4502:
1686:
2282:
4496:
144:
4512:
4518:
4507:
123:
374:. The ribs of the animal, which functioned as gliding organs, were found in a half-open position, which indicates a post-mortem relaxation of the folded wing. So far this is the only known fossil gliding lizard, though there are other unrelated animals that also use their ribs to glide.
419:. Despite the 11-centimetre (4.3 in) "rib-span", the lizard might have been quite agile in the air, possibly to escape the feathered dinosaurs that coexisted with it.
354:
was 15.5 centimetres (6.1 in) long, 9.5 centimetres (3.7 in) of which was tail, although the describers say it was probably a juvenile. This is indicated by the
4733:
1741:
4797:
4501:
1699:
4720:
395:, have a different membrane attachment, toes to toes or limb to limb. Two creatures use the same way to glide, the present day Flying Lizard (genus
366:
V is greatly elongated. The radius and ulna are distally divergent. The body is covered in small, granular scales, showing little size variation.
4792:
1659:
338:, it was later shown that this was due to misinterpretation of the crushed skull, and its affinities with other lizards remains uncertain.
1734:
310:
is its bizarre oversized ribs, eight on each side, which were attached to a membrane of body tissue and allowed the lizard to
429:, states that it is possible that it could have glided as far as half a football field, much farther than that of the modern
1639:
4495:
2281:
288:
1534:
dentition was actually the crushed, jagged broken edge of the jaw, rendering its identification as an iguanian doubtful.
4787:
1703:
1553:
292:
1727:
311:
143:
20:
4802:
461:. This was based on the strict consensus of the four most parsimonious trees. Below is the tree recovered by Li
422:
4613:
1709:
391:
4511:
4688:
4392:
4807:
4759:
3697:
3301:
3267:
3224:
4506:
4517:
4184:
4100:
3104:
3016:
2390:
2348:
4282:
4128:
4028:
3344:
3233:
2166:
1589:
4008:
4001:
3479:
3316:
3309:
3196:
2976:
2969:
2938:
2486:
2369:
2203:
4598:
4035:
4019:
3974:
3967:
3898:
3472:
3372:
3358:
3294:
3251:
3203:
3161:
2778:
2629:
2617:
2592:
2570:
2547:
2472:
2465:
2397:
2210:
1691:
280:
4065:
4058:
3624:
3617:
3548:
3541:
3365:
3351:
3337:
3330:
3097:
3050:
2945:
2820:
2813:
2750:
2577:
2436:
2362:
2355:
2318:
2189:
4644:
4121:
3795:
3514:
3399:
3379:
3323:
3090:
3023:
2992:
2526:
2519:
2448:
2412:
2408:
2334:
2306:
2302:
2196:
2035:
1993:
698:
138:
4764:
4651:
4461:
4372:
4219:
4212:
4135:
3981:
3659:
3603:
3582:
3260:
3175:
2842:
2799:
2688:
4233:
3905:
3781:
3645:
3274:
2931:
2563:
2533:
4746:
4738:
4198:
4107:
2867:
2852:
2827:
2658:
2584:
1948:
1655:
1617:
284:
4751:
4142:
4072:
2860:
4161:
3912:
3673:
2500:
2158:
1647:
1607:
1597:
982:
715:
303:
300:
261:
37:
4666:
3834:
3719:
3070:
2493:
2254:
2233:
2108:
1963:
1288:
1242:
865:
674:
528:
482:
386:
363:
1593:
4576:
4435:
4268:
4049:
3992:
3856:
3788:
3704:
3210:
3147:
3034:
2874:
2794:
2180:
2007:
1986:
1956:
1939:
1612:
1527:
611:
454:
331:
276:
385:
is one of the few creatures that glide using their ribs. Other creatures, such as the
295:
000666, a single complete skeleton with skin impressions. The specimen comes from the
4781:
4658:
4605:
4468:
4275:
4226:
4205:
4191:
4086:
3818:
3803:
3774:
3727:
3711:
3610:
3596:
3429:
3182:
2757:
2717:
2696:
2681:
2247:
2224:
2069:
2061:
2021:
1820:
816:
626:
458:
411:
397:
335:
73:
4453:
4358:
4351:
4330:
4293:
4079:
3960:
3889:
3842:
3555:
3507:
3465:
3456:
3154:
3043:
3003:
2910:
2708:
2673:
2650:
2422:
2376:
2143:
2117:
1797:
1782:
1574:
1335:
1257:
785:
355:
243:
4591:
4421:
4401:
4309:
4261:
4251:
4240:
4177:
4114:
4093:
3940:
3865:
3755:
3682:
3638:
3445:
3438:
3138:
3127:
3118:
2956:
2922:
2771:
2642:
2555:
2479:
2429:
2383:
2341:
2240:
2150:
2090:
2076:
2055:
1921:
1914:
1907:
1894:
1890:
1834:
1713:
1311:
1186:
1122:
880:
48:
1582:
Proceedings of the
National Academy of Sciences of the United States of America
4554:
4383:
4344:
4316:
3690:
3631:
3589:
3494:
3415:
3407:
3285:
2983:
2666:
2458:
2262:
1977:
1875:
1827:
1681:
1651:
1352:
1105:
1029:
945:
838:
505:
359:
269:
93:
58:
4711:
4561:
4547:
4412:
4365:
4337:
3652:
3189:
3168:
3061:
2730:
2083:
2046:
2028:
2014:
1868:
1841:
1602:
1368:
1155:
1044:
770:
557:
319:
296:
155:
98:
1621:
1719:
122:
4705:
3951:
3927:
2540:
1859:
1770:
1751:
1531:
1075:
596:
450:
446:
406:
371:
347:
327:
315:
195:
175:
88:
83:
68:
63:
53:
19:
This article is about the extinct
Cretaceous lizard. For other uses, see
4725:
4535:
4323:
3878:
3521:
3390:
2764:
2741:
2327:
2097:
1928:
1776:
1203:
967:
323:
225:
185:
103:
78:
4583:
4569:
4428:
3081:
1764:
402:
272:
165:
4682:
415:, but the Triassic look-alikes lived over 100 million years before
3532:
265:
2000:
4686:
4632:
4533:
3753:
2908:
2614:
2297:
1795:
1723:
322:
analysis in the same study suggesting that it was grouped with
1573:
Pi-Peng Li; Ke-Qin Gao; Lian-Hai Hou & Xing Xu (2007).
1646:(1 ed.), Cambridge University Press, pp. 5β110,
314:. While in its original description it was considered to
1530:
said that what the describing authors misinterpreted as
425:, a Chinese paleontologist and one of the describers of
1700:
Photo in the News: Ancient Lizard Glided Using Its Ribs
1575:"A gliding lizard from the Early Cretaceous of China"
4046:
3839:
3831:
3815:
3670:
3426:
3248:
3221:
3115:
2989:
2839:
2299:
2221:
2177:
1974:
362:
IV is shorter than the other metacarpals, and pedal
4695:
4445:
4411:
4391:
4382:
4301:
4292:
4250:
4169:
4160:
4045:
4018:
3991:
3950:
3926:
3888:
3864:
3855:
3814:
3766:
3669:
3570:
3531:
3493:
3455:
3425:
3389:
3284:
3247:
3220:
3137:
3114:
3080:
3060:
3033:
3002:
2955:
2921:
2838:
2792:
2740:
2716:
2707:
2628:
2511:
2446:
2407:
2317:
2220:
2176:
2131:
2107:
2045:
1973:
1938:
1889:
1858:
1808:
1644:
1638:Gower, David J.; Zaher, Hussam, eds. (2022-08-11),
370:had slightly curved claws, indicating that it was
306:, near Yizhou. The most notable feature about
1735:
1549:
1547:
8:
1556:Ker Than, NBC News. Published March 20, 2007
4683:
4629:
4530:
4388:
4298:
4166:
3932:
3870:
3861:
3824:
3763:
3750:
3574:
3499:
3008:
2961:
2918:
2905:
2805:
2722:
2713:
2634:
2625:
2611:
2314:
2294:
2135:
1899:
1812:
1805:
1792:
1742:
1728:
1720:
1568:
1566:
1564:
1562:
121:
27:
1611:
1601:
1710:Ancient Lizard Glided on Stretched Ribs
1554:Ancient lizard glided on stretched skin
1543:
1640:"The Squamate and Snake Fossil Record"
1526:However, in a later 2022 publication,
441:In the original paper describing it,
358:carpals and poorly-ossified tarsals.
275:discovered in the Zhuanchengzi, near
7:
1633:
1631:
1327:
1303:
1281:
1235:
1228:
1178:
1171:
1147:
1097:
1090:
1068:
1022:
1015:
960:
938:
931:
858:
831:
809:
763:
756:
749:
690:
666:
659:
572:
550:
543:
521:
497:
475:
468:
14:
4798:Early Cretaceous reptiles of Asia
4516:
4510:
4505:
4500:
4494:
2280:
1684:
142:
1:
4793:Fossil taxa described in 2007
4446:Other platynotans / varanoids
4170:Basal varanoids / platynotans
389:and the Malabar flying frog,
260:(meaning "flying dragon" in
1704:National Geographic Society
4824:
2512:Other polyglyphanodontians
21:Xianglong (disambiguation)
18:
4639:
4628:
4542:
4529:
4492:
3935:
3873:
3827:
3762:
3749:
3577:
3502:
3011:
2964:
2917:
2904:
2808:
2725:
2637:
2624:
2610:
2313:
2293:
2278:
2138:
1902:
1815:
1804:
1791:
1759:
1716:March 19, 2007 01:00pm ET
1652:10.1017/9781108938891.003
1366:
1349:
1332:
1325:
1308:
1301:
1286:
1279:
1255:
1240:
1233:
1226:
1200:
1183:
1176:
1169:
1152:
1145:
1119:
1102:
1095:
1088:
1073:
1066:
1042:
1027:
1020:
1013:
980:
965:
958:
943:
936:
929:
878:
863:
856:
836:
829:
814:
807:
783:
768:
761:
754:
747:
712:
695:
688:
671:
664:
657:
624:
609:
594:
577:
570:
555:
548:
541:
526:
519:
502:
495:
480:
473:
231:
224:
139:Scientific classification
137:
129:
120:
30:
409:fossil reptiles such as
16:Extinct genus of lizards
4634:Indeterminate squamates
2132:Other extinct iguanians
1603:10.1073/pnas.0609552104
392:Rhacophorus malabaricus
4760:Paleobiology Database
4302:Basal Goannasaurians
2449:Polyglyphanodontines
2413:macrocephalosaurines
2630:Scincogekkonomorpha
2618:Scincogekkonomorpha
1692:Paleontology portal
1594:2007PNAS..104.5507L
445:was recovered in a
291:. It is known from
4788:Cretaceous lizards
3698:Mimobecklesisaurus
3571:Other scincomorphs
3302:Camptognathosaurus
3268:Palaeoscincosaurus
3225:Polyodontobaenidae
2993:Polyglyphanodontia
2303:Polyglyphanodontia
699:Macrocephalosaurus
130:Pencil drawing of
4775:
4774:
4747:Open Tree of Life
4689:Taxon identifiers
4680:
4679:
4676:
4675:
4624:
4623:
4525:
4524:
4490:
4489:
4486:
4485:
4482:
4481:
4478:
4477:
4156:
4155:
4152:
4151:
4108:Paraplacosauriops
3922:
3921:
3851:
3850:
3767:Basal anguimorphs
3745:
3744:
3741:
3740:
3737:
3736:
3566:
3565:
3489:
3488:
3243:
3242:
2900:
2899:
2896:
2895:
2892:
2891:
2888:
2887:
2884:
2883:
2788:
2787:
2659:Eichstaettisaurus
2606:
2605:
2602:
2601:
2289:
2288:
2276:
2275:
2272:
2271:
2127:
2126:
1885:
1884:
1661:978-1-108-93889-1
1588:(13): 5507β5509.
1523:
1522:
1514:
1513:
1505:
1504:
1496:
1495:
1487:
1486:
1478:
1477:
1469:
1468:
1460:
1459:
1451:
1450:
1442:
1441:
1433:
1432:
1424:
1423:
1415:
1414:
1406:
1405:
1397:
1396:
1388:
1387:
1379:
1378:
1268:
1267:
1215:
1214:
1134:
1133:
1055:
1054:
1002:
1001:
993:
992:
918:
917:
909:
908:
900:
899:
891:
890:
796:
795:
736:
735:
727:
726:
646:
645:
637:
636:
285:Liaoning Province
253:
252:
247:
220:
4815:
4803:Fossils of China
4768:
4767:
4755:
4754:
4742:
4741:
4729:
4728:
4716:
4715:
4714:
4684:
4630:
4531:
4520:
4514:
4509:
4504:
4498:
4389:
4299:
4185:Colpodontosaurus
4167:
4162:Paleoanguimorpha
4101:Paraglyptosaurus
4048:
3933:
3871:
3862:
3841:
3833:
3825:
3817:
3764:
3751:
3674:Paramacellodidae
3672:
3575:
3500:
3428:
3250:
3223:
3117:
3105:Louisamphisbaena
3017:Oligodontosaurus
3009:
2991:
2962:
2919:
2906:
2841:
2806:
2723:
2714:
2635:
2626:
2612:
2501:Sineoamphisbaena
2391:Stypodontosaurus
2349:Harmodontosaurus
2315:
2301:
2295:
2284:
2223:
2179:
2159:Huehuecuetzpalli
2136:
1976:
1900:
1813:
1806:
1793:
1744:
1737:
1730:
1721:
1694:
1689:
1688:
1687:
1671:
1670:
1669:
1668:
1635:
1626:
1625:
1615:
1605:
1579:
1570:
1557:
1551:
1328:
1304:
1282:
1236:
1229:
1179:
1172:
1148:
1098:
1091:
1069:
1023:
1016:
983:Gymnophthalmidae
961:
939:
932:
859:
832:
810:
764:
757:
750:
716:Sineoamphisbaena
691:
667:
660:
573:
551:
544:
522:
498:
476:
469:
304:Yixian Formation
301:Lower Cretaceous
237:
214:
207:
147:
146:
125:
115:
45:
38:Early Cretaceous
36:Temporal range:
28:
4823:
4822:
4818:
4817:
4816:
4814:
4813:
4812:
4778:
4777:
4776:
4771:
4763:
4758:
4750:
4745:
4737:
4732:
4724:
4719:
4710:
4709:
4704:
4691:
4681:
4672:
4667:Schillerosaurus
4635:
4620:
4538:
4521:
4474:
4441:
4407:
4378:
4288:
4283:Provaranosaurus
4246:
4148:
4129:Proglyptosaurus
4041:
4029:Paragerrhonotus
4014:
3987:
3946:
3918:
3884:
3847:
3835:Aigialosauridae
3810:
3758:
3733:
3720:Pseudosaurillus
3665:
3562:
3527:
3485:
3451:
3421:
3385:
3345:Maioricalacerta
3280:
3239:
3234:Polyodontobaena
3216:
3133:
3110:
3076:
3071:Anniealexandria
3056:
3029:
2998:
2951:
2913:
2880:
2834:
2797:
2784:
2736:
2703:
2620:
2598:
2507:
2494:Polyglyphanodon
2442:
2403:
2309:
2285:
2268:
2255:Pleurodontagama
2234:Heterodontagama
2216:
2172:
2167:Swainiguanoides
2123:
2109:Phrynosomatidae
2103:
2041:
1969:
1964:Geiseltaliellus
1934:
1881:
1854:
1800:
1787:
1755:
1748:
1690:
1685:
1683:
1680:
1675:
1674:
1666:
1664:
1662:
1637:
1636:
1629:
1577:
1572:
1571:
1560:
1552:
1545:
1540:
1524:
1515:
1506:
1497:
1488:
1479:
1470:
1461:
1452:
1443:
1434:
1425:
1416:
1407:
1398:
1389:
1380:
1289:Dolichosauridae
1269:
1243:Aigialosauridae
1216:
1135:
1056:
1003:
994:
919:
910:
901:
892:
866:Diplodactylinae
797:
737:
728:
675:Polyglyphanodon
647:
638:
529:Rhynchocephalia
483:Kuehneosauridae
439:
387:flying squirrel
380:
378:Gliding ability
344:
318:lizard, with a
213:
205:
141:
132:Xianglong zhaoi
116:
114:
113:
112:
111:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
43:
42:
41:
34:
24:
17:
12:
11:
5:
4821:
4819:
4811:
4810:
4805:
4800:
4795:
4790:
4780:
4779:
4773:
4772:
4770:
4769:
4756:
4743:
4730:
4717:
4701:
4699:
4693:
4692:
4687:
4678:
4677:
4674:
4673:
4671:
4670:
4663:
4655:
4648:
4640:
4637:
4636:
4633:
4626:
4625:
4622:
4621:
4619:
4618:
4609:
4602:
4595:
4588:
4580:
4577:Lapparentophis
4573:
4566:
4558:
4551:
4543:
4540:
4539:
4534:
4527:
4526:
4523:
4522:
4493:
4491:
4488:
4487:
4484:
4483:
4480:
4479:
4476:
4475:
4473:
4472:
4465:
4458:
4449:
4447:
4443:
4442:
4440:
4439:
4436:Archaeovaranus
4432:
4425:
4417:
4415:
4409:
4408:
4406:
4405:
4397:
4395:
4386:
4380:
4379:
4377:
4376:
4369:
4362:
4355:
4348:
4341:
4334:
4327:
4320:
4313:
4305:
4303:
4296:
4290:
4289:
4287:
4286:
4279:
4272:
4269:Dalinghosaurus
4265:
4257:
4255:
4248:
4247:
4245:
4244:
4237:
4230:
4223:
4216:
4209:
4202:
4195:
4188:
4181:
4173:
4171:
4164:
4158:
4157:
4154:
4153:
4150:
4149:
4147:
4146:
4139:
4132:
4125:
4118:
4111:
4104:
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4083:
4076:
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4050:Glyptosaurinae
4043:
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4032:
4024:
4022:
4016:
4015:
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4012:
4009:Eodiploglossus
4005:
4002:Apodosauriscus
3997:
3995:
3993:Diploglossinae
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3857:Neoanguimorpha
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3808:
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3792:
3789:Paramacellodus
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3705:Paramacellodus
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3480:Palaeoxantusia
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3383:
3376:
3369:
3362:
3355:
3348:
3341:
3334:
3327:
3320:
3317:Dracaenosaurus
3313:
3310:Dormaalisaurus
3306:
3298:
3290:
3288:
3282:
3281:
3279:
3278:
3271:
3264:
3256:
3254:
3245:
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3238:
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3229:
3227:
3218:
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3211:Spathorhynchus
3207:
3200:
3197:Plesiorhineura
3193:
3186:
3179:
3172:
3165:
3158:
3151:
3148:Archaerhineura
3143:
3141:
3135:
3134:
3132:
3131:
3123:
3121:
3112:
3111:
3109:
3108:
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3077:
3075:
3074:
3066:
3064:
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3047:
3039:
3037:
3035:Amphisbaenidae
3031:
3030:
3028:
3027:
3020:
3012:
3006:
3000:
2999:
2997:
2996:
2987:
2980:
2977:Paracontogenys
2973:
2970:Cnodontosaurus
2965:
2959:
2953:
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2949:
2942:
2939:Ornatocephalus
2935:
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2898:
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2875:Myrmecodaptria
2871:
2864:
2857:
2848:
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2833:
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2824:
2817:
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2803:
2795:Autarchoglossa
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2497:
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2487:Erdenetesaurus
2483:
2476:
2469:
2462:
2454:
2452:
2444:
2443:
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2440:
2433:
2426:
2418:
2416:
2409:Gilmoreteiidae
2405:
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2394:
2387:
2380:
2373:
2370:Meniscognathus
2366:
2359:
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2321:
2311:
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2307:Borioteiioidea
2298:
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2207:
2204:Ergiliinsaurus
2200:
2193:
2185:
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2181:Arretosauridae
2174:
2173:
2171:
2170:
2163:
2155:
2147:
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2039:
2032:
2025:
2018:
2011:
2008:Isodontosaurus
2004:
1997:
1990:
1987:Anchaurosaurus
1982:
1980:
1971:
1970:
1968:
1967:
1960:
1957:Babibasiliscus
1953:
1944:
1942:
1940:Corytophanidae
1936:
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1706:March 20, 2007
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1678:External links
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1528:Susan E. Evans
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608:
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4700:
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4659:Oculudentavis
4656:
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4641:
4638:
4631:
4627:
4616:
4615:
4610:
4608:
4607:
4606:Tetrapodophis
4603:
4601:
4600:
4599:Portugalophis
4596:
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4469:Dryadissector
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4410:
4404:
4403:
4399:
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4393:Lanthanotidae
4390:
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4381:
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4300:
4297:
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4291:
4285:
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4280:
4278:
4277:
4276:Merkurosaurus
4273:
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4266:
4264:
4263:
4259:
4258:
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4253:
4249:
4243:
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4231:
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4227:Proplatynotia
4224:
4222:
4221:
4217:
4215:
4214:
4210:
4208:
4207:
4206:Palaeovaranus
4203:
4201:
4200:
4196:
4194:
4193:
4192:Dorsetisaurus
4189:
4187:
4186:
4182:
4180:
4179:
4175:
4174:
4172:
4168:
4165:
4163:
4159:
4145:
4144:
4140:
4138:
4137:
4133:
4131:
4130:
4126:
4124:
4123:
4119:
4117:
4116:
4112:
4110:
4109:
4105:
4103:
4102:
4098:
4096:
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4087:Helodermoides
4084:
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4070:
4068:
4067:
4063:
4061:
4060:
4056:
4055:
4053:
4051:
4044:
4038:
4037:
4036:Parophisaurus
4033:
4031:
4030:
4026:
4025:
4023:
4021:
4020:Gerrhonotinae
4017:
4011:
4010:
4006:
4004:
4003:
3999:
3998:
3996:
3994:
3990:
3984:
3983:
3979:
3977:
3976:
3975:Ophisauriscus
3972:
3970:
3969:
3968:Helvetisaurus
3965:
3963:
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3958:
3957:
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3899:Entomophontes
3896:
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3830:
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3819:Mosasauroidea
3813:
3806:
3805:
3804:Yabeinosaurus
3801:
3798:
3797:
3793:
3791:
3790:
3786:
3784:
3783:
3779:
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3776:
3775:Chometokadmon
3772:
3771:
3769:
3765:
3761:
3757:
3752:
3748:
3730:
3729:
3728:Sharovisaurus
3725:
3722:
3721:
3717:
3714:
3713:
3712:Parasaurillus
3709:
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3634:
3633:
3629:
3627:
3626:
3622:
3620:
3619:
3615:
3613:
3612:
3611:Kleskunsaurus
3608:
3606:
3605:
3601:
3599:
3598:
3597:Chromatogenys
3594:
3592:
3591:
3587:
3585:
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3510:
3509:
3505:
3504:
3501:
3498:
3496:
3492:
3482:
3481:
3477:
3475:
3474:
3473:Palepidophyma
3470:
3468:
3467:
3463:
3462:
3460:
3458:
3454:
3448:
3447:
3443:
3441:
3440:
3436:
3435:
3433:
3431:
3430:Barbatteiidae
3424:
3418:
3417:
3413:
3410:
3409:
3405:
3402:
3401:
3397:
3396:
3394:
3392:
3388:
3382:
3381:
3377:
3375:
3374:
3373:Pseudolacerta
3370:
3368:
3367:
3363:
3361:
3360:
3359:Plesiolacerta
3356:
3354:
3353:
3349:
3347:
3346:
3342:
3340:
3339:
3335:
3333:
3332:
3328:
3326:
3325:
3321:
3319:
3318:
3314:
3312:
3311:
3307:
3304:
3303:
3299:
3297:
3296:
3295:Amblyolacerta
3292:
3291:
3289:
3287:
3283:
3277:
3276:
3272:
3270:
3269:
3265:
3263:
3262:
3258:
3257:
3255:
3253:
3252:Contogeniidae
3246:
3236:
3235:
3231:
3230:
3228:
3226:
3219:
3213:
3212:
3208:
3206:
3205:
3204:Protorhineura
3201:
3199:
3198:
3194:
3192:
3191:
3187:
3185:
3184:
3183:Macrorhineura
3180:
3178:
3177:
3173:
3171:
3170:
3166:
3164:
3163:
3162:Hadrorhineura
3159:
3157:
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3152:
3150:
3149:
3145:
3144:
3142:
3140:
3136:
3130:
3129:
3125:
3124:
3122:
3120:
3119:Chthonophidae
3113:
3107:
3106:
3102:
3100:
3099:
3095:
3093:
3092:
3088:
3087:
3085:
3083:
3079:
3073:
3072:
3068:
3067:
3065:
3063:
3059:
3053:
3052:
3048:
3046:
3045:
3041:
3040:
3038:
3036:
3032:
3026:
3025:
3021:
3019:
3018:
3014:
3013:
3010:
3007:
3005:
3001:
2994:
2988:
2986:
2985:
2981:
2979:
2978:
2974:
2972:
2971:
2967:
2966:
2963:
2960:
2958:
2954:
2948:
2947:
2943:
2941:
2940:
2936:
2934:
2933:
2929:
2928:
2926:
2924:
2920:
2916:
2912:
2907:
2903:
2877:
2876:
2872:
2870:
2869:
2865:
2863:
2862:
2858:
2855:
2854:
2850:
2849:
2847:
2844:
2837:
2830:
2829:
2825:
2823:
2822:
2818:
2816:
2815:
2811:
2810:
2807:
2804:
2801:
2796:
2791:
2781:
2780:
2779:Yanatarogecko
2776:
2774:
2773:
2769:
2767:
2766:
2762:
2760:
2759:
2758:Cretaceogekko
2755:
2753:
2752:
2748:
2747:
2745:
2743:
2739:
2733:
2732:
2728:
2727:
2724:
2721:
2719:
2718:Gekkonomorpha
2715:
2712:
2710:
2706:
2699:
2698:
2697:Yabeinosaurus
2694:
2691:
2690:
2686:
2684:
2683:
2682:Pedrerasaurus
2679:
2676:
2675:
2671:
2669:
2668:
2664:
2661:
2660:
2656:
2653:
2652:
2648:
2645:
2644:
2640:
2639:
2636:
2633:
2631:
2627:
2623:
2619:
2613:
2609:
2595:
2594:
2593:Tripennaculus
2590:
2587:
2586:
2582:
2580:
2579:
2575:
2573:
2572:
2571:Tchingisaurus
2568:
2566:
2565:
2561:
2558:
2557:
2553:
2550:
2549:
2548:Olindalacerta
2545:
2543:
2542:
2538:
2536:
2535:
2531:
2529:
2528:
2524:
2522:
2521:
2517:
2516:
2514:
2510:
2503:
2502:
2498:
2496:
2495:
2491:
2489:
2488:
2484:
2482:
2481:
2477:
2475:
2474:
2473:Darchansaurus
2470:
2468:
2467:
2466:Cherminsaurus
2463:
2461:
2460:
2456:
2455:
2453:
2450:
2445:
2439:
2438:
2434:
2432:
2431:
2427:
2425:
2424:
2420:
2419:
2417:
2414:
2410:
2406:
2400:
2399:
2398:Trippenaculus
2395:
2393:
2392:
2388:
2386:
2385:
2381:
2379:
2378:
2374:
2372:
2371:
2367:
2365:
2364:
2360:
2358:
2357:
2353:
2351:
2350:
2346:
2344:
2343:
2339:
2337:
2336:
2332:
2330:
2329:
2325:
2324:
2322:
2320:
2316:
2312:
2308:
2304:
2296:
2292:
2283:
2265:
2264:
2260:
2257:
2256:
2252:
2250:
2249:
2248:Phrynosomimus
2245:
2243:
2242:
2238:
2236:
2235:
2231:
2230:
2228:
2226:
2225:Priscagamidae
2219:
2213:
2212:
2211:Khaichinguana
2208:
2206:
2205:
2201:
2199:
2198:
2194:
2192:
2191:
2187:
2186:
2184:
2182:
2175:
2169:
2168:
2164:
2161:
2160:
2156:
2153:
2152:
2148:
2146:
2145:
2141:
2140:
2137:
2134:
2130:
2120:
2119:
2115:
2114:
2112:
2110:
2106:
2100:
2099:
2095:
2093:
2092:
2088:
2086:
2085:
2081:
2079:
2078:
2074:
2072:
2071:
2070:Armandisaurus
2067:
2063:
2062:Polychrotidae
2060:
2059:
2058:
2057:
2053:
2052:
2050:
2048:
2044:
2038:
2037:
2033:
2031:
2030:
2026:
2024:
2023:
2022:Saichangurvel
2019:
2017:
2016:
2012:
2010:
2009:
2005:
2003:
2002:
1998:
1996:
1995:
1991:
1989:
1988:
1984:
1983:
1981:
1979:
1972:
1966:
1965:
1961:
1959:
1958:
1954:
1951:
1950:
1946:
1945:
1943:
1941:
1937:
1931:
1930:
1926:
1924:
1923:
1919:
1917:
1916:
1912:
1910:
1909:
1905:
1904:
1901:
1898:
1896:
1892:
1888:
1878:
1877:
1873:
1871:
1870:
1866:
1865:
1863:
1861:
1857:
1851:
1850:
1846:
1844:
1843:
1839:
1837:
1836:
1832:
1830:
1829:
1825:
1823:
1822:
1821:Anqingosaurus
1818:
1817:
1814:
1811:
1807:
1803:
1799:
1794:
1790:
1784:
1780:
1778:
1774:
1772:
1768:
1766:
1762:
1761:
1758:
1753:
1745:
1740:
1738:
1733:
1731:
1726:
1725:
1722:
1715:
1712:by Ker Than,
1711:
1708:
1705:
1701:
1698:
1697:
1693:
1682:
1677:
1663:
1657:
1653:
1649:
1645:
1641:
1634:
1632:
1628:
1623:
1619:
1614:
1609:
1604:
1599:
1595:
1591:
1587:
1583:
1576:
1569:
1567:
1565:
1563:
1559:
1555:
1550:
1548:
1544:
1537:
1535:
1533:
1529:
1519:
1518:
1510:
1509:
1501:
1500:
1492:
1491:
1483:
1482:
1474:
1473:
1465:
1464:
1456:
1455:
1447:
1446:
1438:
1437:
1429:
1428:
1420:
1419:
1411:
1410:
1402:
1401:
1393:
1392:
1384:
1383:
1375:
1374:
1371:
1370:
1364:
1363:
1360:
1359:
1356:
1355:
1354:
1347:
1346:
1343:
1342:
1339:
1338:
1337:
1330:
1329:
1323:
1322:
1319:
1318:
1315:
1314:
1313:
1306:
1305:
1299:
1298:
1295:
1294:
1291:
1290:
1284:
1283:
1277:
1276:
1273:
1272:
1264:
1263:
1260:
1259:
1253:
1252:
1249:
1248:
1245:
1244:
1238:
1237:
1231:
1230:
1224:
1223:
1220:
1219:
1211:
1210:
1207:
1206:
1205:
1198:
1197:
1194:
1193:
1190:
1189:
1188:
1181:
1180:
1174:
1173:
1167:
1166:
1163:
1162:
1159:
1158:
1157:
1150:
1149:
1143:
1142:
1139:
1138:
1130:
1129:
1126:
1125:
1124:
1117:
1116:
1113:
1112:
1109:
1108:
1107:
1100:
1099:
1093:
1092:
1086:
1085:
1082:
1081:
1078:
1077:
1071:
1070:
1064:
1063:
1060:
1059:
1051:
1050:
1047:
1046:
1040:
1039:
1036:
1035:
1032:
1031:
1025:
1024:
1018:
1017:
1011:
1010:
1007:
1006:
998:
997:
989:
988:
985:
984:
978:
977:
974:
973:
970:
969:
963:
962:
956:
955:
952:
951:
948:
947:
941:
940:
934:
933:
927:
926:
923:
922:
914:
913:
905:
904:
896:
895:
887:
886:
883:
882:
876:
875:
872:
871:
868:
867:
861:
860:
854:
853:
850:
849:
846:
844:
840:
834:
833:
827:
826:
823:
822:
819:
818:
817:Eublepharinae
812:
811:
805:
804:
801:
800:
792:
791:
788:
787:
781:
780:
777:
776:
773:
772:
766:
765:
759:
758:
752:
751:
745:
744:
741:
740:
732:
731:
723:
722:
719:
718:
717:
710:
709:
706:
705:
702:
701:
700:
693:
692:
686:
685:
682:
681:
678:
677:
676:
669:
668:
662:
661:
655:
654:
651:
650:
642:
641:
633:
632:
629:
628:
627:Leiolepidinae
622:
621:
618:
617:
614:
613:
607:
606:
603:
602:
599:
598:
592:
591:
588:
587:
584:
583:
582:
575:
574:
568:
567:
564:
563:
560:
559:
553:
552:
546:
545:
539:
538:
535:
534:
531:
530:
524:
523:
517:
516:
513:
512:
509:
508:
507:
500:
499:
493:
492:
489:
488:
485:
484:
478:
477:
471:
470:
466:
464:
460:
459:Leiolepidinae
456:
452:
448:
444:
436:
434:
432:
428:
424:
420:
418:
414:
413:
412:Kuehneosaurus
408:
404:
400:
399:
394:
393:
388:
384:
377:
375:
373:
369:
365:
361:
357:
353:
349:
341:
339:
337:
336:leiolepidines
333:
332:chamaeleonids
329:
325:
321:
317:
313:
309:
305:
302:
298:
294:
290:
286:
282:
278:
274:
271:
267:
263:
259:
258:
245:
241:
236:
233:
232:
230:
227:
223:
218:
212:
211:
204:
201:
200:
197:
194:
191:
190:
187:
184:
181:
180:
177:
174:
171:
170:
167:
164:
161:
160:
157:
154:
151:
150:
145:
140:
136:
133:
128:
124:
119:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
39:
33:
29:
26:
22:
4808:Yixian fauna
4696:
4665:
4657:
4650:
4643:
4612:
4604:
4597:
4590:
4582:
4575:
4568:
4560:
4553:
4546:
4499:
4467:
4460:
4454:Arcanosaurus
4452:
4434:
4427:
4420:
4400:
4371:
4364:
4359:Palaeosaniwa
4357:
4352:Morohasaurus
4350:
4343:
4336:
4331:Eurheloderma
4329:
4322:
4315:
4308:
4294:Goannasauria
4281:
4274:
4267:
4260:
4239:
4232:
4225:
4218:
4211:
4204:
4197:
4190:
4183:
4176:
4141:
4134:
4127:
4120:
4113:
4106:
4099:
4092:
4085:
4080:Glyptosaurus
4078:
4071:
4066:Melanosaurus
4064:
4059:Arpadosaurus
4057:
4034:
4027:
4007:
4000:
3980:
3973:
3966:
3961:Headonhillia
3959:
3939:
3911:
3904:
3897:
3890:Xenosauridae
3877:
3843:Mosasauridae
3802:
3794:
3787:
3780:
3773:
3726:
3718:
3710:
3703:
3696:
3689:
3681:
3658:
3651:
3644:
3637:
3630:
3625:Monocnemodon
3623:
3618:Lonchisaurus
3616:
3609:
3602:
3595:
3588:
3581:
3556:Pyrenasaurus
3554:
3549:Orthoscincus
3547:
3542:Axonoscincus
3540:
3520:
3513:
3508:Retinosaurus
3506:
3478:
3471:
3466:Catactegenys
3464:
3444:
3437:
3414:
3406:
3398:
3378:
3371:
3366:Pseudeumeces
3364:
3357:
3352:Mediolacerta
3350:
3343:
3338:Ligerosaurus
3336:
3331:Gracilicerta
3329:
3322:
3315:
3308:
3300:
3293:
3273:
3266:
3259:
3232:
3209:
3202:
3195:
3188:
3181:
3174:
3167:
3160:
3155:Dyticonastis
3153:
3146:
3126:
3103:
3098:Cuvieribaena
3096:
3089:
3069:
3051:Lophocranion
3049:
3044:Listromycter
3042:
3022:
3015:
3004:Amphisbaenia
2982:
2975:
2968:
2946:Sakurasaurus
2944:
2937:
2930:
2911:Scincomorpha
2873:
2866:
2859:
2851:
2826:
2821:Tepexisaurus
2819:
2814:Parmeosaurus
2812:
2777:
2770:
2763:
2756:
2751:Cadurcogekko
2749:
2729:
2709:Scleroglossa
2695:
2687:
2680:
2674:Liushusaurus
2672:
2665:
2657:
2651:Bavarisaurus
2649:
2641:
2591:
2583:
2578:Tianyusaurus
2576:
2569:
2562:
2554:
2546:
2539:
2532:
2525:
2518:
2499:
2492:
2485:
2478:
2471:
2464:
2457:
2437:Yechilacerta
2435:
2428:
2423:Gilmoreteius
2421:
2396:
2389:
2382:
2377:Pelsochamops
2375:
2368:
2363:Leptochamops
2361:
2356:Haptosphenus
2354:
2347:
2340:
2333:
2326:
2319:Chamopsiidae
2261:
2253:
2246:
2239:
2232:
2209:
2202:
2195:
2190:Arretosaurus
2188:
2165:
2157:
2149:
2144:Brasiliguana
2142:
2118:Desertiguana
2116:
2096:
2089:
2082:
2075:
2068:
2054:
2034:
2027:
2020:
2013:
2006:
1999:
1992:
1985:
1962:
1955:
1947:
1927:
1920:
1913:
1906:
1874:
1867:
1848:
1847:
1840:
1833:
1826:
1819:
1783:Lepidosauria
1781:Superorder:
1702:Aalok Mehta
1665:, retrieved
1643:
1585:
1581:
1525:
1367:
1351:
1350:
1336:Pachyrhachis
1334:
1333:
1310:
1309:
1287:
1258:Mosasauridae
1256:
1241:
1202:
1201:
1185:
1184:
1154:
1153:
1121:
1120:
1104:
1103:
1074:
1043:
1028:
981:
966:
944:
879:
864:
842:
837:
815:
786:Amphisbaenia
784:
769:
714:
713:
697:
696:
673:
672:
625:
610:
595:
580:
579:
578:
556:
527:
504:
503:
481:
462:
442:
440:
430:
426:
421:
416:
410:
401:, Latin for
396:
390:
382:
381:
367:
351:
350:specimen of
345:
307:
256:
255:
254:
239:
234:
216:
209:
208:
131:
31:
25:
4645:Costasaurus
4592:Parviraptor
4402:Cherminotus
4310:Asprosaurus
4262:Bahndwivici
4252:Shinisauria
4241:Telmasaurus
4178:Aiolosaurus
4122:Placosaurus
4115:Peltosaurus
4094:Odaxosaurus
3941:Odaxosaurus
3866:Carusioidea
3796:Paravaranus
3756:Anguimorpha
3683:Atokasaurus
3639:Saurillodon
3515:Scincoideus
3457:Xantusiidae
3446:Oardasaurus
3439:Barbatteius
3400:Brevisaurus
3380:Quercycerta
3324:Escampcerta
3139:Rhineuridae
3128:Chthonophis
3091:Blanosaurus
3024:Todrasaurus
2957:Lacertoidea
2923:Cordyloidea
2772:Hoburogekko
2643:Ardeosaurus
2556:Paleochelco
2527:Funiusaurus
2520:Adamisaurus
2480:Distortodon
2430:Kuwajimalla
2384:Socognathus
2342:Glyptogenys
2335:Gerontoseps
2241:Mimeosaurus
2197:Dornosaurus
2151:Hoyalacerta
2091:Pristiguana
2077:Lapitiguana
2056:Afairiguana
2036:Zapsosaurus
1994:Ctenomastax
1922:Uquiasaurus
1915:Suzanniwana
1908:Anolbanolis
1891:Pleurodonta
1835:Bharatagama
1714:LiveScience
1312:Adriosaurus
1187:Lanthanotus
1123:Shinisaurus
881:Pygopodidae
342:Description
4782:Categories
4652:Lamiasaura
4555:Diablophis
4462:Cemeterius
4384:Varanoidea
4373:Primaderma
4345:Lowesaurus
4317:Chianghsia
4220:Parviderma
4213:Parasaniwa
4136:Proxestops
3982:Ragesaurus
3691:Becklesius
3660:Webbsaurus
3632:Pachygenys
3604:Dakotaseps
3590:Calanguban
3583:Apsgnathus
3495:Scincoidea
3416:Ptilotodon
3408:Meyasaurus
3286:Lacertidae
3261:Contogenys
3176:Jepsibaena
2984:Purbicella
2843:Bainguidae
2800:Evansauria
2689:Scandensia
2667:Jucaraseps
2459:Bicuspidon
2263:Priscagama
1978:Gobiguania
1895:Iguanoidea
1876:Suratagama
1828:Barbaturex
1667:2022-11-06
1353:Haasiophis
1106:Xenosaurus
1030:Cordylidae
946:Lacertidae
843:sensu lato
839:Gekkoninae
506:Marmoretta
360:Metacarpal
356:unossified
270:Cretaceous
4697:Xianglong
4614:Serpentes
4562:Dinilysia
4548:Coniophis
4413:Varanidae
4366:Paraderma
4338:Gobiderma
4234:Saniwides
3906:Exostinus
3782:Eolacerta
3653:Sciroseps
3646:Saurillus
3533:Scincidae
3275:Utahgenys
3190:Ototriton
3169:Hyporhina
3062:Bipedidae
2932:Eocordyla
2731:Norellius
2564:Peneteius
2534:Gobinatus
2084:Pariguana
2047:Iguanidae
2029:Temujinia
2015:Polrussia
1869:Indiagama
1849:Xianglong
1842:Gueragama
1809:Acrodonta
1763:Kingdom:
1538:Citations
1369:Serpentes
1156:Heloderma
1045:Scincidae
771:Dibamidae
581:Xianglong
558:Iguanidae
449:with the
443:Xianglong
427:Xianglong
417:Xianglong
383:Xianglong
368:Xianglong
352:Xianglong
324:iguanians
320:cladistic
308:Xianglong
297:Barremian
257:Xianglong
210:Xianglong
162:Kingdom:
156:Eukaryota
44:Barremian
32:Xianglong
4706:Wikidata
4617:(snakes)
4199:Eosaniwa
3952:Anguinae
3928:Anguidae
3082:Blanidae
2868:Globaura
2853:Bainguis
2828:Tijubina
2585:Tijubina
2541:Obamadon
1949:Aciprion
1860:Agamidae
1777:Reptilia
1771:Chordata
1769:Phylum:
1765:Animalia
1752:squamate
1750:Extinct
1622:17376871
1532:acrodont
1076:Anguidae
597:Agaminae
465:(2007):
451:Agaminae
447:polytomy
437:Taxonomy
407:Triassic
372:arboreal
348:holotype
328:agamines
326:such as
316:acrodont
242:, 2007 (
235:X. zhaoi
196:Squamata
186:Reptilia
176:Chordata
172:Phylum:
166:Animalia
152:Domain:
4752:6158148
4739:1479311
4726:8524503
4712:Q148080
4536:Ophidia
4324:Estesia
4143:Xestops
4073:Gaultia
3879:Carusia
3522:Slavoia
3391:Teiidae
2861:Eoxanta
2765:Gobekko
2742:Gekkota
2328:Chamops
2098:Pumilia
1929:Xihaina
1798:Iguania
1775:Class:
1613:1838464
1590:Bibcode
1204:Varanus
968:Teiidae
423:Xu Xing
264:) is a
262:Chinese
226:Species
202:Genus:
192:Order:
182:Class:
110:↓
4765:197005
4584:Najash
4570:Eophis
4429:Saniwa
3913:Restes
1754:genera
1658:
1620:
1610:
405:) and
403:dragon
334:, and
281:Yixian
277:Yizhou
273:lizard
240:et al.
219:, 2007
217:et al.
4734:IRMNG
2793:Stem
2616:Stem
1578:(PDF)
463:et al
431:Draco
398:Draco
364:digit
312:glide
299:-age
289:China
266:genus
4721:GBIF
4611:See
4422:Ovoo
2001:Igua
1656:ISBN
1618:PMID
457:and
346:The
244:type
49:Preκ
2411:/ "
1648:doi
1608:PMC
1598:doi
1586:104
293:LPM
287:of
268:of
238:Li
215:Li
4784::
4762::
4749::
4736::
4723::
4708::
2305:/
1893:/
1654:,
1642:,
1630:^
1616:.
1606:.
1596:.
1584:.
1580:.
1561:^
1546:^
845:)
453:,
433:.
330:,
283:,
279:,
99:Pg
40:,
4662:?
4587:?
4565:?
4457:?
4254:?
4047:β
3840:β
3832:β
3821:?
3816:β
3807:?
3799:?
3723:?
3715:?
3686:?
3671:β
3559:?
3427:β
3411:?
3403:?
3305:?
3249:β
3222:β
3116:β
2995:?
2990:β
2856:?
2845:?
2840:β
2831:?
2802:)
2798:(
2700:?
2692:?
2677:?
2662:?
2654:?
2646:?
2588:?
2559:?
2551:?
2504:?
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2447:"
2415:"
2300:β
2258:?
2222:β
2178:β
2162:?
2154:?
1975:β
1952:?
1743:e
1736:t
1729:v
1650::
1624:.
1600::
1592::
841:(
246:)
206:β
104:N
94:K
89:J
84:T
79:P
74:C
69:D
64:S
59:O
54:κ
23:.
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