24:
1675:
106:
156:
126:
146:
136:
116:
75:
in 1913, " joins
Vermillion River through a magnificent gorge, or box canyon, so narrow that at several places the fissure, for it seems little more than a crack in the rock strata, is bridged by great boulders that have become wedged across it. It was a feature well worth seeing."
92:
fossil site in size and preservation. The report said that 22% of the observed species found in the initial excavation were new to science. Additionally, several species previously known only from
Chinese
1716:
1735:
1085:"Cephalic and Limb Anatomy of a New Isoxyid from the Burgess Shale and the Role of "Stem Bivalved Arthropods" in the Disparity of the Frontalmost Appendage"
1755:
97:, created millions of years earlier, were also found at the site. The exact location of the site was kept confidential to avoid damage to the site.
1709:
1636:
63:
39:
654:
1702:
1750:
23:
17:
1313:"Three new naraoiid species from the Burgess Shale, with a morphometric and phylogenetic reinvestigation of Naraoiidae"
1152:"A large new leanchoiliid from the Burgess Shale and the influence of inapplicable states on stem arthropod phylogeny"
1740:
605:
Caron, Jean-Bernard; Gaines, Robert R.; Aria, CĂ©dric; MĂĄngano, M. Gabriela; Streng, Michael (11 February 2014).
1682:
588:
71:
674:"A new hurdiid radiodont from the Burgess Shale evinces the exploitation of Cambrian infaunal food sources"
1745:
731:"A giant nektobenthic radiodont from the Burgess Shale and the significance of hurdiid carapace diversity"
43:
573:
1420:
1258:
1096:
1039:
974:
870:
807:
742:
618:
524:
105:
1568:
1452:
1389:
1340:
1290:
1173:
943:
839:
1470:
Nanglu, Karma; Caron, Jean-Bernard; Conway Morris, Simon; Cameron, Christopher B. (2016-07-07).
532:
505:
169:(2014) unless otherwise noted. Fossils first described from Marble Canyon are written as bold.
1619:
1560:
1552:
1511:
1493:
1444:
1436:
1381:
1332:
1282:
1274:
1227:
1132:
1114:
1065:
1008:
990:
935:
896:
831:
823:
776:
758:
711:
693:
636:
516:
1686:
1609:
1599:
1542:
1501:
1483:
1428:
1371:
1360:"Phylogenetic response of naraoiid arthropods to earlyâmiddle Cambrian environmental change"
1324:
1266:
1217:
1163:
1122:
1104:
1055:
1047:
998:
982:
927:
886:
878:
815:
766:
750:
701:
685:
626:
51:
47:
1674:
859:"A possible case of inverted lifestyle in a new bivalved arthropod from the Burgess Shale"
1312:
1206:"The Burgess Shale paleocommunity with new insights from Marble Canyon, British Columbia"
963:"Extreme multisegmentation in a giant bivalved arthropod from the Cambrian Burgess Shale"
1614:
1587:
1424:
1408:
1262:
1100:
1043:
978:
915:
874:
811:
746:
622:
1506:
1471:
1127:
1084:
1060:
1027:
1003:
962:
916:"A Burgess Shale mandibulate arthropod with a pygidium: a case of convergent evolution"
891:
858:
771:
730:
706:
673:
482:
475:
155:
94:
1729:
1393:
1344:
1294:
947:
542:
445:
400:
310:
276:
208:
89:
88:
soft-bodied organisms in or near Marble Canyon that rival or even surpass the nearby
1246:
1177:
1028:"The problematic Cambrian arthropod Tuzoia and the origin of mandibulates revisited"
607:"A new phyllopod bed-like assemblage from the Burgess Shale of the Canadian Rockies"
568:
125:
1572:
1456:
1311:
Mayers, Benjamin; Aria, CĂ©dric; Caron, JeanâBernard (2019). Zhang, XiâGuang (ed.).
843:
378:
363:
242:
175:
1150:
Aria, CĂ©dric; Caron, Jean-Bernard; Gaines, Robert (2015). Zhang, Xi-Guang (ed.).
1109:
258:
224:
216:
184:
986:
914:
IzquierdoâLĂłpez, Alejandro; Caron, JeanâBernard (2021). Zhang, XiâGuang (ed.).
1547:
1530:
1488:
1270:
795:
466:
422:
407:
393:
386:
333:
285:
59:
1651:
1638:
1556:
1531:"A New Burgess Shale Polychaete and the Origin of the Annelid Head Revisited"
1497:
1440:
1385:
1336:
1278:
1231:
1118:
994:
939:
827:
762:
697:
437:
356:
233:
1623:
1564:
1515:
1448:
1286:
1136:
1069:
1012:
900:
835:
780:
715:
689:
640:
796:"Burgess Shale fossils illustrate the origin of the mandibulate body plan"
145:
135:
496:
415:
371:
302:
200:
85:
1604:
1432:
1222:
1205:
1051:
882:
819:
754:
1376:
1359:
1328:
1168:
1151:
655:"Mighty Burgess Shale fossil site discovered in Kootenay National Park"
631:
606:
453:
430:
325:
317:
267:
1358:
Bond, Andrew D.; Edgecombe, Gregory D. (2021). Zhang, XiâGuang (ed.).
931:
250:
192:
55:
35:
115:
1247:"A middle Cambrian arthropod with chelicerae and proto-book gills"
460:
1204:
Nanglu, Karma; Caron, Jean-Bernard; Gaines, Robert R. (2020).
961:
Izquierdo-LĂłpez, Alejandro; Caron, Jean-Bernard (2022-07-15).
38:
surrounding Tokumm Creek just above its confluence with the
84:
A major new find was announced in early 2014 of fossilized
1407:
Morris, Simon Conway; Caron, Jean-Bernard (2014-08-01).
1026:
Izquierdo-LĂłpez, Alejandro; Caron, Jean-Bernard (2022).
857:
Izquierdo-LĂłpez, Alejandro; Caron, Jean-Bernard (2019).
1690:
1588:"First record of growth patterns in a Cambrian annelid"
678:
Proceedings of the Royal
Society B: Biological Sciences
1472:"Cambrian suspension-feeding tubicolous hemichordates"
1409:"A primitive fish from the Cambrian of North America"
1529:Nanglu, Karma; Caron, Jean-Bernard (2018-01-22).
1083:Aria, CĂ©dric; Caron, Jean-Bernard (2015-06-03).
1586:Osawa, H.; Caron, J.-B.; Gaines, R. R. (2023).
1710:
8:
58:. South of Marble Canyon on Highway 93 is
1717:
1703:
1245:Aria, CĂ©dric; Caron, Jean-Bernard (2019).
794:Aria, CĂ©dric; Caron, Jean-Bernard (2017).
1613:
1603:
1546:
1505:
1487:
1375:
1221:
1167:
1126:
1108:
1059:
1002:
890:
770:
705:
630:
16:For other places with the same name, see
672:Moysiuk, J.; Caron, J.-B. (2019-08-14).
154:
144:
134:
124:
114:
104:
22:
560:
1736:Canyons and gorges of British Columbia
1306:
1304:
1199:
1197:
1195:
1193:
1191:
1189:
1187:
7:
1671:
1669:
600:
598:
1689:. You can help Knowledge (XXG) by
729:Caron, J.-B.; Moysiuk, J. (2021).
14:
1681:This article about a location in
1756:British Columbia geography stubs
1673:
311:Liangshanella cf. burgessensis
18:Marble Canyon (disambiguation)
1:
491:2014) (833 specimens as 2020)
401:Ptychagnostus cf. praecurrens
1110:10.1371/journal.pone.0124979
544:Haplophrentis cf. carinatus
501:(>580 specimens as 2023)
1772:
1668:
1592:Royal Society Open Science
1032:Royal Society Open Science
987:10.1016/j.isci.2022.104675
863:Royal Society Open Science
735:Royal Society Open Science
592:, vol V, 1913, pp. 100â137
476:Kootenayscolex barbarensis
470:) (3373 specimens as 2020)
423:Primicaris cf. larvaformis
15:
1548:10.1016/j.cub.2017.12.019
1489:10.1186/s12915-016-0271-4
1271:10.1038/s41586-019-1525-4
372:Kootenia cf. burgessensis
1683:British Columbia, Canada
569:"Marble Canyon (canyon)"
547:(3694 specimens as 2020)
411:(2707 specimens as 2020)
394:Itagnostus interstrictus
357:Elrathina cf. marginalis
286:Mollisonia plenovenatrix
920:Papers in Palaeontology
589:Canadian Alpine Journal
449:(235 specimens as 2020)
426:(119 specimens as 2020)
306:(252 specimens as 2020)
272:(178 specimens as 2020)
251:Tuzoia cf. burgessensis
72:Canadian Alpine Journal
1751:Kootenay National Park
1652:51.18583°N 116.12278°W
690:10.1098/rspb.2019.1079
446:Metaspriggina walcotti
162:
160:Metaspriggina walcotti
152:
142:
132:
122:
112:
44:Kootenay National Park
42:, at the north end of
28:
611:Nature Communications
574:BC Geographical Names
176:Cambroraster falcatus
158:
148:
138:
128:
118:
110:Cambroraster falcatus
108:
26:
1657:51.18583; -116.12278
661:. February 11, 2014.
659:Royal Ontario Museum
303:Sidneyia inexpectans
225:Fibulacaris nereidis
130:Fibulacaris nereidis
80:Cambrian Lagerstatte
1648: /
1605:10.1098/rsos.221400
1433:10.1038/nature13414
1425:2014Natur.512..419M
1263:2019Natur.573..586A
1223:10.1017/pab.2019.42
1101:2015PLoSO..1024979A
1052:10.1098/rsos.220933
1044:2022RSOS....920933I
979:2022iSci...25j4675I
883:10.1098/rsos.191350
875:2019RSOS....691350I
820:10.1038/nature22080
812:2017Natur.545...89A
755:10.1098/rsos.210664
747:2021RSOS....810664C
623:2014NatCo...5.3210C
346:M. cf. longicaudata
259:Surusicaris elegans
243:Balhuticaris voltae
201:Tokummia katalepsis
185:Titanokorys gainesi
140:Balhuticaris voltae
120:Tokummia katalepsis
101:Species discoveries
1377:10.1111/pala.12516
1329:10.1111/pala.12383
1169:10.1111/pala.12161
684:(1908): 20191079.
632:10.1038/ncomms4210
416:Marrella splendens
163:
153:
143:
133:
123:
113:
29:
1698:
1697:
1541:(2): 319â326.e1.
1419:(7515): 419â422.
1257:(7775): 586â589.
932:10.1002/spp2.1366
431:Molaria spinifera
268:Yawunik kootenayi
150:Yawunik kootenayi
1763:
1741:Canadian Rockies
1719:
1712:
1705:
1677:
1670:
1663:
1662:
1660:
1659:
1658:
1653:
1649:
1646:
1645:
1644:
1641:
1628:
1627:
1617:
1607:
1583:
1577:
1576:
1550:
1526:
1520:
1519:
1509:
1491:
1467:
1461:
1460:
1404:
1398:
1397:
1379:
1355:
1349:
1348:
1308:
1299:
1298:
1242:
1236:
1235:
1225:
1201:
1182:
1181:
1171:
1147:
1141:
1140:
1130:
1112:
1080:
1074:
1073:
1063:
1023:
1017:
1016:
1006:
958:
952:
951:
926:(4): 1877â1894.
911:
905:
904:
894:
854:
848:
847:
791:
785:
784:
774:
726:
720:
719:
709:
669:
663:
662:
651:
645:
644:
634:
602:
593:
585:
579:
578:
565:
467:Margaretia dorus
318:Naraoia compacta
234:Pakucaris apatis
69:As described by
64:Vermillion River
52:British Columbia
48:Canadian Rockies
1771:
1770:
1766:
1765:
1764:
1762:
1761:
1760:
1726:
1725:
1724:
1723:
1666:
1656:
1654:
1650:
1647:
1642:
1639:
1637:
1635:
1634:
1632:
1631:
1585:
1584:
1580:
1535:Current Biology
1528:
1527:
1523:
1469:
1468:
1464:
1406:
1405:
1401:
1357:
1356:
1352:
1310:
1309:
1302:
1244:
1243:
1239:
1203:
1202:
1185:
1149:
1148:
1144:
1095:(6): e0124979.
1082:
1081:
1077:
1025:
1024:
1020:
960:
959:
955:
913:
912:
908:
856:
855:
851:
806:(7652): 89â92.
793:
792:
788:
728:
727:
723:
671:
670:
666:
653:
652:
648:
604:
603:
596:
586:
582:
567:
566:
562:
557:
551:
497:Ursactis comosa
461:Oesia disjuncta
165:Based on Caron
103:
82:
40:Vermilion River
21:
12:
11:
5:
1769:
1767:
1759:
1758:
1753:
1748:
1743:
1738:
1728:
1727:
1722:
1721:
1714:
1707:
1699:
1696:
1695:
1678:
1630:
1629:
1578:
1521:
1462:
1399:
1370:(1): 161â177.
1350:
1300:
1237:
1183:
1162:(4): 629â660.
1142:
1075:
1038:(12): 220933.
1018:
953:
906:
869:(11): 191350.
849:
786:
721:
664:
646:
617:: ncomms4210.
594:
580:
559:
558:
556:
553:
549:
548:
540:
538:M. georginense
529:
521:
513:
502:
492:
483:Burgessochaeta
471:
457:
450:
442:
434:
427:
419:
412:
404:
397:
390:
383:
375:
368:
360:
353:
321:
314:
307:
299:
281:
273:
263:
254:
247:
238:
229:
220:
213:
205:
196:
189:
180:
102:
99:
81:
78:
13:
10:
9:
6:
4:
3:
2:
1768:
1757:
1754:
1752:
1749:
1747:
1746:East Kootenay
1744:
1742:
1739:
1737:
1734:
1733:
1731:
1720:
1715:
1713:
1708:
1706:
1701:
1700:
1694:
1692:
1688:
1684:
1679:
1676:
1672:
1667:
1664:
1661:
1625:
1621:
1616:
1611:
1606:
1601:
1598:(4). 221400.
1597:
1593:
1589:
1582:
1579:
1574:
1570:
1566:
1562:
1558:
1554:
1549:
1544:
1540:
1536:
1532:
1525:
1522:
1517:
1513:
1508:
1503:
1499:
1495:
1490:
1485:
1481:
1477:
1473:
1466:
1463:
1458:
1454:
1450:
1446:
1442:
1438:
1434:
1430:
1426:
1422:
1418:
1414:
1410:
1403:
1400:
1395:
1391:
1387:
1383:
1378:
1373:
1369:
1365:
1364:Palaeontology
1361:
1354:
1351:
1346:
1342:
1338:
1334:
1330:
1326:
1322:
1318:
1317:Palaeontology
1314:
1307:
1305:
1301:
1296:
1292:
1288:
1284:
1280:
1276:
1272:
1268:
1264:
1260:
1256:
1252:
1248:
1241:
1238:
1233:
1229:
1224:
1219:
1215:
1211:
1207:
1200:
1198:
1196:
1194:
1192:
1190:
1188:
1184:
1179:
1175:
1170:
1165:
1161:
1157:
1156:Palaeontology
1153:
1146:
1143:
1138:
1134:
1129:
1124:
1120:
1116:
1111:
1106:
1102:
1098:
1094:
1090:
1086:
1079:
1076:
1071:
1067:
1062:
1057:
1053:
1049:
1045:
1041:
1037:
1033:
1029:
1022:
1019:
1014:
1010:
1005:
1000:
996:
992:
988:
984:
980:
976:
973:(7): 104675.
972:
968:
964:
957:
954:
949:
945:
941:
937:
933:
929:
925:
921:
917:
910:
907:
902:
898:
893:
888:
884:
880:
876:
872:
868:
864:
860:
853:
850:
845:
841:
837:
833:
829:
825:
821:
817:
813:
809:
805:
801:
797:
790:
787:
782:
778:
773:
768:
764:
760:
756:
752:
748:
744:
741:(9): 210664.
740:
736:
732:
725:
722:
717:
713:
708:
703:
699:
695:
691:
687:
683:
679:
675:
668:
665:
660:
656:
650:
647:
642:
638:
633:
628:
624:
620:
616:
612:
608:
601:
599:
595:
591:
590:
584:
581:
576:
575:
570:
564:
561:
554:
552:
546:
545:
541:
539:
535:
534:
530:
527:
526:
522:
519:
518:
514:
512:
511:P. interrupta
508:
507:
503:
500:
499:
498:
493:
490:
486:
484:
479:
478:
477:
472:
469:
468:
463:
462:
458:
456:
455:
451:
448:
447:
443:
440:
439:
435:
433:
432:
428:
425:
424:
420:
418:
417:
413:
410:
409:
405:
403:
402:
398:
396:
395:
391:
389:
388:
384:
381:
380:
376:
374:
373:
369:
366:
365:
361:
359:
358:
354:
351:
347:
343:
342:
337:
336:
335:
329:
328:
327:
322:
320:
319:
315:
313:
312:
308:
305:
304:
300:
297:
293:
292:M. symmetrica
289:
288:
287:
282:
279:
278:
277:Alalcomenaeus
274:
271:
270:
269:
264:
262:
261:
260:
255:
253:
252:
248:
246:
245:
244:
239:
237:
236:
235:
230:
228:
227:
226:
221:
219:
218:
214:
211:
210:
209:Branchiocaris
206:
204:
203:
202:
197:
195:
194:
190:
188:
187:
186:
181:
179:
178:
177:
172:
171:
170:
168:
161:
157:
151:
147:
141:
137:
131:
127:
121:
117:
111:
107:
100:
98:
96:
91:
90:Burgess Shale
87:
79:
77:
74:
73:
67:
65:
61:
57:
53:
49:
45:
41:
37:
33:
32:Marble Canyon
27:Marble Canyon
25:
19:
1691:expanding it
1680:
1665:
1633:
1595:
1591:
1581:
1538:
1534:
1524:
1479:
1475:
1465:
1416:
1412:
1402:
1367:
1363:
1353:
1323:(1): 19â50.
1320:
1316:
1254:
1250:
1240:
1216:(1): 58â81.
1213:
1210:Paleobiology
1209:
1159:
1155:
1145:
1092:
1088:
1078:
1035:
1031:
1021:
970:
966:
956:
923:
919:
909:
866:
862:
852:
803:
799:
789:
738:
734:
724:
681:
677:
667:
658:
649:
614:
610:
587:
583:
572:
563:
550:
543:
537:
531:
525:Linnarssonia
523:
515:
510:
509:n. sp. aff.
504:
495:
494:
488:
485:cf. setigera
481:
474:
473:
465:
459:
452:
444:
436:
429:
421:
414:
406:
399:
392:
385:
379:Zacanthoides
377:
370:
364:Bathyuriscus
362:
355:
349:
345:
340:
339:
332:
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