132:
112:
904:
240:
482:
and in the
Hoplocloniini from the tenth tergum. This peculiarity means that the secondary ovipositor must have developed independently of one another twice within the Obriminae. The abdomen of the smaller males is round in cross-section and thinnest in the middle of abdomen. Except in
365:
when naming the subfamilies they established – and as such he also considers the tribes described by both of them – not always taking into account the genera described first. At least in the case of the
Obriminae this is not true, since both the genus
400:, to the rank of a family of their own. He divided the new subfamily Obriminae into three tribes. In addition to the Obrimini, these were the Eubulidini and the Miroceramiini. The two new tribes were synonymed in 2016 by
749:
Revision of the
Oriental subfamiliy Heteropteryginae Kirby, 1896, with a re-arrangement of the family Heteropterygidae and the descriptions of five new species of Haaniella Kirby, 1904. (Phasmatodea: Areolatae:
510:. Their characteristics can be very different, and their arrangement is mostly species-specific and is often used to identify and delimit the species. This method, known as
404:
et al. and 2021 by Sarah Bank et al. with the
Obrimini. The 2016 by Hennemann et al. established tribe Tisamenini was synonymous in 2021. With the establishment of the
361:
renamed the
Obrimini or the Obrimidae in 1923 to Therameninae. In the introduction to his work he justified the renaming by saying that Brunner von Wattenwyl and
993:
462:
is widened and significantly increased by the eggs that are permanently and often produced in large numbers. Their abdomen ends in a pointed secondary
783:
A taxonomic review, including new species and new records of
Philippine Obrimini stick insects (Insecta: Phasmatodea: Heteropterygidae: Obriminae)
880:
A new species of
Trachyaretaon Rehn & Rehn, 1939 from the Babuyan Islands, Philippines (Phasmatodea: Heteropterygidae, Obriminae, Obrimini)
803:
716:
823:
630:
362:
1036:
600:
Reconstructing the nonadaptive radiation of an ancient lineage of ground-dwelling stick insects (Phasmatodea: Heteropterygidae)
597:; de Haan, M.; Dittmar, D.; Dräger, H.; Kahar, R. S.; Kang, A.; Kneubühler, B.; Langton-Myers, S. & Bradler, S. (2021).
131:
392:
into the
Obrimini and Datamini tribes. Günther transferred both tribes to the Heteropteryginae subfamily in 1953. In 2004
396:
elevated this subfamily to the rank of a family and the tribe included to the rank of subfamilies or, in the case of the
908:
451:
998:
612:
507:
384:
and is therefore a synonym for
Obriminae. In 1939 the Obriminae, now referred to as a subfamily, were split up by
667:
385:
459:
671:
389:
1008:
769:
709:
Revision of the genera of the
Areolatae, including the status of Timema and Agathemera (Insecta, Phasmatodea)
470:
formed from the eighth abdominal sternite, which is here named subgenital plate, or also called operculum.
920:
485:
450:
fields present in all Heteropterygidae are to be found in pairs in the Obriminae in the front area of the
442:
744:
526:, where both Hoplocloniini and Obrimini can be found. The latter are also widespread to the east on the
495:
312:
208:
694:
357:) (abbreviated there as Obrimi.). Lawrence Bruner raised the Obrimini to the rank of a family in 1915.
816:
Phasmiden – Pflege und Zucht von Gespenstschrecken, Stabschrecken und Wandelnden Blättern im Terrarium
695:Über die taxonomische Gliederung und die geographische Verbreitung der Insektenordnung der Phasmatodea
590:
1031:
980:
594:
736:
651:
567:
740:
689:
646:
397:
381:
372:
358:
335:
262:
634:
562:
732:
401:
350:
329:
252:
126:
888:
866:
843:
757:
967:
884:
862:
839:
819:
799:
753:
712:
599:
436:
511:
471:
447:
413:
317:
288:
285:
203:
296:
958:
859:
Stabschrecken, Gespenstschrecken, Wandelnde Blätter - Phasmidensystematik im Überblick I.
434:
The Obriminae can reach very different sizes with almost 3 centimetres (1.2 in) in
883:, Entomofauna - Zeitschrift für Entomologie, Band 27, Heft 18, pp. 217–228, Ansfelden,
341:
292:
1025:
854:
405:
393:
224:
62:
879:
972:
617:
377:
239:
952:
527:
323:
282:
193:
183:
37:
943:
903:
539:
499:, the only known representatives of their genera, the Obrimini have no wings.
463:
82:
47:
676:
502:
The body is often covered with numerous spines, more or less blunt thorns or
543:
503:
490:
419:
278:
163:
143:
87:
31:
568:
Evolution of Oviposition Techniques in Stick and Leaf Insects (Phasmatodea)
426:
and the other tribus is Obrimini with now 14 genera and 70 valid species.
937:
535:
531:
479:
308:
229:
102:
77:
72:
57:
52:
42:
467:
92:
67:
408:, the anatomical peculiarity of the secondary ovipositor of the genus
523:
475:
173:
153:
98:
914:
782:
455:
412:
was taken into account. This special status was also confirmed by
985:
752:, Zootaxa 4159 (1), Magnolia Press, Auckland, New Zealand 2016,
918:
677:
Proceedings of The Academy of Natural Sciences (Vol. 90, 1938)
635:
Die Insektenfamilie der Phasmiden. Vol. 1. Phasmidae Areolatae
506:, which can be found mainly on the upper side of the head and
836:
Zwergformen der Phasmatodea - die Kleinsten unter den Riesen
637:. Verlag Wilhelm Engelmann, Leipzig, pp. 36–57 & 162–172
514:, was developed in 1939 by Rehn and Rehn for the Obriminae.
110:
728:
726:
724:
418:
Thus the Obriminae consist of two valid tribes. One is the
618:
Annali del Museo Civico di Storia Naturale Giacomo Doria
380:. The name Therameninae was withdrawn again in 1929 by
698:, Beiträge zur Entomologie, Band 3, Nr. 5, pp. 541–563
474:
it consists in the Obrimini by the eleventh abdominal
440:
and up to 13 centimetres (5.1 in) in length in
927:
711:, Goecke & Evers, Keltern-Weiler, pp. 191–240,
585:
583:
581:
579:
577:
768:Brock, P. D.; Büscher, T. H. & Baker, E. W.:
603:, Systematic Entomology, DOI: 10.1111/syen.12472
522:The distribution area of the Obriminae includes
814:Seiler, C.; Bradler, S. & Koch, R. (2000).
796:Stabschrecken - Carausius, Sipyloidea & Co.
663:
661:
8:
877:Hennemann, F. H. & Conle, O. V. (2006).
466:that surrounds the actual ovipositor. It is
772:. Version 5.0./5.0 (accessdate 29 May 2021)
915:
861:, Arthropoda 17 (1), Sungaya-Verlag Kiel.
838:, Arthropoda 16 (3), Sungaya-Verlag Kiel.
589:Bank, S.; Buckley, T. R.; Büscher, T. H.;
422:tribus Hoplocloniini, with the only genus
238:
20:
785:, Faunitaxys, 2023, 11 (71), pp. 1–135.
554:
121:Different species of Heteropterygidae
7:
571:, Frontiers in Ecology and Evolution
798:, Natur und Tier Verlag, Münster,
14:
655:, Treubia Vol. III 2, pp. 230–233
902:
130:
621:. Genova (2) 13 (33):101, p. 98
652:Zur Nomenklatur der Phasmoiden
478:called the supraanal plate or
1:
1006:Phasmida Species File (old):
565:& Whiting, M. F. (2018).
770:Phasmida Species File Online
680:, Philadelphia, pp. 389 ff.
1053:
277:are the most species-rich
613:Brunner von Wattenwyl, C.
386:James Abram Garfield Rehn
295:. It is divided into two
258:
251:
246:
237:
221:
216:
127:Scientific classification
125:
120:
109:
23:
834:Zompro, O. (Nov. 2008).
818:. bede, Ruhmannsfelden,
781:Hennemann, F. H. (2023)
390:John William Holman Rehn
1037:Phasmatodea subfamilies
315:in 1893 for the genera
247:Range of the Obriminae
16:Family of stick insects
794:Fritzsche, I. (2007).
486:Miroceramia westwoodii
443:Trachyaretaon carmelae
115:
496:Pterobrimus depressus
376:was built in 1875 by
313:Brunner von Wattenwyl
209:Brunner von Wattenwyl
114:
707:Zompro, O. (2004).
359:Heinrich Hugo Karny
561:Robertson, J. A.;
402:Frank H. Hennemann
363:Josef Redtenbacher
116:
1019:
1018:
921:Taxon identifiers
804:978-3-937285-84-9
750:Heteropterygidae)
518:Distribution area
437:Tisamenus hebardi
271:
270:
265:
212:
1044:
1012:
1011:
1002:
1001:
989:
988:
976:
975:
963:
962:
961:
948:
947:
946:
916:
907:Data related to
906:
891:
875:
869:
852:
846:
832:
826:
812:
806:
792:
786:
779:
773:
766:
760:
733:Hennemann, F. H.
730:
719:
705:
699:
687:
681:
665:
656:
644:
638:
631:Redtenbacher, J.
628:
622:
610:
604:
587:
572:
559:
414:genetic analysis
289:Heteropterygidae
261:
242:
207:
204:Heteropterygidae
135:
134:
113:
97:
34:
27:Temporal range:
21:
1052:
1051:
1047:
1046:
1045:
1043:
1042:
1041:
1022:
1021:
1020:
1015:
1007:
1005:
997:
992:
984:
979:
971:
966:
957:
956:
951:
942:
941:
936:
923:
899:
894:
876:
872:
853:
849:
833:
829:
813:
809:
793:
789:
780:
776:
767:
763:
731:
722:
706:
702:
688:
684:
666:
659:
645:
641:
629:
625:
611:
607:
588:
575:
560:
556:
552:
520:
432:
417:
416:investigations.
311:was created by
305:
206:
129:
111:
105:
96:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
40:
29:
28:
25:
17:
12:
11:
5:
1050:
1048:
1040:
1039:
1034:
1024:
1023:
1017:
1016:
1014:
1013:
1003:
990:
977:
964:
949:
933:
931:
925:
924:
919:
913:
912:
911:at Wikispecies
898:
897:External links
895:
893:
892:
870:
857:(April 2009).
847:
827:
807:
787:
774:
761:
745:Seow-Choen, F.
720:
717:978-3931374396
700:
682:
672:Rehn, J. W. H.
668:Rehn, J. A. G.
657:
639:
623:
605:
573:
553:
551:
548:
534:, most of the
519:
516:
431:
428:
304:
301:
293:Southeast Asia
269:
268:
256:
255:
249:
248:
244:
243:
235:
234:
233:
232:
227:
219:
218:
214:
213:
201:
197:
196:
191:
187:
186:
181:
177:
176:
171:
167:
166:
161:
157:
156:
151:
147:
146:
141:
137:
136:
123:
122:
118:
117:
107:
106:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
36:
35:
26:
15:
13:
10:
9:
6:
4:
3:
2:
1049:
1038:
1035:
1033:
1030:
1029:
1027:
1010:
1004:
1000:
995:
991:
987:
982:
978:
974:
969:
965:
960:
954:
950:
945:
939:
935:
934:
932:
930:
926:
922:
917:
910:
905:
901:
900:
896:
890:
886:
882:
881:
874:
871:
868:
864:
860:
856:
851:
848:
845:
841:
837:
831:
828:
825:
824:3-933646-89-8
821:
817:
811:
808:
805:
801:
797:
791:
788:
784:
778:
775:
771:
765:
762:
759:
755:
751:
746:
742:
738:
734:
729:
727:
725:
721:
718:
714:
710:
704:
701:
697:
696:
691:
686:
683:
679:
678:
673:
669:
664:
662:
658:
654:
653:
648:
643:
640:
636:
632:
627:
624:
620:
619:
614:
609:
606:
602:
601:
596:
592:
586:
584:
582:
580:
578:
574:
570:
569:
564:
558:
555:
549:
547:
545:
541:
537:
533:
529:
525:
517:
515:
513:
509:
505:
500:
498:
497:
492:
488:
487:
481:
477:
473:
469:
465:
461:
457:
453:
449:
445:
444:
439:
438:
429:
427:
425:
421:
415:
411:
407:
406:Hoplocloniini
403:
399:
398:Anisacanthini
395:
394:Oliver Zompro
391:
387:
383:
382:Klaus Günther
379:
375:
374:
369:
364:
360:
356:
352:
348:
344:
343:
338:
337:
332:
331:
326:
325:
320:
319:
314:
310:
302:
300:
298:
294:
290:
287:
284:
280:
276:
267:
264:
260:Therameninae
257:
254:
250:
245:
241:
236:
231:
228:
226:
225:Hoplocloniini
223:
222:
220:
215:
210:
205:
202:
199:
198:
195:
192:
190:Superfamily:
189:
188:
185:
182:
179:
178:
175:
172:
169:
168:
165:
162:
159:
158:
155:
152:
149:
148:
145:
142:
139:
138:
133:
128:
124:
119:
108:
104:
100:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
39:
33:
22:
19:
928:
878:
873:
858:
850:
835:
830:
815:
810:
795:
790:
777:
764:
748:
741:Brock, P. D.
737:Conle, O. V.
708:
703:
693:
685:
675:
650:
647:Karny, H. H.
642:
626:
616:
608:
598:
595:Constant, J.
591:Bresseel, J.
566:
557:
521:
501:
494:
484:
458:females the
441:
435:
433:
423:
409:
388:and his son
371:
367:
354:
346:
340:
334:
328:
322:
316:
306:
274:
272:
259:
99:Early Eocene
18:
1032:Phasmatodea
953:Wikispecies
690:Günther, K.
563:Bradler, S.
528:Philippines
512:acanthotaxy
430:Description
424:Hoploclonia
420:monotypical
410:Hoploclonia
324:Hoploclonia
283:Phasmatodea
194:Bacilloidea
184:Phasmatodea
1026:Categories
855:Zompro, O.
550:References
540:New Guinea
464:ovipositor
452:prosternum
373:Theramenes
355:Pylaemenes
336:Pylaemenes
307:The tribe
291:native to
164:Arthropoda
30:50–0
959:Obriminae
929:Obriminae
909:Obriminae
889:0250-4413
867:0943-7274
844:0943-7274
758:1175-5326
544:Viti Levu
538:islands,
504:tubercles
491:rudiments
378:Carl Stål
330:Tisamenus
279:subfamily
275:Obriminae
150:Kingdom:
144:Eukaryota
24:Obriminae
986:10523555
944:Q1223047
938:Wikidata
747:(2016).
692:(1953).
674:(1939).
649:(1923).
633:(1906).
536:Moluccas
532:Sulawesi
480:Epiproct
472:Dorsally
309:Obrimini
303:Taxonomy
253:Synonyms
230:Obrimini
200:Family:
160:Phylum:
154:Animalia
140:Domain:
1009:1200282
615:(1893)
489:and in
468:ventral
460:abdomen
448:sensory
368:Obrimus
351:synonym
349:(today
347:Datames
318:Obrimus
281:of the
266:, 1923
217:Tribes
180:Order:
174:Insecta
170:Class:
103:present
999:525305
887:
865:
842:
822:
802:
756:
743:&
715:
670:&
524:Borneo
508:thorax
476:tergum
446:. The
286:family
211:, 1893
973:8Q6Z2
456:adult
454:. In
342:Dares
297:tribe
263:Karny
994:NCBI
885:ISSN
863:ISSN
840:ISSN
820:ISBN
800:ISBN
754:ISSN
713:ISBN
542:and
370:and
345:and
273:The
38:PreꞒ
981:EoL
968:CoL
493:in
353:to
101:to
1028::
996::
983::
970::
955::
940::
739:;
735:;
723:^
660:^
593:;
576:^
546:.
530:,
339:,
333:,
327:,
321:,
299:.
88:Pg
32:Ma
93:N
83:K
78:J
73:T
68:P
63:C
58:D
53:S
48:O
43:Ꞓ
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
