778:
957:
968:
446:
136:
3101:
546:
3404:
695:. The carapace of this species was parabolic (approximately U-shaped) with small intramarginal arcuate eyes. The metastoma (a large plate that is part of the abdomen) was oval. Its operculum (a plate-like segment which contains the genital aperture) had a long, narrow spatulae (compared to most eurypterids). The telson was lanceolate, with a keel on both dorsal and
1086:. These eurypterids shared a series of characteristics such as the almost identical carapace, paddle, eye shapes and eye position. However, its short telson, thin cuticular sculpture (ornamentation consisting of small, minute, scales across the back), very small spatulae and increased spinosity in the appendages suggest a more basal position. Before 2004,
866:
in 1964, named after Clarke, who described the original
Shawangunk eurypterid fauna. This small species of only 4 cm (1.6 in) had a lanceolate carapace with intramarginal eyes located anteriorly. The telson was broad and lanceolate. This species is characterized by the presence of spurs in the genal
507:
adapted to swim crosscurrent. It had epimera (lateral "extensions" of the segment) present in the seventh segment, separating the preadomen (body segments 1 to 7) from the postabodmen (segments 8 to 12). The genital spatulae, a long, flat piece in the genital aperture, was very short. The walking
606:(pincers) were small. It differs from other species in the more convergent form of the carapace and in the large size of the eyes, which were furthest from the margin. This species reached a length of 8 cm (3.1 in) in total. Erik N. Kjellesvig-Waering classified this species as part of the
654:. In addition, the carapace of this species was long and with ocelli placed forward. It has been suggested that this species could represent a separate genus. In fact, notes left by Kjellesvig-Waering in museum drawers indicate that he wanted to erect a new genus for
894:(southern) and Siberia, referring to the place where it was discovered. The carapace was parabolic and surrounded by a narrow marginal rim and with elongated eyes probably reniform. Only the fourth and fifth pair of prosomal appendages is known. These were of
2620:
601:
margin which narrows gradually anteriorly. A thin rim surrounded the carapace. The eyes were narrow, reniform and intramarginal. Its ocelli (light-sensitive simple eyes) were very small and located centrally. As in most eurypterid groups, the
861:
in 1912. They believed that the carapaces with intramarginal eyes were modified by the compression. While this could happen in a single specimen, it is impossible to be in so many. Kjellesvig-Waering realized this and erected the species
740:
margin was slightly concave. There is only one preserved eye, which was reniform, large and rose slightly from the surface of the carapace. At the closest point between both eyes, the ocelli were placed. The fossils were found at the
577:("naked name", a name coined without an adequate or completely missing description of it). The fossils mentioned by Banks consisted of two specimens, the first of which included carapace with the first five
417:. This clade was characterised by their small size, their parabolic (approximately U-shaped) carapaces and the presence of epimera (lateral "extensions" of the segment) on the seventh segment, among others.
2408:
1060:. However, the clade has almost never been used in subsequent studies and lists of eurypterids, and instead, they classify the nanahughmillerids as part of Adelophthalmidae. A basal clade between
1109:), as inferred by O. Erik Tetlie and Markus Poschmann in 2008, based on the results of a 2008 analysis specifically pertaining to the Adelophthalmoidea and a preceding 2004 analysis.
939:
lack eyes, making it difficult to determine their phylogenetic position. However, the prosomal and genital appendages were not typical of the
Adelophthalmoidea, but probably of the
935:). The swimming legs of this species were long and narrow, reaching the sixth segment of the body. The telson was lanceolate and with a keel in the middle. All known specimens of
2452:
3467:
2140:
499:
had long and narrow reniform (bean-shaped) eyes placed in an intramarginal (occurring within the margin) position. The streamlined form of its body indicates that
699:
sides. It had a pair of spines in each podomere of the fifth appendage. The paddle of the swimming leg was relatively wide. It was designated the type species of
585:
parts of the body segments) and the 8th and 9th segments of the swimming leg and the second including another nearly complete carapace. Both were collected in
2406:
V. P. Tollerton (2003). "Summary of a revision of New York State
Ordovician eurypterids: implications for eurypterid palaeoecology, diversity and evolution".
532:
is only distinguished from the more derived members of
Adelophthalmidae in the shorter and smaller spatulae and in the increased spinosity in the appendages.
115:
2444:
644:. This species differs from the others by the anterior position of its eyes placed in a submarginal (almost in the margin) point, unlike other species of
526:(leg segment). The paddle of the swimming leg was relatively wide. As in the other adelophthalmids, the telson was lanceolate and not very expanded.
3454:
3551:
1097:
The cladogram below presents the inferred phylogenetic positions of most of the genera included in the three most derived superfamilies of the
825:
was designated as subgenotype. He assigned a total of seven species to the subgenus, of which only four (two of them tentatively) are still in
3536:
3180:
3170:
2692:
2501:
2493:
2159:
736:. This species is only known by a poorly preserved carapace which was semi-oval and elongated, with a narrow margin surrounding it. The
3132:
3147:
2595:
2570:
2545:
1705:
2190:
Pirozhnikov, L. P. (1957). "Remains of
Gigantostraca from the series of Matakara (Devonian of the North Minusinsk Depression)".
3391:
3175:
3117:
1869:
3541:
3165:
3155:
3109:
135:
3531:
3526:
3127:
3122:
777:
3546:
3160:
3137:
2571:"Eurypterid-Associated Biota of the Shunet Fm. at Khakassia near Lake Shunet: Lochkovian - Eifelian, Russian Federation"
1020:
621:
956:
3521:
2095:
O. Erik Tetlie; Paul A. Selden; Dong Ren (2007). "A new
Silurian eurypterid (Arthropoda: Chelicerata) from China".
967:
906:(the "median part"). The total size of the species could be of 9 cm (3.5 in), being one of the largest species of
2793:
2685:
2071:"A reappraisal of Eurypterus dumonti Stainier, 1917 and its position within the Adelophthalmidae Tollerton, 1989"
1864:
2469:
1899:"Cope's rule and Romer's theory: patterns of diversity and gigantism in eurypterids and Palaeozoic vertebrates"
898:-type and with rather small spines. The telson was small and wedge-shaped, known from fragmentary remains. The
3100:
2258:
Shpinev, E. S. (2012). "On some eurypterids (Eurypterida, Chelicerata) from the
Devonian of South Siberia".
684:
445:
757:
and they assigned it in 1966 to its current genus. It has been suggested that this species is a synonym of
114:
3422:
2228:
Kjellesvig-Waering, Erik N.; Leutze, Willard P. (1966). "Eurypterids from the
Silurian of West Virginia".
586:
3493:
2596:"Eurypterid-Associated Biota of the Temeside Shale, Ludlow and Perton, England: Pridoli, United Kingdom"
2165:
742:
3407:
2678:
1946:
1805:
637:
430:
571:". Although Richard Banks first mentioned this species in 1856, this act is currently considered as
737:
562:
1847:
761:, but the Russian paleontologist Evgeniy S. Shpinev does not agree with this since the prosoma of
2546:"Nanahughmilleria norvegica-Ostracoderm Comm. at Rudstangen, Ringerike Dist.: Rootsikula, Norway"
2519:
2425:
2364:
2335:
2275:
2237:
2122:
2044:
2002:
1797:
1752:
1743:
130:
2646:
1704:
can occur in an environment where eurypterids were abundant or scarce. The
Silurian deposits of
910:. This species differs from the others in the proportions of the body and position of the eyes.
3498:
753:. Kjellesvig-Waering and Willard P. Leutze noticed that the species did not really represent a
3480:
3472:
2996:
2929:
2921:
2507:
2497:
2207:
2199:
1938:
1920:
1219:
1126:
1106:
1000:
921:. This species was originally described by Salter in 1856 as another species of his new genus
854:
414:
3485:
1983:"Phylogeny and palaeoecology of the Adelophthalmoidea (Arthropoda; Chelicerata; Eurypterida)"
465:
was a comparatively small eurypterid. The largest species confidently assigned to the genus,
3336:
3050:
3042:
3032:
3024:
3004:
2975:
2891:
2840:
2461:
2417:
2267:
2169:
2112:
2104:
1994:
1928:
1912:
1822:
1656:
1338:
1281:
1052:
996:
982:
858:
674:
650:
582:
410:
216:
203:
30:
2393:
3365:
3211:
3068:
3009:
2965:
2868:
2855:
2531:
2376:
2293:
1903:
1792:
1659:
1442:
1091:
1035:
519:
422:
1034:
2035:
Kjellesvig-Waering, Erik N. (1961). "The Silurian Eurypterida of the Welsh Borderland".
3281:
3239:
3197:
3060:
2987:
2947:
2939:
2863:
1933:
1748:
1404:
1355:
1164:
1116:
1102:
887:
677:
478:
2070:
545:
3515:
3288:
3260:
3253:
3225:
3204:
2957:
2896:
2886:
2878:
2832:
2822:
2814:
2770:
2734:
2279:
2108:
1982:
1828:
1778:
1724:
1671:
1555:
1233:
940:
641:
633:
603:
590:
426:
384:
72:
2429:
2126:
2006:
1898:
3246:
3078:
3014:
2970:
2845:
1837:
1809:
1739:
1730:
1694:, lived mainly in shallow lakes. In addition, the streamlined shape of the body of
1490:
1418:
1181:
1140:
1075:
1016:
931:
514:
359:
242:
2487:
2465:
2354:
3343:
3320:
3218:
2912:
2805:
2743:
2725:
1841:
1264:
1098:
1062:
926:
732:
727:
573:
482:
177:
47:
3445:
3309:
3295:
3274:
3267:
3232:
3073:
2761:
2752:
2701:
2421:
2271:
1998:
1801:
1718:
1691:
1538:
1514:
1011:, but Kjellesvig-Waering and Leutze considered it sufficiently different from
567:
458:
364:
190:
92:
57:
2511:
2211:
2203:
1924:
1698:
suggests that it was an active swimmer capable of swimming against currents.
3386:
3370:
3360:
2719:
1765:
746:
681:
368:
167:
147:
97:
41:
1942:
1916:
874:, was described by Shpinev. This species is known from two specimens, the
835:
813:) for species with small reniform intramarginal eyes. For the latter one,
3439:
3329:
3302:
1667:
1663:
1652:
1466:
903:
899:
879:
875:
607:
598:
523:
400:
380:
376:
87:
82:
67:
62:
52:
34:
2318:
Kjellesvig-Waering, Erik N. (1964). "Eurypterida: Notes on the Subgenus
3459:
2339:
2241:
2048:
1039:
750:
696:
578:
396:
102:
77:
2713:
2117:
1709:
1675:
718:
In 1957, L. P. Pirozhnikov described two new species of eurypterids,
504:
392:
388:
157:
3416:
2651:, Salter, a large crustacean from the Upper Silurian and Downtonian"
1038:
with no or reduced genital spatulae and the second to fifth pair of
1769:
830:
776:
444:
2445:"Distribution and dispersal history of Eurypterida (Chelicerata)"
902:(the posterior "part" of the body) was notably narrower than the
2621:"Delaware Water Gap Eurypterids (Silurian of the United States)"
2078:
Bulletin de l'Institut Royal des Sciences Naturelles de Belgique
3420:
2674:
437:
at 10 cm (3.9 in), making it a comparatively small eurypterid.
2670:
2409:
Transactions of the Royal Society of Edinburgh: Earth Sciences
1015:
and elevated the subgenus to the genus level. It is the only
2161:
Eurypterid phylogeny with remarks on the origin of arachnids
797:
into two subgenera according to the morphology of the eyes,
433:. The largest species confidently assigned to the genus was
849:
is based on a series of fossils described and assigned to
765:
was longer and its eyes were closer to the margin than in
703:
by Kjellesvig-Waering in his description of the subgenus.
421:
was different from its relatives by the presence of more
477:
reached 16 cm (6.3 in), this species is probably not an
2396:. In World Spider Catalog. Natural History Museum Bern.
2356:
On some new Crustacea from the uppermost Silurian Rocks
1070:
is better supported than the Nanahughmilleridae, while
469:, only reached 10 cm (3.9 in), while the smallest one,
403:, and have been referred to several different species.
1078:(closest relative) of the "derived clade" composed by
870:
It would not be until 2012 when another new species,
2394:
A summary list of fossil spiders and their relatives
3429:
3379:
3353:
3319:
3189:
3146:
3108:
3059:
3041:
3023:
2995:
2984:
2956:
2938:
2920:
2909:
2877:
2854:
2831:
2813:
2802:
2786:
2779:
2192:Vsesojuzuoe Paleontologiceskoe Obchestvo Ezegodnik
2453:Palaeogeography, Palaeoclimatology, Palaeoecology
1981:Erik Tetlie, O.; Poschmann, Markus (2008-06-01).
1764:are the only animals of the Devonian deposits of
1019:genus (with grouped species that do not share an
805:) for species with large ovoid marginal eyes and
711:, measuring 10 cm (3.9 in, with the exception of
557:The first species of what today is recognized as
2392:Dunlop, J. A., Penney, D. & Jekel, D. 2018.
1716:have been discovered, include fossil remains of
773:Creation of the subgenus and further discoveries
1031:probably representing new and different forms.
2686:
1897:Lamsdell, James C.; Braddy, Simon J. (2009).
867:angles (a "flat" part of the carapace edge).
8:
1101:suborder of eurypterids (Adelophthalmoidea,
597:had an elongated carapace rounded along the
1796:, as well as some indeterminate species of
1776:from the Silurian has been associated with
3417:
2992:
2917:
2810:
2783:
2693:
2679:
2671:
1855:fossils have also been found in Scotland.
113:
20:
2116:
1932:
999:, the only family within the superfamily
715:, which may represent a separate genus).
473:, did not exceed 4 cm (1.6 in). Although
2313:
2311:
1007:was originally considered a subgenus of
544:
1880:
1023:) of Adelophthlmidae, with the species
726:, and erroneously assigned them to the
489:reached 9 cm (3.5 in) in length, while
2527:
2517:
2372:
2362:
2253:
2251:
829:. The generic name is composed by the
793:In 1961, Kjellesvig-Waering separated
632:, based on a single carapace from the
2153:
2151:
2149:
2069:Tetlie, O. Erik; van Roy, P. (2006).
1976:
917:is tentatively classified as part of
493:had a total length of 8 cm (3.1 in).
7:
2388:
2386:
2139:Clarke, J. K., Ruedemann R. (1912) "
2064:
2062:
2060:
2058:
1974:
1972:
1970:
1968:
1966:
1964:
1962:
1960:
1958:
1956:
1892:
1890:
1888:
1886:
1884:
995:is classified as part of the family
617:) in 1961 along many other species.
508:legs (first to fifth appendages) of
413:, the only clade in the superfamily
375:have been discovered in deposits of
2494:Joint Nature Conservation Committee
2223:
2221:
2030:
2028:
2026:
2024:
2022:
2020:
2018:
2016:
1987:Journal of Systematic Palaeontology
1772:. In addition, the English species
1530:
1506:
1482:
1458:
1434:
1410:
1330:
1306:
1256:
1249:
1225:
1215:
1156:
1132:
1122:
1112:
549:Size comparison of the unequivocal
2489:Fossil Arthropods of Great Britain
2326:) from the Silurian of New York".
1804:. In the Silurian deposits of the
14:
1738:, among other organisms like the
1678:and Siberia. It is believed that
882:(PIN 1139/2). Its specific name,
3403:
3402:
3099:
2168:. pp. 1–344. Archived from
2109:10.1111/j.1475-4983.2007.00651.x
966:
955:
134:
3392:Timeline of eurypterid research
2359:. Vol. 12. pp. 26–34.
1870:Timeline of eurypterid research
666:, this species was assigned to
518:-type, that is, with a pair of
1823:Ruedemannipterus stylonuroides
1725:Stylonuroides dolichopteroides
980:(top) and the closely related
457:Like the other adelophthalmid
367:, an extinct group of aquatic
1:
3552:Fossil taxa described in 1961
1768:, Russia, together with land
1762:Parahughmilleria matakarensis
1046:-type. This family contained
976:Type A genital appendages of
724:Parahughmilleria matarakensis
624:described another species of
3537:Eurypterids of North America
2486:Jarzembowski, E. A. (2010).
2466:10.1016/j.palaeo.2007.05.011
1758:Nanahughmilleria schiraensis
409:is classified in the family
2141:The Eurypterida of New York
707:was the largest species of
687:described a new species of
429:(limbs) and by its genital
16:Extinct genus of arthropods
3568:
2658:Extract Norsk Geol. Tisskr
978:Nanahughmilleria norvegica
252:Nanahughmilleria norvegica
3400:
3097:
2794:List of eurypterid genera
2708:
2422:10.1017/S026359330000064X
2272:10.1134/S0031030112040119
1999:10.1017/S1477201907002416
1865:List of eurypterid genera
1779:Salteropterus abbreviatus
1651:have been found from the
1552:
1535:
1528:
1511:
1504:
1487:
1480:
1463:
1456:
1439:
1432:
1415:
1408:
1352:
1335:
1328:
1311:
1304:
1278:
1261:
1254:
1247:
1230:
1223:
1213:
1178:
1161:
1154:
1137:
1130:
1120:
1021:immediate common ancestor
983:Pittsfordipterus phelpsae
913:In addition, the species
561:was described in 1859 by
267:
262:
248:
241:
131:Scientific classification
129:
121:
112:
23:
2443:Tetlie, O. Erik (2007).
2353:Salter, John W. (1856).
2158:Tetlie, O. Erik (2004).
1784:Parahughmilleria salteri
485:. Other species such as
277:Kjellesvig-Waering, 1964
236:Kjellesvig-Waering, 1961
2328:Journal of Paleontology
2260:Paleontological Journal
2230:Journal of Paleontology
2037:Journal of Paleontology
1834:Hughmilleria shawangunk
1793:Herefordopterus banksii
1788:Erettopterus spatulatus
1731:Kiaeropterus ruedemanni
851:Hughmilleria shawangunk
787:Hughmilleria shawangunk
785:previously assigned to
2645:Størmer, Leif (1935).
1917:10.1098/rsbl.2009.0700
1851:have also been found.
1818:Parahughmilleria maria
1090:was classified in the
790:
554:
454:
3542:Eurypterids of Europe
3494:Paleobiology Database
2166:University of Bristol
1947:Supplemental material
1756:. On the other hand,
780:
548:
448:
3532:Devonian eurypterids
3527:Silurian eurypterids
2304:latin-dictionary.net
1816:occur together with
1806:Shawangunk Formation
685:Johan Aschehoug Kiær
638:geological formation
3547:Eurypterids of Asia
1829:Hardieopterus myops
1744:Dictyocaris slimoni
1712:, where fossils of
568:Eurypterus pygmaeus
563:John William Salter
536:History of research
2922:Onychopterelloidea
1798:cephalaspidomorphs
1753:Kiaeria limuloides
1736:Erettopterus holmi
1074:is considered the
791:
555:
455:
3522:Adelophthalmoidea
3509:
3508:
3481:Open Tree of Life
3423:Taxon identifiers
3414:
3413:
3095:
3094:
3091:
3090:
3087:
3086:
3043:Adelophthalmoidea
3025:Waeringopteroidea
2997:Carcinosomatoidea
2930:Onychopterellidae
2905:
2904:
2503:978-1-86107-486-7
1836:. Species of the
1719:Mixopterus kiaeri
1639:
1638:
1630:
1629:
1621:
1620:
1612:
1611:
1603:
1602:
1594:
1593:
1585:
1584:
1576:
1575:
1567:
1566:
1394:
1393:
1385:
1384:
1376:
1375:
1367:
1366:
1293:
1292:
1220:Adelophthalmoidea
1202:
1201:
1193:
1192:
1127:Waeringopteroidea
1001:Adelophthalmoidea
886:, comes from the
878:(PIN 1139/1) and
855:John Mason Clarke
743:Matarak Formation
541:First discoveries
512:were probably of
415:Adelophthalmoidea
363:") is a genus of
350:
349:
344:
343:Pirozhnikov, 1957
333:
322:
311:
300:
289:
278:
237:
204:Adelophthalmoidea
3559:
3502:
3501:
3489:
3488:
3476:
3475:
3463:
3462:
3450:
3449:
3448:
3431:Nanahughmilleria
3418:
3406:
3405:
3380:Related articles
3337:Merostomichnites
3103:
3051:Adelophthalmidae
3033:Waeringopteridae
3005:Carcinosomatidae
2993:
2976:Strobilopteridae
2918:
2892:Hibbertopteridae
2841:Parastylonuridae
2811:
2784:
2695:
2688:
2681:
2672:
2666:
2665:
2655:
2642:
2636:
2635:
2633:
2631:
2617:
2611:
2610:
2608:
2606:
2592:
2586:
2585:
2583:
2581:
2567:
2561:
2560:
2558:
2556:
2542:
2536:
2535:
2529:
2525:
2523:
2515:
2492:. Vol. 35.
2483:
2477:
2476:
2474:
2468:. Archived from
2460:(3–4): 557–574.
2449:
2440:
2434:
2433:
2403:
2397:
2390:
2381:
2380:
2374:
2370:
2368:
2360:
2350:
2344:
2343:
2324:Nanahughmilleria
2315:
2306:
2290:
2284:
2283:
2255:
2246:
2245:
2236:(5): 1109–1122.
2225:
2216:
2215:
2187:
2181:
2180:
2178:
2177:
2155:
2144:
2137:
2131:
2130:
2120:
2092:
2086:
2085:
2075:
2066:
2053:
2052:
2032:
2011:
2010:
1978:
1951:
1950:
1936:
1894:
1853:Nanahughmilleria
1702:Nanahughmilleria
1696:Nanahughmilleria
1684:Parahughmilleria
1680:Nanahughmilleria
1666:deposits of the
1655:deposits of the
1649:Nanahughmilleria
1531:
1507:
1483:
1459:
1435:
1411:
1339:Parahughmilleria
1331:
1315:Nanahughmilleria
1307:
1282:Pittsfordipterus
1257:
1250:
1226:
1216:
1157:
1133:
1123:
1113:
1107:waeringopteroids
1088:Nanahughmilleria
1080:Parahughmilleria
1072:Nanahughmilleria
1068:Pittsfordipterus
1058:Parahughmilleria
1053:Pittsfordipterus
1036:adelophthalmoids
1005:Nanahughmilleria
997:Adelophthalmidae
993:Nanahughmilleria
970:
959:
919:Nanahughmilleria
908:Nanahughmilleria
859:Rudolf Ruedemann
827:Nanahughmilleria
811:Nanahughmilleria
709:Nanahughmilleria
701:Nanahughmilleria
668:Nanahughmilleria
651:Parahughmilleria
646:Nanahughmilleria
615:Nanahughmilleria
559:Nanahughmilleria
551:Nanahughmilleria
530:Nanahughmilleria
510:Nanahughmilleria
501:Nanahughmilleria
497:Nanahughmilleria
463:Nanahughmilleria
419:Nanahughmilleria
411:Adelophthalmidae
407:Nanahughmilleria
373:Nanahughmilleria
354:Nanahughmilleria
342:
331:
320:
309:
298:
287:
276:
235:
231:Nanahughmilleria
228:
217:Adelophthalmidae
215:
202:
189:
139:
138:
117:
107:
44:
29:Temporal range:
25:Nanahughmilleria
21:
3567:
3566:
3562:
3561:
3560:
3558:
3557:
3556:
3512:
3511:
3510:
3505:
3497:
3492:
3484:
3479:
3471:
3466:
3458:
3453:
3444:
3443:
3438:
3425:
3415:
3410:
3396:
3375:
3366:Chasmataspidida
3349:
3315:
3212:Campylocephalus
3185:
3142:
3104:
3083:
3069:Hughmilleriidae
3055:
3037:
3019:
3010:Megalograptidae
2986:
2980:
2966:Dolichopteridae
2952:
2940:Moselopteroidea
2934:
2911:
2901:
2887:Drepanopteridae
2873:
2869:Hardieopteridae
2856:Kokomopteroidea
2850:
2827:
2804:
2798:
2775:
2704:
2699:
2669:
2653:
2644:
2643:
2639:
2629:
2627:
2625:fossilworks.org
2619:
2618:
2614:
2604:
2602:
2600:fossilworks.org
2594:
2593:
2589:
2579:
2577:
2575:fossilworks.org
2569:
2568:
2564:
2554:
2552:
2550:fossilworks.org
2544:
2543:
2539:
2526:
2516:
2504:
2496:. p. 294.
2485:
2484:
2480:
2472:
2447:
2442:
2441:
2437:
2405:
2404:
2400:
2391:
2384:
2371:
2361:
2352:
2351:
2347:
2317:
2316:
2309:
2291:
2287:
2257:
2256:
2249:
2227:
2226:
2219:
2189:
2188:
2184:
2175:
2173:
2157:
2156:
2147:
2138:
2134:
2094:
2093:
2089:
2073:
2068:
2067:
2056:
2034:
2033:
2014:
1980:
1979:
1954:
1904:Biology Letters
1896:
1895:
1882:
1878:
1861:
1645:
1640:
1631:
1622:
1613:
1604:
1595:
1586:
1577:
1568:
1443:Herefordopterus
1395:
1386:
1377:
1368:
1294:
1203:
1194:
1092:Hughmilleriidae
1048:Nanahughmileria
990:
989:
988:
987:
973:
972:
971:
962:
961:
960:
949:
872:N. notosibirica
775:
767:P. matarakensis
759:P. matarakensis
543:
538:
487:N. notosibirica
449:Restoration of
443:
306:N. notosibirica
258:
255:
234:
226:
213:
200:
187:
133:
108:
106:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
40:435–387.7
39:
38:
27:
17:
12:
11:
5:
3565:
3563:
3555:
3554:
3549:
3544:
3539:
3534:
3529:
3524:
3514:
3513:
3507:
3506:
3504:
3503:
3490:
3477:
3464:
3451:
3435:
3433:
3427:
3426:
3421:
3412:
3411:
3401:
3398:
3397:
3395:
3394:
3389:
3383:
3381:
3377:
3376:
3374:
3373:
3368:
3363:
3357:
3355:
3354:Related groups
3351:
3350:
3348:
3347:
3340:
3333:
3325:
3323:
3317:
3316:
3314:
3313:
3306:
3299:
3292:
3285:
3282:Onychopterella
3278:
3271:
3264:
3257:
3250:
3243:
3240:Hibbertopterus
3236:
3229:
3222:
3215:
3208:
3201:
3198:Adelophthalmus
3193:
3191:
3190:Notable genera
3187:
3186:
3184:
3183:
3178:
3173:
3168:
3163:
3158:
3152:
3150:
3144:
3143:
3141:
3140:
3135:
3130:
3125:
3120:
3114:
3112:
3106:
3105:
3098:
3096:
3093:
3092:
3089:
3088:
3085:
3084:
3082:
3081:
3076:
3071:
3065:
3063:
3061:Pterygotioidea
3057:
3056:
3054:
3053:
3047:
3045:
3039:
3038:
3036:
3035:
3029:
3027:
3021:
3020:
3018:
3017:
3012:
3007:
3001:
2999:
2990:
2988:Diploperculata
2982:
2981:
2979:
2978:
2973:
2968:
2962:
2960:
2954:
2953:
2951:
2950:
2948:Moselopteridae
2944:
2942:
2936:
2935:
2933:
2932:
2926:
2924:
2915:
2907:
2906:
2903:
2902:
2900:
2899:
2894:
2889:
2883:
2881:
2875:
2874:
2872:
2871:
2866:
2864:Kokomopteridae
2860:
2858:
2852:
2851:
2849:
2848:
2843:
2837:
2835:
2829:
2828:
2826:
2825:
2819:
2817:
2815:Rhenopteroidea
2808:
2800:
2799:
2797:
2796:
2790:
2788:
2781:
2777:
2776:
2774:
2773:
2764:
2755:
2746:
2737:
2728:
2722:
2716:
2709:
2706:
2705:
2700:
2698:
2697:
2690:
2683:
2675:
2668:
2667:
2637:
2612:
2587:
2562:
2537:
2528:|journal=
2502:
2478:
2475:on 2011-07-18.
2435:
2416:(3): 235–242.
2398:
2382:
2345:
2334:(2): 410–412.
2307:
2285:
2266:(4): 370–377.
2247:
2217:
2182:
2145:
2132:
2103:(3): 619–625.
2087:
2054:
2043:(4): 789–835.
2012:
1993:(2): 237–249.
1952:
1911:(2): 265–269.
1879:
1877:
1874:
1873:
1872:
1867:
1860:
1857:
1749:chasmataspidid
1644:
1641:
1637:
1636:
1633:
1632:
1628:
1627:
1624:
1623:
1619:
1618:
1615:
1614:
1610:
1609:
1606:
1605:
1601:
1600:
1597:
1596:
1592:
1591:
1588:
1587:
1583:
1582:
1579:
1578:
1574:
1573:
1570:
1569:
1565:
1564:
1561:
1560:
1551:
1548:
1547:
1544:
1543:
1534:
1529:
1527:
1524:
1523:
1520:
1519:
1510:
1505:
1503:
1500:
1499:
1496:
1495:
1486:
1481:
1479:
1476:
1475:
1472:
1471:
1462:
1457:
1455:
1452:
1451:
1448:
1447:
1438:
1433:
1431:
1428:
1427:
1424:
1423:
1414:
1409:
1407:
1405:Pterygotioidea
1401:
1400:
1397:
1396:
1392:
1391:
1388:
1387:
1383:
1382:
1379:
1378:
1374:
1373:
1370:
1369:
1365:
1364:
1361:
1360:
1356:Adelophthalmus
1351:
1348:
1347:
1344:
1343:
1334:
1329:
1327:
1324:
1323:
1320:
1319:
1310:
1305:
1303:
1300:
1299:
1296:
1295:
1291:
1290:
1287:
1286:
1277:
1274:
1273:
1270:
1269:
1260:
1255:
1253:
1248:
1246:
1243:
1242:
1239:
1238:
1229:
1224:
1222:
1214:
1212:
1209:
1208:
1205:
1204:
1200:
1199:
1196:
1195:
1191:
1190:
1187:
1186:
1177:
1174:
1173:
1170:
1169:
1165:Waeringopterus
1160:
1155:
1153:
1150:
1149:
1146:
1145:
1136:
1131:
1129:
1121:
1119:
1117:Diploperculata
1111:
1103:Pterygotioidea
1084:Adelophthalmus
1042:appendages of
975:
974:
965:
964:
963:
954:
953:
952:
951:
950:
948:
947:Classification
945:
839:("dwarf") and
774:
771:
763:N. schiraensis
720:N. schiraensis
678:paleontologist
660:Clintonipterus
542:
539:
537:
534:
505:active swimmer
479:adelophthalmid
442:
439:
348:
347:
346:
345:
339:N. schiraensis
334:
323:
312:
301:
290:
279:
265:
264:
260:
259:
256:
246:
245:
239:
238:
224:
220:
219:
211:
207:
206:
198:
194:
193:
185:
181:
180:
175:
171:
170:
165:
161:
160:
155:
151:
150:
145:
141:
140:
127:
126:
119:
118:
110:
109:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
45:
28:
15:
13:
10:
9:
6:
4:
3:
2:
3564:
3553:
3550:
3548:
3545:
3543:
3540:
3538:
3535:
3533:
3530:
3528:
3525:
3523:
3520:
3519:
3517:
3500:
3495:
3491:
3487:
3482:
3478:
3474:
3469:
3465:
3461:
3456:
3452:
3447:
3441:
3437:
3436:
3434:
3432:
3428:
3424:
3419:
3409:
3399:
3393:
3390:
3388:
3385:
3384:
3382:
3378:
3372:
3369:
3367:
3364:
3362:
3359:
3358:
3356:
3352:
3346:
3345:
3341:
3339:
3338:
3334:
3332:
3331:
3327:
3326:
3324:
3322:
3318:
3312:
3311:
3307:
3305:
3304:
3300:
3298:
3297:
3293:
3291:
3290:
3289:Pentecopterus
3286:
3284:
3283:
3279:
3277:
3276:
3272:
3270:
3269:
3265:
3263:
3262:
3261:Megalograptus
3258:
3256:
3255:
3254:Jaekelopterus
3251:
3249:
3248:
3244:
3242:
3241:
3237:
3235:
3234:
3230:
3228:
3227:
3226:Drepanopterus
3223:
3221:
3220:
3216:
3214:
3213:
3209:
3207:
3206:
3205:Brachyopterus
3202:
3200:
3199:
3195:
3194:
3192:
3188:
3182:
3181:South America
3179:
3177:
3174:
3172:
3171:North America
3169:
3167:
3164:
3162:
3159:
3157:
3154:
3153:
3151:
3149:
3145:
3139:
3136:
3134:
3133:Carboniferous
3131:
3129:
3126:
3124:
3121:
3119:
3116:
3115:
3113:
3111:
3110:Geochronology
3107:
3102:
3080:
3077:
3075:
3072:
3070:
3067:
3066:
3064:
3062:
3058:
3052:
3049:
3048:
3046:
3044:
3040:
3034:
3031:
3030:
3028:
3026:
3022:
3016:
3013:
3011:
3008:
3006:
3003:
3002:
3000:
2998:
2994:
2991:
2989:
2983:
2977:
2974:
2972:
2969:
2967:
2964:
2963:
2961:
2959:
2958:Eurypteroidea
2955:
2949:
2946:
2945:
2943:
2941:
2937:
2931:
2928:
2927:
2925:
2923:
2919:
2916:
2914:
2908:
2898:
2897:Mycteroptidae
2895:
2893:
2890:
2888:
2885:
2884:
2882:
2880:
2879:Mycteropoidea
2876:
2870:
2867:
2865:
2862:
2861:
2859:
2857:
2853:
2847:
2844:
2842:
2839:
2838:
2836:
2834:
2833:Stylonuroidea
2830:
2824:
2823:Rhenopteridae
2821:
2820:
2818:
2816:
2812:
2809:
2807:
2801:
2795:
2792:
2791:
2789:
2785:
2782:
2778:
2772:
2771:Sclerophorata
2768:
2765:
2763:
2759:
2756:
2754:
2750:
2747:
2745:
2741:
2738:
2736:
2735:Euchelicerata
2732:
2729:
2727:
2723:
2721:
2717:
2715:
2711:
2710:
2707:
2703:
2696:
2691:
2689:
2684:
2682:
2677:
2676:
2673:
2663:
2659:
2652:
2650:
2641:
2638:
2626:
2622:
2616:
2613:
2601:
2597:
2591:
2588:
2576:
2572:
2566:
2563:
2551:
2547:
2541:
2538:
2533:
2521:
2513:
2509:
2505:
2499:
2495:
2491:
2490:
2482:
2479:
2471:
2467:
2463:
2459:
2455:
2454:
2446:
2439:
2436:
2431:
2427:
2423:
2419:
2415:
2411:
2410:
2402:
2399:
2395:
2389:
2387:
2383:
2378:
2366:
2358:
2357:
2349:
2346:
2341:
2337:
2333:
2329:
2325:
2321:
2314:
2312:
2308:
2305:
2301:
2300:
2296:
2289:
2286:
2281:
2277:
2273:
2269:
2265:
2261:
2254:
2252:
2248:
2243:
2239:
2235:
2231:
2224:
2222:
2218:
2213:
2209:
2205:
2201:
2197:
2193:
2186:
2183:
2172:on 2021-07-30
2171:
2167:
2163:
2162:
2154:
2152:
2150:
2146:
2142:
2136:
2133:
2128:
2124:
2119:
2114:
2110:
2106:
2102:
2098:
2097:Palaeontology
2091:
2088:
2083:
2079:
2072:
2065:
2063:
2061:
2059:
2055:
2050:
2046:
2042:
2038:
2031:
2029:
2027:
2025:
2023:
2021:
2019:
2017:
2013:
2008:
2004:
2000:
1996:
1992:
1988:
1984:
1977:
1975:
1973:
1971:
1969:
1967:
1965:
1963:
1961:
1959:
1957:
1953:
1948:
1944:
1940:
1935:
1930:
1926:
1922:
1918:
1914:
1910:
1906:
1905:
1900:
1893:
1891:
1889:
1887:
1885:
1881:
1875:
1871:
1868:
1866:
1863:
1862:
1858:
1856:
1854:
1850:
1849:
1844:
1843:
1839:
1835:
1831:
1830:
1825:
1824:
1819:
1815:
1811:
1807:
1803:
1799:
1795:
1794:
1789:
1785:
1781:
1780:
1775:
1771:
1767:
1763:
1759:
1755:
1754:
1750:
1746:
1745:
1741:
1740:malacostracan
1737:
1733:
1732:
1727:
1726:
1721:
1720:
1715:
1711:
1707:
1703:
1699:
1697:
1693:
1689:
1685:
1682:, along with
1681:
1677:
1673:
1672:North America
1669:
1665:
1661:
1658:
1654:
1650:
1642:
1635:
1634:
1626:
1625:
1617:
1616:
1608:
1607:
1599:
1598:
1590:
1589:
1581:
1580:
1572:
1571:
1563:
1562:
1559:
1558:
1557:
1556:Jaekelopterus
1550:
1549:
1546:
1545:
1542:
1541:
1540:
1533:
1532:
1526:
1525:
1522:
1521:
1518:
1517:
1516:
1509:
1508:
1502:
1501:
1498:
1497:
1494:
1493:
1492:
1485:
1484:
1478:
1477:
1474:
1473:
1470:
1469:
1468:
1461:
1460:
1454:
1453:
1450:
1449:
1446:
1445:
1444:
1437:
1436:
1430:
1429:
1426:
1425:
1422:
1421:
1420:
1413:
1412:
1406:
1403:
1402:
1399:
1398:
1390:
1389:
1381:
1380:
1372:
1371:
1363:
1362:
1359:
1358:
1357:
1350:
1349:
1346:
1345:
1342:
1341:
1340:
1333:
1332:
1326:
1325:
1322:
1321:
1318:
1317:
1316:
1309:
1308:
1302:
1301:
1298:
1297:
1289:
1288:
1285:
1284:
1283:
1276:
1275:
1272:
1271:
1268:
1267:
1266:
1259:
1258:
1252:
1251:
1245:
1244:
1241:
1240:
1237:
1236:
1235:
1234:Eysyslopterus
1228:
1227:
1221:
1218:
1217:
1211:
1210:
1207:
1206:
1198:
1197:
1189:
1188:
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1183:
1176:
1175:
1172:
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1168:
1167:
1166:
1159:
1158:
1152:
1151:
1148:
1147:
1144:
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1142:
1135:
1134:
1128:
1125:
1124:
1118:
1115:
1114:
1110:
1108:
1104:
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1095:
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1077:
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1069:
1065:
1064:
1059:
1055:
1054:
1049:
1045:
1041:
1037:
1032:
1030:
1029:N. lanceolata
1026:
1022:
1018:
1014:
1010:
1006:
1002:
998:
994:
985:
984:
979:
969:
958:
946:
944:
942:
941:Eurypteroidea
938:
937:N. lanceolata
934:
933:
928:
924:
923:Himantopterus
920:
916:
915:N. lanceolata
911:
909:
905:
901:
897:
893:
889:
885:
881:
877:
873:
868:
865:
860:
856:
852:
848:
844:
842:
838:
837:
832:
828:
824:
820:
816:
812:
808:
804:
800:
796:
788:
784:
779:
772:
770:
768:
764:
760:
756:
752:
748:
744:
739:
735:
734:
729:
725:
721:
716:
714:
713:N. lanceolata
710:
706:
702:
698:
694:
690:
686:
683:
679:
676:
673:In 1911, the
671:
669:
665:
661:
657:
653:
652:
647:
643:
642:United States
639:
635:
634:Clinton Group
631:
627:
623:
618:
616:
612:
609:
605:
600:
596:
592:
591:Herefordshire
588:
584:
580:
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575:
570:
569:
564:
560:
552:
547:
540:
535:
533:
531:
527:
525:
521:
517:
516:
511:
506:
502:
498:
494:
492:
488:
484:
480:
476:
475:N. lanceolata
472:
468:
464:
460:
452:
447:
440:
438:
436:
432:
428:
424:
420:
416:
412:
408:
404:
402:
398:
394:
390:
386:
385:United States
382:
378:
374:
371:. Fossils of
370:
366:
362:
361:
356:
355:
341:
340:
335:
330:
329:
324:
319:
318:
313:
310:Shpiney, 2012
308:
307:
302:
297:
296:
291:
286:
285:
284:N. lanceolata
280:
275:
274:
269:
268:
266:
261:
254:
253:
247:
244:
240:
233:
232:
225:
222:
221:
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212:
209:
208:
205:
199:
197:Superfamily:
196:
195:
192:
186:
183:
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179:
176:
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159:
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137:
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99:
94:
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84:
79:
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69:
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59:
54:
49:
43:
36:
32:
26:
22:
19:
3430:
3342:
3335:
3328:
3308:
3301:
3294:
3287:
3280:
3273:
3266:
3259:
3252:
3247:Hughmilleria
3245:
3238:
3231:
3224:
3217:
3210:
3203:
3196:
3079:Pterygotidae
3015:Mixopteridae
2971:Eurypteridae
2846:Stylonuridae
2766:
2757:
2748:
2739:
2730:
2661:
2657:
2648:
2640:
2628:. Retrieved
2624:
2615:
2603:. Retrieved
2599:
2590:
2578:. Retrieved
2574:
2565:
2553:. Retrieved
2549:
2540:
2488:
2481:
2470:the original
2457:
2451:
2438:
2413:
2407:
2401:
2355:
2348:
2331:
2327:
2323:
2320:Hughmilleria
2319:
2303:
2298:
2294:
2288:
2263:
2259:
2233:
2229:
2195:
2191:
2185:
2174:. Retrieved
2170:the original
2160:
2135:
2100:
2096:
2090:
2081:
2077:
2040:
2036:
1990:
1986:
1908:
1902:
1852:
1848:Arthrophycus
1846:
1840:
1838:gnathostomes
1833:
1827:
1821:
1817:
1813:
1810:Pennsylvania
1791:
1787:
1783:
1777:
1773:
1761:
1757:
1751:
1742:
1735:
1729:
1723:
1717:
1714:N. norvegica
1713:
1701:
1700:
1695:
1688:Hughmilleria
1687:
1683:
1679:
1648:
1646:
1643:Paleoecology
1554:
1553:
1537:
1536:
1513:
1512:
1491:Erettopterus
1489:
1488:
1465:
1464:
1441:
1440:
1419:Hughmilleria
1417:
1416:
1354:
1353:
1337:
1336:
1314:
1313:
1312:
1280:
1279:
1263:
1262:
1232:
1231:
1182:Grossopterus
1180:
1179:
1163:
1162:
1141:Orcanopterus
1139:
1138:
1096:
1087:
1083:
1079:
1076:sister taxon
1071:
1067:
1061:
1057:
1056:and perhaps
1051:
1047:
1044:Hughmilleria
1043:
1033:
1028:
1025:N. prominens
1024:
1017:polyphyletic
1013:Hughmilleria
1012:
1009:Hughmilleria
1008:
1004:
992:
991:
981:
977:
936:
932:Erettopterus
930:
922:
918:
914:
912:
907:
896:Hughmilleria
895:
891:
884:notosibirica
883:
871:
869:
863:
850:
846:
845:
841:Hughmilleria
840:
834:
826:
822:
818:
814:
810:
806:
803:Hughmilleria
802:
798:
795:Hughmilleria
794:
792:
786:
782:
766:
762:
758:
754:
731:
723:
719:
717:
712:
708:
705:N. norvegica
704:
700:
693:E. norvegica
692:
688:
672:
667:
663:
659:
656:N. prominens
655:
649:
645:
630:E. prominens
629:
625:
619:
614:
611:Hughmilleria
610:
594:
572:
566:
558:
556:
550:
529:
528:
515:Hughmilleria
513:
509:
500:
496:
495:
490:
486:
474:
470:
467:N. norvegica
466:
462:
456:
451:N. norvegica
450:
435:N. norvegica
434:
418:
406:
405:
372:
360:Hughmilleria
358:
353:
352:
351:
338:
337:
332:Salter, 1859
327:
326:
317:N. prominens
316:
315:
305:
304:
295:N. norvegica
294:
293:
288:Salter, 1856
283:
282:
272:
271:
251:
250:
243:Type species
230:
229:
124:N. prominens
123:
122:Carapace of
24:
18:
3344:Palmichnium
3321:Ichnogenera
3219:Carcinosoma
2913:Eurypterina
2806:Stylonurina
2744:Prosomapoda
2726:Chelicerata
2724:Subphylum:
2702:Eurypterids
2649:Dictyocaris
2373:|work=
2292:Meaning of
2198:: 207–213.
1842:Vernonaspis
1802:coelolepids
1692:pterygotids
1647:Fossils of
1265:Bassipterus
1099:Eurypterina
1063:Bassipterus
927:preoccupied
781:Fossils of
755:Rhenopterus
733:Rhenopterus
728:stylonurine
574:nomen nudum
483:eurypteroid
459:eurypterids
441:Description
383:age in the
191:Eurypterida
178:Chelicerata
174:Subphylum:
3516:Categories
3310:Stylonurus
3296:Pterygotus
3275:Mixopterus
3268:Megarachne
3233:Eurypterus
3118:Ordovician
3074:Slimonidae
2985:Infraorder
2762:Dekatriata
2753:Planaterga
2720:Arthropoda
2664:: 267–298.
2176:2018-09-06
1876:References
1814:N. clarkei
1774:N. pygmaea
1657:Llandovery
1539:Acutiramus
1515:Pterygotus
929:name, now
864:N. clarkei
847:N. clarkei
783:N. clarkei
689:Eurypterus
664:N. pygmaea
626:Eurypterus
622:James Hall
595:N. pygmaea
491:N. pygmaea
471:N. clarkei
431:morphology
427:appendages
369:arthropods
365:eurypterid
328:N. pygmaea
321:Hall, 1884
299:Kiær, 1911
273:N. clarkei
257:Kiær, 1911
168:Arthropoda
31:Llandovery
3387:Metastoma
3371:Xiphosura
3361:Arachnida
3148:Geography
2712:Kingdom:
2630:12 August
2605:12 August
2580:11 August
2555:11 August
2530:ignored (
2520:cite book
2512:0965-9994
2375:ignored (
2365:cite book
2280:128903698
2212:229469975
2204:0201-9280
2118:1808/8354
1925:1744-9561
1766:Khakassia
1706:Ringerike
1670:epoch in
986:(bottom).
823:norvegica
747:Minusinsk
738:posterior
682:geologist
675:Norwegian
670:in 1961.
620:In 1884,
481:, but an
154:Kingdom:
148:Eukaryota
3446:Q6962360
3440:Wikidata
3408:Category
3330:Arcuites
3303:Slimonia
3128:Devonian
3123:Silurian
2910:Suborder
2803:Suborder
2780:Taxonomy
2718:Phylum:
2714:Animalia
2430:83873103
2127:56066870
2084:: 79–90.
2007:59488956
1943:19828493
1859:See also
1690:and the
1668:Eifelian
1664:Devonian
1653:Silurian
1467:Slimonia
1105:and the
1094:family.
1040:prosomal
904:mesosoma
900:metasoma
880:paratype
876:holotype
662:. As in
608:subgenus
599:anterior
579:tergites
553:species.
524:podomere
522:in each
401:Scotland
381:Silurian
377:Devonian
357:("dwarf
263:Species
210:Family:
164:Phylum:
158:Animalia
144:Domain:
35:Eifelian
3486:3581779
3473:1054342
3460:4650089
3176:Oceania
3138:Permian
2340:1301566
2242:1301985
2164:(PhD).
2049:1301214
1934:2865068
1747:or the
1662:to the
751:Siberia
697:ventral
640:of the
587:Kington
503:was an
425:in its
397:England
223:Genus:
184:Order:
3166:Europe
3156:Africa
2787:Genera
2510:
2500:
2428:
2338:
2278:
2240:
2210:
2202:
2125:
2047:
2005:
1941:
1931:
1923:
1770:plants
1710:Norway
1676:Europe
730:genus
604:chelae
583:dorsal
520:spines
423:spines
393:Russia
389:Norway
3499:18985
3468:IRMNG
2767:Clade
2758:Clade
2749:Clade
2740:Clade
2731:Clade
2654:(PDF)
2473:(PDF)
2448:(PDF)
2426:S2CID
2336:JSTOR
2295:nanus
2276:S2CID
2238:JSTOR
2123:S2CID
2074:(PDF)
2045:JSTOR
2003:S2CID
1760:and
1660:epoch
892:notos
890:word
888:Greek
833:word
831:Latin
3455:GBIF
3161:Asia
2632:2018
2607:2018
2582:2018
2557:2018
2532:help
2508:ISSN
2498:ISBN
2377:help
2299:nani
2208:OCLC
2200:ISSN
1939:PMID
1921:ISSN
1845:and
1832:and
1800:and
1790:and
1734:and
1082:and
1066:and
1027:and
857:and
836:nana
722:and
680:and
648:and
636:, a
565:as "
399:and
379:and
48:PreꞒ
2462:doi
2458:252
2418:doi
2268:doi
2113:hdl
2105:doi
1995:doi
1929:PMC
1913:doi
925:(a
853:by
745:in
3518::
3496::
3483::
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2080:.
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1985:.
1955:^
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1927:.
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314:†?
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188:†
103:N
93:K
88:J
83:T
78:P
73:C
68:D
63:S
58:O
53:Ꞓ
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