66:
41:
313:. To this point, only a few observations have been made regarding these behaviors. We learn from a study done by Goodman et al. that in early January, some individuals can be found in reproductive condition while others are not. Some males show no scrotal testes while others show large scrotal testes and convoluted
293:. Loky-Manambato is highly heterogenous in its presence of large forested areas and many river systems. In fact, the Loky-Manambato region is so diverse in its make up that areas of dry forest and humid forest occur closely together only to be separated by mountains and rivers in some cases. The
246:
can be characterized by the following: Overall, the texture of body hair is soft and fine. Cover hairs of the dorsum are bi-colored (proximal two-thirds is dark brown, and the distal one-third is a medium buff). The guard hairs, which have the primary role of protecting the rest of the
259:
has a somewhat large cranium, a relatively short rostrum, moderately long molar rows, and a palatine process that is shorter and stouter in comparison to others within the genus. Likewise, the enamel of the anterior portion of the incisors is a dull yellow-orange color.
317:. Likewise, on another occasion in early April, one male was found in non-reproductive condition while another was found with partially scrotal testes and slightly convoluted epididymides. Within this same period, multiple females were found with large
446:
Rakotoarisoa, Jean-Eric; Bailey, Carolyn A.; Hinger, Paula H.; Brenneman, Rick A.; Louis, Edward E. (2013-03-01). "Isolation and characterization of nine microsatellite loci in a
Malagasy endemic rodent, Eliurus carletoni (Rodentia: Nesomyinae)".
584:
Rakotoarisoa, Jean-Eric (June 2010). "Phylogeny and species boundaries of the endemic species complex, Eliurus antsingy and E. carletoni (Rodentia: Muroidea: Nesomyidae), in
Madagascar using mitochondrial and nuclear DNA sequence data".
625:
Rakotoarisoa, J.-E.; Raheriarisena, M.; Goodman, S. M. (2013-05-01). "Late
Quaternary climatic vegetational shifts in an ecological transition zone of northern Madagascar: insights from genetic analyses of two endemic rodent species".
272:
Members of the genus have been found to inhabit all the natural forest types on the island from humid forestation on the eastern side of the island to the xerophytic spiny bushes located in the southwest portion of the island.
277:
can be found specifically in the dry deciduous forests of
Madagascar. Members of the species are known to inhabit forested areas resting on limestone outcrops. In Northern Madagascar, there are three regions of interest where
324:
As we learn with many species, evolution can be driven by a variety of factors. For some, it is competition of various types, and for others, it is a matter of adapting to the changing climate. We learn in the case of
297:
distribution of the region includes elevations ranging from sea level up to 1170 meters. Though the Loky-Manambato region is very diverse, as previously stated,
1851:
337:
forest fragmenting will cause stress on a species to change. However, historical demographic analyses have proposed that the current genetic patterns we see in
1879:
1812:
695:
333:
event occurred between 18,750 and 7,500 years ago, mainly due to the complexity within the Loky-Manambato region. Clearly, deforestation and other
418:
528:"A Phylogeographic Study of the Endemic Rodent Eliurus carletoni (Rodentia: Nesomyinae) in an Ecological Transition Zone of Northern Madagascar"
1889:
914:
229:
499:
1480:
1150:
1838:
986:
1472:
419:"A new species of Eliurus Milne Edwards, 1885 (Rodentia: Nesomyinae) from the Réserve Spéciale d'Ankarana, northern Madagascar"
334:
232:
for his substantial influence in the study of rodent systematics and of rodents native to the island of
Madagascar. Currently,
1512:
890:
228:
is a relatively new species restricted to
Northern Madagascar. The species was named in honor of Dr. Michael Carleton of the
1464:
1387:
1110:
236:
is endemic to the Réserve Spéciale (RS) d’Ankarana. Madagascar is widely known for its unique biomes and biota. The genus
1628:
1488:
906:
688:
1424:
1324:
898:
1536:
1856:
1504:
1277:
995:
938:
808:
800:
1749:
1416:
1332:
922:
65:
1620:
1612:
1528:
1496:
1447:
358:
341:
are occurring due to prehuman vegetational shifts because of the contraction and fragmentation of dry forests.
1087:
1078:
978:
954:
784:
681:
1765:
1393:
1301:
1134:
1118:
1803:
1674:
1378:
1018:
1009:
946:
330:
160:
1285:
1691:
1591:
1142:
970:
1293:
1254:
1064:
1055:
962:
930:
30:
1582:
1246:
659:
472:
60:
1830:
1268:
844:
835:
825:
651:
643:
602:
555:
547:
495:
464:
1884:
1729:
1202:
1193:
1183:
1126:
792:
635:
594:
539:
456:
251:
from abrasion and moisture, show characteristic colors of dark brown to black. The belly of
214:
1720:
1682:
1520:
527:
1369:
1158:
283:
673:
1873:
1825:
1603:
861:
663:
476:
318:
289:
Ankarana and
Analamerana differ from Loky-Manambato in that they are solely areas of
264:
exhibits a unicolor brown tail, which is also characteristic of others in the genus.
50:
45:
1697:
1651:
1642:
1559:
1550:
1407:
1041:
1032:
526:
Rakotoarisoa, Jean-Eric; Raheriarisena, Martin; Goodman, Steven M. (2013-01-01).
492:
Lemurs of the Lost World: exploring the forests and
Crocodile Caves of Madagascar
1843:
1797:
1315:
1219:
767:
757:
598:
1788:
1573:
1349:
705:
460:
314:
294:
210:
202:
127:
647:
551:
468:
1166:
1101:
871:
741:
543:
290:
77:
655:
606:
559:
379:
1782:
747:
723:
97:
1817:
1711:
1665:
1438:
206:
137:
639:
1237:
735:
729:
717:
198:
117:
107:
87:
1759:
309:
Little is known about the reproductive behavior and seasonality of
1763:
677:
321:
all without embryos present, but one with placental scarring.
248:
240:
is part of the order
Rodentia and the family Nesomyidae.
1772:
1709:
1663:
1640:
1601:
1571:
1548:
1436:
1405:
1367:
1348:
1313:
1266:
1235:
1218:
1191:
1182:
1099:
1076:
1053:
1030:
1007:
869:
860:
833:
824:
765:
756:
301:is found to only inhabit the dry forest areas.
689:
8:
1456:Ankarana Special Reserve tufted-tailed rat (
24:Ankarana Special Reserve tufted-tailed rat
1760:
1364:
1232:
1188:
866:
830:
762:
696:
682:
674:
185:Ankarana Special Reserve tufted-tailed rat
39:
20:
205:. It was first described in 2009. It is
176:Goodman, Raheriarisena & Jansa, 2009
350:
359:"Taxonomy browser (Eliurus carletoni)"
587:Molecular Phylogenetics and Evolution
7:
620:
618:
616:
579:
577:
575:
573:
571:
569:
521:
519:
517:
515:
513:
511:
441:
439:
412:
410:
408:
406:
404:
402:
400:
398:
396:
1880:IUCN Red List least concern species
417:Goodman, Steven (September 2009).
230:National Museum of Natural History
14:
915:Monard's African climbing mouse
64:
1513:White-tipped tufted-tailed rat
1481:Grandidier's tufted-tailed rat
1151:Pousargues's African fat mouse
987:Rupp's African climbing mouse (
891:Montane African climbing mouse
628:Journal of Evolutionary Biology
494:. Impact, London. p. 216.
449:Conservation Genetics Resources
197:sp. SAJ-2009a) is a species of
1:
1473:Ellerman's tufted-tailed rat
1388:Brachytarsomys mahajambaensis
255:is characteristically white.
1425:Gregarious short-tailed rat
1419:(Brachyuromys betsileoensis)
1325:South African pouched mouse
1021:(Dendroprionomys rousseloti)
899:Mount Kahuzi climbing mouse
286:, Ankarana, and Analmerana.
1521:Petter's tufted-tailed rat
1505:Dormouse tufted-tailed rat
1465:Daniel's tufted-tailed rat
1278:Southern giant pouched rat
1111:Bocage's African fat mouse
801:Brukkaros pygmy rock mouse
599:10.1016/j.ympev.2010.05.022
221:Distribution and morphology
1906:
1652:Koopman's montane voalavo
1629:Petter's big-footed mouse
1489:Major's tufted-tailed rat
1427:(Brachyuromys ramirohitra)
1417:Betsileo short-tailed rat
1381:(Brachytarsomys albicauda)
907:Lachaise's climbing mouse
803:(Petromyscus monticularis)
305:Reproduction and evolution
1890:Mammals described in 2009
1746:
1621:Greater big-footed mouse
1613:Bastard big-footed mouse
1537:Webb's tufted-tailed rat
1529:Tanala tufted-tailed rat
1497:Lesser tufted-tailed rat
1448:Tsingy tufted-tailed rat
1067:(Megadendromus nikolausi)
811:(Petromyscus shortridgei)
712:
704:Extant species of family
461:10.1007/s12686-012-9768-5
166:
159:
61:Scientific classification
59:
37:
28:
23:
1615:(Macrotarsomys bastardi)
1396:(Brachytarsomys villosa)
1327:(Saccostomus campestris)
1205:(Mystromys albicaudatus)
996:Vernay's climbing mouse
979:Cameroon climbing mouse
955:Chestnut climbing mouse
809:Shortridge's rock mouse
215:Ankarana Special Reserve
1654:(Monticolomys koopmani)
1631:(Macrotarsomys petteri)
1333:Mearns's pouched mouse
1302:Kivu giant pouched rat
1119:Northwestern fat mouse
939:Brant's climbing mouse
923:Lovat's climbing mouse
917:(Dendromus leucostomus)
901:(Dendromus kahuziensis)
1675:White-bellied nesomys
1623:(Macrotarsomys ingens)
1515:(Eliurus penicillatus)
1394:Hairy-tailed Antsangy
1379:White-tailed antsangy
1304:(Cricetomys kivuensis)
1296:(Cricetomys gambianus)
1019:Velvet climbing mouse
957:(Dendromus mystacalis)
947:Banana climbing mouse
795:(Petromyscus collinus)
787:(Petromyscus barbouri)
1723:Voalavo antsahabensis
1585:(Hypogeomys antimena)
1483:(Eliurus grandidieri)
1335:(Saccostomus mearnsi)
1280:(Cricetomys ansorgei)
1169:(Steatomys pratensis)
1129:(Steatomys cuppedius)
1088:Dollman's tree mouse
971:Nyika climbing mouse
949:(Dendromus messorius)
941:(Dendromus mesomelas)
933:(Dendromus melanotis)
909:(Dendromus lachaisei)
785:Barbour's rock mouse
544:10.1093/jhered/ess083
490:Wilson, Jane (2014).
1692:Nesomys narindaensis
1685:(Nesomys lambertoni)
1592:Hypogeomys australis
1562:(Gymnuromys roberti)
1294:Gambian pouched rat
1255:Greater hamster-rat
1137:(Steatomys jacksoni)
1135:Jackson's fat mouse
1121:(Steatomys caurinus)
1044:(Malacothrix typica)
963:Kivu climbing mouse
931:Gray climbing mouse
893:(Dendromus insignis)
1732:Voalavo gymnocaudus
1677:(Nesomys audeberti)
1583:Malagasy giant rat
1475:(Eliurus ellermani)
1286:Emin's pouched rat
1247:Lesser hamster-rat
1145:(Steatomys krebsii)
1113:(Steatomys bocagei)
998:(Dendromus vernayi)
847:(Delanymys brooksi)
532:Journal of Heredity
426:Zoological Bulletin
380:"Eliurus carletoni"
193:) (synonymous with
31:Conservation status
1730:Northern voalavo (
1507:(Eliurus myoxinus)
1450:(Eliurus antsingy)
1288:(Cricetomys emini)
1161:(Steatomys parvus)
1153:(Steatomys opimus)
1143:Krebs's fat mouse
1090:(Prionomys batesi)
973:(Dendromus nyikae)
965:(Dendromus nyasae)
925:(Dendromus lovati)
361:. Ncbi.nlm.nih.gov
1867:
1866:
1804:Eliurus carletoni
1774:Eliurus carletoni
1766:Taxon identifiers
1757:
1756:
1742:
1741:
1721:Eastern voalavo (
1523:(Eliurus petteri)
1467:(Eliurus danieli)
1458:Eliurus carletoni
1361:
1344:
1343:
1229:
1214:
1213:
1203:White-tailed rat
1178:
1177:
1127:Dainty fat mouse
1065:Nikolaus's mouse
981:(Dendromus oreas)
884:
856:
855:
820:
819:
793:Pygmy rock mouse
778:
640:10.1111/jeb.12116
244:Eliurus carletoni
226:Eliurus carletoni
190:Eliurus carletoni
181:
180:
170:Eliurus carletoni
152:E. carletoni
54:
16:Species of rodent
1897:
1860:
1859:
1847:
1846:
1834:
1833:
1821:
1820:
1808:
1807:
1806:
1793:
1792:
1791:
1761:
1683:Western nesomys
1531:(Eliurus tanala)
1491:(Eliurus majori)
1365:
1360:
1359:
1353:
1233:
1228:
1227:
1223:
1189:
883:
882:
876:
867:
831:
777:
776:
772:
763:
698:
691:
684:
675:
668:
667:
634:(5): 1019–1034.
622:
611:
610:
581:
564:
563:
523:
506:
505:
487:
481:
480:
443:
434:
433:
423:
414:
391:
390:
388:
387:
376:
370:
369:
367:
366:
355:
172:
69:
68:
48:
43:
42:
21:
1905:
1904:
1900:
1899:
1898:
1896:
1895:
1894:
1870:
1869:
1868:
1863:
1855:
1850:
1842:
1837:
1829:
1824:
1816:
1811:
1802:
1801:
1796:
1787:
1786:
1781:
1768:
1758:
1753:
1738:
1705:
1700:(Nesomys rufus)
1659:
1636:
1597:
1567:
1544:
1539:(Eliurus webbi)
1499:(Eliurus minor)
1432:
1401:
1357:
1355:
1354:
1352:
1340:
1309:
1262:
1249:(Beamys hindei)
1225:
1224:
1222:
1210:
1174:
1159:Tiny fat mouse
1095:
1072:
1049:
1026:
1010:Dendroprionomys
1003:
989:Dendromus ruppi
880:
878:
877:
875:
852:
845:Delany's mouse
816:
774:
773:
771:
752:
708:
702:
672:
671:
624:
623:
614:
583:
582:
567:
525:
524:
509:
502:
489:
488:
484:
445:
444:
437:
421:
416:
415:
394:
385:
383:
378:
377:
373:
364:
362:
357:
356:
352:
347:
307:
270:
223:
177:
174:
168:
155:
63:
55:
44:
40:
33:
17:
12:
11:
5:
1903:
1901:
1893:
1892:
1887:
1882:
1872:
1871:
1865:
1864:
1862:
1861:
1848:
1835:
1822:
1809:
1794:
1778:
1776:
1770:
1769:
1764:
1755:
1754:
1747:
1744:
1743:
1740:
1739:
1737:
1736:
1727:
1717:
1715:
1707:
1706:
1704:
1703:
1695:
1688:
1680:
1671:
1669:
1661:
1660:
1658:
1657:
1648:
1646:
1638:
1637:
1635:
1634:
1626:
1618:
1609:
1607:
1599:
1598:
1596:
1595:
1588:
1579:
1577:
1569:
1568:
1566:
1565:
1556:
1554:
1546:
1545:
1543:
1542:
1534:
1526:
1518:
1510:
1502:
1494:
1486:
1478:
1470:
1462:
1453:
1444:
1442:
1434:
1433:
1431:
1430:
1422:
1413:
1411:
1403:
1402:
1400:
1399:
1391:
1384:
1375:
1373:
1370:Brachytarsomys
1362:
1356:(Malagasy rats
1346:
1345:
1342:
1341:
1339:
1338:
1330:
1321:
1319:
1311:
1310:
1308:
1307:
1299:
1291:
1283:
1274:
1272:
1264:
1263:
1261:
1260:
1257:(Beamys major)
1252:
1243:
1241:
1230:
1226:(Pouched rats)
1216:
1215:
1212:
1211:
1209:
1208:
1199:
1197:
1186:
1180:
1179:
1176:
1175:
1173:
1172:
1164:
1156:
1148:
1140:
1132:
1124:
1116:
1107:
1105:
1097:
1096:
1094:
1093:
1084:
1082:
1074:
1073:
1071:
1070:
1061:
1059:
1051:
1050:
1048:
1047:
1038:
1036:
1028:
1027:
1025:
1024:
1015:
1013:
1005:
1004:
1002:
1001:
993:
984:
976:
968:
960:
952:
944:
936:
928:
920:
912:
904:
896:
887:
885:
881:climbing mice)
864:
858:
857:
854:
853:
851:
850:
841:
839:
828:
822:
821:
818:
817:
815:
814:
806:
798:
790:
781:
779:
760:
754:
753:
751:
750:
744:
738:
732:
726:
720:
713:
710:
709:
703:
701:
700:
693:
686:
678:
670:
669:
612:
565:
507:
500:
482:
455:(1): 203–205.
435:
392:
371:
349:
348:
346:
343:
306:
303:
284:Loky-Manambato
269:
266:
222:
219:
201:in the family
179:
178:
175:
164:
163:
157:
156:
149:
147:
143:
142:
135:
131:
130:
125:
121:
120:
115:
111:
110:
105:
101:
100:
95:
91:
90:
85:
81:
80:
75:
71:
70:
57:
56:
38:
35:
34:
29:
26:
25:
15:
13:
10:
9:
6:
4:
3:
2:
1902:
1891:
1888:
1886:
1883:
1881:
1878:
1877:
1875:
1858:
1853:
1849:
1845:
1840:
1836:
1832:
1827:
1823:
1819:
1814:
1810:
1805:
1799:
1795:
1790:
1784:
1780:
1779:
1777:
1775:
1771:
1767:
1762:
1752:
1751:
1745:
1735:
1733:
1728:
1726:
1724:
1719:
1718:
1716:
1714:
1713:
1708:
1702:
1701:
1698:Island mouse
1696:
1694:
1693:
1689:
1687:
1686:
1681:
1679:
1678:
1673:
1672:
1670:
1668:
1667:
1662:
1656:
1655:
1650:
1649:
1647:
1645:
1644:
1639:
1633:
1632:
1627:
1625:
1624:
1619:
1617:
1616:
1611:
1610:
1608:
1606:
1605:
1604:Macrotarsomys
1600:
1594:
1593:
1589:
1587:
1586:
1581:
1580:
1578:
1576:
1575:
1570:
1564:
1563:
1560:Voalavoanala
1558:
1557:
1555:
1553:
1552:
1547:
1541:
1540:
1535:
1533:
1532:
1527:
1525:
1524:
1519:
1517:
1516:
1511:
1509:
1508:
1503:
1501:
1500:
1495:
1493:
1492:
1487:
1485:
1484:
1479:
1477:
1476:
1471:
1469:
1468:
1463:
1461:
1459:
1454:
1452:
1451:
1446:
1445:
1443:
1441:
1440:
1435:
1429:
1428:
1423:
1421:
1420:
1415:
1414:
1412:
1410:
1409:
1404:
1398:
1397:
1392:
1390:
1389:
1385:
1383:
1382:
1377:
1376:
1374:
1372:
1371:
1366:
1363:
1351:
1347:
1337:
1336:
1331:
1329:
1328:
1323:
1322:
1320:
1318:
1317:
1312:
1306:
1305:
1300:
1298:
1297:
1292:
1290:
1289:
1284:
1282:
1281:
1276:
1275:
1273:
1271:
1270:
1265:
1259:
1258:
1253:
1251:
1250:
1245:
1244:
1242:
1240:
1239:
1234:
1231:
1221:
1220:Cricetomyinae
1217:
1207:
1206:
1201:
1200:
1198:
1196:
1195:
1190:
1187:
1185:
1181:
1171:
1170:
1165:
1163:
1162:
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1075:
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1063:
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1057:
1056:Megadendromus
1052:
1046:
1045:
1042:Gerbil mouse
1040:
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1029:
1023:
1022:
1017:
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868:
865:
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862:Dendromurinae
859:
849:
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838:
837:
832:
829:
827:
823:
813:
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789:
788:
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769:
764:
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758:Petromyscinae
755:
749:
746:Superfamily:
745:
743:
739:
737:
733:
731:
727:
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721:
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501:9781874687481
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382:. Uniprot.org
381:
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335:anthropogenic
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161:Binomial name
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67:
62:
58:
52:
47:
46:Least Concern
36:
32:
27:
22:
19:
1773:
1748:
1731:
1722:
1710:
1699:
1690:
1684:
1676:
1664:
1653:
1643:Monticolomys
1641:
1630:
1622:
1614:
1602:
1590:
1584:
1572:
1561:
1549:
1538:
1530:
1522:
1514:
1506:
1498:
1490:
1482:
1474:
1466:
1457:
1455:
1449:
1437:
1426:
1418:
1408:Brachyuromys
1406:
1395:
1386:
1380:
1368:
1334:
1326:
1314:
1303:
1295:
1287:
1279:
1267:
1256:
1248:
1236:
1204:
1192:
1184:Mystromyinae
1168:
1160:
1152:
1144:
1136:
1128:
1120:
1112:
1100:
1089:
1077:
1066:
1054:
1043:
1031:
1020:
1008:
997:
988:
980:
972:
964:
956:
948:
940:
932:
924:
916:
908:
900:
892:
870:
846:
834:
826:Delanymyinae
810:
802:
794:
786:
766:
631:
627:
593:(1): 11–22.
590:
586:
538:(1): 23–35.
535:
531:
491:
485:
452:
448:
429:
425:
384:. Retrieved
374:
363:. Retrieved
353:
339:E. carletoni
338:
327:E. carletoni
326:
323:
315:epididymides
311:E. carletoni
310:
308:
299:E. carletoni
298:
288:
280:E. carletoni
279:
275:E. carletoni
274:
271:
262:E. carletoni
261:
257:E. carletoni
256:
253:E. carletoni
252:
243:
242:
237:
234:E. carletoni
233:
225:
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194:
189:
188:
184:
182:
169:
167:
151:
150:
138:
18:
1798:Wikispecies
1316:Saccostomus
1033:Malacothrix
775:(Rock mice)
768:Petromyscus
295:topographic
1874:Categories
1574:Hypogeomys
1551:Gymnuromys
1350:Nesomyinae
1269:Cricetomys
1167:Fat mouse
740:Suborder:
706:Nesomyidae
432:: 133–149.
386:2010-06-03
365:2010-06-03
345:References
331:bottleneck
282:live: the
211:Madagascar
203:Nesomyidae
128:Nesomyidae
1358:and mice)
1194:Mystromys
1102:Steatomys
1079:Prionomys
872:Dendromus
836:Delanymys
742:Myomorpha
716:Kingdom:
664:206046529
648:1420-9101
552:0022-1503
477:255782305
469:1877-7252
291:limestone
213:, in the
146:Species:
84:Kingdom:
78:Eukaryota
1831:48266951
1818:10712421
1789:Q4765992
1783:Wikidata
1750:Category
879:(African
748:Muroidea
736:Rodentia
730:Mammalia
724:Chordata
722:Phylum:
718:Animalia
656:23621368
607:20685291
560:23132906
124:Family:
118:Rodentia
108:Mammalia
98:Chordata
94:Phylum:
88:Animalia
74:Domain:
51:IUCN 3.1
1885:Eliurus
1844:1003744
1712:Voalavo
1666:Nesomys
1439:Eliurus
734:Order:
728:Class:
329:that a
268:Habitat
238:Eliurus
207:endemic
195:Eliurus
139:Eliurus
134:Genus:
114:Order:
104:Class:
49: (
1857:685572
1238:Beamys
662:
654:
646:
605:
558:
550:
498:
475:
467:
319:mammae
249:pelage
199:rodent
660:S2CID
473:S2CID
422:(PDF)
1852:NCBI
1826:IUCN
1813:GBIF
652:PMID
644:ISSN
603:PMID
556:PMID
548:ISSN
496:ISBN
465:ISSN
183:The
1839:MDD
636:doi
595:doi
540:doi
536:104
457:doi
209:to
1876::
1854::
1841::
1828::
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658:.
650:.
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632:26
630:.
615:^
601:.
591:57
589:.
568:^
554:.
546:.
534:.
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510:^
471:.
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451:.
438:^
430:56
428:.
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395:^
217:.
1734:)
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1460:)
991:)
697:e
690:t
683:v
666:.
638::
609:.
597::
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542::
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479:.
459::
453:5
389:.
368:.
187:(
53:)
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