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Nanahughmilleria

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778: 957: 968: 446: 136: 3101: 546: 3404: 695:. The carapace of this species was parabolic (approximately U-shaped) with small intramarginal arcuate eyes. The metastoma (a large plate that is part of the abdomen) was oval. Its operculum (a plate-like segment which contains the genital aperture) had a long, narrow spatulae (compared to most eurypterids). The telson was lanceolate, with a keel on both dorsal and 1086:. These eurypterids shared a series of characteristics such as the almost identical carapace, paddle, eye shapes and eye position. However, its short telson, thin cuticular sculpture (ornamentation consisting of small, minute, scales across the back), very small spatulae and increased spinosity in the appendages suggest a more basal position. Before 2004, 866:
in 1964, named after Clarke, who described the original Shawangunk eurypterid fauna. This small species of only 4 cm (1.6 in) had a lanceolate carapace with intramarginal eyes located anteriorly. The telson was broad and lanceolate. This species is characterized by the presence of spurs in the genal
507:
adapted to swim crosscurrent. It had epimera (lateral "extensions" of the segment) present in the seventh segment, separating the preadomen (body segments 1 to 7) from the postabodmen (segments 8 to 12). The genital spatulae, a long, flat piece in the genital aperture, was very short. The walking
606:(pincers) were small. It differs from other species in the more convergent form of the carapace and in the large size of the eyes, which were furthest from the margin. This species reached a length of 8 cm (3.1 in) in total. Erik N. Kjellesvig-Waering classified this species as part of the 654:. In addition, the carapace of this species was long and with ocelli placed forward. It has been suggested that this species could represent a separate genus. In fact, notes left by Kjellesvig-Waering in museum drawers indicate that he wanted to erect a new genus for 894:(southern) and Siberia, referring to the place where it was discovered. The carapace was parabolic and surrounded by a narrow marginal rim and with elongated eyes probably reniform. Only the fourth and fifth pair of prosomal appendages is known. These were of 2620: 601:
margin which narrows gradually anteriorly. A thin rim surrounded the carapace. The eyes were narrow, reniform and intramarginal. Its ocelli (light-sensitive simple eyes) were very small and located centrally. As in most eurypterid groups, the
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in 1912. They believed that the carapaces with intramarginal eyes were modified by the compression. While this could happen in a single specimen, it is impossible to be in so many. Kjellesvig-Waering realized this and erected the species
740:
margin was slightly concave. There is only one preserved eye, which was reniform, large and rose slightly from the surface of the carapace. At the closest point between both eyes, the ocelli were placed. The fossils were found at the
577:("naked name", a name coined without an adequate or completely missing description of it). The fossils mentioned by Banks consisted of two specimens, the first of which included carapace with the first five 417:. This clade was characterised by their small size, their parabolic (approximately U-shaped) carapaces and the presence of epimera (lateral "extensions" of the segment) on the seventh segment, among others. 2408: 1060:. However, the clade has almost never been used in subsequent studies and lists of eurypterids, and instead, they classify the nanahughmillerids as part of Adelophthalmidae. A basal clade between 1109:), as inferred by O. Erik Tetlie and Markus Poschmann in 2008, based on the results of a 2008 analysis specifically pertaining to the Adelophthalmoidea and a preceding 2004 analysis. 939:
lack eyes, making it difficult to determine their phylogenetic position. However, the prosomal and genital appendages were not typical of the Adelophthalmoidea, but probably of the
935:). The swimming legs of this species were long and narrow, reaching the sixth segment of the body. The telson was lanceolate and with a keel in the middle. All known specimens of 2452: 3467: 2140: 499:
had long and narrow reniform (bean-shaped) eyes placed in an intramarginal (occurring within the margin) position. The streamlined form of its body indicates that
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sides. It had a pair of spines in each podomere of the fifth appendage. The paddle of the swimming leg was relatively wide. It was designated the type species of
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parts of the body segments) and the 8th and 9th segments of the swimming leg and the second including another nearly complete carapace. Both were collected in
2406:
V. P. Tollerton (2003). "Summary of a revision of New York State Ordovician eurypterids: implications for eurypterid palaeoecology, diversity and evolution".
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is only distinguished from the more derived members of Adelophthalmidae in the shorter and smaller spatulae and in the increased spinosity in the appendages.
115: 2444: 644:. This species differs from the others by the anterior position of its eyes placed in a submarginal (almost in the margin) point, unlike other species of 526:(leg segment). The paddle of the swimming leg was relatively wide. As in the other adelophthalmids, the telson was lanceolate and not very expanded. 3454: 3551: 1097:
The cladogram below presents the inferred phylogenetic positions of most of the genera included in the three most derived superfamilies of the
825:
was designated as subgenotype. He assigned a total of seven species to the subgenus, of which only four (two of them tentatively) are still in
3536: 3180: 3170: 2692: 2501: 2493: 2159: 736:. This species is only known by a poorly preserved carapace which was semi-oval and elongated, with a narrow margin surrounding it. The 3132: 3147: 2595: 2570: 2545: 1705: 2190:
Pirozhnikov, L. P. (1957). "Remains of Gigantostraca from the series of Matakara (Devonian of the North Minusinsk Depression)".
3391: 3175: 3117: 1869: 3541: 3165: 3155: 3109: 135: 3531: 3526: 3127: 3122: 777: 3546: 3160: 3137: 2571:"Eurypterid-Associated Biota of the Shunet Fm. at Khakassia near Lake Shunet: Lochkovian - Eifelian, Russian Federation" 1020: 621: 956: 3521: 2095:
O. Erik Tetlie; Paul A. Selden; Dong Ren (2007). "A new Silurian eurypterid (Arthropoda: Chelicerata) from China".
967: 906:(the "median part"). The total size of the species could be of 9 cm (3.5 in), being one of the largest species of 2793: 2685: 2071:"A reappraisal of Eurypterus dumonti Stainier, 1917 and its position within the Adelophthalmidae Tollerton, 1989" 1864: 2469: 1899:"Cope's rule and Romer's theory: patterns of diversity and gigantism in eurypterids and Palaeozoic vertebrates" 898:-type and with rather small spines. The telson was small and wedge-shaped, known from fragmentary remains. The 3100: 2258:
Shpinev, E. S. (2012). "On some eurypterids (Eurypterida, Chelicerata) from the Devonian of South Siberia".
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and they assigned it in 1966 to its current genus. It has been suggested that this species is a synonym of
114: 3422: 2228:
Kjellesvig-Waering, Erik N.; Leutze, Willard P. (1966). "Eurypterids from the Silurian of West Virginia".
586: 3493: 2596:"Eurypterid-Associated Biota of the Temeside Shale, Ludlow and Perton, England: Pridoli, United Kingdom" 2165: 742: 3407: 2678: 1946: 1805: 637: 430: 571:". Although Richard Banks first mentioned this species in 1856, this act is currently considered as 737: 562: 1847: 761:, but the Russian paleontologist Evgeniy S. Shpinev does not agree with this since the prosoma of 2546:"Nanahughmilleria norvegica-Ostracoderm Comm. at Rudstangen, Ringerike Dist.: Rootsikula, Norway" 2519: 2425: 2364: 2335: 2275: 2237: 2122: 2044: 2002: 1797: 1752: 1743: 130: 2646: 1704:
can occur in an environment where eurypterids were abundant or scarce. The Silurian deposits of
910:. This species differs from the others in the proportions of the body and position of the eyes. 3498: 753:. Kjellesvig-Waering and Willard P. Leutze noticed that the species did not really represent a 3480: 3472: 2996: 2929: 2921: 2507: 2497: 2207: 2199: 1938: 1920: 1219: 1126: 1106: 1000: 921:. This species was originally described by Salter in 1856 as another species of his new genus 854: 414: 3485: 1983:"Phylogeny and palaeoecology of the Adelophthalmoidea (Arthropoda; Chelicerata; Eurypterida)" 465:
was a comparatively small eurypterid. The largest species confidently assigned to the genus,
3336: 3050: 3042: 3032: 3024: 3004: 2975: 2891: 2840: 2461: 2417: 2267: 2169: 2112: 2104: 1994: 1928: 1912: 1822: 1656: 1338: 1281: 1052: 996: 982: 858: 674: 650: 582: 410: 216: 203: 30: 2393: 3365: 3211: 3068: 3009: 2965: 2868: 2855: 2531: 2376: 2293: 1903: 1792: 1659: 1442: 1091: 1035: 519: 422: 1034:
In 2004, O. Erik Tetlie erected the family Nanahughmilleridae in a thesis to contain the
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Kjellesvig-Waering, Erik N. (1961). "The Silurian Eurypterida of the Welsh Borderland".
3281: 3239: 3197: 3060: 2987: 2947: 2939: 2863: 1933: 1748: 1404: 1355: 1164: 1116: 1102: 887: 677: 478: 2070: 545: 3515: 3288: 3260: 3253: 3225: 3204: 2957: 2896: 2886: 2878: 2832: 2822: 2814: 2770: 2734: 2279: 2108: 1982: 1828: 1778: 1724: 1671: 1555: 1233: 940: 641: 633: 603: 590: 426: 384: 72: 2429: 2126: 2006: 1898: 3246: 3078: 3014: 2970: 2845: 1837: 1809: 1739: 1730: 1694:, lived mainly in shallow lakes. In addition, the streamlined shape of the body of 1490: 1418: 1181: 1140: 1075: 1016: 931: 514: 359: 242: 2487: 2465: 2354: 3343: 3320: 3218: 2912: 2805: 2743: 2725: 1841: 1264: 1098: 1062: 926: 732: 727: 573: 482: 177: 47: 3445: 3309: 3295: 3274: 3267: 3232: 3073: 2761: 2752: 2701: 2421: 2271: 1998: 1801: 1718: 1691: 1538: 1514: 1011:, but Kjellesvig-Waering and Leutze considered it sufficiently different from 567: 458: 364: 190: 92: 57: 2511: 2211: 2203: 1924: 1698:
suggests that it was an active swimmer capable of swimming against currents.
3386: 3370: 3360: 2719: 1765: 746: 681: 368: 167: 147: 97: 41: 1942: 1916: 874:, was described by Shpinev. This species is known from two specimens, the 835: 813:) for species with small reniform intramarginal eyes. For the latter one, 3439: 3329: 3302: 1667: 1663: 1652: 1466: 903: 899: 879: 875: 607: 598: 523: 400: 380: 376: 87: 82: 67: 62: 52: 34: 2318:
Kjellesvig-Waering, Erik N. (1964). "Eurypterida: Notes on the Subgenus
3459: 2339: 2241: 2048: 1039: 750: 696: 578: 396: 102: 77: 2713: 2117: 1709: 1675: 718:
In 1957, L. P. Pirozhnikov described two new species of eurypterids,
504: 392: 388: 157: 3416: 2651:, Salter, a large crustacean from the Upper Silurian and Downtonian" 1038:
with no or reduced genital spatulae and the second to fifth pair of
1769: 830: 776: 444: 2445:"Distribution and dispersal history of Eurypterida (Chelicerata)" 902:(the posterior "part" of the body) was notably narrower than the 2621:"Delaware Water Gap Eurypterids (Silurian of the United States)" 2078:
Bulletin de l'Institut Royal des Sciences Naturelles de Belgique
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at 10 cm (3.9 in), making it a comparatively small eurypterid.
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Transactions of the Royal Society of Edinburgh: Earth Sciences
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and elevated the subgenus to the genus level. It is the only
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Eurypterid phylogeny with remarks on the origin of arachnids
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into two subgenera according to the morphology of the eyes,
433:. The largest species confidently assigned to the genus was 849:
is based on a series of fossils described and assigned to
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was longer and its eyes were closer to the margin than in
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by Kjellesvig-Waering in his description of the subgenus.
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was different from its relatives by the presence of more
477:
reached 16 cm (6.3 in), this species is probably not an
2396:. In World Spider Catalog. Natural History Museum Bern. 2356:
On some new Crustacea from the uppermost Silurian Rocks
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is better supported than the Nanahughmilleridae, while
469:, only reached 10 cm (3.9 in), while the smallest one, 403:, and have been referred to several different species. 1078:(closest relative) of the "derived clade" composed by 870:
It would not be until 2012 when another new species,
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A summary list of fossil spiders and their relatives
3429: 3379: 3353: 3319: 3189: 3146: 3108: 3059: 3041: 3023: 2995: 2984: 2956: 2938: 2920: 2909: 2877: 2854: 2831: 2813: 2802: 2786: 2779: 2192:Vsesojuzuoe Paleontologiceskoe Obchestvo Ezegodnik 2453:Palaeogeography, Palaeoclimatology, Palaeoecology 1981:Erik Tetlie, O.; Poschmann, Markus (2008-06-01). 1764:are the only animals of the Devonian deposits of 1019:genus (with grouped species that do not share an 805:) for species with large ovoid marginal eyes and 711:, measuring 10 cm (3.9 in, with the exception of 557:The first species of what today is recognized as 2392:Dunlop, J. A., Penney, D. & Jekel, D. 2018. 1716:have been discovered, include fossil remains of 773:Creation of the subgenus and further discoveries 1031:probably representing new and different forms. 2686: 1897:Lamsdell, James C.; Braddy, Simon J. (2009). 867:angles (a "flat" part of the carapace edge). 8: 1101:suborder of eurypterids (Adelophthalmoidea, 597:had an elongated carapace rounded along the 1796:, as well as some indeterminate species of 1776:from the Silurian has been associated with 3417: 2992: 2917: 2810: 2783: 2693: 2679: 2671: 1855:fossils have also been found in Scotland. 113: 20: 2116: 1932: 999:, the only family within the superfamily 715:, which may represent a separate genus). 473:, did not exceed 4 cm (1.6 in). Although 2313: 2311: 1007:was originally considered a subgenus of 544: 1880: 1023:) of Adelophthlmidae, with the species 726:, and erroneously assigned them to the 489:reached 9 cm (3.5 in) in length, while 2527: 2517: 2372: 2362: 2253: 2251: 829:. The generic name is composed by the 793:In 1961, Kjellesvig-Waering separated 632:, based on a single carapace from the 2153: 2151: 2149: 2069:Tetlie, O. Erik; van Roy, P. (2006). 1976: 917:is tentatively classified as part of 493:had a total length of 8 cm (3.1 in). 7: 2388: 2386: 2139:Clarke, J. K., Ruedemann R. (1912) " 2064: 2062: 2060: 2058: 1974: 1972: 1970: 1968: 1966: 1964: 1962: 1960: 1958: 1956: 1892: 1890: 1888: 1886: 1884: 995:is classified as part of the family 617:) in 1961 along many other species. 508:legs (first to fifth appendages) of 413:, the only clade in the superfamily 375:have been discovered in deposits of 2494:Joint Nature Conservation Committee 2223: 2221: 2030: 2028: 2026: 2024: 2022: 2020: 2018: 2016: 1987:Journal of Systematic Palaeontology 1772:. In addition, the English species 1530: 1506: 1482: 1458: 1434: 1410: 1330: 1306: 1256: 1249: 1225: 1215: 1156: 1132: 1122: 1112: 549:Size comparison of the unequivocal 2489:Fossil Arthropods of Great Britain 2326:) from the Silurian of New York". 1804:. In the Silurian deposits of the 14: 1738:, among other organisms like the 1678:and Siberia. It is believed that 882:(PIN 1139/2). Its specific name, 3403: 3402: 3099: 2168:. pp. 1–344. Archived from 2109:10.1111/j.1475-4983.2007.00651.x 966: 955: 134: 3392:Timeline of eurypterid research 2359:. Vol. 12. pp. 26–34. 1870:Timeline of eurypterid research 666:, this species was assigned to 518:-type, that is, with a pair of 1823:Ruedemannipterus stylonuroides 1725:Stylonuroides dolichopteroides 980:(top) and the closely related 457:Like the other adelophthalmid 367:, an extinct group of aquatic 1: 3552:Fossil taxa described in 1961 1768:, Russia, together with land 1762:Parahughmilleria matakarensis 1046:-type. This family contained 976:Type A genital appendages of 724:Parahughmilleria matarakensis 624:described another species of 3537:Eurypterids of North America 2486:Jarzembowski, E. A. (2010). 2466:10.1016/j.palaeo.2007.05.011 1758:Nanahughmilleria schiraensis 409:is classified in the family 2141:The Eurypterida of New York 707:was the largest species of 687:described a new species of 429:(limbs) and by its genital 16:Extinct genus of arthropods 3568: 2658:Extract Norsk Geol. Tisskr 978:Nanahughmilleria norvegica 252:Nanahughmilleria norvegica 3400: 3097: 2794:List of eurypterid genera 2708: 2422:10.1017/S026359330000064X 2272:10.1134/S0031030112040119 1999:10.1017/S1477201907002416 1865:List of eurypterid genera 1779:Salteropterus abbreviatus 1651:have been found from the 1552: 1535: 1528: 1511: 1504: 1487: 1480: 1463: 1456: 1439: 1432: 1415: 1408: 1352: 1335: 1328: 1311: 1304: 1278: 1261: 1254: 1247: 1230: 1223: 1213: 1178: 1161: 1154: 1137: 1130: 1120: 1021:immediate common ancestor 983:Pittsfordipterus phelpsae 913:In addition, the species 561:was described in 1859 by 267: 262: 248: 241: 131:Scientific classification 129: 121: 112: 23: 2443:Tetlie, O. Erik (2007). 2353:Salter, John W. (1856). 2158:Tetlie, O. Erik (2004). 1784:Parahughmilleria salteri 485:. Other species such as 277:Kjellesvig-Waering, 1964 236:Kjellesvig-Waering, 1961 2328:Journal of Paleontology 2260:Paleontological Journal 2230:Journal of Paleontology 2037:Journal of Paleontology 1834:Hughmilleria shawangunk 1793:Herefordopterus banksii 1788:Erettopterus spatulatus 1731:Kiaeropterus ruedemanni 851:Hughmilleria shawangunk 787:Hughmilleria shawangunk 785:previously assigned to 2645:Størmer, Leif (1935). 1917:10.1098/rsbl.2009.0700 1851:have also been found. 1818:Parahughmilleria maria 1090:was classified in the 790: 554: 454: 3542:Eurypterids of Europe 3494:Paleobiology Database 2166:University of Bristol 1947:Supplemental material 1756:. On the other hand, 780: 548: 448: 3532:Devonian eurypterids 3527:Silurian eurypterids 2304:latin-dictionary.net 1816:occur together with 1806:Shawangunk Formation 685:Johan Aschehoug Kiær 638:geological formation 3547:Eurypterids of Asia 1829:Hardieopterus myops 1744:Dictyocaris slimoni 1712:, where fossils of 568:Eurypterus pygmaeus 563:John William Salter 536:History of research 2922:Onychopterelloidea 1798:cephalaspidomorphs 1753:Kiaeria limuloides 1736:Erettopterus holmi 1074:is considered the 791: 555: 455: 3522:Adelophthalmoidea 3509: 3508: 3481:Open Tree of Life 3423:Taxon identifiers 3414: 3413: 3095: 3094: 3091: 3090: 3087: 3086: 3043:Adelophthalmoidea 3025:Waeringopteroidea 2997:Carcinosomatoidea 2930:Onychopterellidae 2905: 2904: 2503:978-1-86107-486-7 1836:. Species of the 1719:Mixopterus kiaeri 1639: 1638: 1630: 1629: 1621: 1620: 1612: 1611: 1603: 1602: 1594: 1593: 1585: 1584: 1576: 1575: 1567: 1566: 1394: 1393: 1385: 1384: 1376: 1375: 1367: 1366: 1293: 1292: 1220:Adelophthalmoidea 1202: 1201: 1193: 1192: 1127:Waeringopteroidea 1001:Adelophthalmoidea 886:, comes from the 878:(PIN 1139/1) and 855:John Mason Clarke 743:Matarak Formation 541:First discoveries 512:were probably of 415:Adelophthalmoidea 363:") is a genus of 350: 349: 344: 343:Pirozhnikov, 1957 333: 322: 311: 300: 289: 278: 237: 204:Adelophthalmoidea 3559: 3502: 3501: 3489: 3488: 3476: 3475: 3463: 3462: 3450: 3449: 3448: 3431:Nanahughmilleria 3418: 3406: 3405: 3380:Related articles 3337:Merostomichnites 3103: 3051:Adelophthalmidae 3033:Waeringopteridae 3005:Carcinosomatidae 2993: 2976:Strobilopteridae 2918: 2892:Hibbertopteridae 2841:Parastylonuridae 2811: 2784: 2695: 2688: 2681: 2672: 2666: 2665: 2655: 2642: 2636: 2635: 2633: 2631: 2617: 2611: 2610: 2608: 2606: 2592: 2586: 2585: 2583: 2581: 2567: 2561: 2560: 2558: 2556: 2542: 2536: 2535: 2529: 2525: 2523: 2515: 2492:. Vol. 35. 2483: 2477: 2476: 2474: 2468:. Archived from 2460:(3–4): 557–574. 2449: 2440: 2434: 2433: 2403: 2397: 2390: 2381: 2380: 2374: 2370: 2368: 2360: 2350: 2344: 2343: 2324:Nanahughmilleria 2315: 2306: 2290: 2284: 2283: 2255: 2246: 2245: 2236:(5): 1109–1122. 2225: 2216: 2215: 2187: 2181: 2180: 2178: 2177: 2155: 2144: 2137: 2131: 2130: 2120: 2092: 2086: 2085: 2075: 2066: 2053: 2052: 2032: 2011: 2010: 1978: 1951: 1950: 1936: 1894: 1853:Nanahughmilleria 1702:Nanahughmilleria 1696:Nanahughmilleria 1684:Parahughmilleria 1680:Nanahughmilleria 1666:deposits of the 1655:deposits of the 1649:Nanahughmilleria 1531: 1507: 1483: 1459: 1435: 1411: 1339:Parahughmilleria 1331: 1315:Nanahughmilleria 1307: 1282:Pittsfordipterus 1257: 1250: 1226: 1216: 1157: 1133: 1123: 1113: 1107:waeringopteroids 1088:Nanahughmilleria 1080:Parahughmilleria 1072:Nanahughmilleria 1068:Pittsfordipterus 1058:Parahughmilleria 1053:Pittsfordipterus 1036:adelophthalmoids 1005:Nanahughmilleria 997:Adelophthalmidae 993:Nanahughmilleria 970: 959: 919:Nanahughmilleria 908:Nanahughmilleria 859:Rudolf Ruedemann 827:Nanahughmilleria 811:Nanahughmilleria 709:Nanahughmilleria 701:Nanahughmilleria 668:Nanahughmilleria 651:Parahughmilleria 646:Nanahughmilleria 615:Nanahughmilleria 559:Nanahughmilleria 551:Nanahughmilleria 530:Nanahughmilleria 510:Nanahughmilleria 501:Nanahughmilleria 497:Nanahughmilleria 463:Nanahughmilleria 419:Nanahughmilleria 411:Adelophthalmidae 407:Nanahughmilleria 373:Nanahughmilleria 354:Nanahughmilleria 342: 331: 320: 309: 298: 287: 276: 235: 231:Nanahughmilleria 228: 217:Adelophthalmidae 215: 202: 189: 139: 138: 117: 107: 44: 29:Temporal range: 25:Nanahughmilleria 21: 3567: 3566: 3562: 3561: 3560: 3558: 3557: 3556: 3512: 3511: 3510: 3505: 3497: 3492: 3484: 3479: 3471: 3466: 3458: 3453: 3444: 3443: 3438: 3425: 3415: 3410: 3396: 3375: 3366:Chasmataspidida 3349: 3315: 3212:Campylocephalus 3185: 3142: 3104: 3083: 3069:Hughmilleriidae 3055: 3037: 3019: 3010:Megalograptidae 2986: 2980: 2966:Dolichopteridae 2952: 2940:Moselopteroidea 2934: 2911: 2901: 2887:Drepanopteridae 2873: 2869:Hardieopteridae 2856:Kokomopteroidea 2850: 2827: 2804: 2798: 2775: 2704: 2699: 2669: 2653: 2644: 2643: 2639: 2629: 2627: 2625:fossilworks.org 2619: 2618: 2614: 2604: 2602: 2600:fossilworks.org 2594: 2593: 2589: 2579: 2577: 2575:fossilworks.org 2569: 2568: 2564: 2554: 2552: 2550:fossilworks.org 2544: 2543: 2539: 2526: 2516: 2504: 2496:. p. 294. 2485: 2484: 2480: 2472: 2447: 2442: 2441: 2437: 2405: 2404: 2400: 2391: 2384: 2371: 2361: 2352: 2351: 2347: 2317: 2316: 2309: 2291: 2287: 2257: 2256: 2249: 2227: 2226: 2219: 2189: 2188: 2184: 2175: 2173: 2157: 2156: 2147: 2138: 2134: 2094: 2093: 2089: 2073: 2068: 2067: 2056: 2034: 2033: 2014: 1980: 1979: 1954: 1904:Biology Letters 1896: 1895: 1882: 1878: 1861: 1645: 1640: 1631: 1622: 1613: 1604: 1595: 1586: 1577: 1568: 1443:Herefordopterus 1395: 1386: 1377: 1368: 1294: 1203: 1194: 1092:Hughmilleriidae 1048:Nanahughmileria 990: 989: 988: 987: 973: 972: 971: 962: 961: 960: 949: 872:N. notosibirica 775: 767:P. matarakensis 759:P. matarakensis 543: 538: 487:N. notosibirica 449:Restoration of 443: 306:N. notosibirica 258: 255: 234: 226: 213: 200: 187: 133: 108: 106: 105: 100: 95: 90: 85: 80: 75: 70: 65: 60: 55: 50: 40:435–387.7  39: 38: 27: 17: 12: 11: 5: 3565: 3563: 3555: 3554: 3549: 3544: 3539: 3534: 3529: 3524: 3514: 3513: 3507: 3506: 3504: 3503: 3490: 3477: 3464: 3451: 3435: 3433: 3427: 3426: 3421: 3412: 3411: 3401: 3398: 3397: 3395: 3394: 3389: 3383: 3381: 3377: 3376: 3374: 3373: 3368: 3363: 3357: 3355: 3354:Related groups 3351: 3350: 3348: 3347: 3340: 3333: 3325: 3323: 3317: 3316: 3314: 3313: 3306: 3299: 3292: 3285: 3282:Onychopterella 3278: 3271: 3264: 3257: 3250: 3243: 3240:Hibbertopterus 3236: 3229: 3222: 3215: 3208: 3201: 3198:Adelophthalmus 3193: 3191: 3190:Notable genera 3187: 3186: 3184: 3183: 3178: 3173: 3168: 3163: 3158: 3152: 3150: 3144: 3143: 3141: 3140: 3135: 3130: 3125: 3120: 3114: 3112: 3106: 3105: 3098: 3096: 3093: 3092: 3089: 3088: 3085: 3084: 3082: 3081: 3076: 3071: 3065: 3063: 3061:Pterygotioidea 3057: 3056: 3054: 3053: 3047: 3045: 3039: 3038: 3036: 3035: 3029: 3027: 3021: 3020: 3018: 3017: 3012: 3007: 3001: 2999: 2990: 2988:Diploperculata 2982: 2981: 2979: 2978: 2973: 2968: 2962: 2960: 2954: 2953: 2951: 2950: 2948:Moselopteridae 2944: 2942: 2936: 2935: 2933: 2932: 2926: 2924: 2915: 2907: 2906: 2903: 2902: 2900: 2899: 2894: 2889: 2883: 2881: 2875: 2874: 2872: 2871: 2866: 2864:Kokomopteridae 2860: 2858: 2852: 2851: 2849: 2848: 2843: 2837: 2835: 2829: 2828: 2826: 2825: 2819: 2817: 2815:Rhenopteroidea 2808: 2800: 2799: 2797: 2796: 2790: 2788: 2781: 2777: 2776: 2774: 2773: 2764: 2755: 2746: 2737: 2728: 2722: 2716: 2709: 2706: 2705: 2700: 2698: 2697: 2690: 2683: 2675: 2668: 2667: 2637: 2612: 2587: 2562: 2537: 2528:|journal= 2502: 2478: 2475:on 2011-07-18. 2435: 2416:(3): 235–242. 2398: 2382: 2345: 2334:(2): 410–412. 2307: 2285: 2266:(4): 370–377. 2247: 2217: 2182: 2145: 2132: 2103:(3): 619–625. 2087: 2054: 2043:(4): 789–835. 2012: 1993:(2): 237–249. 1952: 1911:(2): 265–269. 1879: 1877: 1874: 1873: 1872: 1867: 1860: 1857: 1749:chasmataspidid 1644: 1641: 1637: 1636: 1633: 1632: 1628: 1627: 1624: 1623: 1619: 1618: 1615: 1614: 1610: 1609: 1606: 1605: 1601: 1600: 1597: 1596: 1592: 1591: 1588: 1587: 1583: 1582: 1579: 1578: 1574: 1573: 1570: 1569: 1565: 1564: 1561: 1560: 1551: 1548: 1547: 1544: 1543: 1534: 1529: 1527: 1524: 1523: 1520: 1519: 1510: 1505: 1503: 1500: 1499: 1496: 1495: 1486: 1481: 1479: 1476: 1475: 1472: 1471: 1462: 1457: 1455: 1452: 1451: 1448: 1447: 1438: 1433: 1431: 1428: 1427: 1424: 1423: 1414: 1409: 1407: 1405:Pterygotioidea 1401: 1400: 1397: 1396: 1392: 1391: 1388: 1387: 1383: 1382: 1379: 1378: 1374: 1373: 1370: 1369: 1365: 1364: 1361: 1360: 1356:Adelophthalmus 1351: 1348: 1347: 1344: 1343: 1334: 1329: 1327: 1324: 1323: 1320: 1319: 1310: 1305: 1303: 1300: 1299: 1296: 1295: 1291: 1290: 1287: 1286: 1277: 1274: 1273: 1270: 1269: 1260: 1255: 1253: 1248: 1246: 1243: 1242: 1239: 1238: 1229: 1224: 1222: 1214: 1212: 1209: 1208: 1205: 1204: 1200: 1199: 1196: 1195: 1191: 1190: 1187: 1186: 1177: 1174: 1173: 1170: 1169: 1165:Waeringopterus 1160: 1155: 1153: 1150: 1149: 1146: 1145: 1136: 1131: 1129: 1121: 1119: 1117:Diploperculata 1111: 1103:Pterygotioidea 1084:Adelophthalmus 1042:appendages of 975: 974: 965: 964: 963: 954: 953: 952: 951: 950: 948: 947:Classification 945: 839:("dwarf") and 774: 771: 763:N. schiraensis 720:N. schiraensis 678:paleontologist 660:Clintonipterus 542: 539: 537: 534: 505:active swimmer 479:adelophthalmid 442: 439: 348: 347: 346: 345: 339:N. schiraensis 334: 323: 312: 301: 290: 279: 265: 264: 260: 259: 256: 246: 245: 239: 238: 224: 220: 219: 211: 207: 206: 198: 194: 193: 185: 181: 180: 175: 171: 170: 165: 161: 160: 155: 151: 150: 145: 141: 140: 127: 126: 119: 118: 110: 109: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 51: 46: 45: 28: 15: 13: 10: 9: 6: 4: 3: 2: 3564: 3553: 3550: 3548: 3545: 3543: 3540: 3538: 3535: 3533: 3530: 3528: 3525: 3523: 3520: 3519: 3517: 3500: 3495: 3491: 3487: 3482: 3478: 3474: 3469: 3465: 3461: 3456: 3452: 3447: 3441: 3437: 3436: 3434: 3432: 3428: 3424: 3419: 3409: 3399: 3393: 3390: 3388: 3385: 3384: 3382: 3378: 3372: 3369: 3367: 3364: 3362: 3359: 3358: 3356: 3352: 3346: 3345: 3341: 3339: 3338: 3334: 3332: 3331: 3327: 3326: 3324: 3322: 3318: 3312: 3311: 3307: 3305: 3304: 3300: 3298: 3297: 3293: 3291: 3290: 3289:Pentecopterus 3286: 3284: 3283: 3279: 3277: 3276: 3272: 3270: 3269: 3265: 3263: 3262: 3261:Megalograptus 3258: 3256: 3255: 3254:Jaekelopterus 3251: 3249: 3248: 3244: 3242: 3241: 3237: 3235: 3234: 3230: 3228: 3227: 3226:Drepanopterus 3223: 3221: 3220: 3216: 3214: 3213: 3209: 3207: 3206: 3205:Brachyopterus 3202: 3200: 3199: 3195: 3194: 3192: 3188: 3182: 3181:South America 3179: 3177: 3174: 3172: 3171:North America 3169: 3167: 3164: 3162: 3159: 3157: 3154: 3153: 3151: 3149: 3145: 3139: 3136: 3134: 3133:Carboniferous 3131: 3129: 3126: 3124: 3121: 3119: 3116: 3115: 3113: 3111: 3110:Geochronology 3107: 3102: 3080: 3077: 3075: 3072: 3070: 3067: 3066: 3064: 3062: 3058: 3052: 3049: 3048: 3046: 3044: 3040: 3034: 3031: 3030: 3028: 3026: 3022: 3016: 3013: 3011: 3008: 3006: 3003: 3002: 3000: 2998: 2994: 2991: 2989: 2983: 2977: 2974: 2972: 2969: 2967: 2964: 2963: 2961: 2959: 2958:Eurypteroidea 2955: 2949: 2946: 2945: 2943: 2941: 2937: 2931: 2928: 2927: 2925: 2923: 2919: 2916: 2914: 2908: 2898: 2897:Mycteroptidae 2895: 2893: 2890: 2888: 2885: 2884: 2882: 2880: 2879:Mycteropoidea 2876: 2870: 2867: 2865: 2862: 2861: 2859: 2857: 2853: 2847: 2844: 2842: 2839: 2838: 2836: 2834: 2833:Stylonuroidea 2830: 2824: 2823:Rhenopteridae 2821: 2820: 2818: 2816: 2812: 2809: 2807: 2801: 2795: 2792: 2791: 2789: 2785: 2782: 2778: 2772: 2771:Sclerophorata 2768: 2765: 2763: 2759: 2756: 2754: 2750: 2747: 2745: 2741: 2738: 2736: 2735:Euchelicerata 2732: 2729: 2727: 2723: 2721: 2717: 2715: 2711: 2710: 2707: 2703: 2696: 2691: 2689: 2684: 2682: 2677: 2676: 2673: 2663: 2659: 2652: 2650: 2641: 2638: 2626: 2622: 2616: 2613: 2601: 2597: 2591: 2588: 2576: 2572: 2566: 2563: 2551: 2547: 2541: 2538: 2533: 2521: 2513: 2509: 2505: 2499: 2495: 2491: 2490: 2482: 2479: 2471: 2467: 2463: 2459: 2455: 2454: 2446: 2439: 2436: 2431: 2427: 2423: 2419: 2415: 2411: 2410: 2402: 2399: 2395: 2389: 2387: 2383: 2378: 2366: 2358: 2357: 2349: 2346: 2341: 2337: 2333: 2329: 2325: 2321: 2314: 2312: 2308: 2305: 2301: 2300: 2296: 2289: 2286: 2281: 2277: 2273: 2269: 2265: 2261: 2254: 2252: 2248: 2243: 2239: 2235: 2231: 2224: 2222: 2218: 2213: 2209: 2205: 2201: 2197: 2193: 2186: 2183: 2172:on 2021-07-30 2171: 2167: 2163: 2162: 2154: 2152: 2150: 2146: 2142: 2136: 2133: 2128: 2124: 2119: 2114: 2110: 2106: 2102: 2098: 2097:Palaeontology 2091: 2088: 2083: 2079: 2072: 2065: 2063: 2061: 2059: 2055: 2050: 2046: 2042: 2038: 2031: 2029: 2027: 2025: 2023: 2021: 2019: 2017: 2013: 2008: 2004: 2000: 1996: 1992: 1988: 1984: 1977: 1975: 1973: 1971: 1969: 1967: 1965: 1963: 1961: 1959: 1957: 1953: 1948: 1944: 1940: 1935: 1930: 1926: 1922: 1918: 1914: 1910: 1906: 1905: 1900: 1893: 1891: 1889: 1887: 1885: 1881: 1875: 1871: 1868: 1866: 1863: 1862: 1858: 1856: 1854: 1850: 1849: 1844: 1843: 1839: 1835: 1831: 1830: 1825: 1824: 1819: 1815: 1811: 1807: 1803: 1799: 1795: 1794: 1789: 1785: 1781: 1780: 1775: 1771: 1767: 1763: 1759: 1755: 1754: 1750: 1746: 1745: 1741: 1740:malacostracan 1737: 1733: 1732: 1727: 1726: 1721: 1720: 1715: 1711: 1707: 1703: 1699: 1697: 1693: 1689: 1685: 1682:, along with 1681: 1677: 1673: 1672:North America 1669: 1665: 1661: 1658: 1654: 1650: 1642: 1635: 1634: 1626: 1625: 1617: 1616: 1608: 1607: 1599: 1598: 1590: 1589: 1581: 1580: 1572: 1571: 1563: 1562: 1559: 1558: 1557: 1556:Jaekelopterus 1550: 1549: 1546: 1545: 1542: 1541: 1540: 1533: 1532: 1526: 1525: 1522: 1521: 1518: 1517: 1516: 1509: 1508: 1502: 1501: 1498: 1497: 1494: 1493: 1492: 1485: 1484: 1478: 1477: 1474: 1473: 1470: 1469: 1468: 1461: 1460: 1454: 1453: 1450: 1449: 1446: 1445: 1444: 1437: 1436: 1430: 1429: 1426: 1425: 1422: 1421: 1420: 1413: 1412: 1406: 1403: 1402: 1399: 1398: 1390: 1389: 1381: 1380: 1372: 1371: 1363: 1362: 1359: 1358: 1357: 1350: 1349: 1346: 1345: 1342: 1341: 1340: 1333: 1332: 1326: 1325: 1322: 1321: 1318: 1317: 1316: 1309: 1308: 1302: 1301: 1298: 1297: 1289: 1288: 1285: 1284: 1283: 1276: 1275: 1272: 1271: 1268: 1267: 1266: 1259: 1258: 1252: 1251: 1245: 1244: 1241: 1240: 1237: 1236: 1235: 1234:Eysyslopterus 1228: 1227: 1221: 1218: 1217: 1211: 1210: 1207: 1206: 1198: 1197: 1189: 1188: 1185: 1184: 1183: 1176: 1175: 1172: 1171: 1168: 1167: 1166: 1159: 1158: 1152: 1151: 1148: 1147: 1144: 1143: 1142: 1135: 1134: 1128: 1125: 1124: 1118: 1115: 1114: 1110: 1108: 1104: 1100: 1095: 1093: 1089: 1085: 1081: 1077: 1073: 1069: 1065: 1064: 1059: 1055: 1054: 1049: 1045: 1041: 1037: 1032: 1030: 1029:N. lanceolata 1026: 1022: 1018: 1014: 1010: 1006: 1002: 998: 994: 985: 984: 979: 969: 958: 946: 944: 942: 941:Eurypteroidea 938: 937:N. lanceolata 934: 933: 928: 924: 923:Himantopterus 920: 916: 915:N. lanceolata 911: 909: 905: 901: 897: 893: 889: 885: 881: 877: 873: 868: 865: 860: 856: 852: 848: 844: 842: 838: 837: 832: 828: 824: 820: 816: 812: 808: 804: 800: 796: 788: 784: 779: 772: 770: 768: 764: 760: 756: 752: 748: 744: 739: 735: 734: 729: 725: 721: 716: 714: 713:N. lanceolata 710: 706: 702: 698: 694: 690: 686: 683: 679: 676: 673:In 1911, the 671: 669: 665: 661: 657: 653: 652: 647: 643: 642:United States 639: 635: 634:Clinton Group 631: 627: 623: 618: 616: 612: 609: 605: 600: 596: 592: 591:Herefordshire 588: 584: 580: 576: 575: 570: 569: 564: 560: 552: 547: 540: 535: 533: 531: 527: 525: 521: 517: 516: 511: 506: 502: 498: 494: 492: 488: 484: 480: 476: 475:N. lanceolata 472: 468: 464: 460: 452: 447: 440: 438: 436: 432: 428: 424: 420: 416: 412: 408: 404: 402: 398: 394: 390: 386: 385:United States 382: 378: 374: 371:. Fossils of 370: 366: 362: 361: 356: 355: 341: 340: 335: 330: 329: 324: 319: 318: 313: 310:Shpiney, 2012 308: 307: 302: 297: 296: 291: 286: 285: 284:N. lanceolata 280: 275: 274: 269: 268: 266: 261: 254: 253: 247: 244: 240: 233: 232: 225: 222: 221: 218: 212: 209: 208: 205: 199: 197:Superfamily: 196: 195: 192: 186: 183: 182: 179: 176: 173: 172: 169: 166: 163: 162: 159: 156: 153: 152: 149: 146: 143: 142: 137: 132: 128: 125: 120: 116: 111: 104: 99: 94: 89: 84: 79: 74: 69: 64: 59: 54: 49: 43: 36: 32: 26: 22: 19: 3430: 3342: 3335: 3328: 3308: 3301: 3294: 3287: 3280: 3273: 3266: 3259: 3252: 3247:Hughmilleria 3245: 3238: 3231: 3224: 3217: 3210: 3203: 3196: 3079:Pterygotidae 3015:Mixopteridae 2971:Eurypteridae 2846:Stylonuridae 2766: 2757: 2748: 2739: 2730: 2661: 2657: 2648: 2640: 2628:. Retrieved 2624: 2615: 2603:. Retrieved 2599: 2590: 2578:. Retrieved 2574: 2565: 2553:. Retrieved 2549: 2540: 2488: 2481: 2470:the original 2457: 2451: 2438: 2413: 2407: 2401: 2355: 2348: 2331: 2327: 2323: 2320:Hughmilleria 2319: 2303: 2298: 2294: 2288: 2263: 2259: 2233: 2229: 2195: 2191: 2185: 2174:. Retrieved 2170:the original 2160: 2135: 2100: 2096: 2090: 2081: 2077: 2040: 2036: 1990: 1986: 1908: 1902: 1852: 1848:Arthrophycus 1846: 1840: 1838:gnathostomes 1833: 1827: 1821: 1817: 1813: 1810:Pennsylvania 1791: 1787: 1783: 1777: 1773: 1761: 1757: 1751: 1742: 1735: 1729: 1723: 1717: 1714:N. norvegica 1713: 1701: 1700: 1695: 1688:Hughmilleria 1687: 1683: 1679: 1648: 1646: 1643:Paleoecology 1554: 1553: 1537: 1536: 1513: 1512: 1491:Erettopterus 1489: 1488: 1465: 1464: 1441: 1440: 1419:Hughmilleria 1417: 1416: 1354: 1353: 1337: 1336: 1314: 1313: 1312: 1280: 1279: 1263: 1262: 1232: 1231: 1182:Grossopterus 1180: 1179: 1163: 1162: 1141:Orcanopterus 1139: 1138: 1096: 1087: 1083: 1079: 1076:sister taxon 1071: 1067: 1061: 1057: 1056:and perhaps 1051: 1047: 1044:Hughmilleria 1043: 1033: 1028: 1025:N. prominens 1024: 1017:polyphyletic 1013:Hughmilleria 1012: 1009:Hughmilleria 1008: 1004: 992: 991: 981: 977: 936: 932:Erettopterus 930: 922: 918: 914: 912: 907: 896:Hughmilleria 895: 891: 884:notosibirica 883: 871: 869: 863: 850: 846: 845: 841:Hughmilleria 840: 834: 826: 822: 818: 814: 810: 806: 803:Hughmilleria 802: 798: 795:Hughmilleria 794: 792: 786: 782: 766: 762: 758: 754: 731: 723: 719: 717: 712: 708: 705:N. norvegica 704: 700: 693:E. norvegica 692: 688: 672: 667: 663: 659: 656:N. prominens 655: 649: 645: 630:E. prominens 629: 625: 619: 614: 611:Hughmilleria 610: 594: 572: 566: 558: 556: 550: 529: 528: 515:Hughmilleria 513: 509: 500: 496: 495: 490: 486: 474: 470: 467:N. norvegica 466: 462: 456: 451:N. norvegica 450: 435:N. norvegica 434: 418: 406: 405: 372: 360:Hughmilleria 358: 353: 352: 351: 338: 337: 332:Salter, 1859 327: 326: 317:N. prominens 316: 315: 305: 304: 295:N. norvegica 294: 293: 288:Salter, 1856 283: 282: 272: 271: 251: 250: 243:Type species 230: 229: 124:N. prominens 123: 122:Carapace of 24: 18: 3344:Palmichnium 3321:Ichnogenera 3219:Carcinosoma 2913:Eurypterina 2806:Stylonurina 2744:Prosomapoda 2726:Chelicerata 2724:Subphylum: 2702:Eurypterids 2649:Dictyocaris 2373:|work= 2292:Meaning of 2198:: 207–213. 1842:Vernonaspis 1802:coelolepids 1692:pterygotids 1647:Fossils of 1265:Bassipterus 1099:Eurypterina 1063:Bassipterus 927:preoccupied 781:Fossils of 755:Rhenopterus 733:Rhenopterus 728:stylonurine 574:nomen nudum 483:eurypteroid 459:eurypterids 441:Description 383:age in the 191:Eurypterida 178:Chelicerata 174:Subphylum: 3516:Categories 3310:Stylonurus 3296:Pterygotus 3275:Mixopterus 3268:Megarachne 3233:Eurypterus 3118:Ordovician 3074:Slimonidae 2985:Infraorder 2762:Dekatriata 2753:Planaterga 2720:Arthropoda 2664:: 267–298. 2176:2018-09-06 1876:References 1814:N. clarkei 1774:N. pygmaea 1657:Llandovery 1539:Acutiramus 1515:Pterygotus 929:name, now 864:N. clarkei 847:N. clarkei 783:N. clarkei 689:Eurypterus 664:N. pygmaea 626:Eurypterus 622:James Hall 595:N. pygmaea 491:N. pygmaea 471:N. clarkei 431:morphology 427:appendages 369:arthropods 365:eurypterid 328:N. pygmaea 321:Hall, 1884 299:Kiær, 1911 273:N. clarkei 257:Kiær, 1911 168:Arthropoda 31:Llandovery 3387:Metastoma 3371:Xiphosura 3361:Arachnida 3148:Geography 2712:Kingdom: 2630:12 August 2605:12 August 2580:11 August 2555:11 August 2530:ignored ( 2520:cite book 2512:0965-9994 2375:ignored ( 2365:cite book 2280:128903698 2212:229469975 2204:0201-9280 2118:1808/8354 1925:1744-9561 1766:Khakassia 1706:Ringerike 1670:epoch in 986:(bottom). 823:norvegica 747:Minusinsk 738:posterior 682:geologist 675:Norwegian 670:in 1961. 620:In 1884, 481:, but an 154:Kingdom: 148:Eukaryota 3446:Q6962360 3440:Wikidata 3408:Category 3330:Arcuites 3303:Slimonia 3128:Devonian 3123:Silurian 2910:Suborder 2803:Suborder 2780:Taxonomy 2718:Phylum: 2714:Animalia 2430:83873103 2127:56066870 2084:: 79–90. 2007:59488956 1943:19828493 1859:See also 1690:and the 1668:Eifelian 1664:Devonian 1653:Silurian 1467:Slimonia 1105:and the 1094:family. 1040:prosomal 904:mesosoma 900:metasoma 880:paratype 876:holotype 662:. As in 608:subgenus 599:anterior 579:tergites 553:species. 524:podomere 522:in each 401:Scotland 381:Silurian 377:Devonian 357:("dwarf 263:Species 210:Family: 164:Phylum: 158:Animalia 144:Domain: 35:Eifelian 3486:3581779 3473:1054342 3460:4650089 3176:Oceania 3138:Permian 2340:1301566 2242:1301985 2164:(PhD). 2049:1301214 1934:2865068 1747:or the 1662:to the 751:Siberia 697:ventral 640:of the 587:Kington 503:was an 425:in its 397:England 223:Genus: 184:Order: 3166:Europe 3156:Africa 2787:Genera 2510:  2500:  2428:  2338:  2278:  2240:  2210:  2202:  2125:  2047:  2005:  1941:  1931:  1923:  1770:plants 1710:Norway 1676:Europe 730:genus 604:chelae 583:dorsal 520:spines 423:spines 393:Russia 389:Norway 3499:18985 3468:IRMNG 2767:Clade 2758:Clade 2749:Clade 2740:Clade 2731:Clade 2654:(PDF) 2473:(PDF) 2448:(PDF) 2426:S2CID 2336:JSTOR 2295:nanus 2276:S2CID 2238:JSTOR 2123:S2CID 2074:(PDF) 2045:JSTOR 2003:S2CID 1760:and 1660:epoch 892:notos 890:word 888:Greek 833:word 831:Latin 3455:GBIF 3161:Asia 2632:2018 2607:2018 2582:2018 2557:2018 2532:help 2508:ISSN 2498:ISBN 2377:help 2299:nani 2208:OCLC 2200:ISSN 1939:PMID 1921:ISSN 1845:and 1832:and 1800:and 1790:and 1734:and 1082:and 1066:and 1027:and 857:and 836:nana 722:and 680:and 648:and 636:, a 565:as " 399:and 379:and 48:PreꞒ 2462:doi 2458:252 2418:doi 2268:doi 2113:hdl 2105:doi 1995:doi 1929:PMC 1913:doi 925:(a 853:by 745:in 3518:: 3496:: 3483:: 3470:: 3457:: 3442:: 2769:: 2760:: 2751:: 2742:: 2733:: 2662:15 2660:. 2656:. 2623:. 2598:. 2573:. 2548:. 2524:: 2522:}} 2518:{{ 2506:. 2456:. 2450:. 2424:. 2414:94 2412:. 2385:^ 2369:: 2367:}} 2363:{{ 2332:38 2330:. 2310:^ 2302:. 2297:, 2274:. 2264:46 2262:. 2250:^ 2234:40 2232:. 2220:^ 2206:. 2196:16 2194:. 2148:^ 2121:. 2111:. 2101:50 2099:. 2082:76 2080:. 2076:. 2057:^ 2041:35 2039:. 2015:^ 2001:. 1989:. 1985:. 1955:^ 1945:. 1937:. 1927:. 1919:. 1907:. 1901:. 1883:^ 1826:, 1820:, 1812:, 1808:, 1786:, 1782:, 1728:, 1722:, 1708:, 1686:, 1674:, 1050:, 1003:. 943:. 843:. 821:) 819:N. 815:H. 807:H. 799:H. 769:. 749:, 691:, 658:, 628:, 593:. 589:, 461:, 395:, 391:, 387:, 336:†? 314:†? 281:†? 98:Pg 42:Ma 37:, 2694:e 2687:t 2680:v 2647:" 2634:. 2609:. 2584:. 2559:. 2534:) 2514:. 2464:: 2432:. 2420:: 2379:) 2342:. 2322:( 2282:. 2270:: 2244:. 2214:. 2179:. 2143:" 2129:. 2115:: 2107:: 2051:. 2009:. 1997:: 1991:6 1949:. 1915:: 1909:6 817:( 809:( 801:( 789:. 613:( 581:( 453:. 325:† 303:† 292:† 270:† 249:† 227:† 214:† 201:† 188:† 103:N 93:K 88:J 83:T 78:P 73:C 68:D 63:S 58:O 53:Ꞓ 33:-

Index

Llandovery
Eifelian
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Scientific classification
Edit this classification
Eukaryota
Animalia
Arthropoda
Chelicerata
Eurypterida
Adelophthalmoidea
Adelophthalmidae
Nanahughmilleria
Type species
Hughmilleria
eurypterid
arthropods

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