3886:
3892:
554:
127:
4442:
542:
149:
442:
863:
617:
4472:
807:
4448:
724:
1482:
873:
568:
skeleton (MPC-D 100/535) from the Barun Goyot
Formation at the Hermiin Tsav locality. In 2010 and 2011 this specimen was examined to analyze several borings (tunnel-like holes) left by invertebrae scavengers on joint areas. As of 2019, MPC-D 100/535 remains largely undescribed. In 2019 a partial
781:
and therefore, junior synonyms. Czepiński reexamined many of the specimens originally described by Maryańska and Osmólska, as well as the respective holotypes of these taxa, providing evidence that all traits used to separate them are, in fact, indistinguishably present on
738:. Its holotype (PIN 3142/299) is represented by a very small and juvenile skull that was collected from the Hermiin Tsav locality near the end of the 1970s by the Joint Soviet–Mongolian Paleontological Expedition. Though Alifanov used this skull to erect the new
1392:
field paleoenvironments, under a semi-arid climate. The formation is known for its vertebrate fossils in life-like poses, most of which are preserved in unstructured sandstone, indicating a catastrophic rapid burial.
2715:
3015:
Pei, R.; Qin, Yuying; Wen, Aishu; Zhao, Q.; Wang, Z.; Liu, Z.; Guo, W.; Liu, P.; Ye, W.; Wang, L.; Yin, Z.; Dai, R.; Xu, X. (2022). "A new troodontid from the Upper
Cretaceous Gobi Basin of inner Mongolia, China".
596:, and evaluated the implications of these specimens for correlation of fossil sites of the latter formation. He considered one of these specimens in particular, MPC-D 100/551B, as a potential evidence of an
2741:
456:
During the large field work of the Polish-Mongolian
Palaeontological Expeditions in the 1970s, abundant protoceratopsid specimens were discovered on eroded surfaces of the Hermiin Tsav locality of the
2157:
Averianov, A. O.; Lopatin, A. V. (2022). "A re-appraisal of
Parvicursor remotus from the Late Cretaceous of Mongolia: implications for the phylogeny and taxonomy of alvarezsaurid theropod dinosaurs".
415:
was among the smallest ceratopsians, growing up to 1–1.5 m (3.3–4.9 ft) in length, with a weight about 22.7–45 kg (50–99 lb). Although emerging late in the reign of the dinosaurs,
2642:"Djadokhta Formation correlative strata in Chinese Inner Mongolia: an overview of the stratigraphy, sedimentary geology, and paleontology and comparisons with the type locality in the pre-Altai Gobi"
581:. This specimen was discovered in 2007 also from the Hermiin Tsav locality of the Barun Goyot Formation, and includes a partially preserved skull with partial skeleton of an adult individual.
926:, were long considered valid and distinct taxa, and sometimes placed within Protoceratopsidae, Czepiński found the diagnostic features used to distinguish these taxa to be largely present in
1547:
Watabe, M., Suzuki, S., 2000a. Report on the Japan-Mongolia Joint
Paleontological Expedition to the Gobi desert, 1993. Hayashibara Museum of Natural SciencesResearch Bulletin 1, 17 29.
2679:
Eberth, D. A. (1993). "Depositional environments and facies transitions of dinosaur-bearing Upper
Cretaceous redbeds at Bayan Mandahu (Inner Mongolia, People's Republic of China)".
3059:
Gao, K.; Hou, L. (1996). "Systematics and taxonomic diversity of squamates from the Upper
Cretaceous Djadochta Formation, Bayan Mandahu, Gobi Desert, People's Republic of China".
2869:
Godefroit, P.; Currie, P. J.; Li, H.; Shang, C. Y.; Dong, Z.-M. (2008). "A new species of
Velociraptor (Dinosauria: Dromaeosauridae) from the Upper Cretaceous of northern China".
1995:
Eberth, D. A. (2018). "Stratigraphy and paleoenvironmental evolution of the dinosaur-rich
Baruungoyot-Nemegt succession (Upper Cretaceous), Nemegt Basin, southern Mongolia".
1751:
Alifanov, V. R. (2003). "Two new dinosaurs of the infraorder
Neoceratopsia (Ornithischia) from the Upper Cretaceous of the Nemegt Depression, Mongolian People's Republic".
1636:"The postcranial skeleton of Bagaceratops (Ornithischia: Neoceratopsia) from the Baruungoyot Formation (Upper Cretaceous) in Hermiin Tsav of southwestern Gobi, Mongolia"
820:
was a small-sized protoceratopsid, reaching adult dimensions of about 1–1.5 m (3.3–4.9 ft) in length and 22.7–45 kg (50–99 lb) in body mass based on
720:
based on the reasoning that all exhibit anatomical traits already seen on other specimens of this protoceratopsid, and some of them are likely products of preservation.
465:
712:
region of a large individual and was collected during expeditions led by the Sino-Canadian Dinosaur Project. In 2006 Mackoviky regarded all of these ceratopsians as
1806:
1592:"Trace fossils on dinosaur bones from Upper Cretaceous eolian deposits in Mongolia: Taphonomic interpretation of paleoecosystems in ancient desert environments"
2560:
Gradziński, R.; Jerzykiewicz, T. (1974). "Dinosaur- and mammal-bearing aeolian and associated deposits of the Upper Cretaceous in the Gobi Desert (Mongolia)".
4593:
2079:
Arbour, V. M.; Currie, P. J.; Badamgarav, D. (2014). "The ankylosaurid dinosaurs of the Upper Cretaceous Baruungoyot and Nemegt formations of Mongolia".
4613:
4608:
2476:
Funston, G. F.; Mendonca, S. E.; Currie, P. J.; Barsbold, R.; Barsbold, R. (2018). "Oviraptorosaur anatomy, diversity and ecology in the Nemegt Basin".
3096:"New Genus and Species of Djadochtatheriid Multituberculate (Allotheria, Mammalia) from the Upper Cretaceous Bayan Mandahu Formation of Inner Mongolia"
4552:
2912:
Xing, X.; Choinere, J. N.; Pittman, M.; Tan, Q. W.; Xiao, D.; Li, Z. Q.; Tan, L.; Clark, J. M.; Norell, M. A.; Hone, D. W. E; Sullivan, C. (2010).
2108:"A new ankylosaurid skeleton from the Upper Cretaceous Baruungoyot Formation of Mongolia: its implications for ankylosaurid postcranial evolution"
4603:
3135:"New exceptionally well-preserved specimens of "Zangerlia" neimongolensis from Bayan Mandahu, Inner Mongolia, and their taxonomic significance"
530:
452:
fossils have been reported from the Bayan Mandahu (bottom right), Hermiin Tsav, Khulsan (both left), and possibly Udyn Sayr (center) localities
2597:"Taxonomic composition and systematics of late Cretaceous lizard assemblages from Ukhaa Tolgod and adjacent localities, Mongolian Gobi Desert"
3885:
1955:
3207:
2810:
Xing, X.; Sullivan, Corwin; Pittman, M.; Choiniere, J. N.; Hone, D. W. E.; Upchurch, P.; Tan, Q.; Xiao, Dong; Lin, Tan; Han, F. (2011).
2106:
Park, J.-Y.; Lee, Y. N.; Currie, P. J.; Ryan, M. J.; Bell, P.; Sissons, R.; Koppelhus, E. B.; Barsbold, R.; Lee, S.; Kim, S.-H. (2021).
500:, which consists of a relatively medium-sized skull, and a vast majority of the specimens collected by the expeditions were assigned to
2512:
1847:"The Tiny Horned Dinosaur Gobiceratops minutus gen. et sp. nov. (Bagaceratopidae, Neoceratopsia) from the Upper Cretaceous of Mongolia"
3891:
4465:
2288:
2232:
1770:
You, H.; Dong, Z. (2003). "A New Protoceratopsid (Dinosauria: Neoceratopsia) from the Late Cretaceous of Inner Mongolia, China".
1495:
4530:
758:
1635:
4618:
1002:
743:
1164:
in sands are also common across the unit. In addition, structureless, medium-grained, fine-grained and very fine-grained
4441:
2058:"Upper Cretaceous Djadokhta, Barun Goyot and Nemegt formations of Mongolia, including remarks on previous subdivisions"
148:
2914:"A new dromaeosaurid (Dinosauria: Theropoda) from the Upper Cretaceous Wulansuhai Formation of Inner Mongolia, China"
2716:"A new Species of Protoceratops (Dinosauria, Neoceratopsia) from the Late Cretaceous of Inner Mongolia (P. R. China)"
2989:
Xing, X.; Qing-Wei, T.; Shuo, W.; Sullivan, C.; Hone, D. W. E.; Feng-Lu, H.; Qing-Yu, M.; Lin, T.; Dong, T. (2013).
1807:"Yamaceratops dorngobiensis, a New Primitive Ceratopsian (Dinosauria: Ornithischia) from the Cretaceous of Mongolia"
838:
were very similar (mostly in the postcranial skeleton), the former had a much derived (advanced) skull morphology.
469:
2742:"A new species of the ankylosaurid dinosaur Pinacosaurus from the Late Cretaceous of Inner Mongolia (P.R. China)"
1730:
2608:
2345:"Redescription and affinities of Hulsanpes perlei (Dinosauria, Theropoda) from the Upper Cretaceous of Mongolia"
3298:
3200:
2194:"Description and ecologic analysis of Hollanda luceria, a Late Cretaceous bird from the Gobi Desert (Mongolia)"
1707:
950:
the derived-most one. Below are the proposed phylogenetic relationships within Protoceratopsidae by Czepiński:
824:. It had a smaller frill, about ten grinding teeth per jaw, and more triangular skull than its close relative,
676:
of the Hermiin Tsav locality. He also coined the family Bagaceratopidae in order to contain these new taxa and
633:
2990:
2882:
1821:
1384:-deposited) sediments. It is likely that sediments at Bayan Mandahu were deposited by short-lived rivers and
3416:
2192:
Bell, A. K.; Chiappe, L. M.; Erickson, G. M.; Suzuki, S.; Watabe, M.; Barsbold, R.; Tsogtbaatar, K. (2010).
1977:
1349:
942:. Based on cranial characters such as presence or absence of premaxillary teeth and an antorbital fenestra,
693:
398:
4492:
1157:
978:
573:
was described by Bitnara Kim and colleagues, who noted no significant differences between the skeleton of
564:
In 1993, the Japan-Mongolia Joint Paleontological Expedition collected an articulated and nearly complete
553:
126:
1168:
predominate in sediments of the Barun Goyot Formation. The sediments of this formation were deposited in
4598:
4565:
4475:
3325:
3177:
1558:
1121:
457:
394:
4447:
488:
dedicated to describe this material where they named the new genus and type species of protoceratopsid
2950:"A new oviraptorid (Dinosauria: Theropoda) from the Upper Cretaceous of Bayan Mandahu, Inner Mongolia"
1675:"New protoceratopsid specimens improve the age correlation of the Upper Cretaceous Gobi Desert strata"
4539:
3146:
3068:
3025:
2991:"A new oviraptorid from the Upper Cretaceous of Nei Mongol,China, and its stratigraphic implications"
2961:
2823:
2688:
2653:
2640:
Jerzykiewicz, T.; Currie, P. J.; Eberth, D. A.; Johnston, P. A.; Koster, E. H.; Zheng, J.-J. (1993).
2569:
2485:
2430:
2205:
2031:
2004:
1858:
1779:
1603:
1523:
1161:
668:(PIN 4487/26; a partial small-sized skull), recovered from the Khulsan locality, and the holotype of
3581:
3193:
851:
593:
481:
477:
406:
2535:
4301:
4251:
3683:
3133:
Brinkman, D. B.; Tong, H.-Y.; Li, H.; Sun, Y.; Zhang, J.-S.; Godefroit, P.; Zhang, Z.-M. (2015).
3115:
3041:
2894:
2886:
2792:
2622:
2325:
2269:
2174:
1926:
1874:
1655:
1293:
577:
and the former, with the exception of the anatomy of the skull and the shape and location of the
509:
319:
143:
2767:
541:
423:—and kept the small body size that characterized early ceratopsians. Unlike its close relative,
4570:
846:
teeth, had paired (fused) nasal bones, and an oval-shaped fenestra (hole) was developed in the
734:
In 2008 Alifanov described and named another ceratopsian taxon from the Barun Goyot Formation,
681:
4384:
4322:
4285:
3759:
3646:
2851:
2458:
2376:
2317:
2261:
2139:
1980:
1951:
1731:"The fossil record, systematics and evolution of pachycephalosaurs and ceratopsians from Asia"
1181:
957:
930:
and thus becoming synonyms of this genus. Under this reasoning, Protoceratopsidae consists of
270:
1198:
is the most common taxon across the Barun Goyot Formation, which was also home to many other
664:
from the Barun Goyot Formation. Material assigned by Alifanov corresponds to the holotype of
4414:
4212:
4037:
3739:
3434:
3154:
3107:
3076:
3033:
2969:
2928:
2878:
2841:
2831:
2782:
2696:
2661:
2612:
2604:
2577:
2493:
2448:
2438:
2366:
2356:
2307:
2299:
2251:
2243:
2213:
2166:
2129:
2119:
2088:
2057:
2012:
1918:
1866:
1825:
1817:
1787:
1686:
1647:
1611:
1570:
1487:
1464:
1284:
1189:
2287:
Napoli, J. G.; Ruebenstahl, A. A.; Bhullar, B.-A. S.; Turner, A. H.; Norell, M. A. (2021).
4308:
4260:
4219:
4155:
4058:
4009:
3711:
3635:
3590:
3464:
3450:
3409:
3283:
3267:
1260:
1129:
862:
386:
375:
37:
786:
and they fall within the large intraspecific variation of this taxon. He also considered
3150:
3072:
3029:
2965:
2913:
2827:
2766:
Currie, P. J.; Badamgarav, D.; Koppelhus, E. B.; Sissons, R.; Vickaryous, M. K. (2011).
2692:
2657:
2573:
2489:
2434:
2209:
2008:
1862:
1783:
1607:
441:
4315:
4274:
4205:
4192:
4163:
4148:
4118:
4079:
4065:
3982:
3943:
3822:
3732:
3697:
3690:
3570:
3520:
3443:
3423:
2846:
2811:
2453:
2414:
2371:
2344:
2134:
2107:
1791:
1453:
1432:
1328:
1185:
1169:
713:
697:
527:
225:
2749:
Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre
2723:
Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre
1368:
is reddish-brown, poorly cemented, fine grained sandstone with some conglomerate, and
616:
476:). In 1975, two of the expedition's leading scientists, namely Polish paleontologists
4587:
4393:
4372:
4342:
4333:
4233:
4127:
4072:
4051:
4023:
4016:
4002:
3995:
3801:
3790:
3783:
3725:
3718:
3704:
3662:
3597:
3563:
3534:
3487:
3119:
3095:
3045:
2974:
2949:
2898:
2812:"A monodactyl nonavian dinosaur and the complex evolution of the alvarezsauroid hand"
2796:
2641:
2626:
2596:
2581:
2395:
2329:
2273:
2193:
2178:
1973:
Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages
1945:
1930:
1878:
1846:
1659:
1402:
1269:
1264:
1224:
1026:
902:
In 2019 Czepiński analyzed a vast majority of referred specimens to the ceratopsians
892:
826:
516:
425:
77:
1463:. Other paleofauna from this unit comprises a variety of squamates and mammals, and
910:, and concluded that most were in fact specimens of the former. Although the genera
433:
lacked premaxillary teeth (cylindrical, blunt teeth near the tip of the upper jaw).
4407:
4358:
4351:
4267:
4240:
4226:
4134:
4107:
4100:
4093:
4044:
4030:
3922:
3909:
3840:
3829:
3814:
3768:
3750:
3655:
3621:
3604:
3556:
3549:
3505:
3310:
3258:
1459:
1447:
1426:
1408:
1306:
1203:
798:, as it features a combination of basal (primitive) and derived (advanced) traits.
685:
296:
257:
209:
20:
3037:
2536:"Multituberculate succession in the Late Cretaceous of the Gobi Desert (Mongolia)"
2217:
2170:
1922:
2497:
2443:
2016:
1615:
4524:
4400:
4294:
4177:
4141:
3964:
3676:
3541:
2289:"A New Dromaeosaurid (Dinosauria: Coelurosauria) from Khulsan, Central Mongolia"
1651:
1420:
1318:
1302:
1297:
1234:
1228:
806:
642:
461:
420:
52:
3159:
3134:
2816:
Proceedings of the National Academy of Sciences of the United States of America
2124:
4170:
4086:
3936:
3669:
3457:
3391:
3378:
3337:
3216:
2933:
1870:
1575:
1477:
1443:
1438:
1414:
1273:
1249:
1219:
1199:
1177:
884:
843:
709:
597:
241:
97:
62:
4515:
2321:
2265:
1148:(yellowish, grey-brown, and rarely reddish) that are locally cemented. Sandy
2836:
1905:
Czepiński, Ł. (2019). "Ontogeny and variation of a protoceratopsid dinosaur
1365:
1361:
1323:
1312:
1288:
1279:
1243:
1207:
1165:
1153:
1149:
1133:
723:
485:
160:
102:
46:
2855:
2462:
2396:"A taxonomic review of the Pachycephalosauridae (Dinosauria: Ornithischia)"
2380:
2233:"A New Dromaeosaurid from the Late Cretaceous Khulsan Locality of Mongolia"
2143:
1691:
1674:
2787:
872:
4509:
3971:
3957:
3950:
3513:
3249:
3240:
3111:
2419:
from the Baruungoyot and Nemegt Formations (Late Cretaceous) of Mongolia"
1397:
1373:
1337:
1255:
1141:
1125:
866:
673:
578:
493:
378:
193:
180:
92:
87:
72:
67:
57:
2890:
2768:"Hands, feet and behaviour in Pinacosaurus (Dinosauria: Ankylosauridae)"
899:
during the Cretaceous Period, which ended roughly 66 million years ago.
4557:
4365:
3527:
3471:
2361:
1369:
1213:
847:
656:
In 2003 Russian paleontologist Vladimir R. Alifanov named the new taxa
107:
82:
2092:
753:
A comprehensive study on the intraspecific variation in morphology of
3234:
2617:
2312:
2256:
1830:
1467:
1377:
1333:
888:
473:
445:
170:
4486:
3080:
2700:
2665:
1710:[A new Late Cretaceous protoceratopsid genus from Mongolia]
1591:
592:
from the Udyn Sayr and Zamyn Khond localities, respectively, of the
134:
Skeletal reconstruction (including elements previously described as
2714:
Lambert, O.; Godefroit, P.; Li, H.; Shang, C.-Y.; Dong, Z. (2001).
2303:
2247:
2740:
Godefroit, P.; Pereda-Suberbiola, X.; Li, H.; Dong, Z. M. (1999).
2413:
Fanti, F.; Currie, P. J.; Badamgarav, D.; Lalueza-Fox, C. (2012).
1173:
871:
861:
805:
790:
to be a distinct and separate genus of protoceratopsid, from both
722:
701:
672:(PIN 3142/4; an almost complete medium-sized skull), found in the
615:
560:
specimen MPC-D 100/551B, which belongs to an indeterminate species
552:
540:
440:
371:
688:
described and named the new genus and species of protoceratopsid
464:. This newly collected and rich fossil material is stored in the
419:
had a fairly primitive anatomy—when compared to the much derived
4544:
1389:
1385:
1381:
1239:
1145:
896:
382:
4490:
3907:
3376:
3227:
3189:
1590:
Saneyoshi, M.; Watabe, M.; Suzuki, S.; Tsogtbaatar, K. (2011).
1140:
is known. This formation is mostly characterized by series of
708:, IVPP V12513, represents a nearly complete skull lacking the
3185:
504:, including juvenile and sub-adult skulls. The generic name,
2609:
10.1206/0003-0090(2000)249<0001:TCASOL>2.0.CO;2
1136:) and has virtually yielded the bulk of material for which
813:
specimens compared to a 1.8 m (5.9 ft) tall human
757:
was conducted by Polish paleontologist Łukasz Czepiński in
645:
in 2000 explained this by extrapolating that the juvenile
448:
localities of Mongolia and the location of Bayan Mandahu;
2883:
10.1671/0272-4634(2008)28[432:ANSOVD]2.0.CO;2
2400:
New Mexico Museum of Natural History and Science Bulletin
2056:
Gradziński, R.; Jaworowska, Z. K.; Maryańska, T. (1977).
1822:
10.1206/0003-0082(2006)3530[1:YDANPC]2.0.CO;2
1172:(flat land consisting of sediments deposited by highland
742:, it had already been displayed for several years at the
1360:), is considered to be Late Cretaceous in age, roughly
16:
Protoceratopsid dinosaur genus from the Late Cretaceous
2513:"A new sauropod from the Upper Cretaceous of Mongolia"
1708:"Новый род протоцератопсид из позднего мела Монголии"
2231:
Turner, A. H.; Montanari, S.; Norell, M. A. (2021).
777:
represent additional specimens and growth stages of
692:from red beds at the Bayan Mandahu locality of the
637:
by Kurzanov in 1990, were considered to be juvenile
508:, means "small-horned face" and is derived from the
303:
4499:
4383:
4341:
4332:
4284:
4250:
4191:
4117:
3981:
3920:
3839:
3800:
3758:
3749:
3645:
3619:
3580:
3485:
3433:
3389:
3336:
3309:
3282:
401:. One specimen may argue the possible presence of
2948:Longrich, N. R.; Currie, P. J.; Dong, Z. (2010).
2478:Palaeogeography, Palaeoclimatology, Palaeoecology
1997:Palaeogeography, Palaeoclimatology, Palaeoecology
1596:Palaeogeography, Palaeoclimatology, Palaeoecology
727:Skeletal diagram featuring the known elements of
1944:Dodson, P. (1993). "Dinosaurs Rule The World".
1740:. Cambridge University Press. pp. 488–489.
761:, where he concluded that the previously named
1990:
1988:
1909:from the Late Cretaceous of the Gobi Desert".
1563:The Journal of the Geological Society of Japan
584:In 2020, Czepiński described new specimens of
3201:
3094:Wible, J. R.; Shelley, S. L.; Bi, S. (2019).
627:Juvenile remains, at first tentatively named
8:
2032:"Sedimentation of the Barun Goyot Formation"
1950:. Publications International. p. 132.
1738:The Age of Dinosaurs in Russia and Mongolia
1388:on an alluvial plain with a combination of
4487:
4338:
4197:
3987:
3928:
3917:
3904:
3806:
3755:
3627:
3497:
3493:
3401:
3397:
3386:
3373:
3315:
3288:
3279:
3224:
3208:
3194:
3186:
1900:
1898:
1896:
1894:
1892:
1890:
1888:
1640:Journal of the Geological Society of Korea
1517:
1515:
1513:
1511:
1400:, such as closely related protoceratopsid
125:
27:
3158:
2973:
2932:
2845:
2835:
2786:
2616:
2511:Kurzanov, S. M.; Bannikov, A. F. (1983).
2452:
2442:
2370:
2360:
2311:
2255:
2133:
2123:
2081:Zoological Journal of the Linnean Society
2030:Gradziński, R.; Jerzykiewicz, T. (1974).
1829:
1690:
1629:
1627:
1625:
1574:
1805:Makovicky, P. J.; Norell, M. A. (2006).
1524:"Protoceratopsidae (Dinosauria) of Asia"
887:, a group of herbivorous dinosaurs with
522:= meaning horn face. The specific name,
1772:Acta Geologica Sinica (English Edition)
1507:
946:is the basal-most protoceratopsid and
531:Anatoly Konstantinovich Rozhdestvensky
1634:Kim, B.; Yun, H.; Lee, Y.-N. (2019).
1557:Matsumoto, Y.; Saneyoshi, M. (2010).
1396:This formation has produced numerous
533:for his notorious work on dinosaurs.
389:, around 72 to 71 million years ago.
7:
1522:Maryańska, T.; Osmólska, H. (1975).
1322:. Other taxa are represented by the
1184:paleoenvironments, under relatively
393:remains have been reported from the
2159:Journal of Systematic Palaeontology
1042:
1018:
994:
970:
963:
953:
526:, is named in honor of the Russian
370:(meaning "small-horned face") is a
3061:Canadian Journal of Earth Sciences
2871:Journal of Vertebrate Paleontology
2681:Canadian Journal of Earth Sciences
2646:Canadian Journal of Earth Sciences
1792:10.1111/j.1755-6724.2003.tb00745.x
14:
4594:Late Cretaceous dinosaurs of Asia
2595:Keqin, G.; Norell, M. A. (2000).
1971:Holtz, T. R.; Rey, L. V. (2007).
876:Skull cast of specimen PIN 3142/1
4471:
4470:
4446:
4440:
3890:
3884:
2975:10.1111/j.1475-4983.2010.00968.x
1496:Timeline of ceratopsian research
1480:
147:
569:skeleton (specimen KID 196) of
4614:Taxa named by Teresa Maryańska
4609:Taxa named by Halszka Osmólska
2534:Kielan-Jaworowska, Z. (1974).
1978:Supplementary Information 2012
850:—otherwise known as accessory
313:Maryanska & Osmolska, 1975
290:Maryanska & Osmolska, 1975
1:
4604:Fossil taxa described in 1975
3038:10.1016/j.cretres.2021.105052
2775:Acta Palaeontologica Polonica
2218:10.1016/j.cretres.2009.09.001
2171:10.1080/14772019.2021.2013965
1923:10.1080/08912963.2019.1593404
1679:Acta Palaeontologica Polonica
1003:Protoceratops hellenikorhinus
891:-like beaks which thrived in
744:Moscow Paleontological Museum
4531:Bagaceratops rozhdestvenskyi
4501:Bagaceratops rozhdestvenskyi
2582:10.1016/0037-0738(74)90021-9
2498:10.1016/j.palaeo.2017.10.023
2444:10.1371/journal.pone.0031330
2343:Cau, A.; Madzia, D. (2018).
2017:10.1016/j.palaeo.2017.11.018
1907:Bagaceratops rozhdestvenskyi
1616:10.1016/j.palaeo.2011.07.024
748:Bagaceratops rozhdestvenskyi
606:Bagaceratops rozhdestvenskyi
490:Bagaceratops rozhdestvenskyi
308:Bagaceratops rozhdestvenskyi
1652:10.14770/jgsk.2019.55.2.179
4635:
3160:10.1016/j.crpv.2014.12.005
2125:10.1038/s41598-021-83568-4
1559:"Bored dinosaur skeletons"
1332:, and a wide diversity of
470:Polish Academy of Sciences
18:
4460:
4438:
4200:
3990:
3931:
3916:
3903:
3881:
3809:
3630:
3500:
3496:
3404:
3400:
3385:
3372:
3318:
3291:
3278:
3223:
3100:Annals of Carnegie Museum
2934:10.11646/zootaxa.2403.1.1
2601:American Museum Novitates
2296:American Museum Novitates
2240:American Museum Novitates
1871:10.1134/S0031030108060087
1814:American Museum Novitates
1576:10.5575/geosoc.116.1.I_II
1380:-deposited) and aeolian (
1160:, and large-scale trough
1047:
1040:
1023:
1016:
999:
992:
975:
968:
961:
767:Lamaceratops tereschenkoi
658:Lamaceratops tereschenkoi
649:would grow into a mature
466:Institute of Paleobiology
337:Lamaceratops tereschenkoi
325:
318:
302:
295:
144:Scientific classification
142:
133:
124:
30:
2394:Sullivan, R. M. (2006).
1845:Alifanov, V. R. (2008).
1718:(in Russian) (4): 91−97.
1706:Kurzanov, S. M. (1990).
1144:, mostly light-coloured
771:Platyceratops tatarinovi
662:Platyceratops tatarinovi
634:Breviceratops kozlowskii
629:Protoceratops kozlowskii
353:Platyceratops tatarinovi
19:Not to be confused with
3417:Micropachycephalosaurus
2837:10.1073/pnas.1011052108
2543:Palaeontologia Polonica
2517:Paleontological Journal
2062:Acta Geologica Polonica
2039:Palaeontologia Polonica
1851:Paleontological Journal
1753:Paleontological Journal
1716:Paleontological Journal
1531:Palaeontologia Polonica
1372:. Other facies include
1350:Bayan Mandahu Formation
1344:Bayan Mandahu Formation
694:Bayan Mandahu Formation
399:Bayan Mandahu Formation
3139:Comptes Rendus Palevol
2998:Vertebrata PalAsiatica
1729:Sereno, P. C. (2000).
1692:10.4202/app.00701.2019
1673:Czepiński, Ł. (2020).
1152:(often red-coloured),
979:Protoceratops andrewsi
877:
869:
814:
731:
624:
602:Protoceratops andrewsi
561:
550:
453:
4566:Paleobiology Database
2788:10.4202/app.2010.0055
1132:in age (Middle-Upper
1122:Barun Goyot Formation
1116:Barun Goyot Formation
875:
865:
809:
726:
619:
556:
544:
515:= meaning small; and
458:Barun Goyot Formation
444:
395:Barun Goyot Formation
4619:Ornithischian genera
3112:10.2992/007.085.0401
1947:The Age of Dinosaurs
1162:cross-stratification
775:Magnirostris dodsoni
763:Gobiceratops minutus
736:Gobiceratops minitus
690:Magnirostris dodsoni
680:. Also during 2003,
537:Additional specimens
484:, published a large
437:History of discovery
345:Magnirostris dodsoni
329:Gobiceratops minutus
3582:Archaeoceratopsidae
3151:2015CRPal..14..577B
3073:1996CaJES..33..578G
3030:2022CrRes.13005052P
3018:Cretaceous Research
2966:2010Palgy..53..945L
2828:2011PNAS..108.2338X
2693:1993CaJES..30.2196E
2658:1993CaJES..30.2180J
2574:1974SedG...12..249G
2562:Sedimentary Geology
2490:2018PPP...494..101F
2435:2012PLoSO...731330F
2210:2010CrRes..31...16B
2198:Cretaceous Research
2009:2018PPP...494...29E
1863:2008PalJ...42..621A
1784:2003AcGlS..77..299Y
1608:2011PPP...311...38S
852:antorbital fenestra
704:). The holotype of
631:, and then renamed
594:Djadochta Formation
407:Djadochta Formation
348:You & Dong 2003
4302:Crittendenceratops
4252:Albertaceratopsini
3024:(105052): 105052.
2415:"New specimens of
2362:10.7717/peerj.4868
2112:Scientific Reports
1981:Weight Information
1911:Historical Biology
1294:pachycephalosaurid
878:
870:
815:
779:B. rozhdestvenskyi
755:B. rozhdestvenskyi
732:
625:
620:Holotype skull of
562:
551:
524:B. rozhdestvenskyi
454:
4581:
4580:
4493:Taxon identifiers
4484:
4483:
4456:
4455:
4436:
4435:
4432:
4431:
4428:
4427:
4424:
4423:
4385:Pachyrhinosaurini
4323:Yehuecauhceratops
4286:Nasutoceratopsini
4187:
4186:
3899:
3898:
3879:
3878:
3875:
3874:
3871:
3870:
3867:
3866:
3863:
3862:
3859:
3858:
3760:Protoceratopsidae
3647:Leptoceratopsidae
3615:
3614:
3481:
3480:
3368:
3367:
3364:
3363:
3360:
3359:
3356:
3355:
2755:(supp. B): 17–36.
2687:(10): 2196−2213.
2652:(10): 2180−2195.
2165:(16): 1097–1128.
2093:10.1111/zoj.12185
1957:978-0-7853-0443-2
1917:(10): 1394–1421.
1465:nanhsiungchelyids
1418:; dromaeosaurids
1190:semiarid climates
1128:, is regarded as
1107:
1106:
1098:
1097:
1089:
1088:
1080:
1079:
1071:
1070:
1062:
1061:
958:Protoceratopsidae
842:lacked primitive
363:
362:
357:
349:
341:
333:
291:
271:Protoceratopsidae
4626:
4574:
4573:
4561:
4560:
4548:
4547:
4535:
4534:
4533:
4520:
4519:
4518:
4488:
4474:
4473:
4451:
4450:
4444:
4415:Pachyrhinosaurus
4339:
4213:Machairoceratops
4198:
4038:Coahuilaceratops
3988:
3929:
3918:
3905:
3894:
3889:
3888:
3807:
3756:
3740:Zhuchengceratops
3628:
3498:
3494:
3435:Chaoyangsauridae
3402:
3398:
3387:
3374:
3316:
3289:
3280:
3273:
3272:
3225:
3210:
3203:
3196:
3187:
3165:
3164:
3162:
3145:(6–7): 577−587.
3130:
3124:
3123:
3091:
3085:
3084:
3056:
3050:
3049:
3012:
3006:
3005:
2995:
2986:
2980:
2979:
2977:
2945:
2939:
2938:
2936:
2918:
2909:
2903:
2902:
2866:
2860:
2859:
2849:
2839:
2822:(6): 2338–2342.
2807:
2801:
2800:
2790:
2772:
2763:
2757:
2756:
2746:
2737:
2731:
2730:
2720:
2711:
2705:
2704:
2676:
2670:
2669:
2637:
2631:
2630:
2620:
2592:
2586:
2585:
2557:
2551:
2550:
2540:
2531:
2525:
2524:
2508:
2502:
2501:
2473:
2467:
2466:
2456:
2446:
2410:
2404:
2403:
2391:
2385:
2384:
2374:
2364:
2340:
2334:
2333:
2315:
2293:
2284:
2278:
2277:
2259:
2237:
2228:
2222:
2221:
2189:
2183:
2182:
2154:
2148:
2147:
2137:
2127:
2103:
2097:
2096:
2076:
2070:
2069:
2053:
2047:
2046:
2036:
2027:
2021:
2020:
1992:
1983:
1976:
1968:
1962:
1961:
1941:
1935:
1934:
1902:
1883:
1882:
1842:
1836:
1835:
1833:
1811:
1802:
1796:
1795:
1767:
1761:
1760:
1748:
1742:
1741:
1735:
1726:
1720:
1719:
1713:
1703:
1697:
1696:
1694:
1670:
1664:
1663:
1631:
1620:
1619:
1587:
1581:
1580:
1578:
1554:
1548:
1545:
1539:
1538:
1528:
1519:
1490:
1488:Dinosaurs portal
1485:
1484:
1483:
1412:; alvarezsaurid
1356:(now synonym of
1352:, which yielded
1285:halszkaraptorine
1202:, including the
1111:Paleoenvironment
1043:
1019:
995:
971:
964:
954:
867:Life restoration
830:. Although both
600:transition from
482:Halszka Osmólska
478:Teresa Maryańska
355:
347:
339:
331:
305:
289:
282:
269:
256:
240:
224:
208:
152:
151:
129:
119:
49:
36:Temporal range:
28:
4634:
4633:
4629:
4628:
4627:
4625:
4624:
4623:
4584:
4583:
4582:
4577:
4569:
4564:
4556:
4551:
4543:
4538:
4529:
4528:
4523:
4514:
4513:
4508:
4495:
4485:
4480:
4452:
4445:
4420:
4379:
4328:
4309:Furcatoceratops
4280:
4261:Albertaceratops
4246:
4220:Menefeeceratops
4183:
4156:Tatankaceratops
4113:
4059:Mercuriceratops
4010:Arrhinoceratops
3977:
3925:
3912:
3895:
3883:
3855:
3835:
3796:
3745:
3712:Montanoceratops
3641:
3636:Graciliceratops
3624:
3611:
3591:Archaeoceratops
3576:
3490:
3477:
3465:Xuanhuaceratops
3451:Hualianceratops
3429:
3410:Albalophosaurus
3394:
3381:
3352:
3332:
3305:
3299:Avemetatarsalia
3284:Avemetatarsalia
3274:
3268:Marginocephalia
3230:
3229:
3219:
3214:
3174:
3169:
3168:
3132:
3131:
3127:
3093:
3092:
3088:
3081:10.1139/e96-043
3058:
3057:
3053:
3014:
3013:
3009:
2993:
2988:
2987:
2983:
2947:
2946:
2942:
2916:
2911:
2910:
2906:
2868:
2867:
2863:
2809:
2808:
2804:
2770:
2765:
2764:
2760:
2744:
2739:
2738:
2734:
2718:
2713:
2712:
2708:
2701:10.1139/e93-191
2678:
2677:
2673:
2666:10.1139/e93-190
2639:
2638:
2634:
2594:
2593:
2589:
2559:
2558:
2554:
2538:
2533:
2532:
2528:
2510:
2509:
2505:
2475:
2474:
2470:
2412:
2411:
2407:
2393:
2392:
2388:
2342:
2341:
2337:
2291:
2286:
2285:
2281:
2235:
2230:
2229:
2225:
2191:
2190:
2186:
2156:
2155:
2151:
2105:
2104:
2100:
2078:
2077:
2073:
2055:
2054:
2050:
2034:
2029:
2028:
2024:
1994:
1993:
1986:
1975:. Random House.
1970:
1969:
1965:
1958:
1943:
1942:
1938:
1904:
1903:
1886:
1844:
1843:
1839:
1809:
1804:
1803:
1799:
1769:
1768:
1764:
1750:
1749:
1745:
1733:
1728:
1727:
1723:
1711:
1705:
1704:
1700:
1672:
1671:
1667:
1633:
1632:
1623:
1589:
1588:
1584:
1556:
1555:
1551:
1546:
1542:
1526:
1521:
1520:
1509:
1504:
1486:
1481:
1479:
1476:
1430:; oviraptorids
1406:; ankylosaurid
1364:. The dominant
1346:
1261:protoceratopsid
1130:Late Cretaceous
1118:
1113:
1108:
1099:
1090:
1081:
1072:
1063:
860:
804:
746:under the name
714:junior synonyms
614:
539:
492:. The selected
439:
387:Late Cretaceous
376:protoceratopsid
314:
311:
288:
280:
267:
254:
238:
222:
206:
146:
120:
118:
117:
116:
115:
110:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
44:
43:
41:
38:Late Cretaceous
34:
24:
17:
12:
11:
5:
4632:
4630:
4622:
4621:
4616:
4611:
4606:
4601:
4596:
4586:
4585:
4579:
4578:
4576:
4575:
4562:
4549:
4536:
4521:
4505:
4503:
4497:
4496:
4491:
4482:
4481:
4479:
4478:
4468:
4461:
4458:
4457:
4454:
4453:
4439:
4437:
4434:
4433:
4430:
4429:
4426:
4425:
4422:
4421:
4419:
4418:
4411:
4404:
4397:
4389:
4387:
4381:
4380:
4378:
4377:
4369:
4362:
4355:
4347:
4345:
4336:
4330:
4329:
4327:
4326:
4319:
4316:Nasutoceratops
4312:
4305:
4298:
4290:
4288:
4282:
4281:
4279:
4278:
4275:Medusaceratops
4271:
4264:
4256:
4254:
4248:
4247:
4245:
4244:
4237:
4230:
4223:
4216:
4209:
4206:Diabloceratops
4201:
4195:
4193:Centrosaurinae
4189:
4188:
4185:
4184:
4182:
4181:
4174:
4167:
4164:Titanoceratops
4160:
4152:
4149:Regaliceratops
4145:
4138:
4131:
4123:
4121:
4119:Triceratopsini
4115:
4114:
4112:
4111:
4104:
4097:
4090:
4083:
4080:Sierraceratops
4076:
4069:
4066:Navajoceratops
4062:
4055:
4048:
4041:
4034:
4027:
4020:
4013:
4006:
3999:
3991:
3985:
3983:Chasmosaurinae
3979:
3978:
3976:
3975:
3968:
3961:
3954:
3947:
3944:Brachyceratops
3940:
3932:
3926:
3921:
3914:
3913:
3908:
3901:
3900:
3897:
3896:
3882:
3880:
3877:
3876:
3873:
3872:
3869:
3868:
3865:
3864:
3861:
3860:
3857:
3856:
3854:
3853:
3852:
3851:
3845:
3843:
3837:
3836:
3834:
3833:
3826:
3823:Turanoceratops
3819:
3810:
3804:
3798:
3797:
3795:
3794:
3787:
3780:
3773:
3764:
3762:
3753:
3747:
3746:
3744:
3743:
3736:
3733:Unescoceratops
3729:
3722:
3715:
3708:
3701:
3698:Ischioceratops
3694:
3691:Gryphoceratops
3687:
3680:
3673:
3666:
3659:
3651:
3649:
3643:
3642:
3640:
3639:
3631:
3625:
3620:
3617:
3616:
3613:
3612:
3610:
3609:
3601:
3594:
3586:
3584:
3578:
3577:
3575:
3574:
3571:Sasayamagnomus
3567:
3560:
3553:
3546:
3538:
3531:
3524:
3521:Auroraceratops
3517:
3510:
3501:
3491:
3486:
3483:
3482:
3479:
3478:
3476:
3475:
3468:
3461:
3454:
3447:
3444:Chaoyangsaurus
3439:
3437:
3431:
3430:
3428:
3427:
3424:Psittacosaurus
3420:
3413:
3405:
3395:
3390:
3383:
3382:
3377:
3370:
3369:
3366:
3365:
3362:
3361:
3358:
3357:
3354:
3353:
3351:
3350:
3349:
3348:
3342:
3340:
3334:
3333:
3331:
3330:
3329:
3328:
3319:
3313:
3307:
3306:
3304:
3303:
3302:
3301:
3292:
3286:
3276:
3275:
3271:
3270:
3261:
3252:
3243:
3237:
3228:
3221:
3220:
3215:
3213:
3212:
3205:
3198:
3190:
3184:
3183:
3173:
3172:External links
3170:
3167:
3166:
3125:
3106:(4): 285–327.
3086:
3067:(4): 578−598.
3051:
3007:
2981:
2960:(5): 945−960.
2940:
2904:
2877:(2): 432–438.
2861:
2802:
2781:(3): 489–504.
2758:
2732:
2706:
2671:
2632:
2603:(249): 1−118.
2587:
2568:(4): 249–278.
2552:
2526:
2503:
2468:
2405:
2402:(35): 347–365.
2386:
2335:
2304:10.1206/3982.1
2298:(3982): 1–47.
2279:
2248:10.1206/3965.1
2242:(3965): 1–48.
2223:
2184:
2149:
2118:(4101): 4101.
2098:
2087:(3): 631−652.
2071:
2048:
2022:
1984:
1963:
1956:
1936:
1884:
1857:(6): 621–633.
1837:
1816:(3530): 1–42.
1797:
1778:(3): 299−303.
1762:
1743:
1721:
1698:
1685:(3): 481−497.
1665:
1646:(2): 179−190.
1621:
1602:(1–2): 38−47.
1582:
1549:
1540:
1506:
1505:
1503:
1500:
1499:
1498:
1492:
1491:
1475:
1472:
1454:Papiliovenator
1433:Machairasaurus
1345:
1342:
1329:Quaesitosaurus
1270:dromaeosaurids
1225:alvarezsaurids
1170:alluvial plain
1117:
1114:
1112:
1109:
1105:
1104:
1101:
1100:
1096:
1095:
1092:
1091:
1087:
1086:
1083:
1082:
1078:
1077:
1074:
1073:
1069:
1068:
1065:
1064:
1060:
1059:
1056:
1055:
1046:
1041:
1039:
1036:
1035:
1032:
1031:
1022:
1017:
1015:
1012:
1011:
1008:
1007:
998:
993:
991:
988:
987:
984:
983:
974:
969:
967:
962:
960:
952:
859:
858:Classification
856:
803:
800:
698:Inner Mongolia
613:
610:
538:
535:
528:paleontologist
498:ZPAL MgD-I/126
438:
435:
381:that lived in
361:
360:
359:
358:
356:Alifanov, 2003
350:
342:
340:Alifanov, 2003
334:
332:Alifanov, 2008
323:
322:
316:
315:
312:
300:
299:
293:
292:
278:
274:
273:
265:
261:
260:
252:
245:
244:
236:
229:
228:
226:Neornithischia
220:
213:
212:
204:
197:
196:
191:
184:
183:
178:
174:
173:
168:
164:
163:
158:
154:
153:
140:
139:
131:
130:
122:
121:
113:
112:
111:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
50:
35:
15:
13:
10:
9:
6:
4:
3:
2:
4631:
4620:
4617:
4615:
4612:
4610:
4607:
4605:
4602:
4600:
4597:
4595:
4592:
4591:
4589:
4572:
4567:
4563:
4559:
4554:
4550:
4546:
4541:
4537:
4532:
4526:
4522:
4517:
4511:
4507:
4506:
4504:
4502:
4498:
4494:
4489:
4477:
4469:
4467:
4463:
4462:
4459:
4449:
4443:
4417:
4416:
4412:
4410:
4409:
4405:
4403:
4402:
4398:
4396:
4395:
4394:Achelousaurus
4391:
4390:
4388:
4386:
4382:
4375:
4374:
4373:Styracosaurus
4370:
4368:
4367:
4363:
4361:
4360:
4356:
4354:
4353:
4349:
4348:
4346:
4344:
4343:Centrosaurini
4340:
4337:
4335:
4334:Eucentrosaura
4331:
4325:
4324:
4320:
4318:
4317:
4313:
4311:
4310:
4306:
4304:
4303:
4299:
4297:
4296:
4292:
4291:
4289:
4287:
4283:
4277:
4276:
4272:
4270:
4269:
4265:
4263:
4262:
4258:
4257:
4255:
4253:
4249:
4243:
4242:
4238:
4236:
4235:
4234:Wendiceratops
4231:
4229:
4228:
4224:
4222:
4221:
4217:
4215:
4214:
4210:
4208:
4207:
4203:
4202:
4199:
4196:
4194:
4190:
4180:
4179:
4175:
4173:
4172:
4168:
4166:
4165:
4161:
4158:
4157:
4153:
4151:
4150:
4146:
4144:
4143:
4139:
4137:
4136:
4132:
4130:
4129:
4128:Eotriceratops
4125:
4124:
4122:
4120:
4116:
4110:
4109:
4105:
4103:
4102:
4098:
4096:
4095:
4091:
4089:
4088:
4084:
4082:
4081:
4077:
4075:
4074:
4073:Pentaceratops
4070:
4068:
4067:
4063:
4061:
4060:
4056:
4054:
4053:
4052:Kosmoceratops
4049:
4047:
4046:
4042:
4040:
4039:
4035:
4033:
4032:
4028:
4026:
4025:
4024:Bravoceratops
4021:
4019:
4018:
4017:Bisticeratops
4014:
4012:
4011:
4007:
4005:
4004:
4003:Anchiceratops
4000:
3998:
3997:
3996:Agujaceratops
3993:
3992:
3989:
3986:
3984:
3980:
3974:
3973:
3969:
3967:
3966:
3962:
3960:
3959:
3955:
3953:
3952:
3948:
3946:
3945:
3941:
3939:
3938:
3934:
3933:
3930:
3927:
3924:
3919:
3915:
3911:
3906:
3902:
3893:
3887:
3849:
3848:
3847:
3846:
3844:
3842:
3838:
3832:
3831:
3827:
3825:
3824:
3820:
3817:
3816:
3812:
3811:
3808:
3805:
3803:
3802:Ceratopsoidea
3799:
3793:
3792:
3791:Protoceratops
3788:
3786:
3785:
3784:Breviceratops
3781:
3779:
3778:
3774:
3771:
3770:
3766:
3765:
3763:
3761:
3757:
3754:
3752:
3748:
3742:
3741:
3737:
3735:
3734:
3730:
3728:
3727:
3726:Udanoceratops
3723:
3721:
3720:
3719:Prenoceratops
3716:
3714:
3713:
3709:
3707:
3706:
3705:Leptoceratops
3702:
3700:
3699:
3695:
3693:
3692:
3688:
3686:
3685:
3681:
3679:
3678:
3674:
3672:
3671:
3667:
3665:
3664:
3663:Bainoceratops
3660:
3658:
3657:
3653:
3652:
3650:
3648:
3644:
3638:
3637:
3633:
3632:
3629:
3626:
3623:
3618:
3607:
3606:
3602:
3600:
3599:
3598:Helioceratops
3595:
3593:
3592:
3588:
3587:
3585:
3583:
3579:
3573:
3572:
3568:
3566:
3565:
3564:Mosaiceratops
3561:
3559:
3558:
3554:
3552:
3551:
3547:
3544:
3543:
3539:
3537:
3536:
3535:Koreaceratops
3532:
3530:
3529:
3525:
3523:
3522:
3518:
3516:
3515:
3511:
3508:
3507:
3503:
3502:
3499:
3495:
3492:
3489:
3488:Neoceratopsia
3484:
3474:
3473:
3469:
3467:
3466:
3462:
3460:
3459:
3455:
3453:
3452:
3448:
3446:
3445:
3441:
3440:
3438:
3436:
3432:
3426:
3425:
3421:
3419:
3418:
3414:
3412:
3411:
3407:
3406:
3403:
3399:
3396:
3393:
3388:
3384:
3380:
3375:
3371:
3346:
3345:
3344:
3343:
3341:
3339:
3335:
3327:
3323:
3322:
3321:
3320:
3317:
3314:
3312:
3308:
3300:
3296:
3295:
3294:
3293:
3290:
3287:
3285:
3281:
3277:
3269:
3265:
3262:
3260:
3256:
3253:
3251:
3247:
3244:
3242:
3238:
3236:
3232:
3231:
3226:
3222:
3218:
3211:
3206:
3204:
3199:
3197:
3192:
3191:
3188:
3182:
3181:
3176:
3175:
3171:
3161:
3156:
3152:
3148:
3144:
3140:
3136:
3129:
3126:
3121:
3117:
3113:
3109:
3105:
3101:
3097:
3090:
3087:
3082:
3078:
3074:
3070:
3066:
3062:
3055:
3052:
3047:
3043:
3039:
3035:
3031:
3027:
3023:
3019:
3011:
3008:
3003:
2999:
2992:
2985:
2982:
2976:
2971:
2967:
2963:
2959:
2955:
2954:Palaeontology
2951:
2944:
2941:
2935:
2930:
2926:
2922:
2915:
2908:
2905:
2900:
2896:
2892:
2888:
2884:
2880:
2876:
2872:
2865:
2862:
2857:
2853:
2848:
2843:
2838:
2833:
2829:
2825:
2821:
2817:
2813:
2806:
2803:
2798:
2794:
2789:
2784:
2780:
2776:
2769:
2762:
2759:
2754:
2750:
2743:
2736:
2733:
2728:
2724:
2717:
2710:
2707:
2702:
2698:
2694:
2690:
2686:
2682:
2675:
2672:
2667:
2663:
2659:
2655:
2651:
2647:
2643:
2636:
2633:
2628:
2624:
2619:
2614:
2610:
2606:
2602:
2598:
2591:
2588:
2583:
2579:
2575:
2571:
2567:
2563:
2556:
2553:
2548:
2544:
2537:
2530:
2527:
2522:
2518:
2514:
2507:
2504:
2499:
2495:
2491:
2487:
2483:
2479:
2472:
2469:
2464:
2460:
2455:
2450:
2445:
2440:
2436:
2432:
2429:(2): e31330.
2428:
2424:
2420:
2418:
2409:
2406:
2401:
2397:
2390:
2387:
2382:
2378:
2373:
2368:
2363:
2358:
2354:
2350:
2346:
2339:
2336:
2331:
2327:
2323:
2319:
2314:
2309:
2305:
2301:
2297:
2290:
2283:
2280:
2275:
2271:
2267:
2263:
2258:
2253:
2249:
2245:
2241:
2234:
2227:
2224:
2219:
2215:
2211:
2207:
2203:
2199:
2195:
2188:
2185:
2180:
2176:
2172:
2168:
2164:
2160:
2153:
2150:
2145:
2141:
2136:
2131:
2126:
2121:
2117:
2113:
2109:
2102:
2099:
2094:
2090:
2086:
2082:
2075:
2072:
2068:(3): 281–326.
2067:
2063:
2059:
2052:
2049:
2044:
2040:
2033:
2026:
2023:
2018:
2014:
2010:
2006:
2002:
1998:
1991:
1989:
1985:
1982:
1979:
1974:
1967:
1964:
1959:
1953:
1949:
1948:
1940:
1937:
1932:
1928:
1924:
1920:
1916:
1912:
1908:
1901:
1899:
1897:
1895:
1893:
1891:
1889:
1885:
1880:
1876:
1872:
1868:
1864:
1860:
1856:
1852:
1848:
1841:
1838:
1832:
1827:
1823:
1819:
1815:
1808:
1801:
1798:
1793:
1789:
1785:
1781:
1777:
1773:
1766:
1763:
1759:(5): 524–534.
1758:
1754:
1747:
1744:
1739:
1732:
1725:
1722:
1717:
1709:
1702:
1699:
1693:
1688:
1684:
1680:
1676:
1669:
1666:
1661:
1657:
1653:
1649:
1645:
1641:
1637:
1630:
1628:
1626:
1622:
1617:
1613:
1609:
1605:
1601:
1597:
1593:
1586:
1583:
1577:
1572:
1568:
1564:
1560:
1553:
1550:
1544:
1541:
1536:
1532:
1525:
1518:
1516:
1514:
1512:
1508:
1501:
1497:
1494:
1493:
1489:
1478:
1473:
1471:
1469:
1466:
1462:
1461:
1456:
1455:
1450:
1449:
1445:
1441:
1440:
1435:
1434:
1429:
1428:
1423:
1422:
1417:
1416:
1411:
1410:
1405:
1404:
1403:Protoceratops
1399:
1394:
1391:
1387:
1383:
1379:
1375:
1371:
1367:
1363:
1359:
1355:
1351:
1343:
1341:
1339:
1335:
1331:
1330:
1325:
1321:
1320:
1315:
1314:
1309:
1308:
1304:
1300:
1299:
1295:
1291:
1290:
1286:
1282:
1281:
1276:
1275:
1271:
1267:
1266:
1265:Breviceratops
1262:
1258:
1257:
1252:
1251:
1246:
1245:
1241:
1237:
1236:
1231:
1230:
1226:
1222:
1221:
1216:
1215:
1210:
1209:
1205:
1204:ankylosaurids
1201:
1197:
1193:
1191:
1187:
1183:
1179:
1175:
1171:
1167:
1163:
1159:
1158:conglomerates
1155:
1151:
1147:
1143:
1139:
1135:
1131:
1127:
1123:
1115:
1110:
1103:
1102:
1094:
1093:
1085:
1084:
1076:
1075:
1067:
1066:
1058:
1057:
1054:
1053:
1052:
1045:
1044:
1038:
1037:
1034:
1033:
1030:
1029:
1028:
1027:Breviceratops
1021:
1020:
1014:
1013:
1010:
1009:
1006:
1005:
1004:
997:
996:
990:
989:
986:
985:
982:
981:
980:
973:
972:
966:
965:
959:
956:
955:
951:
949:
945:
941:
940:Protoceratops
937:
936:Breviceratops
933:
929:
925:
924:Platyceratops
921:
917:
913:
909:
908:Breviceratops
905:
900:
898:
894:
893:North America
890:
886:
882:
874:
868:
864:
857:
855:
853:
849:
845:
841:
837:
836:Protoceratops
833:
829:
828:
827:Protoceratops
823:
819:
812:
808:
801:
799:
797:
796:Protoceratops
793:
789:
788:Breviceratops
785:
780:
776:
772:
768:
764:
760:
756:
751:
749:
745:
741:
737:
730:
725:
721:
719:
715:
711:
707:
703:
699:
695:
691:
687:
683:
679:
675:
671:
670:Platyceratops
667:
663:
659:
654:
652:
648:
647:Breviceratops
644:
640:
636:
635:
630:
623:
618:
611:
609:
607:
603:
599:
595:
591:
590:Protoceratops
587:
582:
580:
576:
575:Protoceratops
572:
567:
559:
555:
548:
543:
536:
534:
532:
529:
525:
521:
518:
514:
511:
507:
503:
499:
495:
491:
487:
483:
479:
475:
471:
467:
463:
459:
451:
447:
443:
436:
434:
432:
428:
427:
426:Protoceratops
422:
418:
414:
410:
408:
404:
400:
396:
392:
388:
384:
380:
377:
373:
369:
368:
354:
351:
346:
343:
338:
335:
330:
327:
326:
324:
321:
317:
310:
309:
301:
298:
294:
287:
286:
279:
276:
275:
272:
266:
263:
262:
259:
253:
250:
247:
246:
243:
237:
234:
231:
230:
227:
221:
218:
215:
214:
211:
205:
202:
199:
198:
195:
192:
189:
186:
185:
182:
179:
176:
175:
172:
169:
166:
165:
162:
159:
156:
155:
150:
145:
141:
137:
132:
128:
123:
109:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
48:
39:
33:
29:
26:
22:
4599:Ceratopsians
4500:
4413:
4408:Stellasaurus
4406:
4399:
4392:
4371:
4364:
4359:Coronosaurus
4357:
4352:Centrosaurus
4350:
4321:
4314:
4307:
4300:
4293:
4273:
4268:Lokiceratops
4266:
4259:
4241:Xenoceratops
4239:
4232:
4227:Sinoceratops
4225:
4218:
4211:
4204:
4176:
4169:
4162:
4154:
4147:
4140:
4135:Nedoceratops
4133:
4126:
4108:Vagaceratops
4106:
4101:Utahceratops
4099:
4094:Terminocavus
4092:
4085:
4078:
4071:
4064:
4057:
4050:
4045:Judiceratops
4043:
4036:
4031:Chasmosaurus
4029:
4022:
4015:
4008:
4001:
3994:
3970:
3963:
3956:
3949:
3942:
3935:
3923:Ceratopsidae
3910:Ceratopsidae
3841:Ceratopsidae
3830:Zuniceratops
3828:
3821:
3815:Ajkaceratops
3813:
3789:
3782:
3777:Bagaceratops
3776:
3775:
3769:Ajkaceratops
3767:
3751:Coronosauria
3738:
3731:
3724:
3717:
3710:
3703:
3696:
3689:
3682:
3675:
3668:
3661:
3656:Asiaceratops
3654:
3634:
3622:Euceratopsia
3605:Yamaceratops
3603:
3596:
3589:
3569:
3562:
3557:Microceratus
3555:
3550:Liaoceratops
3548:
3540:
3533:
3526:
3519:
3512:
3506:Ajkaceratops
3504:
3470:
3463:
3456:
3449:
3442:
3422:
3415:
3408:
3326:Ornithischia
3311:Ornithischia
3263:
3259:Ornithischia
3254:
3245:
3180:Bagaceratops
3179:
3142:
3138:
3128:
3103:
3099:
3089:
3064:
3060:
3054:
3021:
3017:
3010:
3004:(2): 85–101.
3001:
2997:
2984:
2957:
2953:
2943:
2924:
2920:
2907:
2874:
2870:
2864:
2819:
2815:
2805:
2778:
2774:
2761:
2752:
2748:
2735:
2726:
2722:
2709:
2684:
2680:
2674:
2649:
2645:
2635:
2600:
2590:
2565:
2561:
2555:
2546:
2542:
2529:
2520:
2516:
2506:
2481:
2477:
2471:
2426:
2422:
2416:
2408:
2399:
2389:
2352:
2348:
2338:
2295:
2282:
2239:
2226:
2204:(1): 16−26.
2201:
2197:
2187:
2162:
2158:
2152:
2115:
2111:
2101:
2084:
2080:
2074:
2065:
2061:
2051:
2042:
2038:
2025:
2000:
1996:
1972:
1966:
1946:
1939:
1914:
1910:
1906:
1854:
1850:
1840:
1813:
1800:
1775:
1771:
1765:
1756:
1752:
1746:
1737:
1724:
1715:
1701:
1682:
1678:
1668:
1643:
1639:
1599:
1595:
1585:
1566:
1562:
1552:
1543:
1534:
1530:
1460:Philovenator
1458:
1452:
1448:Linhevenator
1446:
1437:
1431:
1427:Velociraptor
1425:
1419:
1413:
1409:Pinacosaurus
1407:
1401:
1395:
1358:Bagaceratops
1357:
1354:Magnirostris
1353:
1347:
1327:
1317:
1311:
1307:Conchoraptor
1305:
1303:oviraptorids
1296:
1287:
1278:
1272:
1263:
1254:
1248:
1242:
1233:
1227:
1218:
1212:
1206:
1196:Bagaceratops
1195:
1194:
1138:Bagaceratops
1137:
1119:
1051:Bagaceratops
1050:
1049:
1048:
1025:
1024:
1001:
1000:
977:
976:
948:Bagaceratops
947:
943:
939:
935:
932:Bagaceratops
931:
928:Bagaceratops
927:
923:
920:Magnirostris
919:
916:Lamaceratops
915:
912:Gobiceratops
911:
907:
904:Bagaceratops
903:
901:
883:belonged to
881:Bagaceratops
880:
879:
844:premaxillary
840:Bagaceratops
839:
835:
832:Bagaceratops
831:
825:
821:
818:Bagaceratops
817:
816:
811:Bagaceratops
810:
795:
792:Bagaceratops
791:
787:
784:Bagaceratops
783:
778:
774:
770:
766:
762:
754:
752:
747:
740:Gobiceratops
739:
735:
733:
729:Gobiceratops
728:
718:Bagaceratops
717:
706:Magnirostris
705:
689:
686:Dong Zhiming
678:Bagaceratops
677:
669:
666:Lamaceratops
665:
661:
657:
655:
651:Bagaceratops
650:
646:
639:Bagaceratops
638:
632:
628:
626:
622:Magnirostris
621:
605:
601:
589:
586:Bagaceratops
585:
583:
574:
571:Bagaceratops
570:
566:Bagaceratops
565:
563:
558:Bagaceratops
557:
547:Bagaceratops
546:
523:
519:
512:
506:Bagaceratops
505:
502:Bagaceratops
501:
497:
489:
455:
450:Bagaceratops
449:
431:Bagaceratops
430:
424:
417:Bagaceratops
416:
413:Bagaceratops
412:
411:
403:Bagaceratops
402:
391:Bagaceratops
390:
367:Bagaceratops
366:
365:
364:
352:
344:
336:
328:
307:
306:
297:Type species
285:Bagaceratops
284:
283:
258:Coronosauria
248:
232:
216:
210:Ornithischia
200:
187:
136:Magnirostris
135:
32:Bagaceratops
31:
25:
21:Vagaceratops
4525:Wikispecies
4401:Einiosaurus
4295:Avaceratops
4178:Triceratops
4142:Ojoceratops
3965:Monoclonius
3677:Ferrisaurus
3542:Kulceratops
2484:: 101–120.
2417:Nemegtomaia
1569:(1): I–II.
1444:troodontids
1421:Linheraptor
1326:titanosaur
1319:Nemegtomaia
1298:Tylocephale
1235:Parvicursor
1229:Khulsanurus
1200:vertebrates
1124:, based on
944:P. andrewsi
822:Maginostris
802:Description
643:Paul Sereno
545:Diagram of
462:Gobi Desert
421:ceratopsids
385:during the
45:72–71
4588:Categories
4464:See also:
4171:Torosaurus
4087:Spiclypeus
3937:Agathaumas
3850:see below↓
3670:Cerasinops
3458:Stenopelix
3392:Ceratopsia
3379:Ceratopsia
3347:see below↓
3338:Ceratopsia
3250:Dinosauria
3217:Ceratopsia
2927:(1): 1–9.
2045:: 111−146.
1537:: 134−143.
1502:References
1439:Wulatelong
1415:Linhenykus
1250:Gobipteryx
1220:Zaraapelta
1178:lacustrine
1166:sandstones
1154:siltstones
1150:claystones
885:Ceratopsia
598:anagenetic
242:Ceratopsia
194:Dinosauria
3233:Kingdom:
3178:HVNHM on
3120:210840006
3046:244186762
2899:129414074
2797:129291148
2627:129367764
2618:2246/1596
2355:: e4868.
2330:243849373
2322:0003-0082
2313:2246/7286
2274:231597229
2266:0003-0082
2257:2246/7251
2179:247222017
2003:: 29–50.
1931:132780322
1879:129893753
1831:2246/5808
1660:150321203
1398:dinosaurs
1366:lithology
1362:Campanian
1338:squamates
1313:Heyuannia
1289:Hulsanpes
1259:; fellow
1244:Gobipipus
1208:Saichania
1134:Campanian
1126:sediments
682:You Hailu
579:clavicles
510:Mongolian
486:monograph
379:dinosaurs
374:of small
167:Kingdom:
161:Eukaryota
4510:Wikidata
4476:Category
4466:Timeline
3972:Polyonax
3958:Dysganus
3951:Ceratops
3514:Aquilops
3241:Chordata
3239:Phylum:
3235:Animalia
2891:20490961
2856:21262806
2549:: 23–44.
2523:: 90−96.
2463:22347465
2423:PLOS ONE
2381:29868277
2144:33737515
1474:See also
1374:alluvial
1256:Hollanda
1142:red beds
674:red beds
612:Synonyms
520:ceratops
494:holotype
320:Synonyms
264:Family:
181:Chordata
177:Phylum:
171:Animalia
157:Domain:
4558:4965778
4545:4531819
4516:Q131256
4366:Spinops
3684:Gremlin
3472:Yinlong
3147:Bibcode
3069:Bibcode
3026:Bibcode
2962:Bibcode
2921:Zootaxa
2847:3038769
2824:Bibcode
2729:: 5−28.
2689:Bibcode
2654:Bibcode
2570:Bibcode
2486:Bibcode
2454:3275628
2431:Bibcode
2372:5978397
2206:Bibcode
2135:7973727
2005:Bibcode
1859:Bibcode
1780:Bibcode
1604:Bibcode
1468:turtles
1370:caliche
1334:mammals
1214:Tarchia
1182:aeolian
848:maxilla
549:KID 196
468:of the
405:in the
277:Genus:
114:↓
3118:
3044:
2897:
2889:
2854:
2844:
2795:
2625:
2461:
2451:
2379:
2369:
2328:
2320:
2272:
2264:
2177:
2142:
2132:
1954:
1929:
1877:
1658:
1457:, and
1442:; and
1378:stream
1301:; and
1180:, and
1174:rivers
938:, and
922:, and
889:parrot
474:Poland
446:Fossil
4571:52826
3264:Clade
3255:Clade
3246:Clade
3116:S2CID
3042:S2CID
2994:(PDF)
2917:(PDF)
2895:S2CID
2887:JSTOR
2793:S2CID
2771:(PDF)
2745:(PDF)
2719:(PDF)
2623:S2CID
2539:(PDF)
2349:PeerJ
2326:S2CID
2292:(PDF)
2270:S2CID
2236:(PDF)
2175:S2CID
2035:(PDF)
1927:S2CID
1875:S2CID
1810:(PDF)
1734:(PDF)
1712:(PDF)
1656:S2CID
1527:(PDF)
1386:lakes
1324:large
1240:birds
1146:sands
710:frill
702:China
517:Greek
372:genus
249:Clade
233:Clade
217:Clade
201:Clade
188:Clade
4553:GBIF
3324:see
3297:see
2925:2403
2852:PMID
2459:PMID
2377:PMID
2318:ISSN
2262:ISSN
2140:PMID
1952:ISBN
1436:and
1424:and
1390:dune
1382:wind
1348:The
1336:and
1316:and
1280:Shri
1277:and
1274:Kuru
1253:and
1232:and
1217:and
1186:arid
1120:The
906:and
897:Asia
895:and
834:and
794:and
773:and
759:2019
684:and
660:and
588:and
513:baga
480:and
397:and
383:Asia
53:PreꞒ
4540:EoL
3528:Beg
3155:doi
3108:doi
3077:doi
3034:doi
3022:130
2970:doi
2929:doi
2879:doi
2842:PMC
2832:doi
2820:108
2783:doi
2697:doi
2662:doi
2613:hdl
2605:doi
2578:doi
2494:doi
2482:494
2449:PMC
2439:doi
2367:PMC
2357:doi
2308:hdl
2300:doi
2252:hdl
2244:doi
2214:doi
2167:doi
2130:PMC
2120:doi
2089:doi
2085:172
2013:doi
2001:494
1919:doi
1867:doi
1826:hdl
1818:doi
1788:doi
1687:doi
1648:doi
1612:doi
1600:311
1571:doi
1567:116
1188:to
1176:),
716:of
604:to
496:is
4590::
4568::
4555::
4542::
4527::
4512::
3266::
3257::
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3153:.
3143:14
3141:.
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3114:.
3104:85
3102:.
3098:.
3075:.
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3000:.
2996:.
2968:.
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2956:.
2952:.
2923:.
2919:.
2893:.
2885:.
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2830:.
2818:.
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2791:.
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2727:71
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251::
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47:Ma
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108:N
98:K
93:J
88:T
83:P
78:C
73:D
68:S
63:O
58:Ꞓ
42:~
40:,
23:.
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