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Cereal growth staging scales

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growth regulators are typically applied at Z30. Disease control is most critical in the stem extension and heading stage (Z31, Z32, Z35), in particular as soon as the flag leaf is out (Z37). The crop is also more sensitive to heat or frost at some stages than others (for example, during the meiosis
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Critical stages according to BBCH (Lancashire et al., 1991) and their deviations from the Zadok's scale (in brackets). The spacing between the stages follows calendar time for the temperate climate of Switzerland (sowing in November, harvest
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that is widely used in cereal research and agriculture. Knowing the stages of development of a crop is critical in many management decisions that growers make. They are represented on a scale from 10 to 92. For example, in some countries,
903:
stage the crop is very sensitive to low temperature). Knowing the growth stage of the crop when checking for problems is essential for deciding which control measures should be followed.
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Lancashire, P.D.; H. Bleiholder; P. Langeluddecke; R. Stauss; T. van den Boom; E. Weber; A. Witzen-Berger (1991). "A uniform decimal code for growth stages of crops and weeds".
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Lancashire, P.D.; H. Bleiholder; P. Langeluddecke; R. Stauss; T. van den Boom; E. Weber; A. Witzen-Berger (1991). "A uniform decimal code for growth stages of crops and weeds".
613:. Like other scales of crop development, the Feekes scale is useful in planning management strategies that incorporate plant growth information for the use of 261:
Beginning of stem elongation: pseudostem and tillers erect, first internode begins to elongate, top of inflorescence at least 1 cm above tillering node
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Witzenberger, A.; H. Hack; T. van den Boom (1989). "Erläuterungen zum BBCH-Dezimal-Code für die Entwicklungsstadien des Getreides - mit Abbildungen".
2016: 675:
Tillers formed; leaves often twisted spirally. Main shoot and six tillers. In some varieties of winter wheat, plant may be "creeping," or prostrate.
1983: 898:, the recommendation for the first nitrogen application on wheat is 6 weeks before Z30, with the second application on Z30. 2124: 2024: 610: 2144: 454:
End of flowering: all spikelets have completed flowering but some dehydrated anthers may remain
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Large, E. C. (1 December 1954). "GROWTH STAGES IN CEREALS ILLUSTRATION OF THE FEEKES SCALE".
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Beginning of heading: tip of inflorescence emerged from sheath, first spikelet just visible
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J.C. Zadoks, T.T. Chang, C.F. Konzak, "A Decimal Code for the Growth Stages of Cereals",
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A leaf is unfolded when its ligule is visible or the tip of the next leaf is visible
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Tillering or stem elongation may occur earlier than stage 13; in this case continue
1984:"Growth Stages of Wheat: Identification and Understanding Improve Crop Management" 683:
Beginning of the erection of the pseudo-stem; leaf sheaths beginning to lengthen.
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If stem elongation begins before the end of tillering continue with stage 30
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The phenological growth stages and BBCH-identification keys of cereals are:
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Sheath of flag leaf completely grown out; ear swollen but not yet visible.
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Flag leaf (last leaf) visible but still rolled up; ear beginning to swell.
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One shoot (number of leaves can be added), first leaf through coleoptile.
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Willem Feekes (1907-1979) in 1941. This scale is more widely used in the
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Soft dough: grain content soft but dry. Fingernail impression not held
895: 871: 594: 58: 34: 899: 67: 2023:. University of Kentucky College of Agriculture. Archived from 309:
Flag leaf stage: flag leaf fully unrolled, ligule just visible
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Emergence: coleoptile penetrates soil surface (cracking stage)
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Medium milk: grain content milky, grains reached final size,
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Watery ripe: first grains have reached half their final size
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Second node of stem formed; next-to-last leaf just visible.
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Pseudo-stem (formed by sheaths of leaves) strongly erected.
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Hard dough: grain content solid. Fingernail impression held
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than other similar and more descriptive scales such as the
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Fully ripe: grain hard, difficult to divide with thumbnail
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Over-ripe: grain very hard, cannot be dented by thumbnail
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Mealy ripe; contents of kernel soft but dry. Soft dough.
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Radicle elongated, root hairs and/or side roots visible
1989:. Texas Agricultural Extension Service. Archived from 593:
is a system to identify the growth and development of
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Mid boot stage: flag leaf sheath just visibly swollen
2021:
A Comprehensive Guide to Wheat Management in Kentucky
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End of tillering. Maximum no. of tillers detectable
1907:Feekes, Willem (1941). "De tarwe en haar milieu ". 667:Beginning of tillering; main shoot and one tiller. 269:First node at least 1 cm above tillering node 847:Kernel hard (difficult to divide with thumbnail). 1789:Kernel hard (difficult to divide with thumbnail) 395:Middle of heading: half of inflorescence emerged 27:Scales used to measure the growth of cereal crops 218:Beginning of tillering: first tiller detectable 1803:Kernel hard (no longer dented with thumbnail) 438:Beginning of flowering: first anthers visible 773:Three-quarters of heading process completed. 704:First node of stem visible at base of shoot. 320:Early boot stage: flag leaf sheath extending 33:attempt to objectively measure the growth of 8: 1909:Verslagen van de Technische Tarwe Commissie. 427:End of heading: inflorescence fully emerged 1924: 1922: 632:Cereal growth stages using the Feekes scale 621:to avoid damaging the crop and/or maximize 1956:Wise K, Johnson B, Mansfield C, Krupke C. 890:applications must be completed during the 757:One-quarter of heading process completed. 336:Late boot stage: flag leaf sheath swollen 818:Flowering completed; kernel watery ripe. 352:First awns visible (in awned forms only) 1534:First spikelet of inflorescence visible 954: 924:Z30: ear is one centimeter long in wheat 874:development scale proposed by the Dutch 627: 79: 1899: 285:Node 3 at least 2 cm above node 2 277:Node 2 at least 2 cm above node 1 446:Full flowering: 50% of anthers mature 1589:Emergence of inflorescence completed 1446:Flag leaf ligule/collar just visible 810:Flowering completed at base of head. 749:First spikelet of head just visible. 301:Flag leaf just visible, still rolled 7: 1827:Overripe, straw dead and collapsing 357:5: Inflorescence emergence, heading 2017:"Section 2. Growth and Development" 1851:Viable seed giving 50% germination 802:Flowering complete to top of head. 765:Half of heading process completed. 2081:10.1111/j.1744-7348.1991.tb04895.x 2058:10.1111/j.1744-7348.1991.tb04895.x 1943:10.1111/j.1365-3059.1954.tb00716.x 728:Ligule of flag leaf just visible. 140:Coleoptile emerged from caryopsis 25: 1328:Main shoot and 9 or more tillers 959:Comparison of growth stage scales 1958:"Managing Wheat by Growth Stage" 124:Radicle emerged from caryopsis 1085:First leaf through coleoptile 855:Ripe for cutting. Straw dead. 159:First leaf through coleoptile 1: 2015:Herbek J, Lee C (July 2009). 1963:. Purdue University Extension 1575:3/4 of inflorescence emerged 1561:1/2 of inflorescence emerged 1547:1/4 of inflorescence emerged 1054:Coleoptile emerged from seed 941:Z55: the head is 1/2 emerged. 546:Grains loosening in day-time 419:80% of inflorescence emerged 411:70% of inflorescence emerged 403:60% of inflorescence emerged 387:40% of inflorescence emerged 379:30% of inflorescence emerged 371:20% of inflorescence emerged 108:Beginning of seed imbibition 1815:Kernel loosening in daytime 1067:Leaf just at coleoptile tip 31:Cereal growth staging scales 1875:Secondary dormancy induced 1476:Flag leaf sheath extending 906:Examples of typical stages 293:Stages continuous till ... 242:Stages continuous till ... 191:Stages continuous till ... 2166: 1201:9 or more leaves unfolded 1042:Radicle emerged from seed 554:Plant dead and collapsing 199:9 or more leaves unfolded 1763: 1707: 1638: 1593: 1519: 1502:Flag leaf sheath opening 1450: 1332: 1316:Main shoot and 8 tillers 1304:Main shoot and 7 tillers 1292:Main shoot and 6 tillers 1279:Main shoot and 5 tillers 1267:Main shoot and 4 tillers 1255:Main shoot and 3 tillers 1243:Main shoot and 2 tillers 1205: 1071: 993: 957: 822: 785: 781:All heads out of sheath. 740: 695: 650: 630: 531: 496: 458: 431: 356: 344:Flag leaf sheath opening 313: 254: 203: 152: 116:Seed imbibition complete 93: 2125:Citation Classic of 1985 1911:(in Dutch and English). 1887:Secondary dormancy lost 1520:Inflorescence emergence 1231:Main shoot and 1 tiller 930:Z32: second node visible 794:Beginning of flowering. 597:introduced by the Dutch 459:7: Development of fruit 1433:Flag leaf just visible 927:Z31: first node visible 432:6: Flowering, anthesis 1608:Beginning on anthesis 921:during stem extension 153:1: Leaf development, 74: 51:BBCH-scale for cereals 1346:Pseudo stem erection 916:Z21: tillering begins 234:3 tillers detectable 226:2 tillers detectable 100:Dry seed (caryopsis) 71: 1420:6th node detectable 1408:5th node detectable 1396:4th node detectable 1384:3rd node detectable 1372:2nd node detectable 1359:1st node detectable 1098:First leaf unfolded 1030:Imbibition complete 1018:Start of imbibition 949:Z92: grains are ripe 167:First leaf unfolded 41:BBCH-scale (cereals) 1996:on 25 November 2011 1665:Kernel watery ripe 1634:Anthesis completed 1515:First awns visible 1490:Boots just swollen 255:3: Stem elongation 1708:Dough development 1621:Anthesis half-way 1189:8 leaves unfolded 1176:7 leaves unfolded 1163:6 leaves unfolded 1150:5 leaves unfolded 1137:4 leaves unfolded 1124:3 leaves unfolded 1111:2 leaves unfolded 562:Harvested product 183:3 leaves unfolded 175:2 leaves unfolded 75: 18:BBCH-scale (maize) 1891: 1890: 1863:Seed not dormant 1639:Milk development 910:during tillering 859: 858: 566: 565: 16:(Redirected from 2157: 2097: 2090:Gesunde Pflanzen 2084: 2061: 2036: 2035: 2033: 2032: 2012: 2006: 2005: 2003: 2001: 1995: 1988: 1982:Miller, Travis. 1979: 1973: 1972: 1970: 1968: 1962: 1953: 1947: 1946: 1926: 1917: 1916: 1904: 1333:Stem Elongation 1218:Main shoot only 1072:Seedling growth 955: 946:during ripening 876:phytopathologist 628: 80: 21: 2165: 2164: 2160: 2159: 2158: 2156: 2155: 2154: 2130: 2129: 2121: 2104: 2102:Further reading 2087: 2069:Ann. 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Retrieved 2025:the original 2020: 2010: 1998:. Retrieved 1991:the original 1977: 1965:. Retrieved 1951: 1934: 1930: 1912: 1908: 1902: 1733:Early dough 1691:Medium milk 994:Germination 987: 980: 973: 966: 958: 905: 868:Zadoks scale 867: 865: 862:Zadoks scale 831:Milky ripe. 644: 639: 631: 607:Zadoks scale 595:cereal crops 591:Feekes scale 590: 588: 585:Feekes scale 575: 503:Early dough 497:8: Ripening 483:still green 482: 89:Description 83:Growth stage 76: 55:phenological 50: 44: 30: 29: 2114:14:415-421. 2000:29 February 1967:29 February 1759:Hard dough 1746:Soft dough 1678:Early milk 988:Description 645:Description 619:fertilizers 599:agronomists 473:Early milk 314:4: Booting 210:No tillers 2145:BBCH-scale 2134:Categories 2096:: 384–388. 2031:2016-05-14 1915:: 523–888. 1894:References 1703:Late milk 1206:Tillering 894:stage. In 879:Jan Zadoks 786:Flowering 651:Tillering 623:crop yield 615:pesticides 611:BBCH scale 492:Late milk 63:BBCH-scale 61:using the 1764:Ripening 1594:Anthesis 1006:Dry seed 892:tillering 888:herbicide 823:Ripening 73:mid-July) 2140:Agronomy 1451:Booting 884:nitrogen 741:Heading 47:agronomy 2150:Cereals 609:or the 59:cereals 35:cereals 974:Feekes 896:France 872:cereal 49:, the 1994:(PDF) 1987:(PDF) 1961:(PDF) 1659:10.54 1602:10.51 983:Scale 976:Scale 969:Scale 967:Zadok 900:Wheat 870:is a 815:10.54 807:10.53 799:10.52 791:10.51 640:Stage 2112:1974 2002:2012 1969:2012 1800:16.0 1797:11.4 1784:11.3 1756:15.0 1741:11.2 1730:14.0 1686:11.1 1675:13.0 1662:12.1 1631:11.6 1618:11.5 1605:11.4 1586:11.0 1583:10.5 1572:10.7 1569:10.4 1558:10.5 1555:10.3 1542:10.2 1531:10.2 1528:10.1 1512:10.1 981:Haun 886:and 866:The 852:11.4 844:11.3 836:11.2 828:11.1 778:10.5 770:10.4 762:10.3 754:10.2 746:10.1 617:and 589:The 86:Code 2077:doi 2073:119 2054:doi 2050:119 1939:doi 1487:9.2 1473:8-9 1341:4-5 1186:7.+ 1173:6.+ 1160:5.+ 1147:4.+ 1134:3.+ 1121:2.+ 1108:1.+ 1095:1.+ 1064:0.0 290:3 . 239:2 . 188:1 . 45:In 2136:: 2094:41 2092:. 2071:. 2048:. 2019:. 1933:. 1921:^ 1913:17 1880:99 1868:98 1856:97 1844:96 1832:95 1820:94 1808:93 1794:92 1781:91 1776:- 1769:90 1751:87 1738:85 1725:83 1720:- 1713:80 1696:77 1683:75 1670:73 1656:71 1651:- 1644:70 1626:69 1613:65 1599:60 1580:59 1566:57 1552:55 1539:53 1525:50 1507:49 1495:47 1484:10 1481:45 1468:41 1463:- 1456:40 1438:39 1425:37 1413:36 1401:35 1389:34 1377:33 1364:32 1351:31 1338:30 1321:29 1309:28 1297:27 1284:26 1272:25 1260:24 1248:23 1236:22 1223:21 1211:20 1194:19 1181:18 1168:17 1155:16 1142:15 1129:14 1116:13 1103:12 1090:11 1077:10 1059:09 1047:07 1035:05 1023:03 1011:01 999:00 733:10 625:. 559:99 551:97 543:93 535:92 524:89 516:87 508:85 500:83 489:77 478:75 470:73 462:71 451:69 443:65 435:61 424:59 416:58 408:57 400:56 392:55 384:54 376:53 368:52 360:51 349:49 341:47 333:45 325:43 317:41 306:39 298:37 282:33 274:32 266:31 258:30 247:29 231:23 223:22 215:21 207:20 196:19 180:13 172:12 164:11 156:10 145:09 137:07 129:06 121:05 113:03 105:01 97:00 65:. 37:. 2083:. 2079:: 2060:. 2056:: 2034:. 2004:. 1971:. 1945:. 1941:: 1935:3 1441:9 1428:8 1367:7 1354:6 1287:3 1226:2 1080:1 725:9 717:8 709:7 701:6 688:5 680:4 672:3 664:2 656:1 20:)

Index

BBCH-scale (maize)
cereals
agronomy
phenological
cereals
BBCH-scale

cereal crops
agronomists
United States
Zadoks scale
BBCH scale
pesticides
fertilizers
crop yield
cereal
phytopathologist
Jan Zadoks
nitrogen
herbicide
tillering
France
Wheat


doi
10.1111/j.1365-3059.1954.tb00716.x
"Managing Wheat by Growth Stage"
"Growth Stages of Wheat: Identification and Understanding Improve Crop Management"
the original

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