143:
481:
644:
522:
was broadened considerably and separated from the rest of the skull by paired notches on both sides, at the level of the fourth mandibular teeth, similar to the condition in many other crocodylomorphs. The widest portion of the broadened part is 7.1 centimetres (2.8 inches) wide and the width at the
118:
573:
was considered a problematic taxon by French paleontologist Jeremy E. Martin and
Italian paleontologist Massimo Delfino in 2010, though they noted, like Brochu, that it was likely to have been a eusuchian. Although the narrow snout of
443:
In 2017, Greenlandic paleontologist Jan S. Adolfssen, Danish paleontologist Jesper Milàn and
American paleontologist Matt Friedman noted that a single, rather blunt and wide crocodylomorph tooth from the Faxe quarry in the Middle
523:
point of the notches is just 4.7 centimetres (1.85 inches). The foremost part of the lower jaw, which is generally narrow similar to the upper jaw, was broadened as well, though not to the same extent as the snout.
428:, including some detached teeth, belonging to a single individual. Persson considered this material to be enough to clearly differentiate the fossil animal from all other known long-snouted
1097:"Palaeoenvironments, palaeoecology and palaeobiogeography of Late Cretaceous (Campanian) faunas from the Kristianstad Basin, southern Sweden, with applications for science education"
503:
on both sides of the skull extended forwards until reaching the margin of this skull opening, this being the main diagnostic feature of the genus. Another diagnostic feature is the
530:
were stout and short. The teeth were also somewhat recurved and had cylindrical roots. The surface of the teeth was striated (covered in ridges) densely from the base to the tip.
1235:"A new eusuchian (Crocodylia) tooth from the Early or Middle Paleocene, with a description of the Early–Middle Paleocene boundary succession at Gemmas Allé, Copenhagen, Denmark"
1379:
616:
In 2014, French paleontologists Jeremy E. Martin, Romain Amiot and
Christophe Lécuyer and English paleontologist Michael J. Benton noted that Persson's description of
385:
since its fossils have been discovered in marine deposits. Its teeth were short and stout, possibly an adaptation towards feeding on large fish and invertebrates.
762:
could also have lived in the littoral zone or in a river adjacent to the mainland. Within the
Kristianstad Basin, the fossil site Ivö Klack has yielded the most
377:
In the
Cretaceous, southern Scandinavia was covered by shallow sea and the Ivö Klack site within the Kristianstad Basin, where most of the fossils referred to
366:
in
Denmark. The fragmentary nature of these remains means that the precise classification of the genus remains uncertain. Though typically classified as an
1325:
789:
were stout and short, meaning that it would probably have been adapted to some other form of feeding. According to
Einarsson, the robust teeth of
1409:
609:
significantly predated the earliest similar crocodyloid genera. Schwarz-Wings, Milàn and
Gravesen noted that until a taxonomic revision of the
1265:"Food resources and habitat selection of a diverse vertebrate fauna from the upperlower Campanian of the Kristianstad Basin, southern Sweden"
597:
In 2014, German paleontologist
Daniela Schwarz-Wings and Danish paleontologists Jesper Milàn and Palle Gravesen considered the features of
679:
shallow inland sea home to a diverse marine fauna characteristic of shallow marine life of an inner shelf community and included abundant
754:
described by
Persson in 1959 were recovered from marine sediments, though Persson noted that this was not necessarily an indicator that
774:
preferred to live in coastal waters, where it could lay its eggs on adjacent land, rest and heat up, similar to modern crocodilians.
396:
was described by Swedish paleontologist Per-Ove Persson in 1959 based on fossil material recovered at the Ivö Klack locality in the
495:
was a long- and narrow-snouted crocodylomorph. The bony nostrils on both sides of the head were fused to form a single larger
142:
1404:
1399:
1394:
1389:
1117:
Mesozoic Biotas of Scandinavia and Its Arctic Territories. Special Publications of the Geological Society of London
381:
have been found, was a small and rocky island. Many other dyrosaurids were marine, a lifestyle possibly shared by
731:) were also common and fossils of many species of reptiles, most of them marine, have also been found, including
1096:
613:
material is carried out, its precise systematic position within the entire Crocodylomorpha will remain unclear.
561:
as a crocodyline is no longer considered likely. In 2001, American paleontologist Christopher Brochu noted that
362:
consists of a partial skull and isolated teeth from southern Sweden, with possible additional teeth found on
1384:
1214:
472:
teeth, though it was not referred to the genus due to the lack of a formal comparison to the type material.
1293:
1351:
290:
437:
1183:
1124:
504:
464:. A similar tooth also discovered in Early to Middle Paleocene deposits, this time at Gemmas Allé in
449:
1109:
635:
was also classified as a dyrosaurid in a 2018 paper by Swedish paleontologist Elisabeth Einarsson.
1155:
1156:"Early Campanian mosasaurs (Reptilia; Mosasauridae) from the Kristianstad Basin, southern Sweden"
1142:
668:
397:
352:
137:
130:
1356:
1338:
1234:
1201:
526:
In contrast to modern crocodylians, which typically have long and slender teeth, the teeth of
1343:
1168:
1276:
1249:
1191:
1132:
1081:
1054:
740:
657:
429:
330:
322:
236:
223:
1070:"Crocodylian Snouts in Space and Time: Phylogenetic Approaches Toward Adaptive Radiation"
1187:
1128:
1069:
569:(the group that contains all living crocodilians). Because of the fragmentary material,
1108:
Kear, Benjamin P.; Lindgren, Johan; Hurum, Jørn H.; Milàn, Jesper; Vajda, Vivi (2016).
543:
461:
550:. He based this on the distinct notch in the upper jaw. Persson also noted that since
1373:
1146:
602:
554:
is quite poorly known, detailed comparisons with other crocodylines were impossible.
68:
766:
fossils. Ivö Klack was a small, rocky island during the Cretaceous. The presence of
408:), meaning "crocodile". The species name of the only species referred to the genus,
654:
628:
620:
accorded well with the known material of known contemporary freshwater eusuchians.
547:
480:
371:
249:
210:
117:
1280:
1167:
Martin, Jeremy E.; Amiot, Romain; Lécuyer, Christophe; Benton, Michael J. (2014).
1110:"An introduction to the Mesozoic biotas of Scandinavia and its Arctic territories"
704:
672:
579:
43:
1316:
736:
716:
712:
700:
684:
649:
511:
being unusually long, 13.8 centimetres (5.4 inches), and being reached by the
500:
465:
88:
53:
1254:
1059:
803:
is now believed to have been a marine animal, similar to other dyrosaurids.
782:
676:
351:, described in 1959 by Per Ove Persson based on material recovered from the
326:
197:
154:
93:
37:
30:
1205:
1086:
643:
623:
In a 2016 paper by Australian paleontologist Benjamin Kear and colleagues,
871:
869:
867:
1310:
795:
778:
744:
732:
720:
708:
696:
692:
566:
512:
499:(skull opening), dubbed by Persson as a fenestra exonarina communis. The
496:
425:
421:
413:
367:
174:
83:
78:
63:
58:
48:
1169:"Sea surface temperature contributes to marine crocodylomorph evolution"
370:, since 2016 it has been repeatedly placed within the more basal family
1330:
1196:
1039:
688:
583:
363:
184:
98:
73:
799:, and larger invertebrates. Contrary to Persson's initial assessment,
1137:
724:
445:
164:
1287:
432:, noting that the main distinguishing feature was the nasal bone of
1264:
728:
680:
642:
479:
793:
indicate that it was adapted for feeding on larger fish, such as
777:
Most recent and modern long-snouted crocodylomorphs (notably the
667:
have only been recovered from the Campanian-age deposits in the
453:
1291:
1233:
Schwarz-Wings, Daniela; Milàn, Jesper; Gravesen, Palle (2014).
1040:"Review of the Danian vertebrate fauna of southern Scandinavia"
671:
in Sweden. During the Campanian, the Kristianstad Basin was a
488:(top view), a fragmentary skull. Preserved portions in orange.
436:
extending to the fenestra exonarina communis (a fusion of the
1263:
Sørensen, Anne Mehlin; Surlyk, Finn; Lindgren, Johan (2013).
1215:"Reptiles from the Senonian (U. Cret.) of Scania (S. Sweden)"
345:), meaning "crocodile". The genus contains a single species,
758:
itself would have been purely marine. According to Persson,
468:, in 2014, also accorded well with Persson's description of
594:
contributing to the posterior margin of the bony nostrils.
416:, where the fossils were found. The material Persson based
875:
1038:
Adolfssen, Jan S.; Milàn, Jesper; Friedman, Matt (2017).
903:
901:
888:
886:
884:
578:
is similar to the narrow snouts within the genera in the
962:
934:
932:
930:
928:
842:
840:
838:
836:
823:
821:
819:
817:
815:
507:(the connection between the left and right mandible) of
590:
is part of that superfamily on account of the nasal of
1022:
974:
858:
400:. The generic name derives from the Greek αἰγιαλός (
1300:
440:, or bony nostrils, of both sides of the skull).
727:. Additionally, fish (including a vast array of
565:was an enigmatic crocodyliform, but probably a
1242:Bulletin of the Geological Society of Denmark
1047:Bulletin of the Geological Society of Denmark
8:
325:that lived in what is now Sweden during the
1160:Dissertations in Geology at Lund University
1288:
605:than with the Gavialoidea, but noted that
586:), they considered it to be unlikely that
116:
20:
1380:Late Cretaceous crocodylomorphs of Europe
1253:
1195:
1136:
1085:
1058:
1010:
998:
907:
892:
484:Illustration of the holotype specimen of
456:, Denmark, might be referrable to either
986:
876:Schwarz-Wings, Milàn & Gravesen 2014
938:
919:
846:
827:
811:
412:, derives from the Villand district of
950:
781:) have slender and long teeth, being
7:
1023:Sørensen, Surlyk & Lindgren 2013
859:Adolfssen, Milàn & Friedman 2017
321:is an extinct genus of long-snouted
404:), meaning "seashore", and σοῦχος (
341:), meaning "seashore", and σοῦχος (
653:, another long-snouted and marine
546:, placing it within the subfamily
518:The foremost part of the skull of
14:
1222:Arkiv för Mineralogi och Geologi
770:at the site might indicate that
141:
1068:Brochu, Christopher A. (2001).
337:comes from the Greek αἰγιαλός (
627:was considered to have been a
1:
1410:Fossil taxa described in 1959
1281:10.1016/j.cretres.2013.02.002
1095:Einarsson, Elisabeth (2018).
424:part of the skull and of the
358:The known fossil material of
557:Persson's classification of
460:or to some genus within the
582:(today containing only the
420:on were the remains of the
1426:
601:to accord better with the
303:Aigialosuchus villandensis
1213:Persson, Per-Ove (1959).
296:
289:
138:Scientific classification
136:
124:
115:
23:
16:Extinct genus of reptiles
1255:10.37570/bgsd-2014-62-02
1154:Lindgren, Johan (1998).
1060:10.37570/bgsd-2017-65-01
660:
489:
29:Temporal range: Early
1352:Paleobiology Database
1176:Nature Communications
646:
483:
1087:10.1093/icb/41.3.564
631:, not an eusuchian.
505:mandibular symphysis
438:fenestrae exonarinae
355:in southern Sweden.
282:A. villandensis
125:Teeth attributed to
1269:Cretaceous Research
1188:2014NatCo...5.4658M
1129:2016GSLSP.434....1K
922:, pp. 470–471.
663:Certain fossils of
538:Persson classified
389:History of research
1405:Fossils of Denmark
1400:Cretaceous Denmark
1197:10.1038/ncomms5658
1074:American Zoologist
963:Martin et al. 2014
669:Kristianstad Basin
661:
647:Reconstruction of
490:
398:Kristianstad Basin
353:Kristianstad Basin
131:Kristianstad Basin
1395:Fossils of Sweden
1390:Cretaceous Sweden
1367:
1366:
1339:Open Tree of Life
1294:Taxon identifiers
333:period. The name
314:
313:
270:
129:sp. found in the
1417:
1360:
1359:
1347:
1346:
1334:
1333:
1321:
1320:
1319:
1289:
1284:
1259:
1257:
1239:
1229:
1219:
1209:
1199:
1173:
1163:
1150:
1140:
1138:10.1144/SP434.18
1114:
1104:
1101:Litholund Theses
1091:
1089:
1064:
1062:
1044:
1026:
1020:
1014:
1008:
1002:
1001:, p. 27–30.
996:
990:
984:
978:
975:Kear et al. 2016
972:
966:
960:
954:
948:
942:
936:
923:
917:
911:
905:
896:
890:
879:
873:
862:
856:
850:
844:
831:
825:
305:
301:
268:
261:
248:
146:
145:
120:
110:
40:
21:
1425:
1424:
1420:
1419:
1418:
1416:
1415:
1414:
1370:
1369:
1368:
1363:
1355:
1350:
1342:
1337:
1329:
1324:
1315:
1314:
1309:
1296:
1262:
1237:
1232:
1217:
1212:
1171:
1166:
1153:
1112:
1107:
1094:
1067:
1042:
1037:
1034:
1029:
1021:
1017:
1009:
1005:
997:
993:
985:
981:
973:
969:
961:
957:
949:
945:
937:
926:
918:
914:
906:
899:
891:
882:
874:
865:
857:
853:
845:
834:
826:
813:
809:
785:. The teeth of
750:The fossils of
741:crocodylomorphs
641:
536:
486:A. villandensis
478:
430:crocodylomorphs
410:A. villandensis
391:
348:A. villandensis
331:Late Cretaceous
310:
307:
299:
298:
285:
267:
259:
246:
237:Crocodyliformes
224:Crocodylomorpha
140:
111:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
36:83.5–80.5
35:
34:
33:
27:
17:
12:
11:
5:
1423:
1421:
1413:
1412:
1407:
1402:
1397:
1392:
1387:
1385:Campanian life
1382:
1372:
1371:
1365:
1364:
1362:
1361:
1348:
1335:
1322:
1306:
1304:
1298:
1297:
1292:
1286:
1285:
1260:
1230:
1228:(35): 431–519.
1210:
1164:
1151:
1105:
1092:
1080:(3): 564–585.
1065:
1033:
1030:
1028:
1027:
1015:
1011:Einarsson 2018
1003:
999:Einarsson 2018
991:
979:
967:
955:
953:, p. 566.
943:
941:, p. 473.
924:
912:
908:Einarsson 2018
897:
893:Einarsson 2018
880:
863:
851:
849:, p. 471.
832:
830:, p. 470.
810:
808:
805:
658:crocodylomorph
640:
637:
544:true crocodile
535:
534:Classification
532:
477:
474:
462:Alligatoroidea
450:Faxe Formation
390:
387:
323:crocodylomorph
312:
311:
308:
294:
293:
287:
286:
278:
276:
272:
271:
257:
253:
252:
244:
240:
239:
234:
227:
226:
221:
214:
213:
208:
201:
200:
195:
188:
187:
182:
178:
177:
172:
168:
167:
162:
158:
157:
152:
148:
147:
134:
133:
122:
121:
113:
112:
104:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
41:
28:
15:
13:
10:
9:
6:
4:
3:
2:
1422:
1411:
1408:
1406:
1403:
1401:
1398:
1396:
1393:
1391:
1388:
1386:
1383:
1381:
1378:
1377:
1375:
1358:
1353:
1349:
1345:
1340:
1336:
1332:
1327:
1323:
1318:
1312:
1308:
1307:
1305:
1303:
1302:Aigialosuchus
1299:
1295:
1290:
1282:
1278:
1274:
1270:
1266:
1261:
1256:
1251:
1247:
1243:
1236:
1231:
1227:
1223:
1216:
1211:
1207:
1203:
1198:
1193:
1189:
1185:
1181:
1177:
1170:
1165:
1161:
1157:
1152:
1148:
1144:
1139:
1134:
1130:
1126:
1122:
1118:
1111:
1106:
1102:
1098:
1093:
1088:
1083:
1079:
1075:
1071:
1066:
1061:
1056:
1052:
1048:
1041:
1036:
1035:
1031:
1025:, p. 90.
1024:
1019:
1016:
1012:
1007:
1004:
1000:
995:
992:
988:
987:Lindgren 1998
983:
980:
976:
971:
968:
964:
959:
956:
952:
947:
944:
940:
935:
933:
931:
929:
925:
921:
916:
913:
910:, p. 28.
909:
904:
902:
898:
895:, p. 37.
894:
889:
887:
885:
881:
878:, p. 23.
877:
872:
870:
868:
864:
861:, p. 11.
860:
855:
852:
848:
843:
841:
839:
837:
833:
829:
824:
822:
820:
818:
816:
812:
806:
804:
802:
801:Aigialosuchus
798:
797:
792:
791:Aigialosuchus
788:
787:Aigialosuchus
784:
780:
775:
773:
772:Aigialosuchus
769:
768:Aigialosuchus
765:
764:Aigialosuchus
761:
760:Aigialosuchus
757:
756:Aigialosuchus
753:
752:Aigialosuchus
748:
746:
742:
738:
734:
730:
726:
722:
718:
714:
710:
706:
702:
698:
694:
690:
686:
682:
678:
674:
670:
666:
665:Aigialosuchus
659:
656:
652:
651:
645:
638:
636:
634:
633:Aigialosuchus
630:
626:
625:Aigialosuchus
621:
619:
618:Aigialosuchus
614:
612:
611:Aigialosuchus
608:
607:Aigialosuchus
604:
603:Crocodyloidea
600:
599:Aigialosuchus
595:
593:
592:Aigialosuchus
589:
588:Aigialosuchus
585:
581:
577:
576:Aigialosuchus
572:
571:Aigialosuchus
568:
564:
563:Aigialosuchus
560:
559:Aigialosuchus
555:
553:
552:Aigialosuchus
549:
545:
541:
540:Aigialosuchus
533:
531:
529:
528:Aigialosuchus
524:
521:
520:Aigialosuchus
516:
514:
510:
509:Aigialosuchus
506:
502:
498:
494:
493:Aigialosaurus
487:
482:
475:
473:
471:
470:Aigialosuchus
467:
463:
459:
458:Aigialosuchus
455:
451:
447:
441:
439:
435:
434:Aigialosuchus
431:
427:
423:
419:
418:Aigialosuchus
415:
411:
407:
403:
399:
395:
394:Aigialosuchus
388:
386:
384:
383:Aigialosuchus
380:
379:Aigialosuchus
375:
373:
369:
365:
361:
360:Aigialosuchus
356:
354:
350:
349:
344:
340:
336:
335:Aigialosuchus
332:
329:stage of the
328:
324:
320:
319:
318:Aigialosuchus
309:Persson, 1959
306:
304:
295:
292:
291:Binomial name
288:
284:
283:
277:
274:
273:
269:Persson, 1959
266:
265:
264:Aigialosuchus
258:
255:
254:
251:
245:
242:
241:
238:
235:
232:
229:
228:
225:
222:
219:
216:
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206:
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202:
199:
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189:
186:
183:
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170:
169:
166:
163:
160:
159:
156:
153:
150:
149:
144:
139:
135:
132:
128:
127:Aigialosuchus
123:
119:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
26:
25:Aigialosuchus
22:
19:
1301:
1272:
1268:
1245:
1241:
1225:
1221:
1179:
1175:
1159:
1120:
1116:
1100:
1077:
1073:
1050:
1046:
1032:Bibliography
1018:
1013:, p. 8.
1006:
994:
989:, p. 5.
982:
977:, p. 6.
970:
965:, p. 5.
958:
946:
939:Persson 1959
920:Persson 1959
915:
854:
847:Persson 1959
828:Persson 1959
800:
794:
790:
786:
776:
771:
767:
763:
759:
755:
751:
749:
664:
662:
648:
639:Paleoecology
632:
624:
622:
617:
615:
610:
606:
598:
596:
591:
587:
575:
570:
562:
558:
556:
551:
548:Crocodylinae
539:
537:
527:
525:
519:
517:
508:
492:
491:
485:
469:
457:
442:
433:
417:
409:
405:
401:
393:
392:
382:
378:
376:
372:Dyrosauridae
359:
357:
347:
346:
342:
338:
334:
317:
316:
315:
302:
297:
281:
280:
263:
262:
250:Dyrosauridae
230:
217:
211:Pseudosuchia
204:
191:
126:
24:
18:
1123:(1): 1–14.
951:Brochu 2001
737:sea turtles
713:sea urchins
695:(including
685:brachiopods
673:subtropical
580:Gavialoidea
501:nasal bones
476:Description
198:Archosauria
1374:Categories
807:References
783:piscivores
743:and a few
705:belemnites
701:gastropods
655:dyrosaurid
650:Dyrosaurus
629:dyrosaurid
466:Copenhagen
1275:: 85–92.
1248:: 17–26.
1147:131680084
745:dinosaurs
733:mosasaurs
717:serpulids
709:ammonites
689:bryozoans
677:temperate
567:eusuchian
426:mandibles
368:eusuchian
327:Campanian
275:Species:
161:Kingdom:
155:Eukaryota
31:Campanian
1317:Q4696836
1311:Wikidata
1206:25130564
1182:: 4658.
1053:: 1–23.
796:Enchodus
779:gharials
721:decapods
707:and the
697:bivalves
693:molluscs
513:splenial
497:fenestra
422:anterior
402:aigialos
339:aigialos
243:Family:
185:Reptilia
175:Chordata
171:Phylum:
165:Animalia
151:Domain:
1344:3605992
1331:4967683
1184:Bibcode
1125:Bibcode
725:sponges
584:gharial
515:bone.
406:souchus
364:Zealand
343:souchus
300:†
279:†
256:Genus:
181:Class:
105:↓
1204:
1145:
729:sharks
448:-aged
446:Danian
1357:38436
1238:(PDF)
1218:(PDF)
1172:(PDF)
1143:S2CID
1113:(PDF)
1043:(PDF)
681:algae
542:as a
414:Skåne
231:Clade
218:Clade
205:Clade
192:Clade
1326:GBIF
1202:PMID
723:and
454:Faxe
44:PreꞒ
1277:doi
1250:doi
1192:doi
1133:doi
1121:434
1082:doi
1055:doi
747:.
711:),
675:to
452:at
1376::
1354::
1341::
1328::
1313::
1273:42
1271:.
1267:.
1246:62
1244:.
1240:.
1224:.
1220:.
1200:.
1190:.
1178:.
1174:.
1158:.
1141:.
1131:.
1119:.
1115:.
1099:.
1078:41
1076:.
1072:.
1051:65
1049:.
1045:.
927:^
900:^
883:^
866:^
835:^
814:^
739:,
735:,
719:,
715:,
703:,
699:,
691:,
687:,
683:,
374:.
233::
220::
207::
194::
94:Pg
38:Ma
1283:.
1279::
1258:.
1252::
1226:2
1208:.
1194::
1186::
1180:5
1162:.
1149:.
1135::
1127::
1103:.
1090:.
1084::
1063:.
1057::
260:†
247:†
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:Ꞓ
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