Journal
1777:Smith, Andrew (1829).
1492:
1482:
1432:
1423:
1376:Polar Ocean World and
1318:
1298:
1179:, islands off eastern
1100:
1084:
1071:Behavior toward divers
979:
967:
956:
814:in Western Australia,
638:Edward Perceval Wright
435:and the largest known
398:Rhinodon pentalineatus
5750:Fish of North America
5625:Paleobiology Database
4977:on Sealife Collection
4655:. Wild Singapore News
3125:Katkar, B.N. (1996).
2526:: 2021.10.01.462724.
2163:"Species Fact Sheet,
2084:Norman, Brad (2002).
1421:
1312:
1294:A whale shark in the
1293:
1123:, Western Australia (
1098:
1078:
1032:cross-flow filtration
982:The whale shark is a
973:
962:
947:
374:Micristodus punctatus
359:Range of whale shark
3940:Animal Diversity Web
3757:(17 November 2008).
2542:– via bioRxiv.
2261:Smithsonian Magazine
1653:"Appendices | CITES"
1624:: e.T19488A2365291.
1313:Whale sharks at the
1165:BahĂa de los Ăngeles
1018:life, such as small
535:. The shark has two
467:into Rhinodontidae.
428:) is a slow-moving,
4910:on 16 January 2010.
4871:1997JFBio..51.1219C
4537:"什ćäșćčŽćșŠ æČçžçŸăæ”·æ°Žæ通 ćčŽć ±"
4216:2022PNAS..11917440W
4210:(20): e2117440119.
3889:on 5 September 2006
3827:2010Zool..113..199M
3596:1996EnvBF..46..219J
3584:Environ. Biol. Fish
3463:2011PLoSO...618994D
3232:The Express Tribune
3189:2011EnvBF..92..559B
3088:2001EnvBF..60..299E
2983:1925Sci....62..438S
2897:2011JFBio..78..378R
2850:2007FishR..84...81N
2787:2012JFBio..80.1019R
2739:2007FishR..84....4S
2643:2000EnvBF..59..441W
2450:2023PNAS..12020728Y
2444:(13): e2220728120.
2206:2020PLoSO..1535342T
2161:Compagno, L. J. V.
2044:1997JFBio..51.1219C
1996:2009JFBio..74..706B
1380:Aquarium), Taiwan (
1223:Conservation status
1181:peninsular Malaysia
1131:), Taiwan, Panama (
797:Victoria, Australia
151:Conservation status
123:Whale shark in the
5730:Ovoviviparous fish
5108:Cladoselachiformes
4816:. 5 February 2022.
3859:Martin, R. Aidan.
3604:10.1007/BF00004997
3514:on 7 November 2017
3395:The Times of India
3290:. Annual Reviews.
3284:"Call of the deep"
2838:Fisheries Research
2727:Fisheries Research
1963:Martin, R. Aidan.
1424:
1386:Atlantis, The Palm
1338:Kagoshima Aquarium
1319:
1299:
1101:
1085:
1081:Oslob, Philippines
980:
968:
957:
818:in the GalĂĄpagos,
671:satellite tracking
5760:Fish of Australia
5705:
5704:
5612:Open Tree of Life
5298:Taxon identifiers
5289:
5288:
5186:Shark cage diving
5051:Threatened sharks
5026:Outline of sharks
4486:. The Telegraph.
4464:on 25 March 2010.
4171:978-92-5-104543-5
3861:"Elasmo Research"
3790:Discovery Channel
3288:Knowable Magazine
3060:978-0-00-219960-5
2708:20.500.11815/2381
1817:10.7717/peerj.715
1545:Sharks portal
1521:Outline of sharks
1419:
1346:Notojima Aquarium
1014:larvae and small
921:In a report from
850:coasts of India,
791:, Canada and the
609:sexual dimorphism
416:
415:
410:
402:
394:
386:
378:
305:
191:
174:
16:(Redirected from
5792:
5735:Pantropical fish
5698:
5697:
5685:
5684:
5672:
5671:
5659:
5658:
5646:
5645:
5633:
5632:
5620:
5619:
5607:
5606:
5594:
5593:
5581:
5580:
5571:
5570:
5558:
5557:
5545:
5544:
5532:
5531:
5519:
5518:
5506:
5505:
5493:
5492:
5480:
5479:
5467:
5466:
5454:
5453:
5441:
5440:
5428:
5427:
5415:
5414:
5402:
5401:
5389:
5388:
5379:
5378:
5366:
5365:
5353:
5352:
5340:
5339:
5338:
5325:
5324:
5323:
5293:
5277:
5276:
5265:
5264:
5005:
4998:
4991:
4982:
4950:at marinebio.org
4911:
4890:
4844:
4843:
4841:
4839:
4824:
4818:
4817:
4810:
4804:
4803:
4801:
4799:
4783:
4777:
4771:
4765:
4764:
4762:
4760:
4745:
4739:
4738:
4736:
4734:
4718:
4709:
4708:
4706:
4704:
4689:
4683:
4682:
4671:
4665:
4664:
4662:
4660:
4648:
4642:
4641:
4614:
4608:
4607:
4596:
4590:
4589:
4563:
4552:
4551:
4549:
4547:
4541:
4533:
4527:
4526:
4524:
4522:
4511:
4500:
4499:
4497:
4495:
4485:
4477:
4466:
4465:
4454:
4448:
4447:
4421:
4406:
4405:
4393:
4387:
4381:
4375:
4368:
4362:
4356:
4350:
4344:
4338:
4325:
4319:
4318:
4316:
4314:
4308:
4300:
4294:
4293:
4273:
4262:
4256:
4255:
4245:
4227:
4195:
4189:
4182:
4176:
4175:
4155:
4149:
4148:
4146:
4138:
4132:
4131:
4129:
4127:
4112:
4106:
4099:
4093:
4092:
4090:
4082:
4076:
4075:
4073:
4071:
4057:
4051:
4050:
4048:
4046:
4031:
4025:
4015:
4006:
4005:
3995:
3985:
3957:
3951:
3950:
3948:
3946:
3932:
3926:
3925:
3923:
3921:
3905:
3899:
3898:
3896:
3894:
3879:
3873:
3872:
3870:
3868:
3856:
3847:
3846:
3812:
3799:
3793:
3783:
3774:
3773:
3771:
3769:
3755:Morelle, Rebecca
3751:
3745:
3744:
3742:
3740:
3729:
3723:
3722:
3720:
3718:
3703:
3697:
3696:
3685:
3679:
3678:
3676:
3674:10.3354/esr00300
3652:
3646:
3645:
3643:
3641:
3622:
3616:
3615:
3575:
3569:
3568:
3557:
3551:
3550:
3540:
3522:
3516:
3515:
3510:, archived from
3501:
3495:
3494:
3484:
3474:
3438:
3432:
3431:
3429:
3427:
3412:
3406:
3405:
3403:
3401:
3386:
3380:
3379:
3368:
3362:
3361:
3359:
3357:
3341:
3335:
3334:
3332:
3330:
3315:
3309:
3308:
3306:
3304:
3299:
3279:
3273:
3272:
3270:
3268:
3249:
3236:
3235:
3223:
3217:
3216:
3171:
3165:
3164:
3154:
3145:
3139:
3138:
3122:
3116:
3115:
3071:
3065:
3064:
3044:
3038:
3037:
3017:
3011:
3010:
2966:
2960:
2959:
2933:
2927:
2926:
2908:
2876:
2870:
2869:
2829:
2823:
2822:
2781:(5): 1019â1056.
2770:
2759:
2758:
2722:
2713:
2712:
2710:
2700:
2676:
2663:
2662:
2626:
2617:
2616:
2598:
2574:
2565:
2564:
2558:
2553:
2551:
2543:
2519:
2513:
2512:
2510:
2508:
2494:
2488:
2487:
2477:
2429:
2423:
2422:
2415:
2409:
2408:
2397:
2391:
2390:
2379:
2373:
2372:
2362:
2344:
2320:
2314:
2313:
2303:
2271:
2265:
2264:
2255:McGreevy, Nora.
2252:
2246:
2245:
2235:
2217:
2185:
2179:
2178:
2176:
2174:
2158:
2152:
2151:
2139:
2133:
2132:
2130:
2128:
2114:
2108:
2107:
2081:
2072:
2071:
2038:(6): 1219â1234.
2027:
2016:
2015:
1975:
1969:
1968:
1960:
1954:
1953:
1921:
1904:
1903:
1901:
1899:
1883:
1866:
1865:
1833:
1820:
1804:
1791:
1790:
1774:
1768:
1767:
1747:
1738:
1737:
1709:
1703:
1697:
1685:
1677:
1668:
1667:
1665:
1663:
1649:
1643:
1642:
1640:
1638:
1633:
1603:
1590:
1589:
1587:
1585:
1576:. Archived from
1560:
1543:
1542:
1495:
1485:
1479:
1467:
1454:
1448:
1435:
1420:
1397:Georgia Aquarium
1390:Marine Life Park
1358:Aqua Planet Jeju
1356:), South Korea (
1303:public aquariums
1296:Georgia Aquarium
1209:Gulf of Tadjoura
1129:Christmas Island
1044:The BBC program
750:
738:
726:
714:
702:
560:dermal denticles
490:, South Africa.
408:
406:Rhinodon typicus
401:Kishinouye, 1901
400:
392:
384:
376:
354:
336:
300:
275:Orectolobiformes
206:
205:
185:
168:
163:
162:
139:
118:
108:
45:
38:Temporal range:
32:
21:
5800:
5799:
5795:
5794:
5793:
5791:
5790:
5789:
5710:
5709:
5706:
5701:
5693:
5688:
5680:
5675:
5667:
5662:
5654:
5649:
5641:
5636:
5628:
5623:
5615:
5610:
5602:
5597:
5589:
5586:Observation.org
5584:
5576:
5574:
5566:
5561:
5553:
5548:
5540:
5535:
5527:
5522:
5514:
5509:
5501:
5496:
5488:
5483:
5475:
5470:
5462:
5457:
5449:
5444:
5436:
5431:
5425:rhincodon-typus
5423:
5418:
5410:
5405:
5397:
5392:
5384:
5382:
5376:rhincodon-typus
5374:
5369:
5363:Rhincodon_typus
5361:
5356:
5350:Rhincodon_typus
5348:
5343:
5336:Rhincodon typus
5334:
5333:
5328:
5319:
5318:
5313:
5306:Rhincodon typus
5300:
5290:
5285:
5253:
5229:Shark sanctuary
5212:
5169:South Australia
5141:
5135:
5114:Hybodontiformes
5055:
5014:
5009:
4919:
4914:
4898:
4856:
4852:
4850:Further reading
4847:
4837:
4835:
4834:on 4 March 2009
4826:
4825:
4821:
4812:
4811:
4807:
4797:
4795:
4785:
4784:
4780:
4772:
4768:
4758:
4756:
4747:
4746:
4742:
4732:
4730:
4729:on 8 March 2021
4720:
4719:
4712:
4702:
4700:
4691:
4690:
4686:
4681:on 9 June 2013.
4673:
4672:
4668:
4658:
4656:
4650:
4649:
4645:
4630:
4616:
4615:
4611:
4598:
4597:
4593:
4578:
4565:
4564:
4555:
4545:
4543:
4539:
4535:
4534:
4530:
4520:
4518:
4513:
4512:
4503:
4493:
4491:
4479:
4478:
4469:
4456:
4455:
4451:
4436:
4423:
4422:
4409:
4395:
4394:
4390:
4382:
4378:
4369:
4365:
4357:
4353:
4345:
4341:
4335:Wayback Machine
4326:
4322:
4312:
4310:
4306:
4302:
4301:
4297:
4282:
4271:
4264:
4263:
4259:
4197:
4196:
4192:
4183:
4179:
4172:
4157:
4156:
4152:
4144:
4140:
4139:
4135:
4125:
4123:
4114:
4113:
4109:
4100:
4096:
4088:
4084:
4083:
4079:
4069:
4067:
4059:
4058:
4054:
4044:
4042:
4033:
4032:
4028:
4016:
4009:
3964:Rhincodon typus
3959:
3958:
3954:
3944:
3942:
3934:
3933:
3929:
3919:
3917:
3907:
3906:
3902:
3892:
3890:
3881:
3880:
3876:
3866:
3864:
3858:
3857:
3850:
3810:
3806:Rhincodon typus
3801:
3800:
3796:
3792:, 5 August 2006
3784:
3777:
3767:
3765:
3753:
3752:
3748:
3738:
3736:
3731:
3730:
3726:
3716:
3714:
3705:
3704:
3700:
3687:
3686:
3682:
3654:
3653:
3649:
3639:
3637:
3636:on 5 March 2001
3624:
3623:
3619:
3580:Rhincodon typus
3577:
3576:
3572:
3567:. 24 June 2019.
3565:livescience.com
3559:
3558:
3554:
3524:
3523:
3519:
3503:
3502:
3498:
3445:Rhincodon typus
3440:
3439:
3435:
3425:
3423:
3414:
3413:
3409:
3399:
3397:
3388:
3387:
3383:
3370:
3369:
3365:
3355:
3353:
3352:on 28 June 2016
3343:
3342:
3338:
3328:
3326:
3317:
3316:
3312:
3302:
3300:
3281:
3280:
3276:
3266:
3264:
3251:
3250:
3239:
3225:
3224:
3220:
3173:
3172:
3168:
3152:
3147:
3146:
3142:
3124:
3123:
3119:
3073:
3072:
3068:
3061:
3046:
3045:
3041:
3019:
3018:
3014:
2968:
2967:
2963:
2948:
2935:
2934:
2930:
2878:
2877:
2873:
2831:
2830:
2826:
2772:
2771:
2762:
2724:
2723:
2716:
2678:
2677:
2666:
2628:
2627:
2620:
2576:
2575:
2568:
2554:
2544:
2521:
2520:
2516:
2506:
2504:
2502:www.science.org
2496:
2495:
2491:
2431:
2430:
2426:
2417:
2416:
2412:
2407:. 9 April 2023.
2405:The Japan Times
2399:
2398:
2394:
2381:
2380:
2376:
2322:
2321:
2317:
2273:
2272:
2268:
2254:
2253:
2249:
2200:(6): e0235342.
2187:
2186:
2182:
2172:
2170:
2165:Rhincodon typus
2160:
2159:
2155:
2141:
2140:
2136:
2126:
2124:
2116:
2115:
2111:
2096:
2083:
2082:
2075:
2029:
2028:
2019:
1980:Rhincodon typus
1977:
1976:
1972:
1962:
1961:
1957:
1942:10.1071/MF17393
1923:
1922:
1907:
1897:
1895:
1890:Rhincodon typus
1885:
1884:
1869:
1854:10.1071/MF13330
1835:
1834:
1823:
1805:
1794:
1776:
1775:
1771:
1749:
1748:
1741:
1726:10.2307/1442501
1711:
1710:
1706:
1679:
1678:
1671:
1661:
1659:
1651:
1650:
1646:
1636:
1634:
1610:Rhincodon typus
1605:
1604:
1593:
1583:
1581:
1566:Rhincodon typus
1562:
1561:
1557:
1553:
1537:
1524:
1517:
1411:
1409:
1288:
1225:
1113:Cendrawasih Bay
1073:
1068:
994:). It feeds on
992:megamouth shark
942:
916:Pilar, Sorsogon
885:nuclear testing
876:
761:
754:
751:
742:
739:
730:
727:
718:
715:
706:
703:
680:standard length
631:
614:sexual maturity
601:
582:cloudy catshark
533:caudal peduncle
516:
473:megamouth shark
425:Rhincodon typus
390:Rhiniodon typus
345:
338:
334:Rhincodon typus
332:
319:
299:
200:
192:
175:
164:
160:
153:
129:Similan Islands
109:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
40:
39:
36:
28:
23:
22:
15:
12:
11:
5:
5798:
5796:
5788:
5787:
5782:
5777:
5772:
5770:Fish of Africa
5767:
5762:
5757:
5752:
5747:
5745:Apex predators
5742:
5737:
5732:
5727:
5725:Rhincodontidae
5722:
5712:
5711:
5703:
5702:
5700:
5699:
5686:
5673:
5660:
5647:
5634:
5621:
5608:
5595:
5582:
5572:
5559:
5546:
5533:
5520:
5507:
5494:
5481:
5468:
5455:
5442:
5429:
5416:
5403:
5390:
5380:
5367:
5354:
5341:
5326:
5310:
5308:
5302:
5301:
5296:
5287:
5286:
5284:
5283:
5271:
5258:
5255:
5254:
5252:
5251:
5246:
5241:
5236:
5234:Shark Alliance
5231:
5226:
5220:
5218:
5214:
5213:
5211:
5210:
5205:
5204:
5203:
5198:
5188:
5183:
5178:
5177:
5176:
5171:
5166:
5161:
5156:
5145:
5143:
5137:
5136:
5134:
5133:
5131:List of sharks
5128:
5122:
5116:
5110:
5104:
5099:
5094:
5089:
5084:
5079:
5077:Hexanchiformes
5074:
5069:
5063:
5061:
5057:
5056:
5054:
5053:
5048:
5046:Threat display
5043:
5038:
5033:
5028:
5022:
5020:
5016:
5015:
5010:
5008:
5007:
5000:
4993:
4985:
4979:
4978:
4971:
4966:
4961:
4956:
4951:
4945:
4940:
4935:
4930:
4925:
4918:
4917:External links
4915:
4913:
4912:
4896:
4891:
4865:(6): 1219â34.
4853:
4851:
4848:
4846:
4845:
4819:
4805:
4778:
4766:
4740:
4710:
4699:on 3 June 2007
4684:
4666:
4643:
4628:
4609:
4591:
4576:
4553:
4528:
4501:
4467:
4449:
4434:
4407:
4388:
4376:
4363:
4351:
4339:
4320:
4295:
4280:
4257:
4190:
4188:. gulfnews.com
4177:
4170:
4150:
4133:
4107:
4094:
4077:
4052:
4026:
4007:
3952:
3927:
3900:
3874:
3848:
3821:(4): 199â212.
3794:
3786:Jurassic Shark
3775:
3746:
3724:
3713:on 18 May 2023
3698:
3680:
3667:(2): 117â124.
3647:
3626:Clark, Eugenie
3617:
3590:(3): 219â223.
3570:
3552:
3517:
3496:
3433:
3407:
3381:
3363:
3336:
3310:
3274:
3237:
3218:
3183:(4): 559â567.
3166:
3140:
3117:
3082:(1): 299â308.
3066:
3059:
3039:
3012:
2961:
2946:
2928:
2891:(1): 378â385.
2871:
2824:
2760:
2714:
2664:
2637:(4): 441â451.
2618:
2566:
2557:|journal=
2514:
2489:
2424:
2410:
2392:
2374:
2315:
2286:(1): coaa120.
2266:
2247:
2180:
2153:
2134:
2109:
2094:
2073:
2017:
1970:
1955:
1905:
1867:
1821:
1792:
1769:
1739:
1704:
1700:Penrith (1972)
1669:
1644:
1591:
1580:on 27 May 2022
1554:
1552:
1549:
1548:
1547:
1535:
1530:
1528:List of sharks
1516:
1513:
1483:gurano bintang
1408:
1405:
1344:Marine World,
1331:Osaka Aquarium
1287:
1284:
1224:
1221:
1217:Horn of Africa
1072:
1069:
1067:
1064:
941:
938:
875:
872:
793:Sea of Okhotsk
760:
757:
756:
755:
752:
745:
743:
740:
733:
731:
728:
721:
719:
716:
709:
707:
704:
697:
630:
627:
623:photogrammetry
600:
597:
515:
512:
461:Chondrichthyes
457:Elasmobranchii
453:Rhincodontidae
430:filter-feeding
414:
413:
412:
411:
403:
395:
393:A. Smith, 1828
387:
379:
368:
367:
361:
360:
356:
355:
347:
346:
339:
328:
327:
321:
320:
313:
311:
307:
306:
292:
288:
287:
285:Rhincodontidae
282:
278:
277:
272:
268:
267:
262:
258:
257:
255:Elasmobranchii
252:
248:
247:
245:Chondrichthyes
242:
238:
237:
232:
228:
227:
222:
218:
217:
212:
208:
207:
194:
193:
180:
177:
176:
158:
155:
154:
149:
146:
145:
141:
140:
132:
131:
120:
119:
111:
110:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
46:
37:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
5797:
5786:
5783:
5781:
5778:
5776:
5773:
5771:
5768:
5766:
5763:
5761:
5758:
5756:
5753:
5751:
5748:
5746:
5743:
5741:
5738:
5736:
5733:
5731:
5728:
5726:
5723:
5721:
5718:
5717:
5715:
5708:
5696:
5691:
5687:
5683:
5678:
5674:
5670:
5665:
5661:
5657:
5652:
5648:
5644:
5639:
5635:
5631:
5626:
5622:
5618:
5613:
5609:
5605:
5600:
5596:
5592:
5587:
5583:
5579:
5573:
5569:
5564:
5560:
5556:
5551:
5547:
5543:
5538:
5534:
5530:
5525:
5521:
5517:
5512:
5508:
5504:
5499:
5495:
5491:
5486:
5482:
5478:
5473:
5469:
5465:
5460:
5456:
5452:
5447:
5443:
5439:
5434:
5430:
5426:
5421:
5417:
5413:
5408:
5404:
5400:
5395:
5391:
5387:
5381:
5377:
5372:
5368:
5364:
5359:
5355:
5351:
5346:
5342:
5337:
5331:
5327:
5322:
5316:
5312:
5311:
5309:
5307:
5303:
5299:
5294:
5282:
5281:
5272:
5270:
5269:
5260:
5259:
5256:
5250:
5247:
5245:
5242:
5240:
5237:
5235:
5232:
5230:
5227:
5225:
5222:
5221:
5219:
5215:
5209:
5208:Shark tourism
5206:
5202:
5199:
5197:
5194:
5193:
5192:
5191:Shark culling
5189:
5187:
5184:
5182:
5179:
5175:
5174:United States
5172:
5170:
5167:
5165:
5162:
5160:
5157:
5155:
5152:
5151:
5150:
5147:
5146:
5144:
5138:
5132:
5129:
5127:
5123:
5121:
5117:
5115:
5111:
5109:
5105:
5103:
5100:
5098:
5095:
5093:
5090:
5088:
5085:
5083:
5080:
5078:
5075:
5073:
5070:
5068:
5065:
5064:
5062:
5058:
5052:
5049:
5047:
5044:
5042:
5039:
5037:
5034:
5032:
5031:Shark anatomy
5029:
5027:
5024:
5023:
5021:
5017:
5013:
5006:
5001:
4999:
4994:
4992:
4987:
4986:
4983:
4976:
4972:
4970:
4967:
4965:
4962:
4960:
4957:
4955:
4952:
4949:
4946:
4944:
4941:
4939:
4936:
4934:
4931:
4929:
4926:
4924:
4921:
4920:
4916:
4909:
4905:
4901:
4897:
4895:
4892:
4888:
4884:
4880:
4876:
4872:
4868:
4864:
4860:
4855:
4854:
4849:
4833:
4829:
4828:"Whale Shark"
4823:
4820:
4815:
4809:
4806:
4793:
4789:
4782:
4779:
4775:
4770:
4767:
4755:
4751:
4744:
4741:
4728:
4724:
4717:
4715:
4711:
4698:
4694:
4688:
4685:
4680:
4676:
4670:
4667:
4654:
4647:
4644:
4639:
4635:
4631:
4629:9780867271676
4625:
4621:
4613:
4610:
4605:
4601:
4595:
4592:
4587:
4583:
4579:
4577:9780867271676
4573:
4569:
4562:
4560:
4558:
4554:
4538:
4532:
4529:
4517:
4510:
4508:
4506:
4502:
4489:
4484:
4476:
4474:
4472:
4468:
4463:
4459:
4453:
4450:
4445:
4441:
4437:
4435:9780867271676
4431:
4427:
4420:
4418:
4416:
4414:
4412:
4408:
4403:
4399:
4392:
4389:
4385:
4380:
4377:
4373:
4367:
4364:
4360:
4355:
4352:
4348:
4343:
4340:
4336:
4332:
4329:
4324:
4321:
4305:
4299:
4296:
4291:
4287:
4283:
4281:9781988514628
4277:
4270:
4269:
4261:
4258:
4253:
4249:
4244:
4239:
4235:
4231:
4226:
4221:
4217:
4213:
4209:
4205:
4201:
4194:
4191:
4187:
4181:
4178:
4173:
4167:
4163:
4162:
4154:
4151:
4143:
4137:
4134:
4122:
4118:
4111:
4108:
4104:
4098:
4095:
4087:
4081:
4078:
4066:
4062:
4056:
4053:
4041:
4040:DIVE Magazine
4037:
4030:
4027:
4023:
4022:Time magazine
4019:
4014:
4012:
4008:
4003:
3999:
3994:
3989:
3984:
3979:
3975:
3971:
3967:
3965:
3956:
3953:
3941:
3937:
3931:
3928:
3916:on 5 May 2015
3915:
3911:
3904:
3901:
3888:
3884:
3883:"Whale shark"
3878:
3875:
3862:
3855:
3853:
3849:
3844:
3840:
3836:
3832:
3828:
3824:
3820:
3816:
3809:
3807:
3798:
3795:
3791:
3787:
3782:
3780:
3776:
3764:
3760:
3756:
3750:
3747:
3734:
3728:
3725:
3712:
3708:
3702:
3699:
3694:
3690:
3684:
3681:
3675:
3670:
3666:
3662:
3658:
3651:
3648:
3635:
3631:
3627:
3621:
3618:
3613:
3609:
3605:
3601:
3597:
3593:
3589:
3585:
3581:
3574:
3571:
3566:
3562:
3556:
3553:
3548:
3544:
3539:
3534:
3530:
3529:
3521:
3518:
3513:
3509:
3508:
3500:
3497:
3492:
3488:
3483:
3478:
3473:
3468:
3464:
3460:
3457:(4): e18994.
3456:
3452:
3448:
3446:
3437:
3434:
3422:
3418:
3411:
3408:
3396:
3392:
3385:
3382:
3377:
3373:
3367:
3364:
3351:
3347:
3340:
3337:
3325:
3321:
3314:
3311:
3298:
3293:
3289:
3285:
3278:
3275:
3263:
3259:
3255:
3254:"Whale Shark"
3248:
3246:
3244:
3242:
3238:
3233:
3229:
3222:
3219:
3214:
3210:
3206:
3202:
3198:
3194:
3190:
3186:
3182:
3178:
3170:
3167:
3162:
3158:
3151:
3144:
3141:
3136:
3132:
3128:
3121:
3118:
3113:
3109:
3105:
3101:
3097:
3093:
3089:
3085:
3081:
3077:
3070:
3067:
3062:
3056:
3052:
3051:
3043:
3040:
3035:
3031:
3027:
3023:
3016:
3013:
3008:
3004:
3000:
2996:
2992:
2988:
2984:
2980:
2977:(1611): 438.
2976:
2972:
2965:
2962:
2957:
2953:
2949:
2947:9781241491611
2943:
2939:
2932:
2929:
2924:
2920:
2916:
2912:
2907:
2902:
2898:
2894:
2890:
2886:
2882:
2875:
2872:
2867:
2863:
2859:
2855:
2851:
2847:
2843:
2839:
2835:
2828:
2825:
2820:
2816:
2812:
2808:
2804:
2800:
2796:
2792:
2788:
2784:
2780:
2776:
2769:
2767:
2765:
2761:
2756:
2752:
2748:
2744:
2740:
2736:
2732:
2728:
2721:
2719:
2715:
2709:
2704:
2699:
2694:
2690:
2686:
2682:
2675:
2673:
2671:
2669:
2665:
2660:
2656:
2652:
2648:
2644:
2640:
2636:
2632:
2625:
2623:
2619:
2614:
2610:
2606:
2602:
2597:
2592:
2588:
2584:
2580:
2573:
2571:
2567:
2562:
2549:
2541:
2537:
2533:
2529:
2525:
2518:
2515:
2503:
2499:
2493:
2490:
2485:
2481:
2476:
2471:
2467:
2463:
2459:
2455:
2451:
2447:
2443:
2439:
2435:
2428:
2425:
2420:
2414:
2411:
2406:
2402:
2396:
2393:
2388:
2384:
2378:
2375:
2370:
2366:
2361:
2356:
2352:
2348:
2343:
2338:
2334:
2330:
2326:
2319:
2316:
2311:
2307:
2302:
2297:
2293:
2289:
2285:
2281:
2277:
2270:
2267:
2262:
2258:
2251:
2248:
2243:
2239:
2234:
2229:
2225:
2221:
2216:
2211:
2207:
2203:
2199:
2195:
2191:
2184:
2181:
2168:
2166:
2157:
2154:
2149:
2145:
2138:
2135:
2123:
2122:MarineBio.org
2119:
2113:
2110:
2105:
2101:
2097:
2095:0-642-54900-1
2091:
2087:
2080:
2078:
2074:
2069:
2065:
2061:
2057:
2053:
2049:
2045:
2041:
2037:
2033:
2026:
2024:
2022:
2018:
2013:
2009:
2005:
2001:
1997:
1993:
1990:(3): 706â14.
1989:
1985:
1981:
1974:
1971:
1966:
1959:
1956:
1951:
1947:
1943:
1939:
1935:
1931:
1927:
1920:
1918:
1916:
1914:
1912:
1910:
1906:
1893:
1891:
1882:
1880:
1878:
1876:
1874:
1872:
1868:
1863:
1859:
1855:
1851:
1847:
1843:
1839:
1832:
1830:
1828:
1826:
1822:
1818:
1814:
1810:
1803:
1801:
1799:
1797:
1793:
1788:
1784:
1780:
1773:
1770:
1766:(4): 215â217.
1765:
1761:
1757:
1755:
1746:
1744:
1740:
1735:
1731:
1727:
1723:
1719:
1715:
1708:
1705:
1701:
1698:Reprinted in
1695:
1691:
1690:
1684:
1676:
1674:
1670:
1658:
1654:
1648:
1645:
1632:
1627:
1623:
1619:
1618:
1613:
1611:
1602:
1600:
1598:
1596:
1592:
1579:
1575:
1574:
1569:
1567:
1559:
1556:
1550:
1546:
1541:
1536:
1534:
1531:
1529:
1526:
1525:
1522:
1514:
1512:
1510:
1506:
1501:
1499:
1494:
1489:
1484:
1478:
1477:
1471:
1466:
1460:
1458:
1453:
1447:
1441:
1439:
1434:
1429:
1407:Human culture
1406:
1404:
1402:
1398:
1393:
1391:
1387:
1383:
1379:
1375:
1371:
1367:
1363:
1359:
1355:
1351:
1347:
1343:
1339:
1334:
1332:
1328:
1324:
1316:
1311:
1307:
1304:
1297:
1292:
1285:
1283:
1280:
1278:
1274:
1269:
1267:
1263:
1260:In 2010, the
1258:
1256:
1253:
1248:
1246:
1242:
1238:
1234:
1230:
1222:
1220:
1218:
1214:
1210:
1206:
1202:
1198:
1194:
1190:
1186:
1182:
1178:
1177:West Malaysia
1174:
1170:
1166:
1162:
1158:
1157:Caribbean Sea
1154:
1150:
1146:
1142:
1138:
1134:
1130:
1126:
1125:Ningaloo Reef
1122:
1118:
1114:
1110:
1106:
1097:
1093:
1090:
1082:
1077:
1070:
1065:
1063:
1061:
1060:microplastics
1056:
1052:
1049:
1048:
1042:
1039:
1035:
1033:
1029:
1025:
1021:
1017:
1013:
1009:
1005:
1001:
997:
993:
989:
988:basking shark
985:
984:filter feeder
977:
972:
966:
961:
954:
951:
946:
939:
937:
935:
931:
928:
924:
919:
917:
911:
909:
908:ovoviviparous
904:
902:
901:Ningaloo Reef
898:
893:
889:
886:
882:
873:
871:
868:
863:
861:
857:
853:
849:
845:
841:
837:
833:
830:in Tanzania,
829:
825:
821:
817:
816:Darwin Island
813:
812:Ningaloo Reef
809:
805:
800:
798:
794:
790:
786:
782:
778:
773:
770:
766:
758:
749:
744:
737:
732:
725:
720:
713:
708:
701:
696:
694:
692:
687:
685:
681:
677:
672:
667:
664:
659:
657:
653:
649:
648:Hugh M. Smith
645:
643:
639:
634:
628:
626:
624:
618:
615:
610:
606:
598:
596:
594:
591:
587:
583:
579:
575:
571:
568:
564:
561:
556:
554:
550:
546:
542:
541:pectoral fins
538:
534:
530:
526:
522:
513:
511:
509:
508:baleen whales
505:
501:
497:
493:
489:
484:
482:
478:
477:basking shark
474:
468:
466:
462:
459:in the class
458:
454:
450:
449:
444:
441:
438:
434:
431:
427:
426:
421:
407:
404:
399:
396:
391:
388:
385:Denison, 1937
383:
380:
375:
372:
371:
369:
366:
362:
357:
353:
348:
343:
337:
335:
329:
326:
325:Binomial name
322:
318:
317:
316:R. typus
312:
309:
308:
303:
298:
297:
293:
290:
289:
286:
283:
280:
279:
276:
273:
270:
269:
266:
265:Selachimorpha
263:
261:Subdivision:
260:
259:
256:
253:
250:
249:
246:
243:
240:
239:
236:
233:
230:
229:
226:
223:
220:
219:
216:
213:
210:
209:
204:
199:
195:
189:
183:
178:
172:
167:
156:
152:
147:
142:
138:
133:
130:
126:
121:
117:
112:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
44:
33:
30:
19:
5765:Fish of Asia
5707:
5305:
5278:
5266:
5164:South Africa
5149:Shark attack
5097:Squaliformes
4908:the original
4903:
4862:
4859:J. Fish Biol
4858:
4836:. Retrieved
4832:the original
4822:
4808:
4796:. Retrieved
4791:
4781:
4769:
4759:12 September
4757:. Retrieved
4753:
4743:
4731:. Retrieved
4727:the original
4701:. Retrieved
4697:the original
4687:
4679:the original
4669:
4657:. Retrieved
4646:
4619:
4612:
4594:
4567:
4544:. Retrieved
4531:
4519:. Retrieved
4492:. Retrieved
4462:the original
4452:
4425:
4401:
4391:
4379:
4366:
4354:
4342:
4323:
4311:. Retrieved
4298:
4267:
4260:
4234:10754/676739
4207:
4203:
4193:
4180:
4160:
4153:
4136:
4124:. Retrieved
4120:
4110:
4097:
4080:
4068:. Retrieved
4064:
4055:
4043:. Retrieved
4039:
4029:
4021:
3973:
3969:
3963:
3955:
3943:. Retrieved
3939:
3930:
3918:. Retrieved
3914:the original
3903:
3893:17 September
3891:. Retrieved
3887:the original
3877:
3867:17 September
3865:. Retrieved
3818:
3814:
3805:
3797:
3785:
3766:. Retrieved
3762:
3749:
3737:. Retrieved
3727:
3715:. Retrieved
3711:the original
3701:
3692:
3683:
3664:
3660:
3650:
3640:26 September
3638:. Retrieved
3634:the original
3620:
3587:
3583:
3579:
3573:
3564:
3555:
3527:
3520:
3512:the original
3506:
3499:
3454:
3450:
3444:
3436:
3424:. Retrieved
3420:
3410:
3398:. Retrieved
3394:
3384:
3375:
3366:
3354:. Retrieved
3350:the original
3339:
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854:, southern
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822:in Mexico,
753:Top of head
537:dorsal fins
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514:Description
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125:Andaman Sea
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5714:Categories
5196:Drum lines
5120:Symmoriida
4973:Photos of
4838:3 November
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