463:
566:, given the rather cold climate of the Yixian with an average annual temperature of 10 °C (50 °F). Alternatively, if they were restricted to the regions in which they were found, they may have served as display structures. In addition, the two adult specimens had distinctive, "wavy" crests on their snouts, on both sides of a high central crest, which were probably used for display. The presence of feathers on a large basal tyrannosauroid suggests the possibility that later tyrannosaurids were also feathered, even when adult, despite their size. However, scaly skin impressions have been reported from various Late Cretaceous tyrannosaurids (such as
3683:
5815:
3676:
118:
2180:
3005:
2991:
5267:
4471:
5253:
2984:
5260:
2186:
3690:
136:
921:
2998:
5828:
4489:
555:
4477:
531:
covered various parts of the body. With the holotype they were present on the pelvis and near the foot. Specimen ZCDM V5000 had feathers on the tail pointing backwards under an angle of 30 degrees with the tail axis. The smallest specimen showed 20-centimetre-long (7.9 in) filaments on the neck and 16-centimetre-long (6.3 in) feathers at the upper arm. While it has been known since 2004, upon the description of
4496:
5822:
368:
5247:
858:
667:
519:
4484:
588:
was feathered. Since there is no positive evidence for plumage in tyrannosaurids, some researchers have suggested they may have evolved scales secondarily. If scaly skin was the dominant epidermal trait of later genera, then the extent and nature of the integumentary covering may have changed over
530:
contain direct evidence of feathers in the form of fossil imprints. The feathers were long, up to 20 centimetres (7.9 in), and filamentous. Because the quality of the preservation was low, it could not be established whether the filaments were simple or compound, broad or narrow. The feathers
485:
gave lower estimations of 7.5 metres (24.6 ft) and 1.1 tonnes (1.2 short tons). Its skull has an estimated length of 905 millimetres (35.6 in). The skulls of the paratypes are 80 centimetres (31 in) and 63 centimetres (25 in) long and their weights have been estimated at 596
639:. Primitive traits relative to advanced tyrannosaurs included long forelimbs with three fingers and a short foot which was not specialized for running. Advanced traits included a large and deep skull, the outer side of the premaxilla having rotated upwards, a large cuneiform horn on the
596:
might not represent true feathers and represent filamentous structures, being ancestral state to feathers. However, this hypothesis is now considered overturned with subsequent research that verified that the structures on various dinosaurs and pterosaurs were true feathers.
441:
are the two other specimens: ZCDM V5001 consisting of a skeleton of a smaller individual and part of the same slab as the holotype; and ELDM V1001, a juvenile estimated to have been eight years younger than the holotype. The fossils are part of the collections of the
505:
has a small secondary process, jutting into the upper hind corner of the eye socket. The outer side of the main body of the postorbital is hollowed out. In the lower jaw, the external mandibular fenestra, the main opening in the outer side, is mainly located in the
1008:
would also have been prevalent. Volcanic eruptions and forest fires appear to have been common in the Yixian, and the environment would have been littered with bodies of water and coniferous plants. The environment would have been comparable to the modern day
957:
did prey on sauropods, it would have been one of two predatory animals known from the Yixian formation capable of doing so, the other being an as-of-yet undescribed large theropod known from a tooth embedded in the rib of a
418:
is known from three nearly complete fossil specimens (an adult, a subadult and a juvenile) acquired from a fossil dealer who claimed all three had their provenance in a single quarry at Batu Yingzi in
1542:
Xing L., Bell, P.R., Currie, P.J., Shibata M., Tseng K. & Dong Z. (2012). "A sauropod rib with an embedded theropod tooth: direct evidence for feeding behaviour in the Jehol group, China."
451:
1169:
1143:
869:, or change during growth, of this species. During growth the lower legs, feet, ilia and forelimbs became relatively smaller. The skull, on the other hand, grew more robust and deeper.
481:
predator. The holotype and oldest-known specimen has an estimated length of 9 metres (29.5 ft) and an estimated weight of about 1,414 kilograms (3,117 lb). In 2016,
1251:
1285:
674:
In 2016, a phylogenetic analysis conducted by Thomas Carr and
Stephen Brusatte re-examined the evolutionary relationships of the Tyrannosauroidea. Their analysis found
5961:
5907:
1314:
949:
may have hunted sauropods, and that the three known individuals may have died while doing so. In addition, other sauropod hunting theropods such as
1179:
430:, approximately 125 million years old. The specimens had been cut into pieces about the size of bath mats, which could be carried by two people.
3682:
1135:
5971:
3675:
1336:"Adaptation to the sky: Defining the feather with integument fossils from Mesozoic China and experimental evidence from molecular laboratories"
941:
were allegedly found together, some paleontologists, including Xu Xing, have interpreted the animal as a pack hunter. Based on the presence of
5814:
437:, ZCDM V5000, is the largest specimen, consisting of a nearly complete skeleton with skull, compressed on a slab, of an adult individual. The
1213:
1334:
Chuong, Cheng-Ming; Wu, Ping; Zhang, Fu-Cheng; Xu, Xing; Yu, Minke; Widelitz, Randall B.; Jiang, Ting-Xin; Hou, Lianhai (15 August 2003).
545:
is currently the largest-known species of dinosaur with direct evidence of feathers, forty times heavier than the previous record holder,
1642:
1070:
978:
originating from the
Lujiatun or the Jianshangou beds of the Yixian, meaning it would have been contemporaneous of such dinosaurs as
3004:
4470:
1022:
2990:
1241:
5252:
1277:
2179:
5981:
2983:
5266:
2185:
5693:
4488:
5966:
3689:
5259:
2997:
1394:
537:, that at least some tyrannosauroids possessed filamentous "stage 1" feathers, according to the feather typology of
135:
5827:
4495:
5976:
447:
953:
are known to have exhibited pack hunting behaviour. The true cause of their death, however, remains unknown. If
1730:
379:
296:
4476:
1069:
Xu, X.; Wang, K.; Zhang, K.; Ma, Q.; Xing, L.; Sullivan, C.; Hu, D.; Cheng, S.; Wang, S.; et al. (2012).
443:
117:
5821:
1635:
865:
The knowledge of specimens representing various different ages has allowed paleontologists to determine the
401:
5847:
1515:
652:
617:. An initial analysis of its relationship to other tyrannosauroids showed that it was more primitive than
1307:
5986:
5933:
4201:
4113:
493:, in which it differs from its direct relatives. The snout features a high midline crest, formed by the
277:
5246:
5894:
5779:
3890:
3858:
1438:
1085:
1010:
1520:
4214:
4001:
1246:
1174:
4483:
5686:
4190:
4050:
3906:
3846:
3281:
3191:
3074:
1938:
1628:
1587:
1109:
648:
130:
5938:
945:
material in the quarry in which the three specimens were found, Xu has further speculated that
355:
Province — are currently the largest-known dinosaur specimens that preserve direct evidence of
5920:
5568:
5485:
5459:
5298:
5285:
5163:
4704:
4656:
3488:
2838:
1917:
1525:
1488:
1466:
1373:
1355:
1219:
1209:
1101:
920:
693:
683:
562:
Based on the distribution of the feathers, they may have covered the whole body and served in
319:
232:
5925:
1013:
of
British Columbia, and would have experienced significant seasonal changes in temperature.
5672:
5429:
5339:
5128:
5044:
4906:
4825:
4797:
4757:
4638:
4608:
4126:
3879:
3743:
3600:
3500:
3349:
3256:
3214:
2781:
2091:
1577:
1547:
1456:
1446:
1409:
1363:
1347:
1093:
900:, it was determined that all archosaurs would have had fixed tongues, with the exception of
614:
563:
533:
502:
423:
397:
386:
322:
219:
30:
5885:
1117:
5735:
5473:
5436:
5368:
5223:
5216:
5028:
4970:
4943:
4933:
4892:
4727:
4586:
4570:
4388:
4360:
4338:
4225:
4070:
3777:
3551:
3434:
3412:
3333:
3162:
3139:
3132:
3110:
3096:
3089:
3050:
2901:
2748:
2675:
2664:
2600:
2488:
2375:
1952:
1861:
1715:
1690:
1608:
1504:"A new carcharodontosaurid (Dinosauria, Theropoda) from the Upper Cretaceous of Argentina"
1398:
1318:
992:
731:
554:
482:
408:(华丽 simplified, 華麗 traditional, "beautiful"), in reference to the perceived beauty of the
643:
in front of the eye socket, a postorbital process on the back rim of the eye socket, the
1442:
1089:
5700:
5663:
5644:
5606:
5595:
5392:
5105:
5064:
5055:
5037:
5017:
4913:
4627:
4372:
4234:
4061:
4043:
3871:
3838:
3824:
3785:
3559:
3420:
3377:
3370:
3292:
3268:
3221:
3180:
3117:
3103:
2831:
2816:
2643:
2477:
2341:
2309:
2298:
2275:
2237:
2098:
2068:
1992:
1931:
1830:
1461:
1426:
1368:
1335:
980:
644:
1566:"Evolutionary radiation of the Jehol Biota: chronological and ecological perspectives"
974:
is uncertain, it is unknown what fauna it coexisted with. Age estimates point towards
5955:
5742:
5588:
5549:
5528:
5407:
5400:
5327:
5121:
5094:
4952:
4868:
4846:
4839:
4832:
4775:
4692:
4555:
4527:
4514:
4438:
4315:
4155:
3989:
3920:
3806:
3793:
3538:
3516:
3509:
3473:
3446:
3398:
3322:
3058:
2849:
2823:
2739:
2720:
2713:
2650:
2629:
2608:
2593:
2579:
2538:
2504:
2497:
2416:
2268:
2155:
2148:
2075:
2045:
2034:
1924:
1895:
1888:
1814:
1551:
1503:
640:
606:
580:
547:
396:(τύραννος, "tyrant"), a reference to its classification as a feathered member of the
68:
1591:
1113:
5714:
5652:
5622:
5542:
5535:
5513:
5505:
5385:
5377:
5357:
5072:
4983:
4921:
4811:
4782:
4734:
4563:
4548:
4323:
4255:
4183:
4162:
4141:
3957:
3913:
3721:
3708:
3638:
3591:
3466:
3356:
3308:
3301:
3238:
3207:
3200:
3081:
2908:
2760:
2657:
2559:
2552:
2447:
2431:
2367:
2318:
2250:
2027:
1910:
1903:
1874:
1837:
1806:
1427:"Convergent evolution of a mobile bony tongue in flighted dinosaurs and pterosaurs"
1237:
986:
930:
909:
796:
656:
635:
538:
367:
659:
with a straight upper rim and an appending lobe, a large pubic foot and a slender
1451:
5879:
5768:
5749:
5679:
5629:
5615:
5557:
5498:
5443:
5320:
5114:
4990:
4959:
4875:
4861:
4593:
4541:
4409:
4331:
4294:
4176:
4148:
4101:
4021:
3899:
3756:
3607:
3580:
3480:
3427:
3363:
3342:
3315:
3066:
2879:
2872:
2858:
2684:
2531:
2522:
2424:
2395:
2325:
2290:
2283:
2083:
2061:
2054:
2013:
2005:
1981:
1966:
1945:
1853:
1845:
1165:
998:
574:
568:
43:
857:
666:
518:
462:
5707:
5636:
5581:
5202:
5001:
4818:
4804:
4789:
4675:
4601:
4423:
4416:
4402:
4308:
4241:
4169:
4133:
4086:
4028:
4014:
3981:
3973:
3966:
3947:
3770:
3735:
3523:
3124:
2915:
2865:
2802:
2773:
2728:
2706:
2699:
2691:
2615:
2511:
2470:
2463:
2455:
2439:
2359:
2230:
2106:
1959:
1822:
1681:
894:
882:
619:
507:
498:
494:
478:
336:
193:
88:
53:
5870:
1529:
1359:
1223:
5728:
5721:
5421:
5414:
5087:
5079:
5010:
4663:
4381:
3927:
3831:
3566:
3151:
3036:
3023:
2950:
2886:
2809:
2788:
2545:
2408:
2216:
2203:
2121:
1973:
1792:
1779:
1742:
1651:
1004:
942:
905:
897:
890:
206:
147:
93:
37:
1470:
1391:
1377:
1105:
486:
kilograms (1,314 lb) and 493 kilograms (1,087 lb), respectively.
5864:
5312:
4720:
4713:
4685:
4578:
4078:
3459:
2795:
2622:
2383:
2020:
1699:
1675:
1615:
866:
755:
707:
629:
623:
in the evolutionary tree, but more advanced than tyrannosauroids such as
438:
434:
419:
352:
180:
167:
83:
78:
63:
58:
48:
1097:
5912:
4853:
4615:
4345:
4248:
3573:
2894:
660:
409:
356:
326:
98:
73:
1413:
5521:
5209:
2636:
1669:
1351:
1203:
886:
427:
344:
157:
5841:
1582:
1565:
1392:
The phylogeny and evolutionary history of tyrannosauroid dinosaurs.
589:
time in response to body size, a warmer climate, or other factors.
4352:
4093:
4035:
1071:"A gigantic feathered dinosaur from the Lower Cretaceous of China"
919:
856:
665:
553:
517:
461:
366:
340:
315:
16:
Genus of proceratosaurid dinosaur from the Early
Cretaceous period
5899:
422:
Province, China. They were thus probably found in a layer of the
5785:
4395:
901:
592:
It was considered possible that the integumentary structures of
5845:
5283:
4512:
3706:
3021:
2201:
1777:
1662:
1624:
1170:"The One-Ton Turkey: Further Adventures in Slow-Cooked Science"
1620:
452:
Institute of
Vertebrate Paleontology and Paleoanthropology
501:
and which is covered by large pneumatic recesses. The
1425:
Li, Zhiheng; Zhou, Zhonghe; Clarke, Julia A. (2018).
1242:"There are giant feathered tyrannosaurs now...right?"
893:. Based on hyoid bone comparisons between living and
489:
The describers established some diagnostic traits of
5854:
5767:
5662:
5605:
5567:
5484:
5457:
5367:
5337:
5296:
5194:
5162:
5104:
5054:
5027:
5000:
4969:
4942:
4931:
4890:
4755:
4703:
4674:
4636:
4625:
4525:
4437:
4371:
4293:
4224:
4200:
4112:
4060:
4000:
3956:
3945:
3889:
3857:
3804:
3754:
3719:
3637:
3590:
3537:
3499:
3445:
3396:
3332:
3291:
3267:
3236:
3190:
3160:
3149:
3034:
2949:
2848:
2759:
2738:
2674:
2577:
2521:
2487:
2394:
2339:
2308:
2248:
2214:
2140:
2120:
2044:
1991:
1872:
1790:
1741:
1714:
378:was named and scientifically described in 2012 by
651:forming a large process on the back rim of the
1482:
1480:
613:relationships have classified it in the group
1636:
1278:"Humongous fuzzy dinosaur unearthed in China"
1064:
1062:
1060:
1058:
125:Restored skeletons mounted in fighting poses
8:
1056:
1054:
1052:
1050:
1048:
1046:
1044:
1042:
1040:
1038:
1271:
1269:
1208:. Princeton University Press. p. 106.
5842:
5573:
5490:
5465:
5349:
5345:
5304:
5293:
5280:
5191:
5159:
4975:
4939:
4898:
4767:
4763:
4648:
4644:
4633:
4533:
4522:
4509:
4300:
4290:
4206:
4118:
4006:
3953:
3863:
3816:
3812:
3762:
3727:
3716:
3703:
3634:
3543:
3451:
3404:
3273:
3248:
3244:
3172:
3168:
3157:
3042:
3031:
3018:
2946:
2765:
2756:
2585:
2400:
2351:
2347:
2260:
2256:
2222:
2211:
2198:
1997:
1880:
1798:
1787:
1774:
1720:
1711:
1659:
1643:
1629:
1621:
1487:Kerri Smith and Adam Levi (5 April 2012).
116:
20:
1612:relative is biggest-ever feathered animal
1581:
1519:
1460:
1450:
1367:
861:Skull of the juvenile specimen ELDM V1001
1136:"Bus-Size Dinosaurs, as Fuzzy as Chicks"
522:Artist's restoration based on ELDM V1001
1034:
937:Because the three known individuals of
325:dinosaur which contains a single known
1205:The Princeton Field Guide to Dinosaurs
335:. This species lived during the early
7:
1502:Coria, R. A.; Currie, P. J. (2006).
1390:Brusatte, S.L. and Carr, T.D. 2016.
885:structure, indicating it had a flat
771:
747:
723:
699:
689:
392:(羽, "feather") and Latinised Greek
339:period in what is now northeastern
5962:Early Cretaceous dinosaurs of Asia
1146:from the original on 24 March 2019
686:. Their cladogram is shown below:
314:(meaning "feathered tyrant") is a
14:
1340:Journal of Experimental Zoology B
1288:from the original on 6 April 2012
1254:from the original on 7 April 2012
558:Tail fragment with feather traces
5826:
5820:
5813:
5265:
5258:
5251:
5245:
4494:
4487:
4482:
4475:
4469:
3688:
3681:
3674:
3003:
2996:
2989:
2982:
2184:
2178:
1552:10.1111/j.1502-3931.2012.00310.x
1546:, (advance online publication).
1276:Welsh, Jennifer (5 April 2012).
1134:Wilford, John N (4 April 2012).
1023:Timeline of tyrannosaur research
351:— all found in the rock beds of
134:
1308:Palaeontology: The truth about
924:Illustration showing a pack of
682:, placing it within the family
450:but have been prepared by the
1:
5972:Fossil taxa described in 2012
1282:The Christian Science Monitor
655:, short dorsal vertebrae, an
584:) on parts of the body where
470:specimens compared to a human
1452:10.1371/journal.pone.0198078
454:, under the guidance of Xu.
877:According to a 2018 study,
526:The described specimens of
448:Erlianhaote Dinosaur Museum
6003:
1250:. Tetrapod Zoology Blogs.
1178:. The Loom. Archived from
5810:
5576:
5493:
5468:
5352:
5348:
5307:
5292:
5279:
5243:
5190:
5158:
4978:
4901:
4770:
4766:
4651:
4647:
4536:
4521:
4508:
4467:
4303:
4289:
4209:
4121:
4009:
3866:
3819:
3815:
3765:
3730:
3715:
3702:
3671:
3633:
3546:
3454:
3407:
3276:
3251:
3247:
3175:
3171:
3045:
3030:
3017:
2979:
2945:
2768:
2588:
2403:
2354:
2350:
2263:
2259:
2225:
2210:
2197:
2175:
2000:
1883:
1801:
1786:
1773:
1723:
1710:
1658:
1532:– via ResearchGate.
1202:Paul, Gregory S. (2016).
793:
776:
769:
752:
745:
728:
721:
704:
697:
404:consists of the Mandarin
385:The name is derived from
283:
276:
131:Scientific classification
129:
124:
115:
23:
970:Because the locality of
444:Zhucheng Dinosaur Museum
1317:8 November 2020 at the
934:
862:
678:to be more basal than
671:
653:infratemporal fenestra
564:regulating temperature
559:
523:
471:
372:
5982:Taxa named by Xu Xing
5934:Paleobiology Database
4202:Carcharodontosaurinae
4114:Carcharodontosauridae
1397:17 April 2019 at the
1011:temperate rainforests
923:
860:
669:
607:phylogenetic analyses
557:
521:
465:
370:
5195:Dubious coelurosaurs
3891:Metriacanthosaurinae
3859:Metriacanthosauridae
2141:Dubious neotheropods
1192:(with illustrations)
363:Discovery and naming
5967:Feathered dinosaurs
4215:Carcharodontosaurus
4002:Carcharodontosauria
1443:2018PLoSO..1398078L
1247:Scientific American
1098:10.1038/nature10906
1090:2012Natur.484...92X
5687:Archaeornithomimus
4191:Veterupristisaurus
4051:Veterupristisaurus
3907:Metriacanthosaurus
3847:Piatnitzkysauridae
3282:Eustreptospondylus
3192:Piatnitzkysauridae
3075:Chuandongocoelurus
1939:Notatesseraeraptor
1570:Geological Journal
1491:(Podcast). Nature.
1402:Scientific Reports
1324:, 23 October 2013.
1140:The New York Times
935:
863:
672:
560:
524:
472:
426:, dating from the
373:
371:Fossils on display
5949:
5948:
5921:Open Tree of Life
5848:Taxon identifiers
5839:
5838:
5835:
5834:
5808:
5807:
5804:
5803:
5800:
5799:
5796:
5795:
5763:
5762:
5759:
5758:
5569:Macrocheiriformes
5486:Ornithomimosauria
5460:Maniraptoriformes
5453:
5452:
5299:Maniraptoromorpha
5286:Maniraptoromorpha
5275:
5274:
5241:
5240:
5237:
5236:
5233:
5232:
5186:
5185:
5182:
5181:
5164:Maniraptoromorpha
5154:
5153:
5150:
5149:
5146:
5145:
5142:
5141:
5138:
5137:
4886:
4885:
4751:
4750:
4705:Proceratosauridae
4657:Chingkankousaurus
4504:
4503:
4465:
4464:
4461:
4460:
4457:
4456:
4433:
4432:
4285:
4284:
4281:
4280:
4277:
4276:
4273:
4272:
4269:
4268:
4265:
4264:
3941:
3940:
3937:
3936:
3698:
3697:
3669:
3668:
3665:
3664:
3661:
3660:
3657:
3656:
3629:
3628:
3625:
3624:
3621:
3620:
3617:
3616:
3533:
3532:
3489:Vallibonavenatrix
3392:
3391:
3388:
3387:
3232:
3231:
3013:
3012:
2977:
2976:
2973:
2972:
2969:
2968:
2941:
2940:
2937:
2936:
2933:
2932:
2929:
2928:
2925:
2924:
2573:
2572:
2569:
2568:
2335:
2334:
2193:
2192:
2173:
2172:
2169:
2168:
2165:
2164:
2116:
2115:
1769:
1768:
1765:
1764:
1761:
1760:
1414:10.1038/srep20252
1306:Switek, B. 2013.
1215:978-1-78684-190-2
1123:on 17 April 2012.
844:
843:
835:
834:
826:
825:
817:
816:
808:
807:
694:Proceratosauridae
684:Proceratosauridae
670:Close-up of snout
307:
306:
257:
233:Proceratosauridae
5994:
5977:Proceratosaurids
5942:
5941:
5929:
5928:
5916:
5915:
5903:
5902:
5890:
5889:
5888:
5875:
5874:
5873:
5843:
5830:
5824:
5817:
5673:Aepyornithomimus
5574:
5491:
5466:
5430:Sinocalliopteryx
5350:
5346:
5340:Neocoelurosauria
5305:
5294:
5281:
5269:
5262:
5255:
5249:
5192:
5160:
5129:Zhuchengtyrannus
5045:Thanatotheristes
4976:
4940:
4907:Appalachiosaurus
4899:
4768:
4764:
4758:Pantyrannosauria
4649:
4645:
4639:Tyrannosauroidea
4634:
4609:Xinjiangovenator
4534:
4523:
4510:
4498:
4491:
4486:
4479:
4473:
4301:
4291:
4207:
4127:Acrocanthosaurus
4119:
4007:
3954:
3880:Yangchuanosaurus
3864:
3817:
3813:
3763:
3744:Lourinhanosaurus
3728:
3717:
3704:
3692:
3685:
3678:
3635:
3601:Sigilmassasaurus
3544:
3501:Ceratosuchopsini
3452:
3405:
3350:Dubreuillosaurus
3274:
3257:Streptospondylus
3249:
3245:
3215:Piatnitzkysaurus
3173:
3169:
3158:
3043:
3032:
3019:
3007:
3000:
2993:
2986:
2947:
2782:Ekrixinatosaurus
2766:
2757:
2586:
2401:
2352:
2348:
2261:
2257:
2223:
2212:
2199:
2188:
2182:
2092:Procompsognathus
1998:
1881:
1799:
1788:
1775:
1721:
1712:
1705:
1704:
1660:
1645:
1638:
1631:
1622:
1596:
1595:
1585:
1576:(3–4): 377–393.
1564:Zhou, Z (2006).
1561:
1555:
1540:
1534:
1533:
1523:
1499:
1493:
1492:
1489:"Nature podcast"
1484:
1475:
1474:
1464:
1454:
1422:
1416:
1388:
1382:
1381:
1371:
1352:10.1002/jez.b.25
1331:
1325:
1304:
1298:
1297:
1295:
1293:
1273:
1264:
1263:
1261:
1259:
1240:(4 April 2012).
1234:
1228:
1227:
1199:
1193:
1191:
1189:
1187:
1168:(4 April 2012).
1162:
1156:
1155:
1153:
1151:
1131:
1125:
1124:
1122:
1116:. Archived from
1075:
1066:
966:Paleoenvironment
772:
748:
724:
700:
690:
615:Tyrannosauroidea
503:postorbital bone
424:Yixian Formation
398:Tyrannosauroidea
387:Mandarin Chinese
376:Yutyrannus huali
349:Yutyrannus huali
332:Yutyrannus huali
292:
290:Yutyrannus huali
288:
251:
244:
231:
220:Tyrannosauroidea
218:
139:
138:
120:
110:
40:
31:Early Cretaceous
29:Temporal range:
21:
6002:
6001:
5997:
5996:
5995:
5993:
5992:
5991:
5952:
5951:
5950:
5945:
5937:
5932:
5924:
5919:
5911:
5906:
5898:
5893:
5884:
5883:
5878:
5869:
5868:
5863:
5850:
5840:
5831:
5818:
5792:
5755:
5736:Sinornithomimus
5658:
5601:
5563:
5480:
5474:Compsognathidae
5462:
5449:
5437:Sinosauropteryx
5369:Compsognathidae
5363:
5342:
5333:
5301:
5288:
5271:
5270:
5256:
5229:
5224:Shanyangosaurus
5217:Phaedrolosaurus
5178:
5134:
5100:
5050:
5029:Daspletosaurini
5023:
4996:
4971:Tyrannosaurinae
4965:
4944:Albertosaurinae
4936:
4934:Tyrannosauridae
4927:
4895:
4893:Eutyrannosauria
4882:
4760:
4747:
4728:Proceratosaurus
4699:
4670:
4641:
4630:
4621:
4587:Richardoestesia
4571:Chilantaisaurus
4530:
4517:
4500:
4499:
4480:
4453:
4429:
4389:Australovenator
4367:
4361:Chilantaisaurus
4339:Phuwiangvenator
4261:
4226:Giganotosaurini
4220:
4196:
4108:
4071:Chilantaisaurus
4056:
3996:
3950:
3933:
3885:
3853:
3809:
3800:
3778:Monolophosaurus
3759:
3750:
3724:
3711:
3694:
3693:
3679:
3653:
3613:
3586:
3552:Camarillasaurus
3529:
3495:
3441:
3435:Ostafrikasaurus
3413:Camarillasaurus
3401:
3384:
3334:Afrovenatorinae
3328:
3287:
3263:
3241:
3228:
3186:
3165:
3163:Megalosauroidea
3154:
3145:
3140:Vectaerovenator
3133:Szechuanosaurus
3111:Monolophosaurus
3097:Kaijiangosaurus
3090:Cryolophosaurus
3051:Calamospondylus
3039:
3026:
3009:
3008:
3001:
2994:
2987:
2965:
2921:
2902:Pycnonemosaurus
2844:
2755:
2749:Majungasaurinae
2734:
2676:Majungasaurinae
2670:
2665:Xenotarsosaurus
2601:Dryptosauroides
2582:
2565:
2517:
2489:Elaphrosaurinae
2483:
2390:
2376:Ornithomimoides
2344:
2331:
2304:
2253:
2244:
2219:
2206:
2189:
2183:
2161:
2136:
2112:
2040:
1987:
1953:Shuangbaisaurus
1877:
1868:
1795:
1782:
1757:
1737:
1731:Avemetatarsalia
1716:Avemetatarsalia
1706:
1691:Avemetatarsalia
1665:
1664:
1654:
1649:
1605:
1600:
1599:
1583:10.1002/gj.1045
1563:
1562:
1558:
1541:
1537:
1521:10.1.1.624.2450
1501:
1500:
1496:
1486:
1485:
1478:
1437:(6): e0198078.
1424:
1423:
1419:
1399:Wayback Machine
1389:
1385:
1333:
1332:
1328:
1319:Wayback Machine
1305:
1301:
1291:
1289:
1275:
1274:
1267:
1257:
1255:
1236:
1235:
1231:
1216:
1201:
1200:
1196:
1185:
1183:
1182:on 8 April 2012
1164:
1163:
1159:
1149:
1147:
1133:
1132:
1128:
1120:
1084:(7392): 92–95.
1073:
1068:
1067:
1036:
1031:
1019:
1002:. Fish such as
993:Sinosauropteryx
968:
918:
916:Social behavior
875:
855:
850:
845:
836:
827:
818:
809:
732:Proceratosaurus
603:
516:
483:Gregory S. Paul
460:
365:
320:proceratosaurid
303:
294:
286:
285:
272:
250:
242:
229:
216:
133:
111:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
35:
34:
27:
17:
12:
11:
5:
6000:
5998:
5990:
5989:
5984:
5979:
5974:
5969:
5964:
5954:
5953:
5947:
5946:
5944:
5943:
5930:
5917:
5904:
5891:
5876:
5860:
5858:
5852:
5851:
5846:
5837:
5836:
5833:
5832:
5812:
5811:
5809:
5806:
5805:
5802:
5801:
5798:
5797:
5794:
5793:
5791:
5790:
5789:
5788:
5782:
5773:
5771:
5765:
5764:
5761:
5760:
5757:
5756:
5754:
5753:
5746:
5739:
5732:
5725:
5718:
5711:
5704:
5701:Dromiceiomimus
5697:
5690:
5683:
5676:
5668:
5666:
5664:Ornithomimidae
5660:
5659:
5657:
5656:
5649:
5645:Paraxenisaurus
5641:
5633:
5626:
5619:
5611:
5609:
5607:Deinocheiridae
5603:
5602:
5600:
5599:
5596:Shenzhousaurus
5592:
5585:
5577:
5571:
5565:
5564:
5562:
5561:
5554:
5546:
5539:
5532:
5525:
5518:
5510:
5502:
5494:
5488:
5482:
5481:
5479:
5478:
5469:
5463:
5458:
5455:
5454:
5451:
5450:
5448:
5447:
5440:
5433:
5426:
5418:
5411:
5404:
5397:
5393:Beipiaognathus
5389:
5382:
5373:
5371:
5365:
5364:
5362:
5361:
5353:
5343:
5338:
5335:
5334:
5332:
5331:
5324:
5317:
5308:
5302:
5297:
5290:
5289:
5284:
5277:
5276:
5273:
5272:
5264:
5250:
5244:
5242:
5239:
5238:
5235:
5234:
5231:
5230:
5228:
5227:
5220:
5213:
5206:
5198:
5196:
5188:
5187:
5184:
5183:
5180:
5179:
5177:
5176:
5175:
5174:
5168:
5166:
5156:
5155:
5152:
5151:
5148:
5147:
5144:
5143:
5140:
5139:
5136:
5135:
5133:
5132:
5125:
5118:
5110:
5108:
5106:Tyrannosaurini
5102:
5101:
5099:
5098:
5091:
5084:
5076:
5069:
5065:Bistahieversor
5060:
5058:
5056:Teratophoneini
5052:
5051:
5049:
5048:
5041:
5038:Daspletosaurus
5033:
5031:
5025:
5024:
5022:
5021:
5018:Qianzhousaurus
5014:
5006:
5004:
4998:
4997:
4995:
4994:
4987:
4979:
4973:
4967:
4966:
4964:
4963:
4956:
4948:
4946:
4937:
4932:
4929:
4928:
4926:
4925:
4918:
4914:Bistahieversor
4910:
4902:
4896:
4891:
4888:
4887:
4884:
4883:
4881:
4880:
4872:
4865:
4858:
4850:
4843:
4836:
4829:
4822:
4815:
4808:
4801:
4794:
4786:
4779:
4771:
4761:
4756:
4753:
4752:
4749:
4748:
4746:
4745:
4738:
4731:
4724:
4717:
4709:
4707:
4701:
4700:
4698:
4697:
4689:
4681:
4679:
4672:
4671:
4669:
4668:
4660:
4652:
4642:
4637:
4631:
4628:Tyrannoraptora
4626:
4623:
4622:
4620:
4619:
4612:
4605:
4598:
4590:
4583:
4575:
4567:
4560:
4552:
4545:
4537:
4531:
4526:
4519:
4518:
4513:
4506:
4505:
4502:
4501:
4493:
4474:
4468:
4466:
4463:
4462:
4459:
4458:
4455:
4454:
4452:
4451:
4450:
4449:
4443:
4441:
4435:
4434:
4431:
4430:
4428:
4427:
4420:
4413:
4406:
4399:
4392:
4385:
4377:
4375:
4373:Megaraptoridae
4369:
4368:
4366:
4365:
4357:
4349:
4342:
4335:
4328:
4320:
4312:
4304:
4298:
4287:
4286:
4283:
4282:
4279:
4278:
4275:
4274:
4271:
4270:
4267:
4266:
4263:
4262:
4260:
4259:
4252:
4245:
4238:
4235:Giganotosaurus
4230:
4228:
4222:
4221:
4219:
4218:
4210:
4204:
4198:
4197:
4195:
4194:
4187:
4180:
4173:
4166:
4159:
4152:
4145:
4138:
4130:
4122:
4116:
4110:
4109:
4107:
4106:
4098:
4090:
4083:
4075:
4066:
4064:
4062:Neovenatoridae
4058:
4057:
4055:
4054:
4047:
4044:Ulughbegsaurus
4040:
4032:
4025:
4018:
4010:
4004:
3998:
3997:
3995:
3994:
3986:
3978:
3970:
3962:
3960:
3951:
3946:
3943:
3942:
3939:
3938:
3935:
3934:
3932:
3931:
3924:
3917:
3910:
3903:
3895:
3893:
3887:
3886:
3884:
3883:
3876:
3872:Xuanhanosaurus
3867:
3861:
3855:
3854:
3852:
3851:
3843:
3839:Xuanhanosaurus
3835:
3828:
3825:Asfaltovenator
3820:
3810:
3805:
3802:
3801:
3799:
3798:
3790:
3786:Megalosauridae
3782:
3774:
3766:
3760:
3755:
3752:
3751:
3749:
3748:
3740:
3731:
3725:
3720:
3713:
3712:
3707:
3700:
3699:
3696:
3695:
3687:
3673:
3672:
3670:
3667:
3666:
3663:
3662:
3659:
3658:
3655:
3654:
3652:
3651:
3650:
3649:
3643:
3641:
3631:
3630:
3627:
3626:
3623:
3622:
3619:
3618:
3615:
3614:
3612:
3611:
3604:
3596:
3594:
3588:
3587:
3585:
3584:
3577:
3570:
3563:
3560:Ichthyovenator
3556:
3547:
3541:
3535:
3534:
3531:
3530:
3528:
3527:
3520:
3513:
3505:
3503:
3497:
3496:
3494:
3493:
3485:
3477:
3470:
3463:
3455:
3449:
3443:
3442:
3440:
3439:
3431:
3424:
3421:Cristatusaurus
3417:
3408:
3402:
3397:
3394:
3393:
3390:
3389:
3386:
3385:
3383:
3382:
3378:Poekilopleuron
3374:
3371:Piveteausaurus
3367:
3360:
3353:
3346:
3338:
3336:
3330:
3329:
3327:
3326:
3319:
3312:
3305:
3297:
3295:
3293:Megalosaurinae
3289:
3288:
3286:
3285:
3277:
3271:
3269:Megalosauridae
3265:
3264:
3262:
3261:
3252:
3242:
3237:
3234:
3233:
3230:
3229:
3227:
3226:
3222:Xuanhanosaurus
3218:
3211:
3204:
3196:
3194:
3188:
3187:
3185:
3184:
3181:Yunyangosaurus
3176:
3166:
3161:
3155:
3150:
3147:
3146:
3144:
3143:
3136:
3129:
3121:
3118:Pandoravenator
3114:
3107:
3104:Kayentavenator
3100:
3093:
3086:
3078:
3071:
3063:
3055:
3046:
3040:
3035:
3028:
3027:
3022:
3015:
3014:
3011:
3010:
3002:
2995:
2988:
2981:
2980:
2978:
2975:
2974:
2971:
2970:
2967:
2966:
2964:
2963:
2962:
2961:
2955:
2953:
2943:
2942:
2939:
2938:
2935:
2934:
2931:
2930:
2927:
2926:
2923:
2922:
2920:
2919:
2912:
2905:
2898:
2891:
2883:
2876:
2869:
2862:
2854:
2852:
2846:
2845:
2843:
2842:
2835:
2832:Skorpiovenator
2828:
2820:
2813:
2806:
2799:
2792:
2785:
2778:
2769:
2763:
2754:
2753:
2744:
2742:
2736:
2735:
2733:
2732:
2725:
2717:
2710:
2703:
2696:
2688:
2680:
2678:
2672:
2671:
2669:
2668:
2661:
2654:
2647:
2644:Spectrovenator
2640:
2633:
2626:
2619:
2612:
2605:
2597:
2589:
2583:
2578:
2575:
2574:
2571:
2570:
2567:
2566:
2564:
2563:
2556:
2549:
2542:
2535:
2527:
2525:
2519:
2518:
2516:
2515:
2508:
2501:
2493:
2491:
2485:
2484:
2482:
2481:
2478:Spinostropheus
2474:
2467:
2460:
2452:
2444:
2436:
2428:
2421:
2413:
2404:
2398:
2392:
2391:
2389:
2388:
2380:
2372:
2364:
2355:
2345:
2342:Abelisauroidea
2340:
2337:
2336:
2333:
2332:
2330:
2329:
2322:
2314:
2312:
2310:Ceratosauridae
2306:
2305:
2303:
2302:
2299:Saltriovenator
2295:
2287:
2280:
2276:Fosterovenator
2272:
2264:
2254:
2249:
2246:
2245:
2243:
2242:
2238:Lophostropheus
2234:
2226:
2220:
2215:
2208:
2207:
2202:
2195:
2194:
2191:
2190:
2177:
2176:
2174:
2171:
2170:
2167:
2166:
2163:
2162:
2160:
2159:
2152:
2144:
2142:
2138:
2137:
2135:
2134:
2133:
2132:
2126:
2124:
2118:
2117:
2114:
2113:
2111:
2110:
2103:
2099:Pterospondylus
2095:
2088:
2080:
2072:
2069:Lucianovenator
2065:
2058:
2050:
2048:
2042:
2041:
2039:
2038:
2031:
2024:
2017:
2010:
2001:
1995:
1993:Coelophysoidea
1989:
1988:
1986:
1985:
1978:
1970:
1963:
1956:
1949:
1942:
1935:
1932:Lophostropheus
1928:
1921:
1914:
1907:
1900:
1892:
1884:
1878:
1873:
1870:
1869:
1867:
1866:
1858:
1850:
1842:
1834:
1831:Erythrovenator
1827:
1819:
1811:
1802:
1796:
1791:
1784:
1783:
1778:
1771:
1770:
1767:
1766:
1763:
1762:
1759:
1758:
1756:
1755:
1754:
1753:
1747:
1745:
1739:
1738:
1736:
1735:
1734:
1733:
1724:
1718:
1708:
1707:
1703:
1702:
1693:
1684:
1678:
1672:
1663:
1656:
1655:
1650:
1648:
1647:
1640:
1633:
1625:
1619:
1618:
1604:
1603:External links
1601:
1598:
1597:
1556:
1535:
1494:
1476:
1417:
1383:
1326:
1299:
1265:
1229:
1214:
1194:
1157:
1126:
1033:
1032:
1030:
1027:
1026:
1025:
1018:
1015:
981:Psittacosaurus
967:
964:
917:
914:
910:ornithischians
874:
871:
854:
851:
849:
846:
842:
841:
838:
837:
833:
832:
829:
828:
824:
823:
820:
819:
815:
814:
811:
810:
806:
805:
802:
801:
792:
789:
788:
785:
784:
775:
770:
768:
765:
764:
761:
760:
751:
746:
744:
741:
740:
737:
736:
727:
722:
720:
717:
716:
713:
712:
703:
698:
696:
688:
602:
601:Classification
599:
515:
512:
466:Size of three
459:
456:
364:
361:
323:tyrannosauroid
305:
304:
295:
281:
280:
274:
273:
265:
263:
259:
258:
240:
236:
235:
227:
223:
222:
214:
210:
209:
204:
197:
196:
191:
184:
183:
178:
171:
170:
165:
161:
160:
155:
151:
150:
145:
141:
140:
127:
126:
122:
121:
113:
112:
104:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
41:
28:
15:
13:
10:
9:
6:
4:
3:
2:
5999:
5988:
5985:
5983:
5980:
5978:
5975:
5973:
5970:
5968:
5965:
5963:
5960:
5959:
5957:
5940:
5935:
5931:
5927:
5922:
5918:
5914:
5909:
5905:
5901:
5896:
5892:
5887:
5881:
5877:
5872:
5866:
5862:
5861:
5859:
5857:
5853:
5849:
5844:
5829:
5825:
5823:
5816:
5787:
5783:
5781:
5777:
5776:
5775:
5774:
5772:
5770:
5766:
5752:
5751:
5747:
5745:
5744:
5743:Struthiomimus
5740:
5738:
5737:
5733:
5731:
5730:
5726:
5724:
5723:
5719:
5717:
5716:
5712:
5710:
5709:
5705:
5703:
5702:
5698:
5696:
5695:
5694:"Coelosaurus"
5691:
5689:
5688:
5684:
5682:
5681:
5677:
5675:
5674:
5670:
5669:
5667:
5665:
5661:
5655:
5654:
5650:
5647:
5646:
5642:
5639:
5638:
5634:
5632:
5631:
5627:
5625:
5624:
5620:
5618:
5617:
5613:
5612:
5610:
5608:
5604:
5598:
5597:
5593:
5591:
5590:
5589:Pelecanimimus
5586:
5584:
5583:
5579:
5578:
5575:
5572:
5570:
5566:
5560:
5559:
5555:
5552:
5551:
5550:Thecocoelurus
5547:
5545:
5544:
5540:
5538:
5537:
5533:
5531:
5530:
5529:Kinnareemimus
5526:
5524:
5523:
5519:
5516:
5515:
5511:
5508:
5507:
5503:
5501:
5500:
5496:
5495:
5492:
5489:
5487:
5483:
5476:
5475:
5471:
5470:
5467:
5464:
5461:
5456:
5446:
5445:
5441:
5439:
5438:
5434:
5432:
5431:
5427:
5424:
5423:
5419:
5417:
5416:
5412:
5410:
5409:
5408:Huaxiagnathus
5405:
5403:
5402:
5401:Compsognathus
5398:
5395:
5394:
5390:
5388:
5387:
5383:
5380:
5379:
5375:
5374:
5372:
5370:
5366:
5360:
5359:
5355:
5354:
5351:
5347:
5344:
5341:
5336:
5330:
5329:
5328:Ornitholestes
5325:
5323:
5322:
5318:
5315:
5314:
5310:
5309:
5306:
5303:
5300:
5295:
5291:
5287:
5282:
5278:
5268:
5263:
5261:
5254:
5248:
5226:
5225:
5221:
5219:
5218:
5214:
5212:
5211:
5207:
5205:
5204:
5200:
5199:
5197:
5193:
5189:
5172:
5171:
5170:
5169:
5167:
5165:
5161:
5157:
5131:
5130:
5126:
5124:
5123:
5122:Tyrannosaurus
5119:
5117:
5116:
5112:
5111:
5109:
5107:
5103:
5097:
5096:
5095:Teratophoneus
5092:
5090:
5089:
5085:
5082:
5081:
5077:
5075:
5074:
5070:
5067:
5066:
5062:
5061:
5059:
5057:
5053:
5047:
5046:
5042:
5040:
5039:
5035:
5034:
5032:
5030:
5026:
5020:
5019:
5015:
5013:
5012:
5008:
5007:
5005:
5003:
4999:
4993:
4992:
4988:
4986:
4985:
4981:
4980:
4977:
4974:
4972:
4968:
4962:
4961:
4957:
4955:
4954:
4953:Albertosaurus
4950:
4949:
4947:
4945:
4941:
4938:
4935:
4930:
4924:
4923:
4919:
4916:
4915:
4911:
4909:
4908:
4904:
4903:
4900:
4897:
4894:
4889:
4878:
4877:
4873:
4871:
4870:
4869:Xiongguanlong
4866:
4864:
4863:
4859:
4856:
4855:
4851:
4849:
4848:
4847:Suskityrannus
4844:
4842:
4841:
4840:Stokesosaurus
4837:
4835:
4834:
4833:Santanaraptor
4830:
4828:
4827:
4823:
4821:
4820:
4816:
4814:
4813:
4809:
4807:
4806:
4802:
4800:
4799:
4795:
4792:
4791:
4787:
4785:
4784:
4780:
4778:
4777:
4776:Alectrosaurus
4773:
4772:
4769:
4765:
4762:
4759:
4754:
4744:
4743:
4739:
4737:
4736:
4732:
4730:
4729:
4725:
4723:
4722:
4718:
4716:
4715:
4711:
4710:
4708:
4706:
4702:
4695:
4694:
4693:Tanycolagreus
4690:
4688:
4687:
4683:
4682:
4680:
4677:
4673:
4666:
4665:
4661:
4659:
4658:
4654:
4653:
4650:
4646:
4643:
4640:
4635:
4632:
4629:
4624:
4618:
4617:
4613:
4611:
4610:
4606:
4604:
4603:
4599:
4596:
4595:
4591:
4589:
4588:
4584:
4581:
4580:
4576:
4573:
4572:
4568:
4566:
4565:
4561:
4558:
4557:
4556:Bahariasaurus
4553:
4551:
4550:
4546:
4544:
4543:
4539:
4538:
4535:
4532:
4529:
4528:Coelurosauria
4524:
4520:
4516:
4515:Coelurosauria
4511:
4507:
4497:
4492:
4490:
4485:
4478:
4472:
4447:
4446:
4445:
4444:
4442:
4440:
4439:Coelurosauria
4436:
4426:
4425:
4421:
4419:
4418:
4414:
4412:
4411:
4407:
4405:
4404:
4400:
4398:
4397:
4393:
4391:
4390:
4386:
4384:
4383:
4379:
4378:
4376:
4374:
4370:
4364:
4362:
4358:
4356:
4354:
4350:
4348:
4347:
4343:
4341:
4340:
4336:
4334:
4333:
4329:
4326:
4325:
4321:
4318:
4317:
4316:Bahariasaurus
4313:
4311:
4310:
4306:
4305:
4302:
4299:
4296:
4292:
4288:
4258:
4257:
4253:
4251:
4250:
4246:
4244:
4243:
4239:
4237:
4236:
4232:
4231:
4229:
4227:
4223:
4217:
4216:
4212:
4211:
4208:
4205:
4203:
4199:
4193:
4192:
4188:
4186:
4185:
4181:
4179:
4178:
4174:
4172:
4171:
4167:
4165:
4164:
4160:
4158:
4157:
4156:Kelmayisaurus
4153:
4151:
4150:
4146:
4144:
4143:
4139:
4136:
4135:
4131:
4129:
4128:
4124:
4123:
4120:
4117:
4115:
4111:
4104:
4103:
4099:
4096:
4095:
4091:
4089:
4088:
4084:
4081:
4080:
4076:
4073:
4072:
4068:
4067:
4065:
4063:
4059:
4053:
4052:
4048:
4046:
4045:
4041:
4038:
4037:
4033:
4031:
4030:
4026:
4024:
4023:
4019:
4017:
4016:
4012:
4011:
4008:
4005:
4003:
3999:
3992:
3991:
3990:Saurophaganax
3987:
3984:
3983:
3979:
3976:
3975:
3971:
3969:
3968:
3964:
3963:
3961:
3959:
3955:
3952:
3949:
3944:
3930:
3929:
3925:
3923:
3922:
3921:Siamotyrannus
3918:
3916:
3915:
3911:
3909:
3908:
3904:
3902:
3901:
3897:
3896:
3894:
3892:
3888:
3882:
3881:
3877:
3874:
3873:
3869:
3868:
3865:
3862:
3860:
3856:
3849:
3848:
3844:
3841:
3840:
3836:
3834:
3833:
3829:
3827:
3826:
3822:
3821:
3818:
3814:
3811:
3808:
3807:Allosauroidea
3803:
3796:
3795:
3794:Spinosauridae
3791:
3788:
3787:
3783:
3780:
3779:
3775:
3773:
3772:
3768:
3767:
3764:
3761:
3758:
3753:
3746:
3745:
3741:
3738:
3737:
3733:
3732:
3729:
3726:
3723:
3718:
3714:
3710:
3705:
3701:
3691:
3686:
3684:
3677:
3647:
3646:
3645:
3644:
3642:
3640:
3636:
3632:
3610:
3609:
3605:
3603:
3602:
3598:
3597:
3595:
3593:
3589:
3583:
3582:
3578:
3576:
3575:
3571:
3569:
3568:
3564:
3562:
3561:
3557:
3554:
3553:
3549:
3548:
3545:
3542:
3540:
3539:Spinosaurinae
3536:
3526:
3525:
3521:
3519:
3518:
3517:Riparovenator
3514:
3512:
3511:
3510:Ceratosuchops
3507:
3506:
3504:
3502:
3498:
3491:
3490:
3486:
3483:
3482:
3478:
3476:
3475:
3474:Riojavenatrix
3471:
3469:
3468:
3464:
3462:
3461:
3457:
3456:
3453:
3450:
3448:
3447:Baryonychinae
3444:
3437:
3436:
3432:
3430:
3429:
3425:
3423:
3422:
3418:
3415:
3414:
3410:
3409:
3406:
3403:
3400:
3399:Spinosauridae
3395:
3380:
3379:
3375:
3373:
3372:
3368:
3366:
3365:
3361:
3359:
3358:
3354:
3352:
3351:
3347:
3345:
3344:
3340:
3339:
3337:
3335:
3331:
3325:
3324:
3323:Wiehenvenator
3320:
3318:
3317:
3313:
3311:
3310:
3306:
3304:
3303:
3299:
3298:
3296:
3294:
3290:
3284:
3283:
3279:
3278:
3275:
3272:
3270:
3266:
3259:
3258:
3254:
3253:
3250:
3246:
3243:
3240:
3235:
3224:
3223:
3219:
3217:
3216:
3212:
3210:
3209:
3205:
3203:
3202:
3198:
3197:
3195:
3193:
3189:
3183:
3182:
3178:
3177:
3174:
3170:
3167:
3164:
3159:
3156:
3153:
3148:
3142:
3141:
3137:
3135:
3134:
3130:
3127:
3126:
3122:
3120:
3119:
3115:
3113:
3112:
3108:
3106:
3105:
3101:
3099:
3098:
3094:
3092:
3091:
3087:
3084:
3083:
3079:
3077:
3076:
3072:
3069:
3068:
3064:
3061:
3060:
3059:Chienkosaurus
3056:
3053:
3052:
3048:
3047:
3044:
3041:
3038:
3033:
3029:
3025:
3020:
3016:
3006:
2999:
2992:
2985:
2959:
2958:
2957:
2956:
2954:
2952:
2948:
2944:
2918:
2917:
2913:
2911:
2910:
2906:
2904:
2903:
2899:
2897:
2896:
2892:
2889:
2888:
2884:
2882:
2881:
2877:
2875:
2874:
2870:
2868:
2867:
2863:
2861:
2860:
2856:
2855:
2853:
2851:
2850:Furileusauria
2847:
2841:
2840:
2836:
2834:
2833:
2829:
2826:
2825:
2824:Rahiolisaurus
2821:
2819:
2818:
2814:
2812:
2811:
2807:
2805:
2804:
2800:
2798:
2797:
2793:
2791:
2790:
2786:
2784:
2783:
2779:
2776:
2775:
2771:
2770:
2767:
2764:
2762:
2758:
2751:
2750:
2746:
2745:
2743:
2741:
2740:Carnotaurinae
2737:
2731:
2730:
2726:
2723:
2722:
2721:Rahiolisaurus
2718:
2716:
2715:
2714:Majungasaurus
2711:
2709:
2708:
2704:
2702:
2701:
2697:
2694:
2693:
2689:
2687:
2686:
2682:
2681:
2679:
2677:
2673:
2667:
2666:
2662:
2660:
2659:
2655:
2653:
2652:
2651:Tarascosaurus
2648:
2646:
2645:
2641:
2639:
2638:
2634:
2632:
2631:
2627:
2625:
2624:
2620:
2618:
2617:
2613:
2611:
2610:
2609:Eoabelisaurus
2606:
2603:
2602:
2598:
2596:
2595:
2594:Chenanisaurus
2591:
2590:
2587:
2584:
2581:
2580:Abelisauridae
2576:
2562:
2561:
2557:
2555:
2554:
2550:
2548:
2547:
2543:
2541:
2540:
2539:Masiakasaurus
2536:
2534:
2533:
2529:
2528:
2526:
2524:
2520:
2514:
2513:
2509:
2507:
2506:
2505:Huinculsaurus
2502:
2500:
2499:
2498:Elaphrosaurus
2495:
2494:
2492:
2490:
2486:
2480:
2479:
2475:
2473:
2472:
2468:
2466:
2465:
2461:
2458:
2457:
2453:
2450:
2449:
2445:
2442:
2441:
2437:
2434:
2433:
2429:
2427:
2426:
2422:
2419:
2418:
2417:Austrocheirus
2414:
2411:
2410:
2406:
2405:
2402:
2399:
2397:
2393:
2386:
2385:
2381:
2378:
2377:
2373:
2370:
2369:
2365:
2362:
2361:
2357:
2356:
2353:
2349:
2346:
2343:
2338:
2328:
2327:
2323:
2321:
2320:
2316:
2315:
2313:
2311:
2307:
2301:
2300:
2296:
2293:
2292:
2288:
2286:
2285:
2281:
2278:
2277:
2273:
2271:
2270:
2269:Berberosaurus
2266:
2265:
2262:
2258:
2255:
2252:
2247:
2240:
2239:
2235:
2233:
2232:
2228:
2227:
2224:
2221:
2218:
2213:
2209:
2205:
2200:
2196:
2187:
2181:
2158:
2157:
2156:Halticosaurus
2153:
2151:
2150:
2149:Teinurosaurus
2146:
2145:
2143:
2139:
2130:
2129:
2128:
2127:
2125:
2123:
2119:
2109:
2108:
2104:
2101:
2100:
2096:
2094:
2093:
2089:
2086:
2085:
2081:
2078:
2077:
2076:Megapnosaurus
2073:
2071:
2070:
2066:
2064:
2063:
2059:
2057:
2056:
2052:
2051:
2049:
2047:
2046:Coelophysidae
2043:
2037:
2036:
2035:Powellvenator
2032:
2030:
2029:
2025:
2023:
2022:
2018:
2016:
2015:
2011:
2008:
2007:
2003:
2002:
1999:
1996:
1994:
1990:
1984:
1983:
1979:
1976:
1975:
1971:
1969:
1968:
1964:
1962:
1961:
1957:
1955:
1954:
1950:
1948:
1947:
1943:
1941:
1940:
1936:
1934:
1933:
1929:
1927:
1926:
1925:Liliensternus
1922:
1920:
1919:
1915:
1913:
1912:
1908:
1906:
1905:
1901:
1898:
1897:
1896:Dolichosuchus
1893:
1891:
1890:
1889:Dilophosaurus
1886:
1885:
1882:
1879:
1876:
1871:
1864:
1863:
1862:Herrerasauria
1859:
1856:
1855:
1851:
1848:
1847:
1843:
1840:
1839:
1835:
1833:
1832:
1828:
1825:
1824:
1820:
1817:
1816:
1815:Daemonosaurus
1812:
1809:
1808:
1804:
1803:
1800:
1797:
1794:
1789:
1785:
1781:
1776:
1772:
1751:
1750:
1749:
1748:
1746:
1744:
1740:
1732:
1728:
1727:
1726:
1725:
1722:
1719:
1717:
1713:
1709:
1701:
1697:
1694:
1692:
1688:
1685:
1683:
1679:
1677:
1673:
1671:
1667:
1666:
1661:
1657:
1653:
1646:
1641:
1639:
1634:
1632:
1627:
1626:
1623:
1617:
1613:
1611:
1607:
1606:
1602:
1593:
1589:
1584:
1579:
1575:
1571:
1567:
1560:
1557:
1553:
1549:
1545:
1539:
1536:
1531:
1527:
1522:
1517:
1514:(1): 71–118.
1513:
1509:
1508:Geodiversitas
1505:
1498:
1495:
1490:
1483:
1481:
1477:
1472:
1468:
1463:
1458:
1453:
1448:
1444:
1440:
1436:
1432:
1428:
1421:
1418:
1415:
1411:
1407:
1403:
1400:
1396:
1393:
1387:
1384:
1379:
1375:
1370:
1365:
1361:
1357:
1353:
1349:
1345:
1341:
1337:
1330:
1327:
1323:
1320:
1316:
1313:
1311:
1303:
1300:
1287:
1283:
1279:
1272:
1270:
1266:
1253:
1249:
1248:
1243:
1239:
1238:Naish, Darren
1233:
1230:
1225:
1221:
1217:
1211:
1207:
1206:
1198:
1195:
1181:
1177:
1176:
1171:
1167:
1161:
1158:
1145:
1141:
1137:
1130:
1127:
1119:
1115:
1111:
1107:
1103:
1099:
1095:
1091:
1087:
1083:
1079:
1072:
1065:
1063:
1061:
1059:
1057:
1055:
1053:
1051:
1049:
1047:
1045:
1043:
1041:
1039:
1035:
1028:
1024:
1021:
1020:
1016:
1014:
1012:
1007:
1006:
1001:
1000:
995:
994:
989:
988:
983:
982:
977:
973:
965:
963:
961:
956:
952:
948:
944:
940:
933:
932:
927:
922:
915:
913:
911:
907:
903:
899:
896:
892:
888:
884:
881:had a simple
880:
872:
870:
868:
859:
852:
847:
840:
839:
831:
830:
822:
821:
813:
812:
804:
803:
800:
799:
798:
791:
790:
787:
786:
783:
782:
781:
774:
773:
767:
766:
763:
762:
759:
758:
757:
750:
749:
743:
742:
739:
738:
735:
734:
733:
726:
725:
719:
718:
715:
714:
711:
710:
709:
702:
701:
695:
692:
691:
687:
685:
681:
677:
668:
664:
662:
658:
654:
650:
649:quadratojugal
646:
642:
638:
637:
632:
631:
626:
622:
621:
616:
612:
608:
605:To date, all
600:
598:
595:
590:
587:
583:
582:
581:Tyrannosaurus
577:
576:
571:
570:
565:
556:
552:
550:
549:
548:Beipiaosaurus
544:
540:
536:
535:
529:
520:
513:
511:
509:
504:
500:
496:
492:
487:
484:
480:
476:
469:
464:
457:
455:
453:
449:
445:
440:
436:
431:
429:
425:
421:
417:
413:
411:
407:
403:
402:specific name
399:
395:
391:
388:
384:
381:
377:
369:
362:
360:
358:
354:
350:
346:
342:
338:
334:
333:
328:
324:
321:
317:
313:
312:
301:
298:
293:
291:
282:
279:
278:Binomial name
275:
271:
270:
269:Y. huali
264:
261:
260:
255:
249:
248:
241:
238:
237:
234:
228:
225:
224:
221:
215:
213:Superfamily:
212:
211:
208:
205:
202:
199:
198:
195:
192:
189:
186:
185:
182:
179:
176:
173:
172:
169:
166:
163:
162:
159:
156:
153:
152:
149:
146:
143:
142:
137:
132:
128:
123:
119:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
26:
22:
19:
5987:Yixian fauna
5855:
5819:
5748:
5741:
5734:
5727:
5720:
5715:Ornithomimus
5713:
5706:
5699:
5692:
5685:
5678:
5671:
5653:Tyrannomimus
5651:
5643:
5635:
5628:
5623:Deinocheirus
5621:
5614:
5594:
5587:
5580:
5556:
5548:
5543:Nqwebasaurus
5541:
5536:Nedcolbertia
5534:
5527:
5520:
5514:Calamosaurus
5512:
5506:Aviatyrannis
5504:
5497:
5472:
5442:
5435:
5428:
5420:
5413:
5406:
5399:
5391:
5386:Aristosuchus
5384:
5378:Aniksosaurus
5376:
5358:Aniksosaurus
5356:
5326:
5319:
5311:
5257:
5222:
5215:
5208:
5201:
5127:
5120:
5113:
5093:
5086:
5078:
5073:Dynamoterror
5071:
5063:
5043:
5036:
5016:
5009:
4989:
4984:Asiatyrannus
4982:
4958:
4951:
4922:Dryptosaurus
4920:
4912:
4905:
4874:
4867:
4860:
4852:
4845:
4838:
4831:
4824:
4817:
4812:Jinbeisaurus
4810:
4803:
4796:
4788:
4783:Aviatyrannis
4781:
4774:
4741:
4740:
4735:Sinotyrannus
4733:
4726:
4719:
4712:
4691:
4684:
4662:
4655:
4614:
4607:
4600:
4592:
4585:
4577:
4569:
4564:Bicentenaria
4562:
4554:
4549:Asiamericana
4547:
4540:
4481:
4422:
4415:
4408:
4401:
4394:
4387:
4380:
4359:
4351:
4344:
4337:
4330:
4324:Deltadromeus
4322:
4314:
4307:
4256:Tyrannotitan
4254:
4247:
4240:
4233:
4213:
4189:
4184:Taurovenator
4182:
4175:
4168:
4163:Lajasvenator
4161:
4154:
4147:
4142:Concavenator
4140:
4132:
4125:
4100:
4092:
4085:
4077:
4069:
4049:
4042:
4034:
4027:
4020:
4013:
3988:
3980:
3972:
3965:
3958:Allosauridae
3926:
3919:
3914:Shidaisaurus
3912:
3905:
3898:
3878:
3870:
3845:
3837:
3830:
3823:
3792:
3784:
3776:
3769:
3742:
3734:
3722:Avetheropoda
3709:Avetheropoda
3680:
3639:Avetheropoda
3606:
3599:
3592:Spinosaurini
3579:
3572:
3565:
3558:
3550:
3522:
3515:
3508:
3487:
3479:
3472:
3467:Protathlitis
3465:
3458:
3433:
3426:
3419:
3411:
3376:
3369:
3362:
3357:Leshansaurus
3355:
3348:
3341:
3321:
3314:
3309:Megalosaurus
3307:
3302:Duriavenator
3300:
3280:
3255:
3239:Megalosauria
3220:
3213:
3208:Marshosaurus
3206:
3201:Condorraptor
3199:
3179:
3138:
3131:
3123:
3116:
3109:
3102:
3095:
3088:
3082:Cruxicheiros
3080:
3073:
3065:
3057:
3049:
2914:
2909:Quilmesaurus
2907:
2900:
2893:
2885:
2878:
2871:
2864:
2857:
2837:
2830:
2822:
2815:
2808:
2801:
2794:
2787:
2780:
2772:
2761:Brachyrostra
2747:
2727:
2719:
2712:
2705:
2698:
2690:
2683:
2663:
2658:Tralkasaurus
2656:
2649:
2642:
2635:
2628:
2621:
2614:
2607:
2599:
2592:
2560:Vespersaurus
2558:
2553:Velocisaurus
2551:
2544:
2537:
2530:
2510:
2503:
2496:
2476:
2469:
2462:
2454:
2448:Deltadromeus
2446:
2438:
2432:Compsosuchus
2430:
2423:
2415:
2407:
2382:
2374:
2368:Lametasaurus
2366:
2358:
2324:
2319:Ceratosaurus
2317:
2297:
2289:
2282:
2274:
2267:
2251:Ceratosauria
2236:
2229:
2154:
2147:
2105:
2097:
2090:
2082:
2074:
2067:
2060:
2053:
2033:
2028:Podokesaurus
2026:
2019:
2012:
2004:
1980:
1972:
1965:
1958:
1951:
1944:
1937:
1930:
1923:
1916:
1911:Gojirasaurus
1909:
1904:Dracovenator
1902:
1894:
1887:
1875:Neotheropoda
1860:
1852:
1844:
1838:Guaibasaurus
1836:
1829:
1821:
1813:
1807:Chindesaurus
1805:
1695:
1686:
1609:
1573:
1569:
1559:
1543:
1538:
1511:
1507:
1497:
1434:
1430:
1420:
1405:
1401:
1386:
1346:(1): 42–56.
1343:
1339:
1329:
1321:
1309:
1302:
1290:. Retrieved
1281:
1256:. Retrieved
1245:
1232:
1204:
1197:
1184:. Retrieved
1180:the original
1173:
1166:Zimmer, Carl
1160:
1148:. Retrieved
1139:
1129:
1118:the original
1081:
1077:
1003:
997:
991:
987:Dongbeititan
985:
979:
975:
971:
969:
960:Dongbeititan
959:
954:
950:
946:
938:
936:
931:Dongbeititan
929:
925:
908:and certain
878:
876:
864:
848:Paleobiology
797:Sinotyrannus
795:
794:
779:
778:
777:
754:
753:
730:
729:
706:
705:
679:
675:
673:
636:Sinotyrannus
634:
628:
624:
618:
610:
604:
593:
591:
585:
579:
573:
567:
561:
546:
542:
539:Richard Prum
532:
527:
525:
490:
488:
477:was a large
474:
473:
467:
432:
415:
414:
405:
393:
389:
382:
375:
374:
348:
331:
330:
310:
309:
308:
299:
289:
284:
268:
267:
253:
246:
245:
200:
187:
174:
24:
18:
5880:Wikispecies
5780:Maniraptora
5769:Maniraptora
5750:Tototlmimus
5680:Anserimimus
5630:Garudimimus
5616:Beishanlong
5558:Valdoraptor
5499:Arkansaurus
5444:Xunmenglong
5321:Juravenator
5115:Tarbosaurus
4991:Nanuqsaurus
4960:Gorgosaurus
4876:Megaraptora
4862:Timurlengia
4594:Sciurumimus
4542:Aratasaurus
4410:Murusraptor
4332:Fukuiraptor
4295:Megaraptora
4177:Shaochilong
4149:Eocarcharia
4102:Megaraptora
4022:Lusovenator
3900:Alpkarakush
3757:Carnosauria
3608:Spinosaurus
3581:Siamosaurus
3481:Suchosaurus
3428:Iberospinus
3364:Magnosaurus
3343:Afrovenator
3316:Torvosaurus
3067:Chilesaurus
2880:Carnotaurus
2873:Caletodraco
2859:Abelisaurus
2685:Arcovenator
2532:Laevisuchus
2523:Noasaurinae
2425:Berthasaura
2396:Noasauridae
2326:Genyodectes
2291:Lukousaurus
2284:Jubbulpuria
2084:Panguraptor
2062:Coelophysis
2055:Camposaurus
2014:Panguraptor
2006:Dracoraptor
1982:Zupaysaurus
1967:Tachiraptor
1946:Sarcosaurus
1846:Nhandumirim
1322:Nature News
1150:28 February
999:Caudipteryx
575:Tarbosaurus
569:Gorgosaurus
499:premaxillae
458:Description
5956:Categories
5886:Yutyrannus
5856:Yutyrannus
5708:Gallimimus
5637:Harpymimus
5582:Harpymimus
5203:Iliosuchus
5173:see below↓
5002:Alioramini
4819:Juratyrant
4805:Eotyrannus
4790:Bagaraatan
4742:Yutyrannus
4676:Coeluridae
4602:Vayuraptor
4448:see below↓
4424:Tratayenia
4417:Orkoraptor
4403:Megaraptor
4309:Aoniraptor
4242:Mapusaurus
4170:Sauroniops
4134:Altispinax
4087:Neovenator
4029:Siamraptor
4015:Datanglong
3982:Epanterias
3974:Antrodemus
3967:Allosaurus
3948:Allosauria
3771:Altispinax
3736:Gasosaurus
3648:see below↓
3524:Suchomimus
3125:Sinosaurus
2960:see below↓
2916:Viavenator
2866:Aucasaurus
2803:Ilokelesia
2774:Dahalokely
2729:Rajasaurus
2707:Indosaurus
2700:Genusaurus
2692:Dahalokely
2616:Indosuchus
2512:Limusaurus
2471:Ligabueino
2464:Kiyacursor
2456:Genusaurus
2440:Dahalokely
2360:Betasuchus
2231:Dornraptor
2131:see below↓
2107:Segisaurus
1960:Sinosaurus
1823:Eodromaeus
1752:see below↓
1700:Dinosauria
1682:Sauropsida
1029:References
976:Yutyrannus
972:Yutyrannus
955:Yutyrannus
951:Mapusaurus
947:Yutyrannus
939:Yutyrannus
926:Yutyrannus
906:pterosaurs
898:archosaurs
879:Yutyrannus
780:Yutyrannus
676:Yutyrannus
620:Eotyrannus
611:Yutyrannus
594:Yutyrannus
586:Yutyrannus
528:Yutyrannus
508:surangular
491:Yutyrannus
475:Yutyrannus
468:Yutyrannus
416:Yutyrannus
337:Cretaceous
311:Yutyrannus
247:Yutyrannus
194:Saurischia
181:Dinosauria
25:Yutyrannus
5784:includes
5729:Rativates
5722:Qiupalong
5422:Scipionyx
5415:Mirischia
5088:Lythronax
5080:Labocania
5011:Alioramus
4664:Labocania
4382:Aerosteon
3928:Sinraptor
3832:Erectopus
3567:Irritator
3152:Orionides
3037:Tetanurae
3024:Tetanurae
2951:Tetanurae
2887:Elemgasem
2810:Llukalkan
2789:Elemgasem
2546:Noasaurus
2409:Afromimus
2217:Averostra
2204:Averostra
2122:Averostra
1974:Velocipes
1793:Theropoda
1780:Theropoda
1743:Theropoda
1668:Kingdom:
1652:Theropoda
1530:1280-9659
1516:CiteSeerX
1408:(20252).
1360:1552-5007
1224:985402380
1005:Lycoptera
891:crocodile
889:, like a
645:squamosal
439:paratypes
262:Species:
207:Theropoda
154:Kingdom:
148:Eukaryota
36:125
5900:52570686
5865:Wikidata
5313:Coelurus
4721:Kileskus
4714:Guanlong
4686:Coelurus
4579:Gualicho
4079:Gualicho
3460:Baryonyx
2796:Guemesia
2623:Kryptops
2384:Ozraptor
2021:Pendraig
1676:Chordata
1674:Phylum:
1670:Animalia
1616:BBC News
1592:54590389
1471:29924798
1431:PLOS ONE
1395:Archived
1378:12949768
1315:Archived
1286:Archived
1252:Archived
1175:Discover
1144:Archived
1114:29689629
1106:22481363
1017:See also
943:sauropod
928:hunting
867:ontogeny
853:Ontogeny
756:Kileskus
708:Guanlong
647:and the
641:lacrimal
630:Guanlong
543:Y. huali
514:Feathers
497:and the
446:and the
435:holotype
420:Liaoning
394:tyrannos
357:feathers
353:Liaoning
343:. Three
226:Family:
168:Chordata
164:Phylum:
158:Animalia
144:Domain:
5926:6158494
5913:8648057
5871:Q131710
4854:Timimus
4616:Zuolong
4346:Rapator
4249:Meraxes
3574:Oxalaia
2895:Koleken
1918:Lepidus
1680:Class:
1614:, from
1544:Lethaia
1462:6010247
1439:Bibcode
1369:4381994
1292:5 April
1258:5 April
1186:7 April
1086:Bibcode
895:extinct
873:Feeding
661:ischium
479:bipedal
410:plumage
380:Xu Xing
345:fossils
327:species
287:†
266:†
239:Genus:
105:↓
5939:230946
5522:Hexing
5210:Kakuru
4798:Dilong
2839:Thanos
2817:Niebla
2637:Rugops
2630:Kurupi
1610:T. rex
1590:
1528:
1518:
1469:
1459:
1376:
1366:
1358:
1310:T. rex
1222:
1212:
1112:
1104:
1078:Nature
996:, and
887:tongue
680:Dilong
625:Dilong
534:Dilong
495:nasals
428:Aptian
400:. The
383:et al.
302:, 2012
300:et al.
256:, 2012
254:et al.
5786:birds
4826:Moros
4353:Siats
4094:Siats
4036:Siats
1696:Clade
1687:Clade
1588:S2CID
1121:(PDF)
1110:S2CID
1074:(PDF)
902:birds
883:hyoid
657:ilium
406:huáli
341:China
316:genus
201:Clade
188:Clade
175:Clade
5908:GBIF
5778:see
4396:Maip
1854:Tawa
1729:see
1526:ISSN
1467:PMID
1374:PMID
1356:ISSN
1294:2012
1260:2012
1220:OCLC
1210:ISBN
1188:2012
1152:2017
1102:PMID
633:and
578:and
551:.
433:The
44:PreꞒ
5895:EoL
1578:doi
1548:doi
1457:PMC
1447:doi
1410:doi
1364:PMC
1348:doi
1344:298
1094:doi
1082:484
609:of
347:of
318:of
252:Xu
33:, ~
5958::
5936::
5923::
5910::
5897::
5882::
5867::
1698::
1689::
1586:.
1574:41
1572:.
1568:.
1524:.
1512:28
1510:.
1506:.
1479:^
1465:.
1455:.
1445:.
1435:13
1433:.
1429:.
1404:,
1372:.
1362:.
1354:.
1342:.
1338:.
1284:.
1280:.
1268:^
1244:.
1218:.
1172:.
1142:.
1138:.
1108:.
1100:.
1092:.
1080:.
1076:.
1037:^
990:,
984:,
962:.
912:.
904:,
663:.
627:,
572:,
541:,
510:.
412:.
390:yǔ
359:.
329:,
297:Xu
203::
190::
177::
94:Pg
38:Ma
5648:?
5640:?
5553:?
5517:?
5509:?
5477:?
5425:?
5396:?
5381:?
5316:?
5083:?
5068:?
4917:?
4879:?
4857:?
4793:?
4696:?
4678:?
4667:?
4597:?
4582:?
4574:?
4559:?
4363:?
4355:?
4327:?
4319:?
4297:?
4137:?
4105:?
4097:?
4082:?
4074:?
4039:?
3993:?
3985:?
3977:?
3875:?
3850:?
3842:?
3797:?
3789:?
3781:?
3747:?
3739:?
3555:?
3492:?
3484:?
3438:?
3416:?
3381:?
3260:?
3225:?
3128:?
3085:?
3070:?
3062:?
3054:?
2890:?
2827:?
2777:?
2752:?
2724:?
2695:?
2604:?
2459:?
2451:?
2443:?
2435:?
2420:?
2412:?
2387:?
2379:?
2371:?
2363:?
2294:?
2279:?
2241:?
2102:?
2087:?
2079:?
2009:?
1977:?
1899:?
1865:?
1857:?
1849:?
1841:?
1826:?
1818:?
1810:?
1644:e
1637:t
1630:v
1594:.
1580::
1554:.
1550::
1473:.
1449::
1441::
1412::
1406:6
1380:.
1350::
1312:.
1296:.
1262:.
1226:.
1190:.
1154:.
1096::
1088::
243:†
230:†
217:†
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:Ꞓ
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
