457:
226:
173:
20:
484:(which are only inherited from father to son) have provided new information, yet it can often be difficult to interpret. The two types of mitochondrial DNA which are dominant among the Sámi, haplogroups U5b1b and V, are believed to originate in Western Europe. Even those Y chromosome variants which are found among the Sámi indicate a European origin. However, no genetic similarity with present-day Siberian peoples has been proved. The research group who published these results in 2004 believed that the distinctive genetic character of the Sámi is best explained by their ancestors having been a small, defined group of Europeans.
348:
537:
inhabitants. Researchers no longer believe, however, that the people who left traces at Komsa lived out the Ice Age on the
Northern Norwegian coast, rather that the coastal area was quickly colonised from the south during the final stages of the Ice Age. When these people followed the melting ice southwards across the Tundra, they eventually encountered the people who had colonised Finland from the east. Thus, the genetic heritage of the Sámi, which is primarily European but is thought to have come from both east and west, can be explained.
83:-speaking farmers could mix with each other during the iron age. This has been interpreted to show that the Sámi culture has been so formed as a result of semi-nomadic reindeer husbandry which began around 2500 years ago, rather than as a strictly isolated group. Nevertheless, the Sámi are a genetically unique population. By this it is meant not that the Sámi have unique genes, rather that certain gene variants are present at a different frequency in Sámi people than in other populations.
1014:
Marina
Bermisheva, Elza Khusnutdinova, Vladislava Gusar, Elena Grechanina, Jüri Parik, Erwan Pennarun, Christelle Richard, Andre Chaventre, Jean-Paul Moisan, Lovorka Barać, Marijana Peričić, Pavao Rudan, Rifat Terzić, Ilia Mikerezi, Astrida Krumina, Viesturs Baumanis, Slawomir Koziel, Olga Rickards, Gian Franco De Stefano, Nicholas Anagnou, Kalliopi I. Pappa, Emmanuel Michalodimitrakis, Vladimir Ferák, Sandor Füredi, Radovan Komel, Lars Beckman, Richard Villems (25 February 2004).
217:
or the
Norwegians. Schefferus concluded that it was most likely that the Sámi derived from the Finns, not least due to the apparent similarity between the Sámi and Finnish languages, but also as both peoples were so alike in temperament as well as appearance: "The Finns have black hair, wide faces and grim expressions, as do the Lapps". As well as question of clothing, Schefferus found that the difference between the Sámi and the Finns was insignificant.
249:, which at the time was a large area of research). According to Wiklund, it was characteristic for the Sámi to have a short head and to be of short stature, to have black hair, brown eyes and a weak chin. In his opinion, nobody had been able to prove an anthropological relationship between the Sámi and any other people. He concluded that "the Lappish race" arose through a long time of total isolation from other groups of people.
106:, which is particularly common among Sámi-speaking populations. These days it is uncommon among other European populations, where hunter-gatherers are thought to have been displaced by later farming communities to the areas on Europe's periphery. Variant U5b1b1 appears in the Nordic countries largely only among those with Sámi roots, but also occurs in North Africa (among
400:. North of this boundary the names for rivers and the oldest place names are often of Sámi or Finnish origin, while the equivalent names south of the boundary are Germanic in their origin. The people who made asbestos-ceramics would, according to this, be the forefathers of the Sámi, while those who lived further south would be the forefathers to the Scandinavians.
102:(passed on via one's maternal grandmother's mother etc.), which is particularly common among the Sámi. Another culture had overwintered in refugia in present-day Russia, and populated Scandinavia and Europe from the northeast after the end of the Ice Age, around 10000 years ago. This migration brought with it the mtDNA
528:, or from both directions. Another type of mitochondrial DNA, haplogroup Z, occurs at low rates both among Sámi and North Asian ethnicities, yet is otherwise absent from Europe. Researchers have interpreted this as a sign that the North Sámi group mixed with another from the east as recently as 2700 years ago.
540:
The time of the later migration, which geneticists believe they can see traces of from 2700 years ago, can be compared with that connection which was made before by many archaeologists between the Sámi and the people who made asbestos ceramics. Asbestos ceramics are found in dwellings from circa 3900
216:
devoted a chapter to the lineage of the Sámi. He opened by arguing at the Sámi couldn't originate from Swedes, "... When nothing is more unlike than a Lapp and a Swede. Not in body shape, temperament, clothing, why nothing between them is alike." Nor were they believed to have come from the
Russians
487:
It has, however, clearly been demonstrated that there is a genetic relationship to
Siberia, insofar as nowhere except north-easternmost Europe and northernmost Scandinavia is there so high a frequency as in western Siberia of a particular genetic marker whose very highest area of frequency is found
1013:
Kristiina
Tambets, Siiri Rootsi, Toomas Kivisild, Hela Help, Piia Serk, Eva-Liis Loogväli, Helle-Viivi Tolk, Maere Reidla, Ene Metspalu, Liana Pliss, Oleg Balanovsky, Andrey Pshenichnov, Elena Balanovska, Marina Gubina, Sergey Zhadanov, Ludmila Osipova, Larisa Damba, Mikhail Voevoda, Ildus Kutuev,
515:
A Swedish study from 2007 has concluded that the haplogroups U5b1b and V (those which dominate mitochondrial DNA among Sámi from northernmost Sweden, Norway and
Finland) likely came to the area very soon after the Ice Age ended. They may have come either from the European continent, or from the
511:
person from northeast
Siberia. That the Fulani and Berbers have had contact with each other was already known, but the results were generally a surprise for the research group, particularly that this variant only seems to have emerged 9000 years ago. One possible explanation could be that the
536:
The
Swedish genetic study of 2007 can be compared with new archaeological discoveries, which are thought to show that northernmost Sweden was populated from the north immediately after the Ice Age. The Komsa culture has thus become central again as the origin of northern Sweden's earliest
151:
During the 6th century, humans from the coastal tracts of
Finland as well as central Sweden, mostly belonging to YDNA Haplogroups I1 and R1a which commonly occur among farmers, made contact with the Sámi. The Sámi numbered very few at that time, and were therefore threatened by the
395:
Even scientists who do not support the idea of a double origin have connected the Sámi to the people of the asbestos-ceramic culture. In Sweden this type of discovery does not occur south of a boundary line which separates Upper Norrland from Central Norrland and
456:
436:, called the Sámi a genetically unique population. His research indicated that the Sámi were not closely related to Asiatic-Mongoloid ethnicities. However, his studies could not explain the origin of the Sámi, but he did exclude the idea that their "
315:
was, according to Wiklund's understanding, also a Paleo-Laplandic relic. As it can be interpreted as "the watercourse that lies east of and underneath the mountain range" he argued that it seemed to have come from the west. Many other place names in
86:
Europe has been populated by four prehistoric waves of migration, of which the first three waves helped form the Germanic and Nordic peoples, whilst a great deal of the Sámi and Finnish population have their roots in the fourth wave.
431:
was developed as a science in the mid-1950s, it became a tool in the research into the origin of the Sámi, and the results suggested that K.B. Wiklund had come closer to the truth than Blumenbach. Lars Beckman, who primarily studied
58:
was developed in the 19th century, Sámi were seen instead to differ from surrounding peoples, which in turn led to the theory that the Sámi had developed as an isolated people during the Ice Age, when they would have overwintered by
339:, he drew the conclusion "racial biology an expression of national prejudices and has nothing to do with science". According to Dahlberg, the differences between the Sámi and others probably depended on environmental factors.
225:
359:
Nor did K.B. Wiklund's opinions last long. Only a few decades after his death, most researchers had abandoned the overwintering theory. The concept of the Paleo-Laplandic language, however, lived on to a certain extent. The
544:
One theory on the origin the Sámi was that they originate from the hunter-gatherer culture known by archaeologists as the Pitted Ware culture. Emellertid However, modern genetic studies have shown this not to be the case.
512:
forefathers of all of these ethnic groups originated in Southwestern Europe, on the border between France and Spain, from where hunter-gatherer tribes spread out in different directions after the last Ice Age.
387:
where they then existed as elk-hunters from 4000 BCE onwards. This group would have spoken Paleo-Laplandic. Around 2000 BCE a finno-ugric speaking people, the asbestos-ceramic culture, would have come to
256:
realised that he could show that relatively large parts of the Norwegian Arctic and Atlantic coasts were ice-free during the last Ice Age, this dovetailed remarkably with Wiklunds theory. Archaeologist
156:, which resulted in uniquely autosomal DNA and an abnormal frequency of haplogroups, but the Sámi traded and, in time, mixed with the resident population of northern Fennoscandia, not least with the
299:
K.B. Wiklund believed that the forefathers of the Sámi, who had overwintered on the Arctic coast, wouldn't have spoken Sámi, rather they would have spoken several unknown languages which he called "
164:
show a somewhat higher frequency of haplogroups commonly found among speakers of Sámi and Finnish languages, and the population in Norrland shows more variety than other Germanic population groups.
440:" was somewhere in Asia or Eastern Europe. His studies of the frequency of so-called Sámi marker-genes indicated that between a quarter and a third of the genetic material of the populations of
375:
and thought the ethnic group of the Sámi actually consisted of two different peoples. The first is suspected to have lived on the Norwegian coast after the Ice Age, and from there traversed the
245:
stated that the Sámi "are in anthropological regards just as removed from the Finns as ever from the Nordic people". (Wiklund meant in terms of physical anthropology, that is the study of
292:
on the edge of the west coast of Sweden betrayed, according to Wiklund, "an unmistakable similarity to our Lapps". Wiklund claimed that right up to modern times there would have been
303:". He thought he found lingering traces of Paleo-Laplandic in the form of words which couldn't be attributed to other languages. For example, this included the Sámi word for water,
284:
in Norway were of an "anthropological type" which were astoundingly similar to the Sámi people, and that archaeologists in precisely this area had found relics of the so-called
128:
A fourth wave, from Siberia, reached Europe c.4000 years ago, constituting a significant addition to finns and Sámi. The YDNA haplogroup (inherited from father to son etc.)
269:, in precisely the area Nordhagen had identified as having been ice-free. Thus Wiklund was finally able to solve "the mystery of the Lappish prehistory". The relics of the
424:. Despite K.B. Wiklund's understanding that the Sámi were just as different from the Finns as from the Scandinavians, this classification remained long into the 1900s.
1063:
328:, could, according to Wiklund, be relics from the Paleo-Laplandic era. Only later did the Sámi switch from speaking their original language to a Finno-Ugric language.
273:
had to be traces of the Sámi people's forefathers, who spent the Ice Age in isolation by the coast. From there they would have spread further south as the ice melted.
229:
Taking different body measurements, in particular the size of the skull, was the basis for the late 19th century and early 20th century's physical anthropology. Here,
75:
research shows that the Sámi group has developed from several different directions at different times from many different hunter-gatherer peoples who moved across the
208:, the Sámi have always lived in Lappland, and did not arrive there from anywhere else. However, the origin of the Sámi has long been a matter of debate. In his book,
491:
While mapping out human mitochondrial DNA in its entirety, one of the aforementioned variants, U5b1b, was found in the mitochondrial DNA of three Sámi people, a
941:
851:. Kungl. Skytteanska samfundets handlingar, 0560-2416 ; 45 (in Swedish) (1. uppl. /3. tr. ed.). Umeå: Skytteanska samf. pp. 110–111.
160:
populations who began the colonisation of central Norrland in the Bronze Age, and northern Norrland since the Middle Ages. Among Swedes, those from
335:
wrote that the thought "that Europeans originate fro a number of pure races is an unsubstantiated hypothesis". Despite he himself working with the
172:
351:
Mellannorrlands The elk-hunters of central Norrland depicted their favourite hunting prey in rock paintings, such as this example at Fångsjön in
331:
It ought to be noted that K.B. Wiklund's discourse about "the Lappish race" soon came to be considered outdate. As early as 1941 the physician
889:
784:
181:
19:
689:
619:
241:
Not everyone agreed, however, on the physical similarities between the Sámi and the Finns. At the turn of the 20th century, the linguist
336:
989:
925:
856:
820:
698:
628:
592:
347:
288:, which strongly resembled the Komsa culture. Wiklund argued that this was no mere coincidence. Even the so-called "pyttar" of
98:
of Southern Europe reached Scandinavia from the South 13000 years ago. Traces of them appear in the Nordic population as mtDNA
392:
from the east. When the two groups met, it is thought they may have merged into a single ethnic group which became the Sámi.
95:
460:
Sámi peoples also stand out as a distinctive people for modern geneticists. Here, Pavva Lars Nilsson Tuorda from Tuorpon
1016:"The Western and Eastern Roots of the Saami—the Story of Genetic "Outliers" Told by Mitochondrial DNA and Y Chromosomes"
118:
114:
285:
417:
413:
300:
253:
948:
80:
28:
1126:
1121:
913:
880:
Niskanen, Markku (2009). "Saami and their origins in light of physical anthropology". In Äikäs Tiina (ed.).
409:
157:
55:
582:
441:
376:
185:
276:
As further evidence for this overwintering theory, K.B Wiklund pointed out that the modern population in
1057:
153:
103:
884:. Publications of the Giellagas Institute, 1458-6282; 12. Oulu: Giellagas-instituhtta. pp. 20–29.
578:
99:
733:. Acta Lapponica, 0348-8993 ; 8 (in Swedish). Uppsala: Almqvist & Wiksell. pp. 79–81.
714:
644:
808:
726:
469:
437:
364:
213:
982:
Norrländsk uppslagsbok: ett uppslagsverk på vetenskaplig grund om den norrländska regionen. Bd 4,
844:
24:
684:
614:
397:
277:
1078:
918:
Genetiska studier av svenska samer. Thule. Kungl. Skytteanska samfundets Årsbok. ISSN 0280-8692
1098:
1045:
985:
921:
885:
852:
816:
780:
704:
694:
634:
624:
588:
477:
445:
129:
47:
136:
from the east at least 1500 years ago. The N1c population brought with it metalwork from the
1090:
1035:
1027:
380:
372:
368:
246:
141:
137:
113:
A second wave of immigration from outside of Europe consisted of Stone Age farmers from the
76:
51:
670:
656:
332:
258:
209:
122:
91:
554:
384:
289:
193:
161:
39:
1040:
1015:
230:
132:
is especially common in Finland and among the Sámi, and is thought to have arrived in
1115:
270:
234:
38:
has been of research interest since at least the early 17th century. Initially, the
521:
481:
389:
262:
242:
177:
133:
60:
964:
Professor Lars Beckman, Umeå: Samerna är en Europeisk genetisk unik urbefolkning!
296:
along the Norwegian and Swedish coasts of a population who survived the Ice Age.
500:
352:
266:
882:
Máttut - máddagat: the roots of Saami ethnicities, societies and spaces/places
433:
321:
317:
205:
708:
638:
1094:
421:
361:
281:
1102:
1049:
559:
541:
BCE to 1300 BCE in Finland, and from 1500 BCE to 1000 CE in Scandinavia.
465:
428:
312:
145:
72:
1079:"A recent genetic link between Sami and the Volga-Ural region of Russia"
504:
496:
492:
107:
94:. A culture which is believed to have overwintered the Ice Age in the
995:
895:
862:
826:
790:
757:
734:
525:
508:
325:
293:
184:
depicts the different peoples of the Nordic region in the form of a
1031:
899:
794:
999:
866:
830:
738:
517:
461:
455:
346:
224:
197:
189:
171:
43:
966:
Samefolket Nr 8/1997 s. 11–13, enligt föredrag av L. Beckman och
761:
252:
But how would such an isolation have occurred? When the botanist
140:, resulting in what is believed to be early Sámi metalwork using
420:, he placed all those who spoke Finno-Ugric languages into the
367:
pointed out in the 1980s the considerable differences between
752:
Brøndum-Nielsen Johannes, Wiklund Karl Bernhard, ed. (1947).
233:
is taking the measurements of the Sámi man Fjellstedt from
754:
Nordisk kultur: samlingsverk. 10, Lapperne ; Lapparna
690:
Svenskarna och deras fäder - de senaste 11 000 åren
620:
Svenskarna och deras fäder - de senaste 11 000 åren
920:(in Swedish). Umeå: Skytteanska Samfundet. p. 58.
984:(in Swedish). Umeå: Norrlands univ.-förl. p. 50.
815:(in Swedish). Stockholm: Norstedt. pp. 136–140.
343:
Relations with elk-hunters and asbestos-ceramicists.
488:in the border regions between Europe and Siberia.
416:(1752-1840) categorised humans into five separate
110:), in Northernmost Asia and in Southern Europe.
79:, and that Sámi as well as Finnish-speaking and
693:(Andra tryckningen ed.). Bonniers förlag.
623:(Andra tryckningen ed.). Bonniers förlag.
980:Edlund Lars-Erik, Frängsmyr Tore, ed. (1996).
480:(which is only passed down by the mother) and
476:Later types of genetic study, particularly of
46:, due to the relative similarity between the
8:
1062:: CS1 maint: multiple names: authors list (
849:Norrlands forntid: ett historiskt perspektiv
1039:
813:Samernas samhälle i tradition och nutid:
452:Studies of mitochondria and Y-chromosomes
204:According to Sámi tradition expressed by
968:Samerna – en genetiskt unik urbefolkning
63:. This theory was eventually abandoned.
18:
777:Stulet land: svensk makt på samisk mark
584:Stulet land: svensk makt på samisk mark
570:
1055:
729:; Granlund John, Manker Ernst (1956).
468:parish, is depicted, photographed by
261:had also found very old dwellings at
182:A Description of the Northern Peoples
7:
220:
1077:Max Ingman; Ulf Gyllensten (2007).
779:(in Swedish). Stockholm: Ordfront.
221:K.B. Wiklund's overwintering theory
1083:European Journal of Human Genetics
1020:American Journal of Human Genetics
756:(in Swedish). Stockholm: Bonnier.
337:State Institute for Racial Biology
307:, which is totally unlike Finnish
14:
117:, and a third wave consisted of
168:Original inhabitants or Finns?
16:Earliest phase in Sámi history
1:
90:The first wave consisted of
414:Johann Friedrich Blumenbach
42:were grouped together with
1143:
775:Lundmark, Lennart (2008).
448:county have Sámi origins.
125:, just before Bronze Age.
687:; Peter Sjölund (2016).
617:; Peter Sjölund (2016).
587:(in Swedish). Ordfront.
532:Genetics and archaeology
29:Johan Fredrik Eckersberg
1095:10.1038/sj.ejhg.5201712
671:"Haplogroup U5 (mtDNA)"
56:biological anthropology
657:"Haplogroup V (mtDNA)"
473:
377:Scandinavian Mountains
356:
238:
201:
36:The origin of the Sámi
32:
970:Umeå universitet 1996
459:
410:physical anthropology
350:
228:
175:
81:Scandinavian language
22:
727:Schefferus, Johannes
942:"Arkiverade kopian"
408:When the father of
214:Johannes Schefferus
176:A woodcarving from
474:
357:
239:
202:
33:
891:978-951-42-9282-8
809:Fjellström, Phebe
786:978-91-7037-353-4
478:mitochondrial DNA
311:. The place-name
247:phenotypic traits
154:bottleneck effect
148:and other areas.
121:herders from the
23:Sámi people from
1134:
1107:
1106:
1074:
1068:
1067:
1061:
1053:
1043:
1010:
1004:
1003:
977:
971:
962:
960:
959:
953:
947:. Archived from
946:
938:
932:
931:
910:
904:
903:
877:
871:
870:
841:
835:
834:
805:
799:
798:
772:
766:
765:
749:
743:
742:
723:
717:
712:
681:
675:
674:
667:
661:
660:
653:
647:
642:
611:
605:
604:
602:
601:
579:Lennart Lundmark
575:
470:Lotten von Düben
365:Phebe Fjellström
142:asbestos-ceramic
138:Ananyino Culture
92:hunter-gatherers
77:Cap of the North
61:the Arctic Ocean
1142:
1141:
1137:
1136:
1135:
1133:
1132:
1131:
1112:
1111:
1110:
1076:
1075:
1071:
1054:
1012:
1011:
1007:
992:
979:
978:
974:
957:
955:
951:
944:
940:
939:
935:
928:
912:
911:
907:
892:
879:
878:
874:
859:
843:
842:
838:
823:
807:
806:
802:
787:
774:
773:
769:
751:
750:
746:
725:
724:
720:
701:
683:
682:
678:
669:
668:
664:
655:
654:
650:
631:
613:
612:
608:
599:
597:
595:
577:
576:
572:
568:
551:
534:
507:, as well as a
454:
406:
404:Genetic studies
345:
333:Gunnar Dahlberg
301:Paleo-Laplandic
259:Anders Nummedal
223:
170:
123:Eurasian Steppe
69:
17:
12:
11:
5:
1140:
1138:
1130:
1129:
1124:
1114:
1113:
1109:
1108:
1089:(1): 115–120.
1069:
1032:10.1086/383203
1026:(4): 661–682.
1005:
990:
972:
933:
926:
905:
890:
872:
857:
836:
821:
800:
785:
767:
744:
718:
699:
676:
662:
648:
629:
606:
593:
581:(2012-04-02).
569:
567:
564:
563:
562:
557:
550:
547:
533:
530:
453:
450:
422:Mongoloid race
405:
402:
344:
341:
254:Rolf Nordhagen
231:Gustaf Retzius
222:
219:
169:
166:
158:Proto-Germanic
68:
65:
48:Sámi languages
15:
13:
10:
9:
6:
4:
3:
2:
1139:
1128:
1125:
1123:
1120:
1119:
1117:
1104:
1100:
1096:
1092:
1088:
1084:
1080:
1073:
1070:
1065:
1059:
1051:
1047:
1042:
1037:
1033:
1029:
1025:
1021:
1017:
1009:
1006:
1001:
997:
993:
991:91-972484-2-8
987:
983:
976:
973:
969:
965:
954:on 2011-08-10
950:
943:
937:
934:
929:
927:91-86438-11-5
923:
919:
915:
914:Beckman, Lars
909:
906:
901:
897:
893:
887:
883:
876:
873:
868:
864:
860:
858:91-7191-227-4
854:
850:
846:
845:Baudou, Evert
840:
837:
832:
828:
824:
822:91-1-853222-5
818:
814:
810:
804:
801:
796:
792:
788:
782:
778:
771:
768:
763:
759:
755:
748:
745:
740:
736:
732:
728:
722:
719:
716:
710:
706:
702:
700:9789100167547
696:
692:
691:
686:
680:
677:
672:
666:
663:
658:
652:
649:
646:
640:
636:
632:
630:9789100167547
626:
622:
621:
616:
610:
607:
596:
594:9789174413823
590:
586:
585:
580:
574:
571:
565:
561:
558:
556:
553:
552:
548:
546:
542:
538:
531:
529:
527:
523:
519:
513:
510:
506:
502:
498:
494:
489:
485:
483:
482:Y chromosomes
479:
471:
467:
463:
458:
451:
449:
447:
443:
439:
435:
430:
425:
423:
419:
415:
411:
403:
401:
399:
393:
391:
386:
383:and northern
382:
378:
374:
373:Southern Sámi
370:
369:Northern Sámi
366:
363:
354:
349:
342:
340:
338:
334:
329:
327:
323:
319:
314:
310:
306:
302:
297:
295:
291:
287:
286:Fosna culture
283:
280:district and
279:
274:
272:
271:Komsa culture
268:
264:
260:
255:
250:
248:
244:
236:
232:
227:
218:
215:
212:, from 1673,
211:
207:
199:
195:
191:
187:
183:
179:
174:
167:
165:
163:
159:
155:
149:
147:
143:
139:
135:
131:
126:
124:
120:
119:Indo-European
116:
111:
109:
105:
104:Haplogroup U5
101:
97:
93:
88:
84:
82:
78:
74:
66:
64:
62:
57:
53:
49:
45:
41:
37:
30:
26:
21:
1127:Sámi peoples
1122:Sámi history
1086:
1082:
1072:
1058:cite journal
1023:
1019:
1008:
981:
975:
967:
963:
956:. Retrieved
949:the original
936:
917:
908:
881:
875:
848:
839:
812:
803:
776:
770:
753:
747:
730:
721:
688:
679:
665:
651:
618:
609:
598:. Retrieved
583:
573:
555:Sámi History
543:
539:
535:
514:
503:person from
490:
486:
475:
442:Västerbotten
426:
407:
394:
390:Fennoscandia
385:Ångermanland
358:
330:
308:
304:
298:
290:Bohus-Malmön
275:
251:
243:K.B. Wiklund
240:
203:
178:Olaus Magnus
162:Västerbotten
150:
134:Fennoscandia
127:
112:
100:Haplogroup V
89:
85:
70:
44:ethnic Finns
35:
34:
444:county and
434:blood types
362:ethnologist
115:Middle East
27:painted by
1116:Categories
958:2011-01-01
685:Karin Bojs
615:Karin Bojs
600:2019-09-25
566:References
524:region of
446:Norrbotten
320:, such as
318:Lappmarken
263:Gurravárri
235:Härjedalen
206:Johan Turi
67:Prehistory
715:inskannad
709:973876808
645:inskannad
639:973876808
322:Sulitelma
282:Sunnfjord
186:Muscovite
1103:16985502
1050:15024688
916:(1990).
900:11766968
847:(1995).
811:(1985).
795:10740653
731:Lappland
549:See also
472:in 1868.
466:Jokkmokk
438:urheimat
429:genetics
398:Jämtland
353:Jämtland
210:Lapponia
146:Norrland
96:refugium
31:in 1852.
25:Karasjok
1041:1181943
1000:1610874
998::
898::
867:7617822
865::
831:8345656
829::
793::
760::
739:1178624
737::
505:Senegal
497:Algeria
381:Lapland
294:relicts
108:Berbers
73:genetic
54:. When
52:Finnish
1101:
1048:
1038:
996:LIBRIS
988:
924:
896:LIBRIS
888:
863:LIBRIS
855:
827:LIBRIS
819:
791:LIBRIS
783:
762:128247
758:LIBRIS
735:LIBRIS
707:
697:
637:
627:
591:
526:Russia
501:Fulani
493:Berber
326:Abisko
196:and a
952:(PDF)
945:(PDF)
560:Sápmi
518:Volga
509:Yakut
495:from
462:siida
427:When
418:races
313:Luleå
305:čáhci
144:, in
1099:PMID
1064:link
1046:PMID
986:ISBN
922:ISBN
886:ISBN
853:ISBN
817:ISBN
781:ISBN
705:OCLC
695:ISBN
635:OCLC
625:ISBN
589:ISBN
522:Ural
499:, a
371:and
324:and
309:vesi
278:Møre
267:Áltá
198:Geat
194:Sámi
192:, a
190:Finn
188:, a
71:New
50:and
40:Sámi
1091:doi
1036:PMC
1028:doi
464:in
379:to
265:in
130:N1c
1118::
1097:.
1087:15
1085:.
1081:.
1060:}}
1056:{{
1044:.
1034:.
1024:74
1022:.
1018:.
994:.
894:.
861:.
825:.
789:.
713:,
703:.
643:,
633:.
412:,
200:.
180:'
1105:.
1093::
1066:)
1052:.
1030::
1002:.
961:.
930:.
902:.
869:.
833:.
797:.
764:.
741:.
711:.
673:.
659:.
641:.
603:.
520:-
355:.
237:.
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.