1880:
111:
368:
940:
132:
574:"A nearly complete skeleton of a new eusphenodontian from the Upper Jurassic Morrison Formation, Wyoming, USA, provides insight into the evolution and diversity of Rhynchocephalia (Reptilia: Lepidosauria)"
647:
Vertebrate paleobiodiversity of the Early
Cretaceous (Berriasian) Angeac-Charente Lagerstätte (southwestern France): implications for continental faunal turnover at the J/K boundary
326:. A undescribed crushed skull (DINO 16454) from the Morrison Formaiton has been attributed to this genus by some sources, though this has been strongly disputed by others.
506:"A re-evaluation of the taxonomy and diversity of the Late Jurassic sphenodontian Opisthias rarus Gilmore 1909 (Lepidosauria: Rhynchocephalia)[oral presentation]"
1924:
1826:
726:
1813:
386:
337:
of the United
Kingdom, which appears to be distinct from the type North American species. A fragmentary dentary possibly attributable to
1917:
1958:
523:
Malafaia, E.; Ortega, F.; Escaso, F.; Dantas, P.; Pimentel, N.; Gasulla, J. M.; Ribeiro, B.; Barriga, F.; Sanz, J. L. (2010).
1910:
1948:
719:
939:
356:
has been interpreted as a generalist, with its tooth morphology indicating had an ability to process plant material.
829:
131:
1953:
712:
1943:
1620:
342:
199:
1781:
1852:
1460:
1412:
319:
536:
524:
1879:
802:
381:
373:
1446:
1089:
334:
299:
126:
110:
1857:
1351:
409:
314:), present in stratigraphic zones 2 and 4–6. Other remains suggested to be closely related to
1839:
1831:
1255:
1133:
916:
814:
690:
605:
554:
1894:
1844:
525:"Vertebrate fauna at the Allosaurus fossil-site of Andrés (Upper Jurassic), Pombal, Portugal"
1706:
1633:
1072:
1046:
971:
680:
672:
595:
585:
544:
469:
1735:
1595:
1533:
1492:
1217:
899:
871:
841:
735:
183:
646:
540:
1690:
1670:
1480:
1436:
1392:
1186:
1143:
1112:
1099:
1056:
1033:
981:
685:
660:
649:. Geodiversitas, Museum National d’Histoire Naturelle Paris, In press. ffhal-03264773f
1937:
1720:
1713:
1640:
1543:
1522:
1512:
1502:
1276:
1266:
1200:
1177:
1163:
1153:
1011:
909:
624:
505:
295:
291:
276:
264:
61:
1573:
1563:
1453:
1358:
1287:
1245:
1021:
991:
950:
771:
645:
Ronan Allain, Romain Vullo, Lee Rozada, Jérémy
Anquetin, Renaud Bourgeais, et al..
549:
268:
225:
590:
573:
661:"The youngest South American rhynchocephalian, a survivor of the K/Pg extinction"
1727:
1584:
1328:
1298:
1193:
923:
36:
1804:
1678:
1663:
1655:
1468:
1422:
1365:
1342:
1318:
1001:
787:
363:
330:
256:
99:
81:
46:
609:
558:
430:
Foster J (2007). "Table 2.1: Fossil
Vertebrates of the Morrison Formation.".
1699:
1605:
1552:
1308:
450:"Redescription of the type specimens for the Late Jurassic rhynchocephalian
272:
143:
86:
694:
676:
474:
449:
1887:
1798:
1234:
759:
323:
303:
163:
76:
71:
56:
51:
41:
1890:
1818:
1402:
765:
600:
311:
173:
91:
66:
572:
DeMar, David G.; Jones, Marc E. H.; Carrano, Matthew T. (2022-12-31).
491:
Jurassic West: The
Dinosaurs of the Morrison Formation and Their World
432:
Jurassic West: The
Dinosaurs of the Morrison Formation and Their World
753:
153:
1775:
629:
Proceedings of the Dorset
Natural History and Archaeological Society
704:
252:
307:
1779:
746:
708:
625:"New insights on an enigmatic sphenodontian jaw from Dorset"
404:
402:
510:
13th
Symposium on Mesozoic Terrestrial Ecosystems and Biota
448:
Herrera-Flores, J.A.; Stubbs, T.L.; Sour-Tovar, F. (2022).
1898:
1617:
1601:
1591:
1580:
1569:
1559:
1539:
1529:
1518:
1508:
1498:
1488:
1433:
1418:
1408:
1398:
1388:
1339:
1324:
1314:
1304:
1294:
1283:
1272:
1262:
1251:
1241:
1230:
1174:
1159:
1149:
1139:
1129:
1095:
1085:
1052:
1042:
1017:
1007:
997:
987:
977:
967:
896:
1788:
1689:
1616:
1479:
1432:
1375:
1338:
1215:
1173:
1110:
1070:
1031:
948:
895:
840:
813:
786:
659:Apesteguía S, Gómez RO, Rougier GW (October 2014).
458:from Quarry 9, Morrison Formation, Wyoming, USA"
1918:
720:
493:. Indiana University Press. pp. 327–329.
290:is primarily known from remains found in the
8:
425:
423:
434:. Indiana University Press. pp. 58–59.
410:"Opisthias Gilmore 1909 (rhynchocephalian)"
1925:
1911:
1776:
1625:
1381:
1223:
1122:
1118:
1078:
960:
956:
892:
888:
819:
792:
783:
743:
727:
713:
705:
623:Herrera-Flores J (2020). Randall C (ed.).
504:Herrera-Flores JA, Stubbs TL (July 2018).
109:
20:
684:
599:
589:
548:
473:
398:
341:is also known from the Berriasian aged
318:are also known from the Late Jurassic
7:
1876:
1874:
443:
441:
387:Paleobiota of the Morrison Formation
578:Journal of Systematic Palaeontology
329:A lower jaw is also known from the
1897:. You can help Knowledge (XXG) by
14:
1878:
938:
665:Proceedings. Biological Sciences
366:
130:
550:10.5209/rev_JIGE.2010.v36.n2.7
1:
591:10.1080/14772019.2022.2093139
489:Foster J (2007). "Appendix".
462:Acta Palaeontologica Polonica
298:, with remains found in the
259:reptile. The type species,
1975:
1886:This article related to a
1873:
529:Journal of Iberian Geology
1628:
1384:
1226:
1125:
1121:
1081:
963:
959:
935:
891:
887:
868:
822:
795:
782:
742:
231:
224:
127:Scientific classification
125:
117:
108:
23:
16:Extinct genus of reptiles
343:Angeac-Charente bonebed
1959:Jurassic reptile stubs
677:10.1098/rspb.2014.0811
475:10.4202/app.00929.2021
454:and a new specimen of
32:Kimmeridgian–Tithonian
1853:Paleobiology Database
414:Paleobiology Database
1949:Jurassic lepidosaurs
456:Theretairus antiquus
263:, is known from the
541:2010JIbG...36..193M
382:Prehistoric reptile
374:Paleontology portal
1447:Palaeopleurosaurus
1090:Planocephalosaurus
671:(1792): 20140811.
335:Lulworth Formation
320:Alcobaça Formation
300:Morrison Formation
1906:
1905:
1868:
1867:
1840:Open Tree of Life
1782:Taxon identifiers
1773:
1772:
1769:
1768:
1765:
1764:
1761:
1760:
1757:
1756:
1753:
1752:
1749:
1748:
1745:
1744:
1256:Lanceirosphenodon
1211:
1210:
1134:Lanceirosphenodon
1066:
1065:
933:
932:
917:Penegephyrosaurus
883:
882:
879:
878:
864:
863:
860:
859:
830:Lepidosauromorpha
815:Lepidosauromorpha
244:
243:
220:
103:
1966:
1927:
1920:
1913:
1882:
1875:
1861:
1860:
1848:
1847:
1835:
1834:
1822:
1821:
1809:
1808:
1807:
1777:
1707:Kaikaifilusaurus
1634:Fraserosphenodon
1626:
1619:
1603:
1593:
1582:
1571:
1561:
1541:
1531:
1520:
1510:
1500:
1490:
1435:
1420:
1410:
1400:
1390:
1382:
1341:
1326:
1316:
1306:
1296:
1285:
1274:
1264:
1253:
1243:
1232:
1224:
1176:
1161:
1151:
1141:
1131:
1123:
1119:
1097:
1087:
1079:
1073:Acrosphenodontia
1054:
1047:Paleollanosaurus
1044:
1019:
1009:
999:
989:
979:
972:Diphydontosaurus
969:
961:
957:
943:
942:
898:
893:
889:
820:
793:
784:
777:
776:
744:
729:
722:
715:
706:
699:
698:
688:
656:
650:
643:
637:
636:
620:
614:
613:
603:
593:
569:
563:
562:
552:
520:
514:
513:
501:
495:
494:
486:
480:
479:
477:
445:
436:
435:
427:
418:
417:
406:
376:
371:
370:
369:
218:
211:
198:
135:
134:
113:
97:
96:
33:
29:Temporal range:
21:
1974:
1973:
1969:
1968:
1967:
1965:
1964:
1963:
1954:Rhynchocephalia
1934:
1933:
1932:
1931:
1871:
1869:
1864:
1856:
1851:
1843:
1838:
1830:
1825:
1817:
1812:
1803:
1802:
1797:
1784:
1774:
1741:
1736:Toxolophosaurus
1685:
1612:
1534:Navajosphenodon
1493:Ankylosphenodon
1475:
1428:
1377:
1371:
1334:
1220:
1218:Neosphenodontia
1207:
1169:
1115:
1106:
1075:
1062:
1027:
953:
944:
937:
929:
900:Gephyrosauridae
875:
872:Rhynchocephalia
856:
842:Rhynchocephalia
836:
809:
778:
749:
748:
738:
736:Rhynchocephalia
733:
703:
702:
658:
657:
653:
644:
640:
622:
621:
617:
571:
570:
566:
522:
521:
517:
503:
502:
498:
488:
487:
483:
452:Opisthias rarus
447:
446:
439:
429:
428:
421:
408:
407:
400:
395:
372:
367:
365:
362:
351:
285:
261:Opisthias rarus
240:
237:
234:Opisthias rarus
217:
209:
196:
184:Rhynchocephalia
129:
120:Opisthias rarus
104:
95:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
39:
31:
30:
27:
17:
12:
11:
5:
1972:
1970:
1962:
1961:
1956:
1951:
1946:
1944:Morrison fauna
1936:
1935:
1930:
1929:
1922:
1915:
1907:
1904:
1903:
1883:
1866:
1865:
1863:
1862:
1849:
1836:
1823:
1810:
1794:
1792:
1786:
1785:
1780:
1771:
1770:
1767:
1766:
1763:
1762:
1759:
1758:
1755:
1754:
1751:
1750:
1747:
1746:
1743:
1742:
1740:
1739:
1732:
1724:
1717:
1710:
1703:
1695:
1693:
1691:Eilenodontinae
1687:
1686:
1684:
1683:
1675:
1671:Pleurosauridae
1667:
1660:
1652:
1645:
1637:
1629:
1623:
1614:
1613:
1611:
1610:
1599:
1589:
1578:
1567:
1557:
1548:
1537:
1527:
1516:
1506:
1496:
1485:
1483:
1481:Sphenodontidae
1477:
1476:
1474:
1473:
1465:
1457:
1450:
1442:
1440:
1437:Pleurosauridae
1430:
1429:
1427:
1426:
1416:
1406:
1396:
1393:Lamarquesaurus
1385:
1379:
1373:
1372:
1370:
1369:
1362:
1355:
1347:
1345:
1343:Sapheosauridae
1336:
1335:
1333:
1332:
1322:
1312:
1302:
1292:
1281:
1270:
1260:
1249:
1239:
1227:
1221:
1216:
1213:
1212:
1209:
1208:
1206:
1205:
1197:
1190:
1187:Brachyrhinodon
1182:
1180:
1171:
1170:
1168:
1167:
1157:
1147:
1144:Microsphenodon
1137:
1126:
1116:
1113:Eusphenodontia
1111:
1108:
1107:
1105:
1104:
1100:Godavarisaurus
1093:
1082:
1076:
1071:
1068:
1067:
1064:
1063:
1061:
1060:
1057:Whitakersaurus
1050:
1039:
1037:
1034:Incertae sedis
1029:
1028:
1026:
1025:
1015:
1005:
995:
985:
982:Godavarisaurus
975:
964:
954:
949:
946:
945:
936:
934:
931:
930:
928:
927:
920:
913:
905:
903:
885:
884:
881:
880:
877:
876:
869:
866:
865:
862:
861:
858:
857:
855:
854:
853:
852:
846:
844:
838:
837:
835:
834:
833:
832:
823:
817:
811:
810:
808:
807:
806:
805:
796:
790:
780:
779:
775:
774:
768:
762:
756:
747:
740:
739:
734:
732:
731:
724:
717:
709:
701:
700:
651:
638:
615:
584:(1): 2093139.
564:
535:(2): 193–204.
515:
496:
481:
468:(3): 623–630.
437:
419:
397:
396:
394:
391:
390:
389:
384:
378:
377:
361:
358:
350:
347:
284:
281:
275:) of western
242:
241:
238:
229:
228:
222:
221:
207:
203:
202:
194:
187:
186:
181:
177:
176:
171:
167:
166:
161:
157:
156:
151:
147:
146:
141:
137:
136:
123:
122:
115:
114:
106:
105:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
40:
35:
34:
28:
15:
13:
10:
9:
6:
4:
3:
2:
1971:
1960:
1957:
1955:
1952:
1950:
1947:
1945:
1942:
1941:
1939:
1928:
1923:
1921:
1916:
1914:
1909:
1908:
1902:
1900:
1896:
1892:
1889:
1884:
1881:
1877:
1872:
1859:
1854:
1850:
1846:
1841:
1837:
1833:
1828:
1824:
1820:
1815:
1811:
1806:
1800:
1796:
1795:
1793:
1791:
1787:
1783:
1778:
1738:
1737:
1733:
1730:
1729:
1725:
1723:
1722:
1721:Priosphenodon
1718:
1716:
1715:
1714:Patagosphenos
1711:
1709:
1708:
1704:
1702:
1701:
1697:
1696:
1694:
1692:
1688:
1681:
1680:
1676:
1673:
1672:
1668:
1666:
1665:
1661:
1658:
1657:
1653:
1651:
1650:
1646:
1643:
1642:
1641:Kawasphenodon
1638:
1636:
1635:
1631:
1630:
1627:
1624:
1622:
1621:Opisthodontia
1615:
1608:
1607:
1600:
1598:
1597:
1590:
1587:
1586:
1579:
1576:
1575:
1568:
1566:
1565:
1558:
1556:
1555:
1554:
1549:
1546:
1545:
1544:Pamizinsaurus
1538:
1536:
1535:
1528:
1525:
1524:
1523:Kawasphenodon
1517:
1515:
1514:
1513:Derasmosaurus
1507:
1505:
1504:
1503:Cynosphenodon
1497:
1495:
1494:
1487:
1486:
1484:
1482:
1478:
1471:
1470:
1466:
1463:
1462:
1458:
1456:
1455:
1451:
1449:
1448:
1444:
1443:
1441:
1438:
1431:
1425:
1424:
1417:
1415:
1414:
1407:
1405:
1404:
1397:
1395:
1394:
1387:
1386:
1383:
1380:
1376:"Eupropalinal
1374:
1368:
1367:
1363:
1361:
1360:
1356:
1354:
1353:
1349:
1348:
1346:
1344:
1337:
1331:
1330:
1323:
1321:
1320:
1313:
1311:
1310:
1303:
1301:
1300:
1293:
1290:
1289:
1282:
1279:
1278:
1277:Rebbanasaurus
1271:
1269:
1268:
1267:Pamizinsaurus
1261:
1258:
1257:
1250:
1248:
1247:
1240:
1237:
1236:
1229:
1228:
1225:
1222:
1219:
1214:
1203:
1202:
1201:Polysphenodon
1198:
1196:
1195:
1191:
1189:
1188:
1184:
1183:
1181:
1179:
1178:Clevosauridae
1172:
1166:
1165:
1164:Polysphenodon
1158:
1156:
1155:
1154:Opisthiamimus
1148:
1146:
1145:
1138:
1136:
1135:
1128:
1127:
1124:
1120:
1117:
1114:
1109:
1102:
1101:
1094:
1092:
1091:
1084:
1083:
1080:
1077:
1074:
1069:
1059:
1058:
1051:
1049:
1048:
1041:
1040:
1038:
1036:
1035:
1030:
1024:
1023:
1016:
1014:
1013:
1012:Rebbanasaurus
1006:
1004:
1003:
996:
994:
993:
986:
984:
983:
976:
974:
973:
966:
965:
962:
958:
955:
952:
947:
941:
926:
925:
921:
919:
918:
914:
912:
911:
910:Gephyrosaurus
907:
906:
904:
901:
894:
890:
886:
874:
873:
867:
850:
849:
848:
847:
845:
843:
839:
831:
827:
826:
825:
824:
821:
818:
816:
812:
804:
800:
799:
798:
797:
794:
791:
789:
785:
781:
773:
769:
767:
763:
761:
757:
755:
751:
750:
745:
741:
737:
730:
725:
723:
718:
716:
711:
710:
707:
696:
692:
687:
682:
678:
674:
670:
666:
662:
655:
652:
648:
642:
639:
634:
630:
626:
619:
616:
611:
607:
602:
597:
592:
587:
583:
579:
575:
568:
565:
560:
556:
551:
546:
542:
538:
534:
530:
526:
519:
516:
511:
507:
500:
497:
492:
485:
482:
476:
471:
467:
463:
459:
457:
453:
444:
442:
438:
433:
426:
424:
420:
415:
411:
405:
403:
399:
392:
388:
385:
383:
380:
379:
375:
364:
359:
357:
355:
348:
346:
344:
340:
336:
332:
327:
325:
321:
317:
313:
309:
305:
301:
297:
296:United States
293:
292:Late Jurassic
289:
282:
280:
278:
277:North America
274:
270:
266:
265:Late Jurassic
262:
258:
254:
250:
249:
239:Gilmore, 1905
236:
235:
230:
227:
223:
216:
215:
208:
205:
204:
201:
200:Opisthodontia
195:
192:
189:
188:
185:
182:
179:
178:
175:
172:
169:
168:
165:
162:
159:
158:
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107:
101:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
26:
22:
19:
1899:expanding it
1885:
1870:
1789:
1734:
1726:
1719:
1712:
1705:
1698:
1677:
1669:
1662:
1654:
1648:
1647:
1639:
1632:
1604:
1594:
1583:
1574:Sphenovipera
1572:
1564:Sphenofontis
1562:
1551:
1550:
1542:
1532:
1521:
1511:
1501:
1491:
1467:
1459:
1454:Pleurosaurus
1452:
1445:
1421:
1411:
1401:
1391:
1378:sphenodonts"
1364:
1359:Sapheosaurus
1357:
1350:
1327:
1317:
1307:
1297:
1288:Sphenocondor
1286:
1275:
1265:
1254:
1246:Homoeosaurus
1244:
1233:
1199:
1192:
1185:
1162:
1152:
1142:
1132:
1098:
1088:
1055:
1045:
1032:
1022:Sphenocondor
1020:
1010:
1000:
992:Micromenodon
990:
980:
970:
951:Sphenodontia
922:
915:
908:
870:
772:Lepidosauria
770:Superorder:
668:
664:
654:
641:
632:
628:
618:
581:
577:
567:
532:
528:
518:
509:
499:
490:
484:
465:
461:
455:
451:
431:
413:
353:
352:
338:
328:
315:
287:
286:
283:Distribution
269:Kimmeridgian
260:
247:
246:
245:
233:
232:
226:Type species
219:Gilmore 1909
213:
212:
190:
119:
24:
18:
1728:Sphenotitan
1679:Sapheosaurs
1585:Theretairus
1329:Leptosaurus
1299:Theretairus
1194:Clevosaurus
924:Deltadectes
601:2440/136608
345:in France.
1938:Categories
1664:Trullidens
1656:Pelecymala
1469:Vadasaurus
1423:Vadasaurus
1366:Oenosaurus
1319:Kallimodon
1002:Pelecymala
851:see below↓
803:Sauropsida
788:Sauropsida
393:References
331:Berriasian
257:sphenodont
100:Berriasian
1790:Opisthias
1700:Eilenodon
1649:Opisthias
1606:Zapatadon
1553:Sphenodon
1461:Tingitana
1413:Tingitana
1352:Piocormus
1309:Zapatadon
752:Kingdom:
610:1477-2019
559:1698-6180
354:Opisthias
339:Opisthias
316:Opisthias
288:Opisthias
273:Tithonian
248:Opisthias
214:Opisthias
150:Kingdom:
144:Eukaryota
98:Possible
25:Opisthias
1888:Jurassic
1805:Q7098296
1799:Wikidata
1235:Colobops
766:Reptilia
760:Chordata
758:Phylum:
754:Animalia
695:25143041
635:: 41–43.
360:See also
324:Portugal
304:Colorado
174:Reptilia
164:Chordata
160:Phylum:
154:Animalia
140:Domain:
1891:reptile
1845:4119531
1832:1111940
1819:4821619
1403:Sigmala
764:Class:
686:4150314
537:Bibcode
312:Wyoming
294:of the
206:Genus:
180:Order:
170:Class:
118:Jaw of
693:
683:
608:
557:
310:, and
102:record
1893:is a
1858:37820
1827:IRMNG
333:aged
253:genus
251:is a
191:Clade
1895:stub
1814:GBIF
1596:Tika
828:see
801:see
691:PMID
606:ISSN
555:ISSN
349:Diet
308:Utah
37:PreꞒ
681:PMC
673:doi
669:281
633:141
596:hdl
586:doi
545:doi
470:doi
322:of
279:.
255:of
1940::
1855::
1842::
1829::
1816::
1801::
689:.
679:.
667:.
663:.
631:.
627:.
604:.
594:.
582:20
580:.
576:.
553:.
543:.
533:36
531:.
527:.
508:.
466:67
464:.
460:.
440:^
422:^
412:.
401:^
306:,
193::
87:Pg
1926:e
1919:t
1912:v
1901:.
1731:?
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988:†
978:†
968:†
902:?
897:†
728:e
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714:v
697:.
675::
612:.
598::
588::
561:.
547::
539::
512:.
478:.
472::
416:.
302:(
271:-
267:(
210:†
197:†
92:N
82:K
77:J
72:T
67:P
62:C
57:D
52:S
47:O
42:Ꞓ
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