140:
1382:
1376:
119:
382:
392:
is the smallest and one of the oldest desmostylians with an estimated body length of 168 cm (66 in). It is known from a rib, a humerus, a femur, and three vertebrae of the single species and holotype,
577:
Hayashi, Shoji; Houssaye, Alexandra; Nakajima, Yasuhisa; Kentaro, Chiba; Ando, Tatsuro; Sawamura, Hiroshi; Inuzuka, Norihisa; Kaneko, Naotomo; Osaki, Tomohiro (2013).
660:
1429:
1508:
1493:
1498:
653:
1311:
646:
127:
1381:
1503:
1076:
139:
1340:
501:
483:
1488:
1483:
1397:
564:
Inuzuka, N. (2000). "Primitive late
Oligocene desmostylians from Japan and Phylogeny of the Desmostylia".
341:
310:
1455:
507:
489:
248:
1032:
997:
467:. as a "shallow water swimmer, either hovering slowly at a preferred depth, or walking on the bottom".
1193:
590:
579:"Bone Inner Structure Suggests Increasing Aquatic Adaptations in Desmostylia (Mammalia, Afrotheria)"
967:
404:
1375:
1251:
1224:
909:
836:
720:
669:
134:
1011:
1460:
953:
826:
796:
1442:
1122:
1039:
626:
608:
296:
207:
1447:
1235:
616:
598:
280:
431:, but not as dense as in sirenians. The recovered long bones lack inner cavities, like in
1271:
1262:
944:
424:
594:
1244:
1186:
1138:
621:
578:
419:
407:(large and dense) bones. The ribs are broader than in other desmostylians, similar to
1477:
1156:
1018:
988:
69:
1285:
1211:
1145:
1069:
1004:
856:
786:
747:
428:
603:
1300:
1179:
1172:
1112:
1099:
1050:
974:
960:
900:
876:
806:
699:
437:
284:
194:
44:
1420:
1278:
1165:
1083:
1059:
930:
886:
866:
846:
816:
737:
705:
413:
89:
54:
612:
356:
343:
325:
312:
1025:
923:
916:
442:
370:
151:
94:
38:
630:
1414:
1317:
687:
300:
171:
84:
79:
64:
59:
49:
30:
118:
1434:
1350:
1329:
757:
408:
99:
74:
1346:
693:
681:
638:
288:
181:
161:
1391:
381:
769:
380:
304:
1395:
642:
546:
534:
519:
452:
503:
Morawan (Lower Hard Shale Member) (Oligocene of Japan)
1208:
1109:
1065:
985:
941:
897:
882:
872:
862:
852:
842:
832:
822:
812:
802:
792:
782:
753:
743:
733:
530:
528:
1404:
1328:
1299:
1261:
1234:
1207:
1155:
1121:
1108:
1097:
1049:
984:
940:
896:
767:
718:
654:
566:Bulletin of the Ashoro Museum of Paleontology
8:
423:; and extant, semi-aquatic mammals such as
1392:
1216:
1118:
1105:
775:
726:
715:
661:
647:
639:
117:
20:
620:
602:
476:
7:
411:ribs, and very dense, like those of
445:pattern is different from that in
14:
1380:
1374:
138:
279:(named after its type locality
1:
1509:Fossil taxa described in 2000
369:) and were dated to the late
1494:Prehistoric placental genera
604:10.1371/journal.pone.0059146
1525:
128:Hokkaido University Museum
16:Species of mammal (fossil)
1499:Oligocene mammals of Asia
1371:
1219:
778:
729:
714:
676:
283:) is an extinct genus of
254:
247:
135:Scientific classification
133:
125:
116:
23:
510:. Retrieved March 2013.
295:have been found in the
287:, aquatic, herbivorous
492:. Retrieved March 2013
386:
1456:Paleobiology Database
508:Paleobiology Database
490:Paleobiology Database
384:
126:Mounted skeleton in
595:2013PLoSO...859146H
547:Hayashi et al. 2013
535:Hayashi et al. 2013
520:Hayashi et al. 2013
453:Hayashi et al. 2013
405:pachyosteosclerotic
353: /
338:, paleocoordinates
322: /
721:Paenungulatomorpha
670:Paenungulatomorpha
549:, pp. 7–9, 15
387:
1471:
1470:
1443:Open Tree of Life
1406:Ashoroa laticosta
1398:Taxon identifiers
1389:
1388:
1369:
1368:
1365:
1364:
1361:
1360:
1295:
1294:
1203:
1202:
1123:Paleoparadoxiidae
1093:
1092:
1033:Postschizotherium
998:Hengduanshanhyrax
485:Ashoroa laticosta
395:Ashoroa laticosta
297:Morawan Formation
272:
271:
261:Ashoroa laticosta
240:A. laticosta
228:
208:Paleoparadoxiidae
1516:
1504:Fossils of Japan
1464:
1463:
1451:
1450:
1438:
1437:
1425:
1424:
1423:
1393:
1384:
1379:
1378:
1236:Arsinoitheriidae
1217:
1210:
1194:Ounalashkastylus
1119:
1111:
1106:
1067:
987:
943:
899:
884:
874:
864:
854:
844:
834:
824:
814:
804:
794:
784:
776:
755:
745:
735:
727:
716:
663:
656:
649:
640:
634:
624:
606:
573:
550:
544:
538:
532:
523:
517:
511:
499:
493:
481:
459:, together with
385:Skeletal diagram
368:
367:
365:
364:
363:
358:
354:
351:
350:
349:
346:
337:
336:
334:
333:
332:
327:
323:
320:
319:
318:
315:
281:Ashoro, Hokkaido
263:
259:
226:
219:
206:
193:
143:
142:
121:
111:
41:
29:Temporal range:
21:
1524:
1523:
1519:
1518:
1517:
1515:
1514:
1513:
1474:
1473:
1472:
1467:
1459:
1454:
1446:
1441:
1433:
1428:
1419:
1418:
1413:
1400:
1390:
1385:
1373:
1357:
1324:
1291:
1272:Crivadiatherium
1263:Palaeoamasiidae
1257:
1230:
1199:
1151:
1102:
1089:
1045:
980:
945:Titanohyracidae
936:
892:
772:
763:
723:
710:
672:
667:
637:
576:
563:
559:
554:
553:
545:
541:
533:
526:
518:
514:
500:
496:
482:
478:
473:
425:Eurasian beaver
379:
361:
359:
355:
352:
347:
344:
342:
340:
339:
330:
328:
324:
321:
316:
313:
311:
309:
308:
268:
265:
257:
256:
243:
225:
217:
204:
191:
137:
112:
110:
109:
108:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
36:
35:
33:
27:
17:
12:
11:
5:
1522:
1520:
1512:
1511:
1506:
1501:
1496:
1491:
1486:
1476:
1475:
1469:
1468:
1466:
1465:
1452:
1439:
1426:
1410:
1408:
1402:
1401:
1396:
1387:
1386:
1372:
1370:
1367:
1366:
1363:
1362:
1359:
1358:
1356:
1355:
1354:
1353:
1343:
1334:
1332:
1326:
1325:
1323:
1322:
1321:
1320:
1314:
1305:
1303:
1297:
1296:
1293:
1292:
1290:
1289:
1282:
1275:
1267:
1265:
1259:
1258:
1256:
1255:
1248:
1245:Arsinoitherium
1240:
1238:
1232:
1231:
1229:
1228:
1220:
1214:
1205:
1204:
1201:
1200:
1198:
1197:
1190:
1187:Kronokotherium
1183:
1176:
1169:
1161:
1159:
1153:
1152:
1150:
1149:
1142:
1139:Paleoparadoxia
1135:
1127:
1125:
1116:
1103:
1098:
1095:
1094:
1091:
1090:
1088:
1087:
1080:
1073:
1063:
1055:
1053:
1047:
1046:
1044:
1043:
1036:
1029:
1022:
1015:
1008:
1001:
993:
991:
982:
981:
979:
978:
971:
964:
957:
949:
947:
938:
937:
935:
934:
927:
920:
913:
905:
903:
894:
893:
891:
890:
880:
870:
860:
850:
840:
830:
820:
810:
800:
790:
779:
773:
768:
765:
764:
762:
761:
751:
741:
730:
724:
719:
712:
711:
709:
708:
702:
696:
690:
684:
677:
674:
673:
668:
666:
665:
658:
651:
643:
636:
635:
574:
560:
558:
555:
552:
551:
539:
524:
512:
494:
475:
474:
472:
469:
465:Paleoparadoxia
433:Paleoparadoxia
420:Paleoparadoxia
378:
375:
357:44.6°N 141.4°E
326:43.3°N 143.8°E
291:. Fossils of
270:
269:
266:
252:
251:
245:
244:
236:
234:
230:
229:
215:
211:
210:
202:
198:
197:
189:
185:
184:
179:
175:
174:
169:
165:
164:
159:
155:
154:
149:
145:
144:
131:
130:
123:
122:
114:
113:
105:
104:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
42:
28:
15:
13:
10:
9:
6:
4:
3:
2:
1521:
1510:
1507:
1505:
1502:
1500:
1497:
1495:
1492:
1490:
1489:Chattian life
1487:
1485:
1484:Desmostylians
1482:
1481:
1479:
1462:
1457:
1453:
1449:
1444:
1440:
1436:
1431:
1427:
1422:
1416:
1412:
1411:
1409:
1407:
1403:
1399:
1394:
1383:
1377:
1352:
1348:
1344:
1342:
1338:
1337:
1336:
1335:
1333:
1331:
1327:
1319:
1315:
1313:
1309:
1308:
1307:
1306:
1304:
1302:
1298:
1288:
1287:
1283:
1281:
1280:
1276:
1274:
1273:
1269:
1268:
1266:
1264:
1260:
1254:
1253:
1249:
1247:
1246:
1242:
1241:
1239:
1237:
1233:
1227:
1226:
1222:
1221:
1218:
1215:
1213:
1206:
1196:
1195:
1191:
1189:
1188:
1184:
1182:
1181:
1177:
1175:
1174:
1170:
1168:
1167:
1163:
1162:
1160:
1158:
1157:Desmostylidae
1154:
1148:
1147:
1143:
1141:
1140:
1136:
1134:
1133:
1129:
1128:
1126:
1124:
1120:
1117:
1114:
1107:
1104:
1101:
1096:
1086:
1085:
1081:
1079:
1078:
1074:
1072:
1071:
1064:
1062:
1061:
1057:
1056:
1054:
1052:
1048:
1042:
1041:
1037:
1035:
1034:
1030:
1028:
1027:
1023:
1021:
1020:
1019:Parapliohyrax
1016:
1014:
1013:
1009:
1007:
1006:
1002:
1000:
999:
995:
994:
992:
990:
989:Pliohyracidae
983:
977:
976:
972:
970:
969:
968:Rupestrohyrax
965:
963:
962:
958:
956:
955:
951:
950:
948:
946:
939:
933:
932:
928:
926:
925:
921:
919:
918:
914:
912:
911:
907:
906:
904:
902:
895:
889:
888:
881:
879:
878:
871:
869:
868:
861:
859:
858:
851:
849:
848:
841:
839:
838:
831:
829:
828:
821:
819:
818:
811:
809:
808:
801:
799:
798:
791:
789:
788:
781:
780:
777:
774:
771:
766:
760:
759:
752:
750:
749:
742:
740:
739:
732:
731:
728:
725:
722:
717:
713:
707:
703:
701:
697:
695:
691:
689:
685:
683:
679:
678:
675:
671:
664:
659:
657:
652:
650:
645:
644:
641:
632:
628:
623:
618:
614:
610:
605:
600:
596:
592:
589:(4): e59146.
588:
584:
580:
575:
571:
567:
562:
561:
556:
548:
543:
540:
536:
531:
529:
525:
521:
516:
513:
509:
505:
504:
498:
495:
491:
487:
486:
480:
477:
470:
468:
466:
462:
458:
454:
450:
448:
444:
440:
439:
434:
430:
426:
422:
421:
416:
415:
410:
406:
402:
398:
396:
391:
383:
376:
374:
372:
366:
335:
306:
302:
298:
294:
290:
286:
282:
278:
277:
267:Inuzuka, 2000
264:
262:
253:
250:
249:Binomial name
246:
242:
241:
235:
232:
231:
227:Inuzuka, 2000
224:
223:
216:
213:
212:
209:
203:
200:
199:
196:
190:
187:
186:
183:
180:
177:
176:
173:
170:
167:
166:
163:
160:
157:
156:
153:
150:
147:
146:
141:
136:
132:
129:
124:
120:
115:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
40:
32:
26:
22:
19:
1405:
1286:Palaeoamasia
1284:
1277:
1270:
1250:
1243:
1223:
1212:Embrithopoda
1192:
1185:
1178:
1171:
1164:
1146:Neoparadoxia
1144:
1137:
1131:
1130:
1082:
1075:
1070:Gigantohyrax
1068:
1058:
1038:
1031:
1024:
1017:
1010:
1005:Kvabebihyrax
1003:
996:
973:
966:
959:
952:
929:
922:
915:
908:
885:
875:
865:
857:Saghatherium
855:
845:
835:
825:
815:
805:
795:
787:Dimaitherium
785:
756:
748:Hadrogeneios
746:
736:
704:Superorder:
698:Infraclass:
586:
582:
569:
565:
557:Bibliography
542:
515:
502:
497:
484:
479:
464:
460:
456:
455:interpreted
451:
446:
436:
432:
429:hippopotamus
418:
412:
400:
399:
394:
389:
388:
292:
285:desmostylian
275:
274:
273:
260:
255:
239:
238:
221:
220:
24:
18:
1312:Proboscidea
1301:Proboscidea
1252:Namatherium
1225:Stylolophus
1180:Desmostylus
1173:Cornwallius
1113:Desmostylia
1100:Tethytheria
1077:Heterohyrax
1060:Dendrohyrax
1051:Procaviidae
975:Titanohyrax
961:Antilohyrax
910:Brachyhyrax
901:Geniohyidae
877:Selenohyrax
837:Regubahyrax
807:Megalohyrax
700:Placentalia
522:, p. 2
438:Desmostylus
377:Description
362:44.6; 141.4
360: /
331:43.3; 143.8
329: /
195:Desmostylia
37:28–23
1478:Categories
1279:Hypsamasia
1166:Behemotops
1012:Meroehyrax
931:Pachyhyrax
887:Thyrohyrax
867:Seggeurius
847:Rukwalorax
817:Microhyrax
770:Hyracoidea
738:Abdounodus
706:Afrotheria
537:, Abstract
471:References
461:Behemotops
447:Behemotops
443:trabecular
441:, and the
414:Behemotops
1345:includes
1318:elephants
1316:includes
1026:Pliohyrax
954:Afrohyrax
924:Geniohyus
917:Bunohyrax
827:Namahyrax
797:Helioseus
680:Kingdom:
613:837402105
572:: 91–123.
371:Oligocene
233:Species:
158:Kingdom:
152:Eukaryota
1421:Q4805827
1415:Wikidata
1351:manatees
1084:Procavia
1040:Prohyrax
694:Mammalia
688:Chordata
686:Phylum:
682:Animalia
631:23565143
583:PLOS ONE
409:sirenian
348:141°24′E
317:143°48′E
301:Hokkaido
201:Family:
182:Mammalia
172:Chordata
168:Phylum:
162:Animalia
148:Domain:
31:Chattian
1448:3617752
1435:4968810
1347:dugongs
1341:Sirenia
1330:Sirenia
1132:Ashoroa
758:Ocepeia
692:Class:
622:3615000
591:Bibcode
506:in the
488:in the
457:Ashoroa
401:Ashoroa
390:Ashoroa
345:44°36′N
314:43°18′N
293:Ashoroa
276:Ashoroa
258:†
237:†
222:Ashoroa
214:Genus:
188:Order:
178:Class:
106:↓
25:Ashoroa
629:
619:
611:
289:mammal
1461:71795
305:Japan
1430:GBIF
1349:and
1339:see
1310:see
627:PMID
609:OCLC
463:and
435:and
427:and
417:and
403:had
45:PreꞒ
617:PMC
599:doi
299:on
1480::
1458::
1445::
1432::
1417::
625:.
615:.
607:.
597:.
585:.
581:.
568:.
527:^
449:.
397:.
373:.
303:,
95:Pg
39:Ma
1209:†
1115:?
1110:†
1066:†
986:†
942:†
898:†
883:†
873:†
863:†
853:†
843:†
833:†
823:†
813:†
803:†
793:†
783:†
754:†
744:†
734:†
662:e
655:t
648:v
633:.
601::
593::
587:8
570:1
307:(
218:†
205:†
192:†
100:N
90:K
85:J
80:T
75:P
70:C
65:D
60:S
55:O
50:Ꞓ
34:~
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.