Knowledge (XXG)

McAbee Fossil Beds

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52: 295: 168: 60: 95:). The McAbee Fossil Beds, comprising 548.23 hectares (1,354.7 acres), were officially designated a Provincial Heritage Site under British Columbia's Heritage Conservation Act on July 19, 2012. The site is part of an old lake bed which was deposited about 52 million years ago and is internationally recognised for the diversity of plant, insect, and fish fossils found there. Similar 248:
and wet, with a mean annual temperature about 11 Â°C (52 Â°F), winters lacking frost (coldest month mean temperature ~5 Â°C), and annual precipitation over 1,000 mm (39 in) a year with little or no seasonality of precipitation. The extraordinary detail preserved in the insect
2922:. Significant collections of fossils from the McAbee Fossil Beds were in private ownership and fossils from the McAbee Fossil Beds are listed for sale on the internet. In August 2018 a substantial private collection of fossils from McAbee was donated to the Royal British Columbia Museum. 1092:
found in the McAbee Fossil Beds based on information in Archibald, Bossert, Greenwood, and Farrell (2010), Archibald, Mathewes, and Greenwood (2013), Archibald, Rasnitsyn and Akhmetiev (2005) and other sources cited in the list below, with extinct taxa denoted with a †.
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Lowe, A. J.; Greenwood, D. R.; West, C. K.; Galloway, J. M.; Sudermann, M.; Reichgelt, T. (2018). "Plant community ecology and climate on an upland volcanic landscape during the Early Eocene Climatic Optimum: McAbee Fossil Beds, British Columbia, Canada".
4058:
Pigg, K.B.; Dillhoff, R.M.; DeVore, M.L.; Wehr, W.C. (2007). "New diversity among the Trochodendraceae from the Early/Middle Eocene Okanogan Highlands of British Columbia, Canada, and Northeastern Washington State, United States".
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Hilton, E.J.; Grande, L. (2008). "Fossil Mooneyes (Teleostei: Hiodontiformes, Hiodontidae) from the Eocene of western North America, with a reassessment of their taxonomy" in "Birth of the modern world: the Tertiary".
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are also Eocene, but primarily preserve an aquatic plant community. A 2016 review of the early Eocene fossil sites from the interior of British Columbia discusses the history of paleobotanical research at McAbee, the
3654: 1069:) was described from the McAbee Fossil Beds. The very high diversity of fossil insects in the McAbee fossil beds is comparable to that of modern-day tropical forest areas. Additionally, fossil palm beetles ( 3609: 3987: 207:. Significant research on the fossil plants and insects has only occurred since the late 1980s. The McAbee Fossil Beds are best known for the abundant and well-preserved insect and fish fossils ( 4127:
Makarkin, V.N.; Archibald, S.B. (2013). "A diverse new assemblage of green lacewings (Insecta, Neuroptera, Chrysopidae) from the early Eocene Okanagan Highlands, western North America".
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Moss, PT; Greenwood, DR; Archibald, SB (2005). "Regional and local vegetation community dynamics of the Eocene Okanagan Highlands (British Columbia - Washington State) from palynology".
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Archibald, S.B.; Rasnitsyn, A.P. (2015). "New early Eocene Siricomorpha (Hymenoptera: Symphyta: Pamphiliidae, Siricidae, Cephidae) from the Okanagan Highlands, western North America".
4162:
Archibald, SB; Bradler, S (2015). "Stem-group stick insects (Phasmatodea) in the early Eocene at McAbee, British Columbia, Canada, and Republic, Washington, United States of America".
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Archibald, S.B.; Greenwood, D.R.; Mathewes, R.W. (2013). "Seasonality, montane beta diversity, and Eocene insects: Testing Janzen's dispersal hypothesis in an equable world".
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Douglas, S.D.; Stockey, R.A. (1996). "Insect fossils in Middle Eocene deposits from British Columbia and Washington State: faunal diversity and geological range extensions".
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Dillhoff, R.M.; Dillhoff, T.A.; Greenwood, D.R.; DeVore, M.L.; Pigg, K.B. (2013). "The Eocene Thomas Ranch flora, Allenby Formation, Princeton, British Columbia, Canada".
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Gushulak, C.A.; West, C.K.; Greenwood, D.R. (2016). "Paleoclimate and precipitation seasonality of the Early Eocene McAbee megaflora, Kamloops Group, British Columbia".
2966: 203:. Thomas Ewing provided a detailed analysis of the geology of the Kamloops Group, including the McAbee beds. More recently, Lowe et al. have assessed in detail the site 2925:
The cessation of fossil collecting at the McAbee Fossil Beds through heritage listing is consistent with British Columbia's Fossil Management Framework which seeks to:
4449: 4483: 3396:"Fossil biotas from the Okanagan Highlands, southern British Columbia and northeastern Washington State: climates and ecosystems across an Eocene landscape" 4523: 2899: 360:
found in the McAbee fossil beds based on the list found in Dillhoff, Leopold & Manchester (2005) and Wilson (2009) with extinct taxa denoted with a †.
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leaves and fruits from the Early-Middle Eocene of northwestern North America: systematics and implications for character evolution within Ulmaceae".
3110: 4528: 3060:"A review of paleobotanical studies of the Early Eocene Okanagan (Okanogan) Highlands floras of British Columbia, Canada and Washington, USA" 2879: 305: 2995: 3846:"The McAbee flora of British Columbia and its relations to the Early-Middle Eocene Okanagan Highlands flora of the Pacific Northwest" 187:. Palaeontological and geological studies of the McAbee Fossil Beds first commenced in the 1960s and early 1970s by Len Hills of the 104: 4503: 4488: 2970: 21: 3438:"Great Canadian Lagerstätten 1. Early Eocene Lagerstätten of the Okanagan Highlands (British Columbia and Washington State)" 3130: 4518: 3221:
Ewing, T.E. (1981). "Regional stratigraphy and structural setting of the Kamloops Group, south-central British Columbia".
176: 149: 124: 84: 76: 4199:"New Eocene crayfish from the McAbee Beds in British Columbia: First record of Parastacoidea in the Northern Hemisphere" 249:
fossils, as well as the high diversity of insects, plants and other organisms means the McAbee Fossil Beds represent a
159:, who have title on the lands. The site is currently closed to the public while interpretive components are installed. 4027:"Morphology, anatomy, phylogenetics and distribution of fossil and extant Trochodendraceae in the Northern Hemisphere" 3150: 2287: 229:
from the McAbee Fossil Beds and other Eocene sites in British Columbia is now considered to belong to the present-day
3685:(Sapindaceae) from the Tertiary of North America and implications for the phytogeographic history of the Aceroideae" 4416:"Bulldog Ants of the Eocene Okanagan Highlands and History of the Subfamily (Hymenoptera: Formicidae: Myrmeciinae)" 4096:"Bulldog Ants of the Eocene Okanagan Highlands and History of the Subfamily (Hymenoptera: Formicidae: Myrmeciinae)" 2907: 108: 92: 4508: 2891: 120: 4346: 4325: 3377: 3184:
Hills, L.V.; Baadsgaard, H. (1967). "Potassium-argon dating of some Lower Tertiary strata in British Columbia".
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Fossils of plant leaves, shoots, seeds, flowers and cones are abundant and well preserved, and include up to 76
950: 924: 4498: 4435: 4112: 4095: 3747:(Betulaceae) from the Middle Eocene Klondike Mountain and Allenby Formations of Northwestern North America". 2990: 1055: 3031:
Cevallos-Ferriz, SRS; Stockey, RA; Pigg, KB (1991). "Princeton chert: evidence for in situ aquatic plants".
2740: 2649: 2578: 1065: 930: 4460: 4493: 3615: 2544: 1016: 294: 156: 51: 3356:
Greenwood, D.R.; Wing, S.L. (1995). "Eocene continental climates and latitudinal temperature gradients".
2398: 257: 200: 188: 167: 59: 3786:(Fagaceae) fruits, foliage, and pollen from the Middle Eocene of Pacific Northwestern North America". 4278: 4239: 3890:, sp. nov., from the Middle Eocene Klondike Mountain Formation, Republic, Ferry County, Washington". 3860: 3576: 3531: 3407: 3365: 3266: 3230: 3071: 2887: 2474: 1471: 936: 765: 742: 184: 88: 3845: 568: 328: 1541: 968: 349: 1506: 791: 4381: 4179: 4144: 4076: 4007: 3965: 3926: 3764: 3706: 3632: 3477: 3282: 2915: 2510: 2436: 1434: 1041: 942: 718: 695: 318: 300: 272: 128: 4306: 3714: 2946:
clarify the rights and obligations of the public, business, government and other stakeholders.
2012: 1288: 1175: 773: 583: 245: 204: 3437: 592: 4430: 4373: 4341: 4296: 4286: 4267:"Fossil palm beetles refine upland winter temperatures in the Early Eocene Climatic Optimum" 4247: 4210: 4171: 4136: 4107: 4068: 4038: 3999: 3957: 3948:, a new juglandaceous winged fruit from the Eocene and Oligocene of Western North America". 3918: 3868: 3826: 3795: 3756: 3696: 3624: 3584: 3547: 3539: 3504: 3469: 3415: 3373: 3335: 3274: 3238: 3148:
Dawson, G. M. (1877). "Report on explorations in the southern portion of British Columbia".
3089: 3079: 3040: 1687: 1085: 915: 648: 343: 145: 136: 112: 80: 3395: 2875: 2706: 2690: 2072: 1660: 1444: 1081: 663: 427: 141: 132: 3988:"Newly recognized diversity in Trochodendraceae from the Eocene of western North America" 4326:"The ecology and distribution of Cenozoic Eomeropidae (Mecoptera), and a new species of 4282: 4243: 3864: 3580: 3535: 3411: 3369: 3309:
Journal of the Faculty of Science, Hokkaido University. Series 4, Geology and Mineralogy
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The fossil insects are particularly diverse and well preserved, and include an extinct
1007: 686: 624: 607: 598: 250: 116: 96: 3436:
Archibald, S.B.; Greenwood, D.R.; Smith, R.Y.; Mathewes, R.W.; Basinger, J.F. (2012).
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raise internal and external awareness of the framework and the importance of fossils;
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of plants. Fossil plants described from the fossil beds include rare fruits such as
1824: 1714: 1676: 1576: 1565: 1380: 1369: 900: 878: 415: 401: 378: 283: 103:, also known for their well preserved plant, insect and fish fossils, are found at 72: 4251: 4043: 4026: 3278: 2943:
build knowledge of the nature and extent of the resource in British Columbia; and
3111:"Fossil bed reopens under new Indigenous management in B.C.'s southern Interior" 3012:
Wilson, M.V.H. (1977). "Middle Eocene freshwater fishes from British Columbia".
2903: 2839: 2612: 2487: 2449: 2411: 2246: 2235: 2222: 2211: 2158: 2056: 2003: 1936: 1786: 1649: 1532: 1497: 1462: 1425: 1407: 1396: 1217: 1187: 1158: 1060: 1050: 1037: 870: 820: 797: 782: 527: 497: 448: 421: 334: 261: 196: 3986:
Manchester, S.; Pigg, K. B.; KvaÄŤek, Z; DeVore, M. L.; Dillhoff, R. M. (2018).
3205:
Paleobotany of the Tertiary (early Middle Eocene) McAbee Beds, British Columbia
2883: 2663: 2626: 2465: 2427: 2389: 2339: 2314: 2253: 2097: 2038: 1911: 1869: 1813: 1759: 1614: 1525: 1490: 1455: 1418: 1281: 1257: 1205: 1046: 894: 847: 757: 548: 323: 322:, a genus of trees now endemic to China, extinct members of the birch family ( 192: 36: 23: 4291: 3303:
Wolfe, J.A.; Tanai, T. (1987). "Systematics, Phylogeny, and Distribution of
2767: 2731: 2364: 2199: 2147: 1806: 1632: 1621: 1603: 1592: 1342: 1335: 1198: 1146: 1074: 1070: 826: 733: 710: 616: 386: 100: 4310: 3718: 3543: 3155:. Vol. 1875–1876. Montreal, Quebec: Dawson Brothers. pp. 233–265. 3084: 3059: 441:
cypress, Chinese fir, dawn redwood, California redwood, red or white cedar
4330:
Cockerell from the Early Eocene McAbee locality, British Columbia, Canada"
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LePage, B.A.; Basinger, J.F. (1995). "Evolutionary history of the genus
3636: 3610:"Seasonality, the latitudinal gradient of diversity, and Eocene insects" 4175: 3969: 3710: 3607:
Archibald, S.B.; Bossert, W.H.; Greenwood, D.R.; Farrell, B.D. (2010).
2828: 2046: 2031: 1877: 1741: 1270: 1139: 855: 533: 230: 221: 4215: 4198: 4140: 3552: 3168:
Palynology and age of early Tertiary basins, interior British Columbia
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Greenwood, D.R.; Archibald, S.B.; Mathewes, R.W.; Moss, P.T. (2005).
2895: 1353: 1089: 832: 562: 357: 313: 237: 215: 4265:
Archibald, S.B.; Morse, G.; Greenwood, D.R.; Mathewes, R.W. (2014).
3961: 3872: 3830: 3799: 3701: 3680: 3628: 3588: 3419: 3339: 3242: 266: 4072: 4003: 3922: 3760: 3058:
Greenwood, D.R.; Pigg, K.B.; Basinger, J.F.; DeVore, M.L. (2016).
991: 671: 556: 503: 491: 485: 479: 433: 339: 293: 233: 166: 58: 50: 1073:) were detailed from the beds, confirming the presence of palms ( 3170:(Ph.D.). Edmonton, Alberta: University of Alberta. p. 189. 3892:
Burke Museum Contributions in Anthropology and Natural History
2931:
clarify the rules governing the management and use of fossils;
271:; however, a 2005 paper provided a radiometric date using the 244:
The climate of the McAbee Eocene lake was reconstructed to be
3207:(M.Sc). Calgary, Alberta: University of Calgary. p. 128. 256:
A volcanic ash exposed in the lake shale beds was originally
175:
Fossil plants from the same area as the McAbee fossil beds (
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10.1603/0013-8746(2005)098[0503:EADOCE]2.0.CO;2
3378:
10.1130/0091-7613(1995)023<1044:eccalt>2.3.co;2
3179: 3177: 4324:
Archibald, S.B.; Rasnitsyn, A.P.; Akhmetiev, M.A. (2005).
195:(spores and pollen) and leaf fossils, and research on the 4436:
10.1603/0013-8746(2006)99[487:BAOTEO]2.0.CO;2
4113:
10.1603/0013-8746(2006)99[487:BAOTEO]2.0.CO;2
3844:
Dillhoff, R.M.; Leopold, E.B.; Manchester, S.R. (2005).
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provide for the stewardship of significant fossil sites;
282:, with both dates placing the McAbee fossil beds in the 4414:
Archibald, S.B.; Cover, S. P.; Moreau, C. S. (2006).
4094:
Archibald, S.B.; Cover, S. P.; Moreau, C. S. (2006).
4197:
Feldmann, R.A.; Schweitzer, C.E.; Leahy, J. (2011).
3307:(maples) in the Cenozoic of Western North America". 2906:; as well as in university collections, principally 908:
maples, buckeyes/horse chestnuts, golden rain trees
83:, Canada, just north of and visible from Provincial 4025:Manchester, S. R.; KvaÄŤek, Z.; Judd, W. S. (2020). 155:The McAbee site is now under the management of the 4461:Fossil Management Framework (accessed May 4, 2015) 4409: 4407: 4405: 4403: 4401: 4399: 4397: 4395: 3735:Pigg, K.B.; Manchester, S.R.; Wehr, W.C. (2003). " 3608: 4359: 4357: 4232:Palaeogeography, Palaeoclimatology, Palaeoecology 3259:Palaeogeography, Palaeoclimatology, Palaeoecology 3014:Life Sciences Contributions, Royal Ontario Museum 2967:"McAbee fossil site receives heritage protection" 3462:Geological Society, London, Special Publications 3431: 3429: 3389: 3387: 3351: 3349: 2934:manage impacts on fossils from other activities; 1077:) in the local environment in the early Eocene. 4479:Cenozoic paleontological sites of North America 4271:Proceedings of the National Academy of Sciences 3981: 3979: 3602: 3600: 3598: 4423:Annals of the Entomological Society of America 4334:Annals of the Entomological Society of America 4100:Annals of the Entomological Society of America 16:Fossil bed in the Interior of British Columbia 280: ± 0.83 million years ago 148:, and related Eocene fossil sites such as at 8: 3298: 3296: 3216: 3214: 3007: 3005: 1063:). A species of fossil freshwater crayfish ( 3730: 3728: 2900:Burke Museum of Natural History and Culture 199:from the fossil beds by Mark Wilson of the 55:McAbee Fossil Beds viewed from the Highway. 3782:Manchester, S.R.; Dillhoff, R.M. (2004). " 1080:Below is an incomplete list of the insect 308:collection. Collected by L. Hills in 1983. 4434: 4345: 4300: 4290: 4214: 4111: 4042: 3700: 3551: 3093: 3083: 2874:Collections of fossils are housed in the 511:fir, spruce, pine, golden larch, hemlock 356:Below is an incomplete list of the plant 4031:Botanical Journal of the Linnean Society 3679:McClain, A.M.; Manchester, S.R. (2001). 3660:(Report). Government of British Columbia 362: 275:method places the McAbee Fossil Beds at 4061:International Journal of Plant Sciences 3992:International Journal of Plant Sciences 3911:International Journal of Plant Sciences 3749:International Journal of Plant Sciences 2958: 3648: 3646: 3186:Canadian Petroleum Geologists Bulletin 3033:Review of Palaeobotany and Palynology 2880:Royal Tyrrell Museum of Palaeontology 2657:Archibald, Cover, & Moreau, 2006 2620:Archibald, Cover, & Moreau, 2006 2586:Archibald, Cover, & Moreau, 2006 2552:Archibald, Cover, & Moreau, 2006 2518:Archibald, Cover, & Moreau, 2006 2482:Archibald, Cover, & Moreau, 2006 2444:Archibald, Cover, & Moreau, 2006 2406:Archibald, Cover, & Moreau, 2006 805:extinct sycamore/plane tree relative 306:Royal Tyrrell Museum of Palaeontology 7: 2996:Canadian Register of Historic Places 541:catkin yew, nutmeg yew or 'torreya' 123:in central British Columbia, and at 71:is a Heritage Site that protects an 4484:Natural history of British Columbia 3653:Wilson, M.V.H. (October 24, 2008). 4524:Heritage sites in British Columbia 3853:Canadian Journal of Earth Sciences 3813:Denk, T.; Dillhoff, R.M. (2005). " 3569:Canadian Journal of Earth Sciences 3524:Canadian Journal of Earth Sciences 3400:Canadian Journal of Earth Sciences 3223:Canadian Journal of Earth Sciences 3131:"McAbee Fossil Beds Heritage Site" 3064:Canadian Journal of Earth Sciences 953:, extinct trochodendron relatives 183:B.C.) were first reported by G.M. 14: 4514:Thompson-Nicola Regional District 3109:Wilson, Deborah (June 21, 2019). 2870:Collections and collecting status 1024:Flower of uncertain affliliation 4534:Paleontology in British Columbia 3886:Schorn, H.; Wehr, W.C. (1986). " 2991:McAbee Fossil Beds Heritage Site 2748:Archibald & Rasnitsyn, 2015 2295:Archibald & Rasnitsyn, 2015 105:Driftwood Canyon Provincial Park 1045:, a species of green lacewing ( 840:serviceberry, hawthorn, cherry 576:alder, birch, extinct hornbeam 191:and his students on the fossil 1: 4450:Royal British Columbia Museum 4203:Journal of Crustacean Biology 3655:McAbee Fossil Site Assessment 3203:Verschoor, K. van R. (1974). 2999:. Retrieved 24 October 2014. 703:extinct witch hazel relative 304:from the McAbee Fossil Beds. 4529:Eocene paleontological sites 4252:10.1016/j.palaeo.2012.10.043 3279:10.1016/j.palaeo.2018.09.010 3135:Province of British Columbia 3045:10.1016/0034-6667(91)90085-H 1032:Insects and other arthropods 338:), fruits and leaves of the 3328:Canadian Journal of Zoology 3151:Geological Survey of Canada 2288:Cuspilongus cachecreekensis 171:Indeterminate bird skeleton 4550: 4044:10.1093/botlinnean/boaa046 3944:Manchester, S.R. (1991). " 3819:Canadian Journal of Botany 3788:Canadian Journal of Botany 3689:American Journal of Botany 2908:Thompson Rivers University 4366:The Canadian Entomologist 4164:The Canadian Entomologist 2892:Canadian Museum of Nature 163:Palaeontology and geology 3153:, Report of Progress for 2083:long-bodied crane flies 1797:tumbling flower beetles 925:Paraconcavistylon wehrii 750:extinct mallow relative 464:ginkgo, maidenhair tree 135:fossil beds in southern 75:fossil locality east of 37:50.797183°N 121.141150°W 4292:10.1073/pnas.1323269111 4129:Journal of Paleontology 2878:in Victoria, B.C.; the 2741:Ypresiosirex orthosemos 2650:Myrmeciites herculeanus 2579:Ypresiomyrma orbiculata 1887:panorpid scorpionflies 1066:Aenigmastacus crandalli 1056:Archaeochrysa profracta 931:Trochodendron drachukii 269: million years ago 4504:Paleontology in Canada 4489:Fossil parks in Canada 3544:10.1139/cjes-2015-0160 3085:10.1139/cjes-2015-0177 2613:Myrmeciites(?) goliath 2545:Ypresiomyrma bartletti 2022:eomeropid mecopterans 1059:), and stick insects ( 309: 251:Konservat-Lagerstätten 172: 157:Bonaparte First Nation 64: 56: 42:50.797183; -121.141150 3509:10.1139/cjb-2012-0313 2399:Avitomyrmex elongatus 2108:limoniid crane flies 297: 258:radiometrically dated 201:University of Alberta 189:University of Calgary 170: 131:, United States. The 62: 54: 4519:2012 in paleontology 3909:Gordon (Pinaceae)". 3166:Hills, L.V. (1965). 2888:Royal Ontario Museum 2475:Avitomyrmex systenus 1102:Super family/family 1084:, superfamilies and 1017:Dillhoffia cashensis 937:Trochodendron nastae 863:cottonwoods/poplars 766:Comptonia columbiana 89:Trans-Canada Highway 63:Heritage status sign 4378:10.4039/tce.2015.55 4283:2014PNAS..111.8095A 4244:2013PPP...371....1A 3865:2005CaJES..42..151D 3581:2005CaJES..42..187M 3536:2016CaJES..53..591G 3412:2005CaJES..42..167G 3370:1995Geo....23.1044G 3271:2018PPP...511..433L 3235:1981CaJES..18.1464E 3076:2016CaJES..53..548G 2803:tenthredinid wasps 2717:proctotrupid wasps 2185:winter crane flies 1688:long horned beetles 1633:reticulated beetles 969:Ulmus okanaganensis 679:currant/gooseberry 350:Ulmus okanaganensis 33: /  4176:10.4039/tce.2015.2 2916:Brandon University 2511:Macabeemyrma ovata 2437:Avitomyrmex mastax 1840:†Cimbrophlebiidae 1247:harvester termites 1042:Macabeemyrma ovata 719:Sassafras hesperia 696:Langeria magnifica 319:Dipteronia brownii 310: 301:Sassafras hesperia 173: 69:McAbee Fossil Beds 65: 57: 4277:(22): 8095–8100. 4216:10.1651/10-3399.1 4141:10.1666/12-052R.1 3950:Systematic Botany 3825:(12): 1663–1681. 3794:(10): 1509–1517. 3442:Geoscience Canada 3364:(11): 1044–1048. 2867: 2866: 1604:osmylid lacewings 1289:Prophalangopsidae 1176:Megapodagrionidae 1029: 1028: 584:Cercidiphyllaceae 369:genus or species 205:lithostratigraphy 4541: 4509:Thompson Country 4463: 4458: 4452: 4447: 4441: 4440: 4438: 4420: 4411: 4390: 4389: 4361: 4352: 4351: 4349: 4321: 4315: 4314: 4304: 4294: 4262: 4256: 4255: 4227: 4221: 4220: 4218: 4194: 4188: 4187: 4159: 4153: 4152: 4124: 4118: 4117: 4115: 4091: 4085: 4084: 4055: 4049: 4048: 4046: 4022: 4016: 4015: 3983: 3974: 3973: 3941: 3935: 3934: 3902: 3896: 3895: 3883: 3877: 3876: 3850: 3841: 3835: 3834: 3810: 3804: 3803: 3779: 3773: 3772: 3732: 3723: 3722: 3704: 3695:(7): 1316–1325. 3676: 3670: 3669: 3667: 3665: 3659: 3650: 3641: 3640: 3612: 3604: 3593: 3592: 3564: 3558: 3557: 3555: 3519: 3513: 3512: 3492: 3486: 3485: 3474:10.1144/SP295.13 3456: 3450: 3449: 3433: 3424: 3423: 3391: 3382: 3381: 3353: 3344: 3343: 3334:(6): 1140–1157. 3323: 3317: 3316: 3300: 3291: 3290: 3253: 3247: 3246: 3229:(9): 1464–1477. 3218: 3209: 3208: 3200: 3194: 3193: 3181: 3172: 3171: 3163: 3157: 3156: 3145: 3139: 3138: 3127: 3121: 3120: 3106: 3100: 3099: 3097: 3087: 3055: 3049: 3048: 3039:(1–2): 173–185. 3028: 3022: 3021: 3009: 3000: 2988: 2982: 2981: 2979: 2978: 2969:. Archived from 2963: 1744:, snout beetles 1551:Green lacewings 1516:Green lacewings 1481:Green lacewings 1099:Arthropod order 1096: 916:Trochodendraceae 649:Fagus langevinii 394:Scouring rushes 363: 344:Fagus langevinii 281: 270: 146:Driftwood Canyon 137:British Columbia 113:British Columbia 81:British Columbia 48: 47: 45: 44: 43: 38: 34: 31: 30: 29: 26: 4549: 4548: 4544: 4543: 4542: 4540: 4539: 4538: 4469: 4468: 4467: 4466: 4459: 4455: 4448: 4444: 4418: 4413: 4412: 4393: 4363: 4362: 4355: 4323: 4322: 4318: 4264: 4263: 4259: 4229: 4228: 4224: 4196: 4195: 4191: 4161: 4160: 4156: 4126: 4125: 4121: 4093: 4092: 4088: 4057: 4056: 4052: 4024: 4023: 4019: 3985: 3984: 3977: 3962:10.2307/2418873 3943: 3942: 3938: 3904: 3903: 3899: 3885: 3884: 3880: 3873:10.1139/e04-084 3848: 3843: 3842: 3838: 3831:10.1139/b05-122 3812: 3811: 3807: 3800:10.1139/b04-112 3781: 3780: 3776: 3734: 3733: 3726: 3702:10.2307/3558343 3678: 3677: 3673: 3663: 3661: 3657: 3652: 3651: 3644: 3629:10.1666/09021.1 3606: 3605: 3596: 3589:10.1139/E04-095 3566: 3565: 3561: 3521: 3520: 3516: 3494: 3493: 3489: 3458: 3457: 3453: 3435: 3434: 3427: 3420:10.1139/E04-100 3393: 3392: 3385: 3355: 3354: 3347: 3340:10.1139/z96-126 3325: 3324: 3320: 3302: 3301: 3294: 3255: 3254: 3250: 3243:10.1139/e81-137 3220: 3219: 3212: 3202: 3201: 3197: 3183: 3182: 3175: 3165: 3164: 3160: 3147: 3146: 3142: 3129: 3128: 3124: 3108: 3107: 3103: 3057: 3056: 3052: 3030: 3029: 3025: 3011: 3010: 3003: 2989: 2985: 2976: 2974: 2965: 2964: 2960: 2955: 2876:Royal BC Museum 2872: 2753:horntail wasps 2691:ichneumon wasps 2350:diapriid wasps 2325:cimbicid wasps 2264:braconid wasps 2073:Cylindrotomidae 1989:extinct family 1955:extinct family 1921:extinct family 1661:soldier beetles 1577:brown lacewings 1472:Okanaganochrysa 1445:Green lacewings 1034: 664:Grossulariaceae 298:Fossil leaf of 292: 276: 265: 165: 142:Princeton Chert 133:Princeton Chert 99:in Eocene lake 41: 39: 35: 32: 27: 24: 22: 20: 19: 17: 12: 11: 5: 4547: 4545: 4537: 4536: 4531: 4526: 4521: 4516: 4511: 4506: 4501: 4496: 4491: 4486: 4481: 4471: 4470: 4465: 4464: 4453: 4442: 4429:(3): 487–523. 4391: 4372:(2): 209–228. 4353: 4340:(4): 503–514. 4316: 4257: 4222: 4209:(2): 320–331. 4189: 4170:(6): 744–753. 4154: 4119: 4106:(3): 487–523. 4086: 4073:10.1086/512104 4067:(4): 521–532. 4050: 4037:(3): 467–484. 4017: 4004:10.1086/699282 3998:(8): 663–676. 3975: 3956:(4): 715–725. 3936: 3923:10.1086/297313 3917:(6): 910–950. 3897: 3878: 3859:(2): 151–166. 3836: 3805: 3774: 3761:10.1086/376816 3755:(5): 807–822. 3745:Palaeocarpinus 3724: 3671: 3642: 3623:(3): 374–398. 3594: 3575:(2): 187–204. 3559: 3530:(6): 591–604. 3514: 3503:(8): 514–529. 3487: 3451: 3425: 3406:(2): 167–185. 3383: 3345: 3318: 3292: 3248: 3210: 3195: 3173: 3158: 3140: 3122: 3101: 3070:(6): 548–564. 3050: 3023: 3001: 2983: 2957: 2956: 2954: 2951: 2950: 2949: 2948: 2947: 2944: 2941: 2938: 2935: 2932: 2871: 2868: 2865: 2864: 2862: 2855: 2853: 2851: 2843: 2835: 2834: 2832: 2825: 2823: 2821: 2814: 2808: 2807: 2805: 2800: 2798: 2796: 2793:Tenthredinidae 2789: 2783: 2782: 2780: 2778:Sphecid wasps 2775: 2773: 2771: 2764: 2758: 2757: 2755: 2750: 2745: 2735: 2728: 2722: 2721: 2719: 2714: 2712: 2710: 2707:Proctotrupidae 2703: 2697: 2696: 2694: 2687: 2685: 2683: 2676: 2670: 2669: 2667: 2659: 2654: 2644: 2639: 2633: 2632: 2630: 2622: 2617: 2607: 2602: 2596: 2595: 2593: 2588: 2583: 2573: 2568: 2562: 2561: 2559: 2554: 2549: 2539: 2534: 2528: 2527: 2525: 2520: 2515: 2505: 2500: 2494: 2493: 2491: 2484: 2479: 2469: 2462: 2456: 2455: 2453: 2446: 2441: 2431: 2424: 2418: 2417: 2415: 2408: 2403: 2393: 2386: 2380: 2379: 2377: 2375:figitid wasps 2372: 2370: 2368: 2361: 2355: 2354: 2352: 2347: 2345: 2343: 2336: 2330: 2329: 2327: 2322: 2320: 2318: 2311: 2305: 2304: 2302: 2297: 2292: 2282: 2275: 2269: 2268: 2266: 2261: 2259: 2257: 2250: 2242: 2241: 2239: 2232: 2230: 2228: 2226: 2218: 2217: 2215: 2210:flower flies, 2207: 2205: 2203: 2196: 2190: 2189: 2187: 2182: 2180: 2178: 2171: 2165: 2164: 2162: 2155: 2153: 2151: 2144: 2138: 2137: 2135: 2130: 2128: 2126: 2123:Mycetophilidae 2119: 2113: 2112: 2110: 2105: 2103: 2101: 2094: 2088: 2087: 2085: 2080: 2078: 2076: 2069: 2063: 2062: 2060: 2053: 2051: 2042: 2035: 2027: 2026: 2024: 2019: 2017: 2007: 2000: 1994: 1993: 1991: 1986: 1984: 1974: 1966: 1960: 1959: 1957: 1952: 1950: 1940: 1932: 1926: 1925: 1923: 1918: 1916: 1906: 1903:Dinopanorpidae 1898: 1892: 1891: 1889: 1884: 1882: 1873: 1866: 1860: 1859: 1857: 1855:extinct group 1852: 1850: 1842: 1837: 1831: 1830: 1828: 1821: 1819: 1817: 1810: 1802: 1801: 1799: 1794: 1792: 1790: 1783: 1777: 1776: 1774: 1767: 1765: 1763: 1755: 1749: 1748: 1746: 1738: 1736: 1734: 1727: 1721: 1720: 1718: 1711: 1709: 1707: 1700: 1694: 1693: 1691: 1684: 1682: 1680: 1673: 1667: 1666: 1664: 1657: 1655: 1653: 1645: 1639: 1638: 1636: 1629: 1627: 1625: 1618: 1610: 1609: 1607: 1600: 1598: 1596: 1589: 1583: 1582: 1580: 1573: 1571: 1569: 1562: 1556: 1555: 1553: 1548: 1546: 1536: 1529: 1521: 1520: 1518: 1513: 1511: 1501: 1494: 1486: 1485: 1483: 1478: 1476: 1466: 1459: 1451: 1450: 1448: 1441: 1439: 1429: 1422: 1414: 1413: 1411: 1404: 1402: 1400: 1393: 1387: 1386: 1384: 1377: 1375: 1373: 1366: 1360: 1359: 1357: 1350: 1348: 1346: 1339: 1331: 1330: 1328: 1321: 1319: 1317: 1310: 1304: 1303: 1301: 1296: 1294: 1292: 1285: 1277: 1276: 1274: 1267: 1265: 1263: 1261: 1253: 1252: 1250: 1243: 1241: 1239: 1236:Hodotermitidae 1232: 1224: 1223: 1221: 1213: 1211: 1209: 1202: 1194: 1193: 1191: 1183: 1181: 1179: 1172: 1166: 1165: 1163: 1154: 1152: 1150: 1143: 1135: 1134: 1132: 1130: 1128: 1126: 1124: 1116: 1115: 1112: 1109: 1106: 1105:Genus/Species 1103: 1100: 1033: 1030: 1027: 1026: 1021: 1012: 1008:incertae sedis 1002: 1001: 996: 987: 979: 978: 973: 964: 956: 955: 947: 919: 911: 910: 905: 874: 866: 865: 860: 851: 843: 842: 837: 816: 808: 807: 802: 786: 778: 777: 770: 761: 753: 752: 747: 737: 729: 728: 723: 714: 706: 705: 700: 690: 687:Hamamelidaceae 682: 681: 676: 667: 659: 658: 653: 643: 635: 634: 629: 620: 612: 611: 603: 599:Cercidiphyllum 587: 579: 578: 573: 569:Palaeocarpinus 552: 544: 543: 538: 522: 514: 513: 508: 475: 467: 466: 461: 452: 444: 443: 438: 405: 397: 396: 391: 382: 374: 373: 370: 367: 329:Palaeocarpinus 291: 288: 227:Eohiodon rosei 164: 161: 117:Horsefly River 15: 13: 10: 9: 6: 4: 3: 2: 4546: 4535: 4532: 4530: 4527: 4525: 4522: 4520: 4517: 4515: 4512: 4510: 4507: 4505: 4502: 4500: 4497: 4495: 4494:Eocene plants 4492: 4490: 4487: 4485: 4482: 4480: 4477: 4476: 4474: 4462: 4457: 4454: 4451: 4446: 4443: 4437: 4432: 4428: 4424: 4417: 4410: 4408: 4406: 4404: 4402: 4400: 4398: 4396: 4392: 4387: 4383: 4379: 4375: 4371: 4367: 4360: 4358: 4354: 4348: 4343: 4339: 4335: 4331: 4329: 4320: 4317: 4312: 4308: 4303: 4298: 4293: 4288: 4284: 4280: 4276: 4272: 4268: 4261: 4258: 4253: 4249: 4245: 4241: 4237: 4233: 4226: 4223: 4217: 4212: 4208: 4204: 4200: 4193: 4190: 4185: 4181: 4177: 4173: 4169: 4165: 4158: 4155: 4150: 4146: 4142: 4138: 4134: 4130: 4123: 4120: 4114: 4109: 4105: 4101: 4097: 4090: 4087: 4082: 4078: 4074: 4070: 4066: 4062: 4054: 4051: 4045: 4040: 4036: 4032: 4028: 4021: 4018: 4013: 4009: 4005: 4001: 3997: 3993: 3989: 3982: 3980: 3976: 3971: 3967: 3963: 3959: 3955: 3951: 3947: 3940: 3937: 3932: 3928: 3924: 3920: 3916: 3912: 3908: 3901: 3898: 3893: 3889: 3888:Abies milleri 3882: 3879: 3874: 3870: 3866: 3862: 3858: 3854: 3847: 3840: 3837: 3832: 3828: 3824: 3820: 3816: 3809: 3806: 3801: 3797: 3793: 3789: 3785: 3778: 3775: 3770: 3766: 3762: 3758: 3754: 3750: 3746: 3742: 3738: 3731: 3729: 3725: 3720: 3716: 3712: 3708: 3703: 3698: 3694: 3690: 3686: 3684: 3675: 3672: 3656: 3649: 3647: 3643: 3638: 3634: 3630: 3626: 3622: 3618: 3617: 3611: 3603: 3601: 3599: 3595: 3590: 3586: 3582: 3578: 3574: 3570: 3563: 3560: 3554: 3549: 3545: 3541: 3537: 3533: 3529: 3525: 3518: 3515: 3510: 3506: 3502: 3498: 3491: 3488: 3483: 3479: 3475: 3471: 3467: 3463: 3455: 3452: 3448:(4): 155–164. 3447: 3443: 3439: 3432: 3430: 3426: 3421: 3417: 3413: 3409: 3405: 3401: 3397: 3390: 3388: 3384: 3379: 3375: 3371: 3367: 3363: 3359: 3352: 3350: 3346: 3341: 3337: 3333: 3329: 3322: 3319: 3314: 3310: 3306: 3299: 3297: 3293: 3288: 3284: 3280: 3276: 3272: 3268: 3264: 3260: 3252: 3249: 3244: 3240: 3236: 3232: 3228: 3224: 3217: 3215: 3211: 3206: 3199: 3196: 3191: 3187: 3180: 3178: 3174: 3169: 3162: 3159: 3154: 3152: 3144: 3141: 3136: 3132: 3126: 3123: 3118: 3117: 3112: 3105: 3102: 3096: 3091: 3086: 3081: 3077: 3073: 3069: 3065: 3061: 3054: 3051: 3046: 3042: 3038: 3034: 3027: 3024: 3019: 3015: 3008: 3006: 3002: 2998: 2997: 2992: 2987: 2984: 2973:on 2013-03-12 2972: 2968: 2962: 2959: 2952: 2945: 2942: 2939: 2936: 2933: 2930: 2929: 2928: 2927: 2926: 2923: 2921: 2917: 2913: 2909: 2905: 2901: 2897: 2893: 2889: 2886:Alberta; the 2885: 2881: 2877: 2869: 2863: 2861: 2860: 2859:stick insects 2856: 2854: 2852: 2850: 2849: 2848:Susumanioidea 2844: 2842: 2841: 2837: 2836: 2833: 2831: 2830: 2826: 2824: 2822: 2820: 2819: 2815: 2813: 2810: 2809: 2806: 2804: 2801: 2799: 2797: 2795: 2794: 2790: 2788: 2785: 2784: 2781: 2779: 2776: 2774: 2772: 2770: 2769: 2765: 2763: 2760: 2759: 2756: 2754: 2751: 2749: 2746: 2744: 2743: 2742: 2736: 2734: 2733: 2729: 2727: 2724: 2723: 2720: 2718: 2715: 2713: 2711: 2709: 2708: 2704: 2702: 2699: 2698: 2695: 2693: 2692: 2688: 2686: 2684: 2682: 2681: 2680:Ichneumonidae 2677: 2675: 2672: 2671: 2668: 2666: 2665: 2660: 2658: 2655: 2653: 2652: 2651: 2645: 2643: 2640: 2638: 2635: 2634: 2631: 2629: 2628: 2623: 2621: 2618: 2616: 2615: 2614: 2608: 2606: 2603: 2601: 2598: 2597: 2594: 2592: 2591:bulldog ants 2589: 2587: 2584: 2582: 2581: 2580: 2574: 2572: 2569: 2567: 2564: 2563: 2560: 2558: 2557:bulldog ants 2555: 2553: 2550: 2548: 2547: 2546: 2540: 2538: 2535: 2533: 2530: 2529: 2526: 2524: 2523:bulldog ants 2521: 2519: 2516: 2514: 2513: 2512: 2506: 2504: 2501: 2499: 2496: 2495: 2492: 2490: 2489: 2485: 2483: 2480: 2478: 2477: 2476: 2470: 2468: 2467: 2463: 2461: 2458: 2457: 2454: 2452: 2451: 2447: 2445: 2442: 2440: 2439: 2438: 2432: 2430: 2429: 2425: 2423: 2420: 2419: 2416: 2414: 2413: 2409: 2407: 2404: 2402: 2401: 2400: 2394: 2392: 2391: 2387: 2385: 2382: 2381: 2378: 2376: 2373: 2371: 2369: 2367: 2366: 2362: 2360: 2357: 2356: 2353: 2351: 2348: 2346: 2344: 2342: 2341: 2337: 2335: 2332: 2331: 2328: 2326: 2323: 2321: 2319: 2317: 2316: 2312: 2310: 2307: 2306: 2303: 2301: 2298: 2296: 2293: 2291: 2290: 2289: 2283: 2281: 2280: 2276: 2274: 2271: 2270: 2267: 2265: 2262: 2260: 2258: 2256: 2255: 2251: 2249: 2248: 2244: 2243: 2240: 2238: 2237: 2233: 2231: 2229: 2227: 2225: 2224: 2220: 2219: 2216: 2214: 2213: 2208: 2206: 2204: 2202: 2201: 2197: 2195: 2192: 2191: 2188: 2186: 2183: 2181: 2179: 2177: 2176: 2175:Trichoceridae 2172: 2170: 2167: 2166: 2163: 2161: 2160: 2156: 2154: 2152: 2150: 2149: 2145: 2143: 2140: 2139: 2136: 2134: 2133:fungus gnats 2131: 2129: 2127: 2125: 2124: 2120: 2118: 2115: 2114: 2111: 2109: 2106: 2104: 2102: 2100: 2099: 2095: 2093: 2090: 2089: 2086: 2084: 2081: 2079: 2077: 2075: 2074: 2070: 2068: 2065: 2064: 2061: 2059: 2058: 2054: 2052: 2050: 2049: 2048: 2043: 2041: 2040: 2036: 2034: 2033: 2029: 2028: 2025: 2023: 2020: 2018: 2016: 2015: 2014: 2008: 2006: 2005: 2001: 1999: 1996: 1995: 1992: 1990: 1987: 1985: 1983: 1982: 1981: 1975: 1973: 1972: 1967: 1965: 1962: 1961: 1958: 1956: 1953: 1951: 1949: 1948: 1947: 1941: 1939: 1938: 1933: 1931: 1928: 1927: 1924: 1922: 1919: 1917: 1915: 1914: 1913: 1907: 1905: 1904: 1899: 1897: 1894: 1893: 1890: 1888: 1885: 1883: 1881: 1880: 1879: 1874: 1872: 1871: 1867: 1865: 1862: 1861: 1858: 1856: 1853: 1851: 1849: 1848: 1847:Cimbrophlebia 1843: 1841: 1838: 1836: 1833: 1832: 1829: 1827: 1826: 1822: 1820: 1818: 1816: 1815: 1811: 1809: 1808: 1804: 1803: 1800: 1798: 1795: 1793: 1791: 1789: 1788: 1784: 1782: 1779: 1778: 1775: 1773: 1772: 1771:click beetles 1768: 1766: 1764: 1762: 1761: 1756: 1754: 1751: 1750: 1747: 1745: 1743: 1739: 1737: 1735: 1733: 1732: 1731:Curculionidae 1728: 1726: 1723: 1722: 1719: 1717: 1716: 1712: 1710: 1708: 1706: 1705: 1704:Chrysomelidae 1701: 1699: 1696: 1695: 1692: 1690: 1689: 1685: 1683: 1681: 1679: 1678: 1674: 1672: 1669: 1668: 1665: 1663: 1662: 1658: 1656: 1654: 1652: 1651: 1646: 1644: 1641: 1640: 1637: 1635: 1634: 1630: 1628: 1626: 1624: 1623: 1619: 1617: 1616: 1612: 1611: 1608: 1606: 1605: 1601: 1599: 1597: 1595: 1594: 1590: 1588: 1585: 1584: 1581: 1579: 1578: 1574: 1572: 1570: 1568: 1567: 1563: 1561: 1558: 1557: 1554: 1552: 1549: 1547: 1545: 1544: 1543: 1542:Archaeochrysa 1537: 1535: 1534: 1530: 1528: 1527: 1523: 1522: 1519: 1517: 1514: 1512: 1510: 1509: 1508: 1502: 1500: 1499: 1495: 1493: 1492: 1488: 1487: 1484: 1482: 1479: 1477: 1475: 1474: 1473: 1467: 1465: 1464: 1460: 1458: 1457: 1453: 1452: 1449: 1447: 1446: 1442: 1440: 1438: 1437: 1436: 1430: 1428: 1427: 1423: 1421: 1420: 1416: 1415: 1412: 1410: 1409: 1405: 1403: 1401: 1399: 1398: 1394: 1392: 1389: 1388: 1385: 1383: 1382: 1378: 1376: 1374: 1372: 1371: 1367: 1365: 1362: 1361: 1358: 1356: 1355: 1351: 1349: 1347: 1345: 1344: 1340: 1338: 1337: 1333: 1332: 1329: 1327: 1326: 1322: 1320: 1318: 1316: 1315: 1314:Tettigoniidae 1311: 1309: 1306: 1305: 1302: 1300: 1297: 1295: 1293: 1291: 1290: 1286: 1284: 1283: 1279: 1278: 1275: 1273: 1272: 1268: 1266: 1264: 1262: 1260: 1259: 1255: 1254: 1251: 1249: 1248: 1244: 1242: 1240: 1238: 1237: 1233: 1231: 1230: 1226: 1225: 1222: 1220: 1219: 1214: 1212: 1210: 1208: 1207: 1203: 1201: 1200: 1196: 1195: 1192: 1190: 1189: 1184: 1182: 1180: 1178: 1177: 1173: 1171: 1168: 1167: 1164: 1162: 1160: 1155: 1153: 1151: 1149: 1148: 1144: 1142: 1141: 1137: 1136: 1133: 1131: 1129: 1127: 1125: 1123: 1122: 1121:Ephemeroptera 1118: 1117: 1113: 1110: 1107: 1104: 1101: 1098: 1097: 1094: 1091: 1087: 1083: 1078: 1076: 1072: 1068: 1067: 1062: 1058: 1057: 1052: 1048: 1044: 1043: 1039: 1031: 1025: 1022: 1020: 1019: 1018: 1013: 1011: 1010: 1009: 1004: 1003: 1000: 997: 995: 994: 993: 988: 986: 985: 981: 980: 977: 974: 972: 971: 970: 965: 963: 962: 958: 957: 954: 952: 948: 946: 945: 944: 939: 938: 933: 932: 927: 926: 920: 918: 917: 913: 912: 909: 906: 904: 903: 902: 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Index

50°47.831′N 121°8.469′W / 50.797183°N 121.141150°W / 50.797183; -121.141150


Eocene Epoch
Cache Creek
British Columbia
Highway 97
Trans-Canada Highway
Highway 1
fossil beds
sediments
Driftwood Canyon Provincial Park
Smithers
British Columbia
Horsefly River
Quesnel
Republic
Washington
Princeton Chert
British Columbia
Princeton Chert
Driftwood Canyon
Republic
Bonaparte First Nation

Cache Creek
Kamloops
Dawson
University of Calgary
palynology

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