2229:
2235:
2785:
651:
139:
779:
2815:
118:
397:
2791:
731:. It became apparent that the 1-3 frill spikes mentioned above are in fact spikes 2–4. The first epiparietal is small and variably procurving and thus was misinterpreted before. At least one more epiparietal pair was also identified (after the fourth), resulting in a total of at least 5 pairs, consistent with centrosaurines like
678:, which totals at several hundred individual elements, upon its original description only these two partial frills have been scientifically described while the rest of the material was being reexamined. Many of the other elements are not diagnosable to a genus level and can only be confidently referable to
701:
in having only three epiparietals (frill spikes) on each side of the frill. The first frill spike pair is large and the second is smaller, and both are uniquely widened at base, pachyostotic, curve down the sides of the frill, and are depressed down the front of the frill. The third frill spike pair
445:
from a nearby area, however, could not refer the bonebed material to any valid existing taxon. The bonebed, known as the
Mansfield Bonebed honoring its landowner, is located on private land and historically has been excavated by several commercial companies. The type material of
747:, a bone separating the two sides of the frill, was also among the newly described material. It is broad, resulting in rounder and smaller frill fenestrae (holes) like in other centrosaurines. Thus, the new study reassigned
1270:
Ryan, Michael J.; Russell, Anthony P., and
Hartman, Scott. (2010). "A New Chasmosaurine Ceratopsid from the Judith River Formation, Montana", In: Michael J. Ryan, Brenda J. Chinnery-Allgeier, and David A. Eberth (eds),
530:
whose "hair" consists of snakes and its gaze could turn men to stone, alluding to a unique trait of this genus - the large, thick snake-like spikes that extend to the sides of the frill, in combination with
2941:
1424:
gen. et sp. nov., a Boldly
Audacious New Chasmosaurine Ceratopsid (Dinosauria: Ornithischia) from the Judith River Formation (Upper Cretaceous: Campanian) of Montana, USA"
458:. Canada Fossils, Ltd., also assembled two composite skeletons using the Mansfield Bonebed material which are in the collections of the Wyoming Dinosaur Center and the
622:
from the
Mansfield Bonebed, found in 2011 and 2012 by David Trexler, indicating the presence of traits that were characteristic of Centrosaurinae in the skeleton of
1481:
Ryan, M. J.; Evans, D. C.; Currie, P. J.; Loewen, M. A. (2014). "A new chasmosaurine from northern
Laramidia expands frill disparity in ceratopsid dinosaurs".
434:, based on misidentified frill spikes. Trexler and Sweeney (1995) reinterpreted the spikes as eye-socket horncores and noted their similarity to those of the
560:, in reference to the years confusion that surrounded the taxonomic designations of the Mansfield Bonebed material before it was given its own name.
2882:
720:
was thought to differ (like all chasmosaurines) in lacking tab-shaped, frequently overlapping fourth to seventh pairs of the frill ornamentation of
626:. They concluded that all the material could be referred to a single species. The phylogenetic analysis conducted by the authors indicated that
2971:
2946:
2966:
2228:
455:
1550:
428:
of
Montana. The material was first reported by Sweeney and Boyden (1993), who considered it to represent the southernmost occurrence of
2976:
2234:
1345:
Kentaro Chiba; Michael J. Ryan; Federico Fanti; Mark A. Loewen; David C. Evans (2018). "New material and systematic re-evaluation of
1280:
614:
However, Ryan had already indicated that part of the
Mansfield material represented not a chasmosaurine, but a centrosaurine. Chiba
459:
413:
454:
from Canada
Fossils, Ltd., of Calgary, Alberta. Additional material from the same excavation was purchased and accessioned by the
2808:
1238:
607:
by
Longrich (2013), also from the Judith River Formation, but from an area equivalent to the lower Oldman Formation or upper
630:
was not a member of
Chasmosaurinae after all, but rather an early centrosaurine ceratopsid that was more closely related to
2961:
138:
2784:
2951:
702:
is small and triangular, unmodified in comparison to early chasmosaurines, but similarly depressed, and borders the
498:
which had been described by Ryan in 2007. Later, Ryan realized that the Mansfield Bonebed fossils did not belong to
1089:
1072:
1041:
519:
658:
Two partial parietals (frills) that are housed at the Wyoming Dinosaur Center were chosen as the type material of
2956:
430:
2873:
1641:
1543:
908:
2936:
1759:
546:
451:
417:
2835:
1115:
801:
564:
369:
2908:
2818:
1668:
995:
891:
317:
2790:
727:
This was challenged in 2018, by the description of additional Mansfield Bonebed material assignable to
1490:
1435:
1358:
951:
507:
425:
294:
1924:
1536:
2644:
2594:
2026:
1514:
1418:
Jordan C. Mallon; Christopher J. Ott; Peter L. Larson; Edward M. Iuliano; David C. Evans (2016).
1393:
1374:
1314:
608:
133:
2913:
2895:
2727:
2665:
2628:
2102:
1989:
1506:
1463:
1276:
1173:
937:
639:
635:
441:
2900:
2757:
2555:
2380:
2082:
1777:
1498:
1453:
1443:
1366:
1306:
572:
506:
was formally described and name by Michael J. Ryan, Anthony P. Russell and Scott Hartman in
2651:
2603:
2562:
2498:
2401:
2352:
2054:
1978:
1933:
1807:
1793:
1752:
1626:
1610:
828:
733:
712:
650:
581:
557:
527:
486:
366:
1494:
1439:
1362:
2658:
2548:
2535:
2506:
2491:
2461:
2422:
2408:
2325:
2286:
2165:
2075:
2040:
2033:
1913:
1863:
1786:
1766:
1458:
1419:
1145:
870:
833:
778:
752:
703:
698:
568:
535:
477:
421:
409:
356:
273:
257:
215:
1297:, a New Horned Dinosaur from the Middle Campanian Judith River Formation of Montana".
2930:
2736:
2715:
2685:
2676:
2576:
2470:
2415:
2394:
2366:
2359:
2345:
2338:
2144:
2133:
2126:
2068:
2061:
2047:
2005:
1940:
1906:
1877:
1830:
1378:
1318:
1158:
784:
739:
721:
690:
671:
631:
598:
491:
381:
68:
1518:
1273:
New Perspectives on Horned Dinosaurs: The Royal Tyrrell Museum Ceratopsian Symposium
117:
2750:
2701:
2694:
2610:
2583:
2569:
2477:
2450:
2443:
2436:
2387:
2373:
2265:
2252:
2183:
2172:
2157:
2119:
2111:
2093:
1998:
1964:
1947:
1899:
1892:
1848:
1653:
1601:
816:
679:
603:
511:
481:
436:
396:
244:
199:
1349:(Dinosauria, Ceratopsidae) from the Judith River Formation (Campanian, Montana)".
1448:
2867:
2743:
2637:
2520:
2484:
2307:
2019:
1884:
1394:"New horned dinosaur: Two-ton plant-eater lived 78 million years ago in Montana"
810:
450:
and other specimens were excavated more recently and have been purchased by the
43:
2858:
2513:
2429:
2279:
2012:
1800:
1734:
1721:
1680:
1559:
1502:
822:
797:
586:
542:
359:
231:
88:
53:
852:
590:
579:. The bonebed is located at approximately the same level as the holotype of
474:
373:
150:
93:
37:
30:
1510:
1467:
2852:
2314:
2300:
2293:
1856:
1592:
1583:
1310:
667:
663:
362:
183:
170:
83:
78:
63:
58:
48:
1370:
563:
The Mansfield Bonebed material was collected from the upper part of the
480:
Michael J. Ryan of the Cleveland Museum of Natural History in 2003 in a
2887:
2708:
1870:
1814:
826:
based on its unique frill ornamentation. It differs from chasmosaurine
495:
377:
349:
98:
73:
1577:
576:
523:
160:
2829:
766:
In 2010, a length of roughly 6 meters (~20 feet) was estimated for
674:
material from the Mansfield Bonebed was thought to be referable to
777:
649:
532:
395:
352:
782:
Reconstruction of parietals of various centrosaurines, including
553:
2833:
2250:
1719:
1570:
1532:
710:
closely resembles the third pair of the frill ornamentation of
462:, however neither of the casts has an exact reconstruction of
1528:
706:. The highly broadened and curved first frill spike pair of
693:. It was suggested that based on its type material alone,
855:
presented below follows a phylogenetic analysis by Chiba
788:(A) showing 5 epiparietal pairs and broad midline ramus.
125:
Skeleton at Wyoming Dinosaur Museum formerly referred to
840:. All material previously assigned to the centrosaurine
408:
came from a bonebed in the badlands on the west side of
484:. Its fossils were subsequently confused with those of
541:, meaning "horned-face", which is a common suffix for
836:, as evident from newly described squamosal bones of
743:, but not with chasmosaurines. The midline ramus of
2842:
2726:
2684:
2675:
2627:
2593:
2534:
2460:
2324:
2263:
2182:
2143:
2101:
2092:
1988:
1962:
1923:
1828:
1776:
1732:
1679:
1652:
1625:
1392:Cleveland Museum of Natural History (30 May 2010).
1299:Bulletin of the Peabody Museum of Natural History
808:can be directly distinguished from centrosaurine
1266:
1264:
1262:
1260:
1258:
1256:
1254:
1544:
597:was considered to represent the oldest known
8:
2942:Late Cretaceous dinosaurs of North America
2830:
2681:
2540:
2330:
2271:
2260:
2247:
2149:
2098:
1970:
1840:
1836:
1744:
1740:
1729:
1716:
1658:
1631:
1622:
1567:
1551:
1537:
1529:
116:
20:
1457:
1447:
670:WDC DJR 002. Although all the
1340:
1338:
1336:
1334:
1332:
1330:
1328:
1250:
844:from the formation is now assigned to
456:Royal Tyrrell Museum of Palaeontology
7:
1275:, Indiana University Press, 656 pp.
755:, among which it is most similar to
593:stage of the Late Cretaceous. Thus,
1151:
1064:
1057:
1033:
943:
933:
883:
876:
866:
618:in 2017 described new material of
14:
460:Fukui Prefectural Dinosaur Museum
336:Ryan, Russell & Hartman, 2010
2814:
2813:
2789:
2783:
2233:
2227:
1239:Timeline of ceratopsian research
384:. It contains a single species,
137:
848:or considered too fragmentary.
792:itself is marked by letter (F).
685:Upon its original description
666:WDC DJR 001 and the
1:
2972:Ceratopsians of North America
2947:Fossil taxa described in 2010
556:, a troublemaking god in the
293:Ryan, Russell & Hartman,
2967:Late Cretaceous ceratopsians
1449:10.1371/journal.pone.0154218
689:was thought to represent a
466:as it was later described.
2993:
1090:Wendiceratops pinhornensis
1073:Sinoceratops zhuchengensis
1042:Xenoceratops foremostensis
567:, in a region where it is
16:Extinct genus of dinosaurs
2977:Monotypic dinosaur genera
2803:
2781:
2543:
2333:
2274:
2259:
2246:
2224:
2152:
1973:
1843:
1839:
1747:
1743:
1728:
1715:
1661:
1634:
1621:
1566:
1503:10.1007/s00114-014-1183-1
1293:Longrich, N. R. (2013). "
1171:
1156:
1149:
1129:
1112:
1086:
1069:
1062:
1055:
1038:
1031:
1009:
992:
979:
966:
948:
941:
931:
905:
888:
881:
874:
832:based on its less unique
431:Styracosaurus albertensis
323:
316:
134:Scientific classification
132:
124:
115:
23:
909:Machairoceratops cronusi
1760:Micropachycephalosaurus
1351:Journal of Paleontology
859:(2017) who redescribed
452:Wyoming Dinosaur Center
418:Milk River Natural Area
29:Temporal range: Middle
1116:Albertaceratops nesmoi
802:Judith River Formation
793:
655:
601:, until the naming of
565:Judith River Formation
404:The known material of
401:
370:Judith River Formation
2909:Paleobiology Database
996:Nasutoceratops titusi
892:Diabloceratops eatoni
814:, and chasmosaurines
781:
653:
400:Side view of skeleton
399:
2962:Ornithischian genera
2874:Medusaceratops lokii
2844:Medusaceratops lokii
1422:Spiclypeus shipporum
1347:Medusaceratops lokii
1311:10.3374/014.054.0103
1133:Medusaceratops lokii
952:Avaceratops lammersi
863:as a centrosaurine:
768:Medusaceratops lokii
516:Medusaceratops lokii
386:Medusaceratops lokii
330:Medusaceratops lokii
1925:Archaeoceratopsidae
1495:2014NW....101..505R
1483:Naturwissenschaften
1440:2016PLoSO..1154218M
1371:10.1017/jpa.2017.62
1363:2018JPal...92..272C
1295:Judiceratops tigris
2952:Fossils of Montana
2645:Crittendenceratops
2595:Albertaceratopsini
794:
656:
609:Foremost Formation
571:equivalent to the
545:genera names. The
402:
274:Albertaceratopsini
2924:
2923:
2896:Open Tree of Life
2836:Taxon identifiers
2827:
2826:
2799:
2798:
2779:
2778:
2775:
2774:
2771:
2770:
2767:
2766:
2728:Pachyrhinosaurini
2666:Yehuecauhceratops
2629:Nasutoceratopsini
2530:
2529:
2242:
2241:
2222:
2221:
2218:
2217:
2214:
2213:
2210:
2209:
2206:
2205:
2202:
2201:
2103:Protoceratopsidae
1990:Leptoceratopsidae
1958:
1957:
1824:
1823:
1711:
1710:
1707:
1706:
1703:
1702:
1699:
1698:
1229:
1228:
1220:
1219:
1211:
1210:
1202:
1201:
1193:
1192:
1184:
1183:
1174:Pachyrhinosaurini
1101:
1100:
1020:
1019:
938:Nasutoceratopsini
920:
919:
640:Nasutoceratopsini
636:Pachyrhinosaurini
587:million years ago
585:, dating to 77.5
526:, a monster from
494:ceratopsian from
442:Ceratops montanus
341:
340:
297:
2984:
2957:Campanian genera
2917:
2916:
2904:
2903:
2891:
2890:
2878:
2877:
2876:
2863:
2862:
2861:
2831:
2817:
2816:
2794:
2793:
2787:
2758:Pachyrhinosaurus
2682:
2556:Machairoceratops
2541:
2381:Coahuilaceratops
2331:
2272:
2261:
2248:
2237:
2232:
2231:
2150:
2099:
2083:Zhuchengceratops
1971:
1841:
1837:
1778:Chaoyangsauridae
1745:
1741:
1730:
1717:
1659:
1632:
1623:
1616:
1615:
1568:
1553:
1546:
1539:
1530:
1523:
1522:
1478:
1472:
1471:
1461:
1451:
1415:
1409:
1408:
1406:
1404:
1389:
1383:
1382:
1342:
1323:
1322:
1290:
1284:
1268:
1152:
1065:
1058:
1034:
1011:Malta new taxon
944:
934:
884:
877:
867:
697:is unique among
662:, including the
589:, to the middle
573:Oldman Formation
412:adjacent to the
332:
328:
292:
285:
272:
256:
243:
230:
214:
198:
142:
141:
120:
110:
40:
21:
2992:
2991:
2987:
2986:
2985:
2983:
2982:
2981:
2927:
2926:
2925:
2920:
2912:
2907:
2899:
2894:
2886:
2881:
2872:
2871:
2866:
2857:
2856:
2851:
2838:
2828:
2823:
2795:
2788:
2763:
2722:
2671:
2652:Furcatoceratops
2623:
2604:Albertaceratops
2589:
2563:Menefeeceratops
2526:
2499:Tatankaceratops
2456:
2402:Mercuriceratops
2353:Arrhinoceratops
2320:
2268:
2255:
2238:
2226:
2198:
2178:
2139:
2088:
2055:Montanoceratops
1984:
1979:Graciliceratops
1967:
1954:
1934:Archaeoceratops
1919:
1833:
1820:
1808:Xuanhuaceratops
1794:Hualianceratops
1772:
1753:Albalophosaurus
1737:
1724:
1695:
1675:
1648:
1642:Avemetatarsalia
1627:Avemetatarsalia
1617:
1611:Marginocephalia
1573:
1572:
1562:
1557:
1527:
1526:
1480:
1479:
1475:
1434:(5): e0154218.
1417:
1416:
1412:
1402:
1400:
1391:
1390:
1386:
1344:
1343:
1326:
1292:
1291:
1287:
1269:
1252:
1247:
1235:
1230:
1221:
1212:
1203:
1194:
1185:
1102:
1021:
921:
842:Albertaceratops
829:Mercuriceratops
776:
757:Albertaceratops
734:Albertaceratops
713:Albertaceratops
648:
582:Albertaceratops
558:Norse mythology
528:Greek mythology
500:Albertaceratops
487:Albertaceratops
394:
367:Late Cretaceous
365:known from the
337:
334:
326:
325:
312:
291:
283:
270:
254:
241:
228:
212:
196:
136:
127:Albertaceratops
111:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
35:
34:
27:
17:
12:
11:
5:
2990:
2988:
2980:
2979:
2974:
2969:
2964:
2959:
2954:
2949:
2944:
2939:
2937:Centrosaurines
2929:
2928:
2922:
2921:
2919:
2918:
2905:
2892:
2879:
2864:
2848:
2846:
2840:
2839:
2834:
2825:
2824:
2822:
2821:
2811:
2804:
2801:
2800:
2797:
2796:
2782:
2780:
2777:
2776:
2773:
2772:
2769:
2768:
2765:
2764:
2762:
2761:
2754:
2747:
2740:
2732:
2730:
2724:
2723:
2721:
2720:
2712:
2705:
2698:
2690:
2688:
2679:
2673:
2672:
2670:
2669:
2662:
2659:Nasutoceratops
2655:
2648:
2641:
2633:
2631:
2625:
2624:
2622:
2621:
2618:Medusaceratops
2614:
2607:
2599:
2597:
2591:
2590:
2588:
2587:
2580:
2573:
2566:
2559:
2552:
2549:Diabloceratops
2544:
2538:
2536:Centrosaurinae
2532:
2531:
2528:
2527:
2525:
2524:
2517:
2510:
2507:Titanoceratops
2503:
2495:
2492:Regaliceratops
2488:
2481:
2474:
2466:
2464:
2462:Triceratopsini
2458:
2457:
2455:
2454:
2447:
2440:
2433:
2426:
2423:Sierraceratops
2419:
2412:
2409:Navajoceratops
2405:
2398:
2391:
2384:
2377:
2370:
2363:
2356:
2349:
2342:
2334:
2328:
2326:Chasmosaurinae
2322:
2321:
2319:
2318:
2311:
2304:
2297:
2290:
2287:Brachyceratops
2283:
2275:
2269:
2264:
2257:
2256:
2251:
2244:
2243:
2240:
2239:
2225:
2223:
2220:
2219:
2216:
2215:
2212:
2211:
2208:
2207:
2204:
2203:
2200:
2199:
2197:
2196:
2195:
2194:
2188:
2186:
2180:
2179:
2177:
2176:
2169:
2166:Turanoceratops
2162:
2153:
2147:
2141:
2140:
2138:
2137:
2130:
2123:
2116:
2107:
2105:
2096:
2090:
2089:
2087:
2086:
2079:
2076:Unescoceratops
2072:
2065:
2058:
2051:
2044:
2041:Ischioceratops
2037:
2034:Gryphoceratops
2030:
2023:
2016:
2009:
2002:
1994:
1992:
1986:
1985:
1983:
1982:
1974:
1968:
1963:
1960:
1959:
1956:
1955:
1953:
1952:
1944:
1937:
1929:
1927:
1921:
1920:
1918:
1917:
1914:Sasayamagnomus
1910:
1903:
1896:
1889:
1881:
1874:
1867:
1864:Auroraceratops
1860:
1853:
1844:
1834:
1829:
1826:
1825:
1822:
1821:
1819:
1818:
1811:
1804:
1797:
1790:
1787:Chaoyangsaurus
1782:
1780:
1774:
1773:
1771:
1770:
1767:Psittacosaurus
1763:
1756:
1748:
1738:
1733:
1726:
1725:
1720:
1713:
1712:
1709:
1708:
1705:
1704:
1701:
1700:
1697:
1696:
1694:
1693:
1692:
1691:
1685:
1683:
1677:
1676:
1674:
1673:
1672:
1671:
1662:
1656:
1650:
1649:
1647:
1646:
1645:
1644:
1635:
1629:
1619:
1618:
1614:
1613:
1604:
1595:
1586:
1580:
1571:
1564:
1563:
1558:
1556:
1555:
1548:
1541:
1533:
1525:
1524:
1489:(6): 505–512.
1473:
1410:
1384:
1357:(2): 272–288.
1324:
1285:
1249:
1248:
1246:
1243:
1242:
1241:
1234:
1231:
1227:
1226:
1223:
1222:
1218:
1217:
1214:
1213:
1209:
1208:
1205:
1204:
1200:
1199:
1196:
1195:
1191:
1190:
1187:
1186:
1182:
1181:
1178:
1177:
1170:
1167:
1166:
1163:
1162:
1155:
1150:
1148:
1142:
1141:
1138:
1137:
1128:
1125:
1124:
1121:
1120:
1111:
1108:
1107:
1104:
1103:
1099:
1098:
1095:
1094:
1085:
1082:
1081:
1078:
1077:
1068:
1063:
1061:
1056:
1054:
1051:
1050:
1047:
1046:
1037:
1032:
1030:
1027:
1026:
1023:
1022:
1018:
1017:
1014:
1013:
1008:
1005:
1004:
1001:
1000:
991:
988:
987:
984:
983:
978:
975:
974:
971:
970:
965:
962:
961:
958:
957:
947:
942:
940:
932:
930:
927:
926:
923:
922:
918:
917:
914:
913:
904:
901:
900:
897:
896:
887:
882:
880:
875:
873:
871:Centrosaurinae
865:
861:Medusaceratops
846:Medusaceratops
838:Medusaceratops
834:squamosal bone
806:Medusaceratops
790:Medusaceratops
775:
774:Classification
772:
753:Centrosaurinae
749:Medusaceratops
745:Medusaceratops
729:Medusaceratops
722:centrosaurines
718:Medusaceratops
708:Medusaceratops
704:squamosal bone
699:Chasmosaurinae
695:Medusaceratops
687:Medusaceratops
676:Medusaceratops
660:Medusaceratops
647:
644:
628:Medusaceratops
620:Medusaceratops
595:Medusaceratops
569:lithologically
504:Medusaceratops
478:paleontologist
473:was coined by
471:Medusaceratops
464:Medusaceratops
448:Medusaceratops
410:Kennedy Coulee
406:Medusaceratops
393:
390:
345:Medusaceratops
339:
338:
335:
321:
320:
314:
313:
305:
303:
299:
298:
288:Medusaceratops
281:
277:
276:
268:
261:
260:
258:Centrosaurinae
252:
248:
247:
239:
235:
234:
226:
219:
218:
216:Neornithischia
210:
203:
202:
194:
187:
186:
181:
174:
173:
168:
164:
163:
158:
154:
153:
148:
144:
143:
130:
129:
122:
121:
113:
112:
104:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
41:
28:
25:Medusaceratops
15:
13:
10:
9:
6:
4:
3:
2:
2989:
2978:
2975:
2973:
2970:
2968:
2965:
2963:
2960:
2958:
2955:
2953:
2950:
2948:
2945:
2943:
2940:
2938:
2935:
2934:
2932:
2915:
2910:
2906:
2902:
2897:
2893:
2889:
2884:
2880:
2875:
2869:
2865:
2860:
2854:
2850:
2849:
2847:
2845:
2841:
2837:
2832:
2820:
2812:
2810:
2806:
2805:
2802:
2792:
2786:
2760:
2759:
2755:
2753:
2752:
2748:
2746:
2745:
2741:
2739:
2738:
2737:Achelousaurus
2734:
2733:
2731:
2729:
2725:
2718:
2717:
2716:Styracosaurus
2713:
2711:
2710:
2706:
2704:
2703:
2699:
2697:
2696:
2692:
2691:
2689:
2687:
2686:Centrosaurini
2683:
2680:
2678:
2677:Eucentrosaura
2674:
2668:
2667:
2663:
2661:
2660:
2656:
2654:
2653:
2649:
2647:
2646:
2642:
2640:
2639:
2635:
2634:
2632:
2630:
2626:
2620:
2619:
2615:
2613:
2612:
2608:
2606:
2605:
2601:
2600:
2598:
2596:
2592:
2586:
2585:
2581:
2579:
2578:
2577:Wendiceratops
2574:
2572:
2571:
2567:
2565:
2564:
2560:
2558:
2557:
2553:
2551:
2550:
2546:
2545:
2542:
2539:
2537:
2533:
2523:
2522:
2518:
2516:
2515:
2511:
2509:
2508:
2504:
2501:
2500:
2496:
2494:
2493:
2489:
2487:
2486:
2482:
2480:
2479:
2475:
2473:
2472:
2471:Eotriceratops
2468:
2467:
2465:
2463:
2459:
2453:
2452:
2448:
2446:
2445:
2441:
2439:
2438:
2434:
2432:
2431:
2427:
2425:
2424:
2420:
2418:
2417:
2416:Pentaceratops
2413:
2411:
2410:
2406:
2404:
2403:
2399:
2397:
2396:
2395:Kosmoceratops
2392:
2390:
2389:
2385:
2383:
2382:
2378:
2376:
2375:
2371:
2369:
2368:
2367:Bravoceratops
2364:
2362:
2361:
2360:Bisticeratops
2357:
2355:
2354:
2350:
2348:
2347:
2346:Anchiceratops
2343:
2341:
2340:
2339:Agujaceratops
2336:
2335:
2332:
2329:
2327:
2323:
2317:
2316:
2312:
2310:
2309:
2305:
2303:
2302:
2298:
2296:
2295:
2291:
2289:
2288:
2284:
2282:
2281:
2277:
2276:
2273:
2270:
2267:
2262:
2258:
2254:
2249:
2245:
2236:
2230:
2192:
2191:
2190:
2189:
2187:
2185:
2181:
2175:
2174:
2170:
2168:
2167:
2163:
2160:
2159:
2155:
2154:
2151:
2148:
2146:
2145:Ceratopsoidea
2142:
2136:
2135:
2134:Protoceratops
2131:
2129:
2128:
2127:Breviceratops
2124:
2122:
2121:
2117:
2114:
2113:
2109:
2108:
2106:
2104:
2100:
2097:
2095:
2091:
2085:
2084:
2080:
2078:
2077:
2073:
2071:
2070:
2069:Udanoceratops
2066:
2064:
2063:
2062:Prenoceratops
2059:
2057:
2056:
2052:
2050:
2049:
2048:Leptoceratops
2045:
2043:
2042:
2038:
2036:
2035:
2031:
2029:
2028:
2024:
2022:
2021:
2017:
2015:
2014:
2010:
2008:
2007:
2006:Bainoceratops
2003:
2001:
2000:
1996:
1995:
1993:
1991:
1987:
1981:
1980:
1976:
1975:
1972:
1969:
1966:
1961:
1950:
1949:
1945:
1943:
1942:
1941:Helioceratops
1938:
1936:
1935:
1931:
1930:
1928:
1926:
1922:
1916:
1915:
1911:
1909:
1908:
1907:Mosaiceratops
1904:
1902:
1901:
1897:
1895:
1894:
1890:
1887:
1886:
1882:
1880:
1879:
1878:Koreaceratops
1875:
1873:
1872:
1868:
1866:
1865:
1861:
1859:
1858:
1854:
1851:
1850:
1846:
1845:
1842:
1838:
1835:
1832:
1831:Neoceratopsia
1827:
1817:
1816:
1812:
1810:
1809:
1805:
1803:
1802:
1798:
1796:
1795:
1791:
1789:
1788:
1784:
1783:
1781:
1779:
1775:
1769:
1768:
1764:
1762:
1761:
1757:
1755:
1754:
1750:
1749:
1746:
1742:
1739:
1736:
1731:
1727:
1723:
1718:
1714:
1689:
1688:
1687:
1686:
1684:
1682:
1678:
1670:
1666:
1665:
1664:
1663:
1660:
1657:
1655:
1651:
1643:
1639:
1638:
1637:
1636:
1633:
1630:
1628:
1624:
1620:
1612:
1608:
1605:
1603:
1599:
1596:
1594:
1590:
1587:
1585:
1581:
1579:
1575:
1574:
1569:
1565:
1561:
1554:
1549:
1547:
1542:
1540:
1535:
1534:
1531:
1520:
1516:
1512:
1508:
1504:
1500:
1496:
1492:
1488:
1484:
1477:
1474:
1469:
1465:
1460:
1455:
1450:
1445:
1441:
1437:
1433:
1429:
1425:
1423:
1414:
1411:
1399:
1395:
1388:
1385:
1380:
1376:
1372:
1368:
1364:
1360:
1356:
1352:
1348:
1341:
1339:
1337:
1335:
1333:
1331:
1329:
1325:
1320:
1316:
1312:
1308:
1304:
1300:
1296:
1289:
1286:
1282:
1281:0-253-35358-0
1278:
1274:
1267:
1265:
1263:
1261:
1259:
1257:
1255:
1251:
1244:
1240:
1237:
1236:
1232:
1225:
1224:
1216:
1215:
1207:
1206:
1198:
1197:
1189:
1188:
1180:
1179:
1176:
1175:
1169:
1168:
1165:
1164:
1161:
1160:
1159:Centrosaurini
1154:
1153:
1147:
1146:Eucentrosaura
1144:
1143:
1140:
1139:
1136:
1135:
1134:
1127:
1126:
1123:
1122:
1119:
1118:
1117:
1110:
1109:
1106:
1105:
1097:
1096:
1093:
1092:
1091:
1084:
1083:
1080:
1079:
1076:
1075:
1074:
1067:
1066:
1060:
1059:
1053:
1052:
1049:
1048:
1045:
1044:
1043:
1036:
1035:
1029:
1028:
1025:
1024:
1016:
1015:
1012:
1007:
1006:
1003:
1002:
999:
998:
997:
990:
989:
986:
985:
982:
977:
976:
973:
972:
969:
964:
963:
960:
959:
956:
955:(ANSP 15800)
954:
953:
946:
945:
939:
936:
935:
929:
928:
925:
924:
916:
915:
912:
911:
910:
903:
902:
899:
898:
895:
894:
893:
886:
885:
879:
878:
872:
869:
868:
864:
862:
858:
854:
849:
847:
843:
839:
835:
831:
830:
825:
824:
819:
818:
813:
812:
807:
803:
799:
791:
787:
786:
785:Wendiceratops
780:
773:
771:
769:
764:
762:
761:Wendiceratops
758:
754:
750:
746:
742:
741:
740:Wendiceratops
736:
735:
730:
725:
723:
719:
715:
714:
709:
705:
700:
696:
692:
691:chasmosaurine
688:
683:
681:
677:
673:
672:chasmosaurine
669:
665:
661:
652:
645:
643:
641:
637:
633:
632:Centrosaurini
629:
625:
621:
617:
612:
610:
606:
605:
600:
599:chasmosaurine
596:
592:
588:
584:
583:
578:
574:
570:
566:
561:
559:
555:
551:
548:
547:specific name
544:
540:
537:
534:
529:
525:
521:
517:
513:
509:
505:
501:
497:
493:
492:centrosaurine
489:
488:
483:
479:
476:
472:
467:
465:
461:
457:
453:
449:
444:
443:
439:
438:
433:
432:
427:
423:
419:
415:
411:
407:
398:
391:
389:
387:
383:
382:United States
379:
375:
371:
368:
364:
361:
358:
357:centrosaurine
354:
351:
347:
346:
333:
331:
322:
319:
318:Binomial name
315:
311:
310:
309:M. lokii
304:
301:
300:
296:
290:
289:
282:
279:
278:
275:
269:
266:
263:
262:
259:
253:
250:
249:
246:
240:
237:
236:
233:
227:
224:
221:
220:
217:
211:
208:
205:
204:
201:
195:
192:
189:
188:
185:
182:
179:
176:
175:
172:
169:
166:
165:
162:
159:
156:
155:
152:
149:
146:
145:
140:
135:
131:
128:
123:
119:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
26:
22:
19:
2843:
2756:
2751:Stellasaurus
2749:
2742:
2735:
2714:
2707:
2702:Coronosaurus
2700:
2695:Centrosaurus
2693:
2664:
2657:
2650:
2643:
2636:
2617:
2616:
2611:Lokiceratops
2609:
2602:
2584:Xenoceratops
2582:
2575:
2570:Sinoceratops
2568:
2561:
2554:
2547:
2519:
2512:
2505:
2497:
2490:
2483:
2478:Nedoceratops
2476:
2469:
2451:Vagaceratops
2449:
2444:Utahceratops
2442:
2437:Terminocavus
2435:
2428:
2421:
2414:
2407:
2400:
2393:
2388:Judiceratops
2386:
2379:
2374:Chasmosaurus
2372:
2365:
2358:
2351:
2344:
2337:
2313:
2306:
2299:
2292:
2285:
2278:
2266:Ceratopsidae
2253:Ceratopsidae
2184:Ceratopsidae
2173:Zuniceratops
2171:
2164:
2158:Ajkaceratops
2156:
2132:
2125:
2120:Bagaceratops
2118:
2112:Ajkaceratops
2110:
2094:Coronosauria
2081:
2074:
2067:
2060:
2053:
2046:
2039:
2032:
2025:
2018:
2011:
2004:
1999:Asiaceratops
1997:
1977:
1965:Euceratopsia
1948:Yamaceratops
1946:
1939:
1932:
1912:
1905:
1900:Microceratus
1898:
1893:Liaoceratops
1891:
1883:
1876:
1869:
1862:
1855:
1849:Ajkaceratops
1847:
1813:
1806:
1799:
1792:
1785:
1765:
1758:
1751:
1669:Ornithischia
1654:Ornithischia
1606:
1602:Ornithischia
1597:
1588:
1486:
1482:
1476:
1431:
1427:
1421:
1413:
1401:. Retrieved
1398:ScienceDaily
1397:
1387:
1354:
1350:
1346:
1302:
1298:
1294:
1288:
1272:
1172:
1157:
1132:
1131:
1130:
1114:
1113:
1088:
1087:
1071:
1070:
1040:
1039:
1010:
994:
993:
980:
967:
950:
949:
907:
906:
890:
889:
860:
856:
850:
845:
841:
837:
827:
821:
817:Judiceratops
815:
809:
805:
796:Among valid
795:
789:
783:
767:
765:
760:
756:
748:
744:
738:
732:
728:
726:
717:
711:
707:
694:
686:
684:
680:Ceratopsidae
675:
659:
657:
627:
623:
619:
615:
613:
604:Judiceratops
602:
594:
580:
562:
549:
538:
520:generic name
515:
512:type species
503:
499:
485:
482:dissertation
470:
468:
463:
447:
440:
437:nomen dubium
435:
429:
405:
403:
385:
344:
343:
342:
329:
324:
308:
307:
287:
286:
264:
245:Ceratopsidae
222:
206:
200:Ornithischia
190:
177:
126:
24:
18:
2868:Wikispecies
2744:Einiosaurus
2638:Avaceratops
2521:Triceratops
2485:Ojoceratops
2308:Monoclonius
2020:Ferrisaurus
1885:Kulceratops
811:Avaceratops
798:ceratopsids
654:Restoration
646:Description
543:ceratopsian
426:Hill County
380:, northern
360:ceratopsian
251:Subfamily:
2931:Categories
2807:See also:
2514:Torosaurus
2430:Spiclypeus
2280:Agathaumas
2193:see below↓
2013:Cerasinops
1801:Stenopelix
1735:Ceratopsia
1722:Ceratopsia
1690:see below↓
1681:Ceratopsia
1593:Dinosauria
1560:Ceratopsia
1403:24 January
1245:References
823:Spiclypeus
716:, however
522:refers to
490:, another
414:Milk River
376:stage) of
232:Ceratopsia
184:Dinosauria
36:77.5
1576:Kingdom:
1379:134031275
1319:129801786
1305:: 51–65.
981:CMN 8804
853:cladogram
800:from the
591:Campanian
533:Latinized
469:The name
416:, in the
392:Discovery
374:Campanian
302:Species:
157:Kingdom:
151:Eukaryota
31:Campanian
2859:Q2272018
2853:Wikidata
2819:Category
2809:Timeline
2315:Polyonax
2301:Dysganus
2294:Ceratops
1857:Aquilops
1584:Chordata
1582:Phylum:
1578:Animalia
1519:13957187
1511:24859020
1468:27191389
1428:PLOS ONE
1233:See also
968:MOR 692
668:paratype
664:holotype
624:M. lokii
539:ceratops
510:and the
475:Canadian
372:(middle
363:dinosaur
238:Family:
171:Chordata
167:Phylum:
161:Animalia
147:Domain:
2901:6150280
2888:8654674
2709:Spinops
2027:Gremlin
1815:Yinlong
1491:Bibcode
1459:4871577
1436:Bibcode
1359:Bibcode
552:honors
496:Alberta
420:, near
378:Montana
350:extinct
327:†
306:†
280:Genus:
105:↓
2914:170632
1517:
1509:
1466:
1456:
1377:
1317:
1279:
857:et al.
616:et al.
577:Canada
524:Medusa
518:. The
348:is an
1607:Clade
1598:Clade
1589:Clade
1515:S2CID
1375:S2CID
1315:S2CID
638:than
550:lokii
536:Greek
422:Havre
353:genus
265:Clade
223:Clade
207:Clade
191:Clade
178:Clade
2883:GBIF
1667:see
1640:see
1507:PMID
1464:PMID
1405:2022
1277:ISBN
851:The
820:and
759:and
737:and
634:and
554:Loki
508:2010
295:2010
44:PreꞒ
1871:Beg
1499:doi
1487:101
1454:PMC
1444:doi
1367:doi
1307:doi
751:to
575:of
514:is
355:of
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1432:11
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1365:.
1355:92
1353:.
1327:^
1313:.
1303:54
1301:.
1253:^
804:,
770:.
763:.
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682:.
642:.
611:.
502:.
424:,
388:.
267::
225::
209::
193::
180::
94:Pg
38:Ma
33:,
2719:?
2502:?
2161:?
2115:?
1951:?
1888:?
1852:?
1552:e
1545:t
1538:v
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284:†
271:†
255:†
242:†
229:†
213:†
197:†
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:Ꞓ
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