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Type (biology)

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96: 33: 867:, isoneotype, isolectotype, etc.) but these are not formally regulated by the Code, and a great many are obsolete and/or idiosyncratic. However, some of these categories can potentially apply to genuine type specimens, such as a neotype; e.g., isotypic/topotypic specimens are preferred to other specimens, when they are available at the time a neotype is chosen (because they are from the same time and/or place as the original type). A topotype is a specimen that was obtained from the same location that the original type specimen came from. 534: 885: 826:, in which the species description included DNA sequences from blood and feather samples. Assuming there is no future question as to the status of such a species, the absence of a type specimen does not invalidate the name, but it may be necessary for the future to designate a neotype for such a taxon, should any questions arise. However, in the case of the bushshrike, ornithologists have argued that the specimen was a rare and hitherto unknown 431: 460:, the type of a species or subspecies is a specimen or series of specimens. The type of a genus or subgenus is a species. The type of a suprageneric taxon (e.g., family, etc.) is a genus. Names higher than superfamily rank do not have types. A "name-bearing type" is a specimen or image that "provides the objective standard of reference whereby the application of the name of a nominal taxon can be determined." 921:
common occurrence), then all of the constituent species must be either moved into the pre-existing genus or disassociated from the original type species and given a new generic name; the old generic name passes into synonymy and is abandoned unless there is a pressing need to make an exception (decided case-by-case, via petition to the International Commission on Zoological Nomenclature).
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taxa. If there is more than one named type that all appear to be the same taxon, then the oldest name takes precedence and is considered to be the correct name of the material in hand. If on the other hand, the taxon appears never to have been named at all, then the scientist or another qualified expert picks a type specimen and publishes a new name and an official description.
703:, based on the 1994 reporting by Louie Psihoyos of an unpublished proposal by Bob Bakker to do so. However, this designation is invalid both because Edward Cope was not one of the specimens described in Systema Naturae 10th Ed., and therefore not being a syntype is not eligible, and because Stearne's designation in 1959 has seniority and invalidates future designations. 663:. When a species was originally described on the basis of a name-bearing type consisting of multiple specimens, one of those may be designated as the lectotype. Having a single name-bearing type reduces the potential for confusion, especially considering that it is not uncommon for a series of syntypes to contain specimens of more than one species. 1173:"Laws of Botanical Nomenclature adopted by the International Botanical Congress held at Paris in August 1867; together with an Historical Introduction and Commentary by Alphonse de Candolle, Translated from the French; Reprinted from the English translation published by L. Reeve and Co., London, 1868 (with three-page commentary by Asa Gray)" 839:
The various types listed above are necessary because many species were described one or two centuries ago, when a single type specimen, a holotype, was often not designated. Also, types were not always carefully preserved, and intervening events such as wars and fires have resulted in the destruction
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A type description must include a diagnosis (typically, a discussion of similarities to and differences from closely related species), and an indication of where the type specimen or specimens are deposited for examination. The geographical location where a type specimen was originally found is known
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When identifying material, a scientist attempts to apply a taxon name to a specimen or group of specimens based on their understanding of the relevant taxa, based on (at least) having read the type description(s), preferably also based on an examination of all the type material of all of the relevant
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Although in reality biologists may examine many specimens (when available) of a new taxon before writing an official published species description, nonetheless, under the formal rules for naming species (the International Code of Zoological Nomenclature), a single type must be designated, as part of
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A detailed drawing, painting, etc., depicting the plant, from the early days of plant taxonomy. A dried plant was difficult to transport and hard to keep safe for the future; many specimens from the early days of botany have since been lost or damaged. Highly skilled botanical artists were sometimes
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or subfamily is formed. As with type species, the type genus is not necessarily the most representative but is usually the earliest described, largest or best-known genus. It is not uncommon for the name of a family to be based upon the name of a type genus that has passed into synonymy; the family
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Ideally, a type species best exemplifies the essential characteristics of the genus to which it belongs, but this is subjective and, ultimately, technically irrelevant, as it is not a requirement of the Code. If the type species proves, upon closer examination, to belong to a pre-existing genus (a
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of a long-known species, using only the available blood and feather samples. While there is still some debate on the need to deposit actual killed individuals as type specimens, it can be observed that given proper vouchering and storage, tissue samples can be just as valuable should dispute about
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A syntype is any one of two or more specimens that is listed in a species description where no holotype was designated; historically, syntypes were often explicitly designated as such, and under the present ICZN this is a requirement, but modern attempts to publish species description based on
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The ICZN has existed only since 1961 when the first edition of the Code was published. The ICZN does not always demand a type specimen for the historical validity of a species, and many "type-less" species do exist. The current edition of the Code, Article 75.3, prohibits the designation of a
294:), "is that element to which the name of a taxon is permanently attached." (article 7.2) In botany a type is either a specimen or an illustration. A specimen is a real plant (or one or more parts of a plant or a lot of small plants), dead and kept safe, "curated", in a 1156:
Lois de la nomenclature botanique adoptées par le Congrès International de Botanique tenu à Paris en août 1867 suivies d'une deuxième édition de l'introduction historique et du commentaire qui accompagnaient la rédaction préparatoire présentée à la
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makes for a poor type: the microscope slide may be lost or damaged, or it may be very difficult to find the "plant" in question among whatever else is on the microscope slide. An illustration makes for a much more reliable type (Art 37.5 of the
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refers, by definition, to the species of that particular specimen. That species was named and described by Jardine and Selby in 1828, and the holotype was placed in the museum collection so that other scientists might refer to it as necessary.
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Zoological collections are maintained by universities and museums. Ensuring that types are kept in good condition and made available for examination by taxonomists are two important functions of such collections. And, while there is only one
1647:"Terms Used in Bionomenclature: The Naming of Organisms and Plant Communities : Including Terms Used in Botanical, Cultivated Plant, Phylogenetic, Phytosociological, Prokaryote (bacteriological), Virus, and Zoological Nomenclature." 917:). The description of a genus is usually based primarily on its type species, modified and expanded by the features of other included species. The generic name is permanently associated with the name-bearing type of its type species. 619:
An allotype is a specimen of the opposite sex to the holotype, designated from among paratypes. The word was also formerly used for a specimen that shows features not seen in the holotype of a fossil. The term is not regulated by the
213:. Type specimens are theoretically even allowed to be aberrant or deformed individuals or color variations, though this is rarely chosen to be the case, as it makes it hard to determine to which population the individual belonged. 687:
He justified his choice by noting that the specimen that Linnaeus, who wrote his own autobiography five times, had most studied was probably himself. This sufficiently and correctly designated Linnaeus to be the lectotype for
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of the original type material. The validity of a species name often rests upon the availability of original type specimens; or, if the type cannot be found, or one has never existed, upon the clarity of the description.
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McNeill, J.; Barrie, F.R.; Buck, W.R.; Demoulin, V.; Greuter, W.; Hawksworth, D.L.; Herendeen, P.S.; Knapp, S.; Marhold, K.; Prado, J.; Prud'homme Van Reine, W.F.; Smith, G.F.; Wiersema, J.H.; Turland, N.J. (2012).
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Frey, Jennifer K.; Yates, Terry L.; Duszynski, Donald W.; Gannon, William L. & Gardner, Scott L. (1992). "Designation and Curatorial Management of Type Host Specimens (Symbiotypes) for New Parasite Species".
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Translation: This single character permits distinguish this type from all other species of the section ... After studying the diverse forms, I came to consider them as belonging to the one and the same specific
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Ce seul caractère permet de distinguer ce type de toutes les autres espèces de la section. ... Après avoir étudié ces diverses formes, j'en arrivai à les considérer comme appartenant à un seul et même type
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and determined all valid syntypes for the species. Crucially, in 1959, Professor William Stearne wrote in a passing remark on Linnaeus's contributions, "Linnaeus himself, must stand as the type of his
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When the original description designated a holotype, there may be additional specimens that the author designates as additional representatives of the same species, termed paratypes. These are not
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is the same and the type of the name is the same, but the extent to which the name actually applies varies greatly. Setting the circumscription of a taxon is done by a taxonomist in a publication.
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Locupletissimi Rerum naturalium Thesauri accurata Descriptio, et Iconibus artificiosissimus Expressio, per universam Physices Historiam. Opus, cui in hoc Rerum Genere, nullum par exstitit
159:, or (under the bacteriological code) a description. Some codes consider a subordinate taxon to be the type, but under the botanical code, the type is always a specimen or illustration. 593:
of that species. The holotype is typically placed in a major museum, or similar well-known public collection, so that it is freely available for later examination by other biologists.
95: 194:, particularly those established by early taxonomists, tend to be named after species that are more "typical" for them, but here too this is not always the case and due to changes in 632:
A neotype is a specimen later selected to serve as the single type specimen when an original holotype has been lost or destroyed or where the original author never cited a specimen.
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lectotype – a specimen or illustration designated from the original material as the nomenclatural type when there was no holotype specified or the holotype has been lost or destroyed
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syntypes are generally frowned upon by practicing taxonomists, and most are gradually being replaced by lectotypes. Those that still exist are still considered name-bearing types.
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research collection, but failing that, an image of an individual of that taxon has sometimes been designated as a type. Describing species and appointing type specimens is part of
1610: 676:, designated in 1959. He published the first book considered to be part of taxonomical nomenclature, the 10th edition of Systema Naturae, which included the first description of 487:, thigh bones, part of a pelvis, some ribs, and some arm and shoulder bones. There may be more than one type specimen, but there is (at least in modern times) only one holotype. 414:
epitype – a specimen or illustration selected to serve as an interpretative type, usually when another kind of type does not show the critical features needed for identification
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Each genus must have a designated type species (the term "genotype" was once used for this but has been abandoned because the word has become much better known as the term for
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syntype – any specimen (or illustration) cited in the original description when there is no holotype, or any one of two or more specimens simultaneously designated as types
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International Code of Nomenclature for algae, fungi, and plants (Melbourne Code) adopted by the Eighteenth International Botanical Congress Melbourne, Australia, July 2011
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employed by a botanist to make faithful and detailed illustrations. Some such illustrations have become the best record and have been chosen to serve as the type of taxon.
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refer to type preparations additional to the hapantotype and designated by the describing author. As with other type designations the use of the prefix "Neo-", such as
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A type specimen is a vernacular term (not a formally defined term) typically used for an individual or fossil that is any of the various name-bearing types for a
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A paralectotype is any additional specimen from among a set of syntypes after a lectotype has been designated from among them. These are not name-bearing types.
80:) and on the provision of type material, which is usually available to scientists for examination in a major museum research collection, or similar institution. 457: 111: 524:
are all those specimens included by the author in a taxon's formal description, unless the author explicitly or implicitly excludes them as part of the series.
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of that organism is formally associated. In other words, a type is an example that serves to anchor or centralizes the defining features of that particular
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is a controversial taxon: some botanists consider it to consist of over a hundred species, and others regard it as a single species. The type of the name
1751: 735:, is employed when a replacement for the original hapantotype is designated, or when an original description did not include a designated type specimen. 723:
where the type consists of two or more specimens of "directly related individuals" within a preparation medium such as a blood smear. The terms
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A hypotype is a specimen whose details have previously been published that is used in a supplementary figure or description of the species.
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paratype – any specimen (or illustration) cited in the original description that is not the holotype nor an isotype, nor one of the syntypes
1708: 1661:"Reappraisal of the parrots (Aves: Psittacidae) from the Mascarene Islands, with comments on their ecology, morphology, and affinities" 411:
neotype – a specimen or illustration selected to serve as nomenclatural type if no material from the original description is available
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provides a listing of the various kinds of types (article 9 and the Glossary), the most important of which is the holotype. These are
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Stearn, W. T. (1 March 1959). "The Background of Linnaeus's Contributions to the Nomenclature and Methods of Systematic Biology".
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The term fixation is used by the Code for the declaration of a name-bearing type, whether by original or subsequent designation.
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A lectotype is a specimen later selected to serve as the single type specimen for species originally described from a set of
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holotype – the single specimen or illustration that the author(s) clearly indicated to be the nomenclatural type of a name
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Bauer, Aaron M. (2002). "Albertus Seba, Cabinet of Natural Curiosities. The Complete Plates in Colour, 1734–1765. 2001".
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In some older taxonomic works the word "type" has sometimes been used differently. The meaning was similar in the first
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Recently, some species have been described where the type specimen was released alive back into the wild, such as the
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in London, there is a bird specimen numbered 1886.6.24.20. This is a specimen of a kind of bird commonly known as the
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or onomatophore is sometimes used, to denote the fact that biological types do not define "typical" individuals or
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is a "big" species) or whether the circumscription is limited to only one small species among the other hundred (
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The word "type" appears in botanical literature as a part of some older terms that have no status under the
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that includes some organisms and excludes others, based on a detailed published description (for example a
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unless there is "an exceptional need" for "clarifying the taxonomic status" of a species (Article 75.2).
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When a single specimen is clearly designated in the original description, this specimen is known as the
475: 362:(published on or after 1 January 2007, article 37) at these ranks, a type should not be an illustration. 799:" informally refers to a type specimen or a part of it that has been stolen, or improperly relocated. 1288: 480: 38: 1596: 1646: 1504:
A taxonomic study of the Haemoproteidae (Apicomplexa: Haemosporina) of the Avian order Strigiformes
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An illustration on which a new species or subspecies was based. For instance, the Burmese python,
1483: 1270: 1262: 1027: 567:(or symbiotype) is used to indicate the host organism from which the type specimen was obtained. 152: 1634:
Kleptotype. (n.d.). Academic Dictionaries and Encyclopedias. Retrieved November 21, 2022, from
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designated, there can be other "type" specimens, the following of which are formally defined:
298:(or the equivalent for fungi). Examples of where an illustration may serve as a type include: 202: 73: 1675: 1542: 1475: 1409: 1254: 1103: 1085: 1019: 936: 775: 484: 447: 377: 311: 191: 1757: 1134:. Australian National Botanic Gardens, Centre for Australian National Biodiversity Research 186:
At least for type specimens there is no requirement for a "typical" individual to be used.
124:, or in some cases specimens. Types are of great significance to biologists, especially to 851:
There are many other permutations and variations on terms using the suffix "-type" (e.g.,
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is the same whether the circumscription of the species includes all those small species (
1108: 1071: 745: 440: 141: 1218: 1202:(2: Comptes-redus des séances de la Société Royale de Botanique de Belgique): 163–217. 1048: 430: 1767: 1355: 897: 751: 667: 99: 72:
A taxon is a scientifically named grouping of organisms with other like organisms, a
1437:"Know Thyself: Responsible Science and the Lectotype of Homo sapiens Linnaeus, 1758" 1371:"Know Thyself: Responsible Science and the Lectotype of Homo sapiens Linnaeus, 1758" 1274: 959: 879: 763: 672: 491: 120:(ICN), the scientific name of every taxon is almost always based on one particular 1007: 69:. In older usage (pre-1900 in botany), a type was a taxon rather than a specimen. 54: 1649: (2010). United Kingdom: Global Biodiversity Information Facility. 1529:
Peirce, M. A.; Bennett, G. F. (1992). "Neohapantotype and paraneohapantotypes of
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Hansen, Hans V.; Seberg, Ole (1984). "Paralectotype, a new type term in botany".
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Terms Used in Bionomenclature. The naming of organisms (and plant communities)
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applied to the organism in question, a type can be a specimen, a culture, an
1289:"The Code Online | International Commission on Zoological Nomenclature" 976: 970: 949: 807:
Type illustrations have also been used by zoologists, as in the case of the
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in 1856 at Feldhofer in the Neander Valley in Germany, consisting of a
470: 370: 225: 46: 1051:. Smithsonian National Museum of Natural History, Department of Botany 811:, which is known only from historical illustrations and descriptions. 1579:
International Society for the History and Bibliography of Herpetology
1315:. Copenhagen: Global Biodiversity Information Facility. p. 216. 762:
An ergatotype is a specimen selected to represent a worker member in
221: 187: 129: 1479: 1258: 1023: 1510:(Doctorate thesis). Memorial University of Newfound land. p. 3 883: 532: 499: 381: 366: 359: 251: 94: 66: 31: 902: 306:
A detailed picture of something that can be seen only through a
206: 1192:"Rosa Synstylae: études sur les roses de la section Synstyleés" 1049:"Botanical nomenclature, types, & standard reference works" 117:
International Code of Nomenclature for algae, fungi, and plants
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Proceedings of the Academy of Natural Sciences of Philadelphia
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Proceedings of the Academy of Natural Sciences of Philadelphia
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that is the name-bearing type of a nominal family-group taxon.
1223:. Vol. Regnum Vegetabile 154. A.R.G. Gantner Verlag KG. 88:"Type specimen" redirects here. For the mineralogy term, see 27:
Specimen(s) to which a scientific name is formally attached
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is one of many species that are based on illustrations by
128:. Types are usually physical specimens that are kept in a 979:– a type of intrasubspecific taxon of pathogenic bacteria 1739: 1613:
Journal of the New York Entomological Society, 201-205.
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https://en_ichthyology.en-academic.com/9763/kleptotype
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Bulletin de la Société Royale de Botanique de Belgique
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name does not need to be changed in such a situation.
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is somewhat complicated by slightly different uses in
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According to a precise set of rules laid down in the
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International Commission on Zoological Nomenclature
1304: 1302: 935:A type genus is a genus from which the name of a 473:. For example, the type specimen for the species 209:, but rather fix a scientific name to a specific 1070:Marshall, Stephen A.; Evenhuis, Neal L. (2015). 1076:Hesse (Diptera, Bombyliidae) from South Africa" 479:was the specimen "Neanderthal-1" discovered by 162:For example, in the research collection of the 57:(or in some cases a group of specimens) of an 1740:International Code of Zoological Nomenclature 1042: 1040: 774:"Hypotype" redirects here. For the moth, see 112:International Code of Zoological Nomenclature 8: 258: 170:, which currently bears the scientific name 102:'s remains constitute the type specimen for 1001: 999: 997: 973:– genetic sequence data from type specimens 498:that is the name-bearing type of a nominal 452:3) dorsal and 4) ventral aspect of paratype 1622:Glime, J. M., & Wagner, D. H. (2013). 1160:. Genève et Bale: J.-B. Baillière et fils. 1501:Bishop, M. A. W. (1989). "introduction". 1107: 1089: 358:can have a type of its own. For most new 234:, type-based definitions are replaced by 1609:Baker, N. T., & Timm, R. M. (1976). 1177:The American Journal of Science and Arts 429: 380:has the same type as that of one of its 369:has the same type as that of one of its 1758:Zoological Type Nomenclature (Evenhuis) 1008:"The Type Concept in Systematic Botany" 993: 659:. In zoology, a lectotype is a kind of 250:, but has a meaning closer to the term 1179:. Series II, Volume 46 (63–74, 75–77). 327:of the taxon. For example, the common 1611:"Modern type concepts in entomology." 1342: 1340: 1338: 1336: 1334: 1332: 1211: 1209: 559:. In the case of parasites, the term 402:isotype – a duplicate of the holotype 7: 1659:Hume, Julian Pender (25 June 2007). 1132:"Plant names – a basic introduction" 900:, is the type species for the genus 1709:"Topotype Definition & Meaning" 1566:. Amsterdam: Janssonio-Waesbergios. 803:Alternatives to preserved specimens 1624:"Herbarium methods and exchanges." 1348:"Who is the type of Homo sapiens?" 450:and 2) ventral aspect of holotype, 174:. This particular specimen is the 25: 831:the validity of a species arise. 835:Formalisation of the type system 695:It has also been suggested that 343:is a "small" species). The name 766:which have polymorphic castes. 323:A type does not determine the 248:Laws of Botanical Nomenclature 1: 955:Glossary of scientific naming 1748:Glossary section (archived) 1247:The Journal of Parasitology 547:the published description. 422:: for example a clonotype. 178:for that species; the name 1805: 1535:Journal of Natural History 1153:de Candolle, A.P. (1867). 1012:American Journal of Botany 928: 877: 788: 773: 639: 600: 582: 211:operational taxonomic unit 90:type specimen (mineralogy) 87: 1680:10.11646/zootaxa.1513.1.1 1547:10.1080/00222939200770431 1435:Spamer, Earle E. (1999). 1369:Spamer, Earle E. (1999). 1309:Hawksworth, D.L. (2010). 983:Principle of typification 436:gossamer-winged butterfly 1091:10.3897/zookeys.525.6143 1006:Hitchcock, A.S. (1921), 540:Mormopterus acetabulosus 236:phylogenetic definitions 1784:Zoological nomenclature 1560:Seba, Albertus (1734). 458:zoological nomenclature 138:scientific nomenclature 1789:Botanical nomenclature 1742:, the official website 1599:. University of Basel. 906: 543: 453: 310:. A tiny "plant" on a 280:botanical nomenclature 271: 265: 259: 216:The usage of the term 164:Natural History Museum 107: 42: 1752:A compendium of terms 1597:"Compendium of Types" 1531:Haemoproteus passeris 887: 699:is the lectotype for 670:is the lectotype for 537:Type illustration of 536: 529:Use of type specimens 476:Homo neanderthalensis 433: 354:Only a species or an 350:Miscellaneous notes: 266: 256: 254:in some other works: 98: 35: 1414:10.2307/sysbio/8.1.4 481:Johann Karl Fuhlrott 345:Taraxacum officinale 341:Taraxacum officinale 337:Taraxacum officinale 333:Taraxacum officinale 201:be. Hence, the term 39:Marocaster coronatus 1774:Biological concepts 1190:CrĂ©pin, F. (1886). 911:a different concept 824:Laniarius liberatus 356:infraspecific taxon 78:species description 1779:Taxonomy (biology) 1402:Systematic Biology 1047:Nicholson, Dan H. 907: 719:A special case in 609:name-bearing types 544: 454: 292:nomenclatural type 108: 43: 36:Type specimen for 1322:978-87-92020-09-3 1230:978-3-87429-425-6 816:Bulo Burti boubou 746:Python bivittatus 661:name-bearing type 242:Older terminology 203:name-bearing type 153:nomenclature code 151:Depending on the 16:(Redirected from 1796: 1724: 1723: 1721: 1719: 1705: 1699: 1698: 1696: 1694: 1665: 1656: 1650: 1644: 1638: 1632: 1626: 1620: 1614: 1607: 1601: 1600: 1593: 1587: 1586: 1574: 1568: 1567: 1557: 1551: 1550: 1526: 1520: 1519: 1517: 1515: 1509: 1498: 1492: 1491: 1463: 1457: 1456: 1432: 1426: 1425: 1397: 1391: 1390: 1366: 1360: 1359: 1358:on 5 March 2016. 1354:. Archived from 1344: 1327: 1326: 1306: 1297: 1296: 1285: 1279: 1278: 1241: 1235: 1234: 1213: 1204: 1203: 1187: 1181: 1180: 1171:Weddell (1868). 1168: 1162: 1161: 1150: 1144: 1143: 1141: 1139: 1128: 1122: 1121: 1111: 1093: 1084:(525): 117–127. 1067: 1061: 1060: 1058: 1056: 1044: 1035: 1034: 1003: 822:), described as 809:RĂ©union parakeet 776:Hypotype (genus) 729:lectohapantotype 565: 564: 557: 556: 312:microscope slide 263: 180:Circus assimilis 172:Circus assimilis 53:is a particular 21: 1804: 1803: 1799: 1798: 1797: 1795: 1794: 1793: 1764: 1763: 1732: 1727: 1717: 1715: 1713:Merriam-Webster 1707: 1706: 1702: 1692: 1690: 1663: 1658: 1657: 1653: 1645: 1641: 1633: 1629: 1621: 1617: 1608: 1604: 1595: 1594: 1590: 1576: 1575: 1571: 1559: 1558: 1554: 1528: 1527: 1523: 1513: 1511: 1507: 1500: 1499: 1495: 1480:10.2307/1220790 1465: 1464: 1460: 1434: 1433: 1429: 1399: 1398: 1394: 1368: 1367: 1363: 1346: 1345: 1330: 1323: 1308: 1307: 1300: 1287: 1286: 1282: 1259:10.2307/3283335 1243: 1242: 1238: 1231: 1215: 1214: 1207: 1189: 1188: 1184: 1170: 1169: 1165: 1152: 1151: 1147: 1137: 1135: 1130: 1129: 1125: 1069: 1068: 1064: 1054: 1052: 1046: 1045: 1038: 1024:10.2307/2434993 1005: 1004: 995: 991: 946: 933: 927: 882: 876: 837: 805: 793: 787: 779: 772: 760: 741: 725:parahapantotype 717: 709: 653: 644: 638: 630: 617: 605: 599: 587: 581: 562: 561: 554: 553: 531: 496:nominal species 466: 451: 445: 428: 325:circumscription 276: 244: 168:spotted harrier 114:(ICZN) and the 93: 86: 63:scientific name 28: 23: 22: 15: 12: 11: 5: 1802: 1800: 1792: 1791: 1786: 1781: 1776: 1766: 1765: 1762: 1761: 1755: 1749: 1743: 1731: 1730:External links 1728: 1726: 1725: 1700: 1674:(1513): 1–76. 1651: 1639: 1627: 1615: 1602: 1588: 1569: 1552: 1541:(3): 689–690. 1533:Kruse, 1890". 1521: 1493: 1474:(4): 707–711. 1458: 1427: 1392: 1361: 1328: 1321: 1298: 1280: 1253:(5): 930–993. 1236: 1229: 1205: 1182: 1163: 1145: 1123: 1062: 1036: 1018:(5): 251–255, 992: 990: 987: 986: 985: 980: 974: 968: 963: 957: 952: 945: 942: 929:Main article: 926: 923: 878:Main article: 875: 872: 836: 833: 804: 801: 789:Main article: 786: 783: 771: 768: 759: 756: 740: 737: 733:Neohapantotype 716: 713: 708: 705: 652: 649: 640:Main article: 637: 634: 629: 626: 616: 613: 601:Main article: 598: 595: 583:Main article: 580: 577: 530: 527: 526: 525: 518: 507: 488: 465: 462: 441:Jamides elioti 427: 424: 416: 415: 412: 409: 406: 403: 400: 397: 386: 385: 374: 363: 321: 320: 304: 275: 272: 243: 240: 142:alpha taxonomy 85: 82: 26: 24: 18:Botanical type 14: 13: 10: 9: 6: 4: 3: 2: 1801: 1790: 1787: 1785: 1782: 1780: 1777: 1775: 1772: 1771: 1769: 1759: 1756: 1753: 1750: 1747: 1744: 1741: 1737: 1734: 1733: 1729: 1714: 1710: 1704: 1701: 1689: 1685: 1681: 1677: 1673: 1669: 1662: 1655: 1652: 1648: 1643: 1640: 1637: 1631: 1628: 1625: 1619: 1616: 1612: 1606: 1603: 1598: 1592: 1589: 1584: 1580: 1573: 1570: 1565: 1564: 1556: 1553: 1548: 1544: 1540: 1536: 1532: 1525: 1522: 1506: 1505: 1497: 1494: 1489: 1485: 1481: 1477: 1473: 1469: 1462: 1459: 1454: 1450: 1446: 1442: 1438: 1431: 1428: 1423: 1419: 1415: 1411: 1407: 1403: 1396: 1393: 1388: 1384: 1380: 1376: 1372: 1365: 1362: 1357: 1353: 1349: 1343: 1341: 1339: 1337: 1335: 1333: 1329: 1324: 1318: 1314: 1313: 1305: 1303: 1299: 1294: 1290: 1284: 1281: 1276: 1272: 1268: 1264: 1260: 1256: 1252: 1248: 1240: 1237: 1232: 1226: 1222: 1221: 1212: 1210: 1206: 1201: 1197: 1193: 1186: 1183: 1178: 1174: 1167: 1164: 1159: 1158: 1149: 1146: 1133: 1127: 1124: 1119: 1115: 1110: 1105: 1101: 1097: 1092: 1087: 1083: 1082: 1077: 1075: 1066: 1063: 1050: 1043: 1041: 1037: 1033: 1029: 1025: 1021: 1017: 1013: 1009: 1002: 1000: 998: 994: 988: 984: 981: 978: 975: 972: 969: 967: 964: 961: 958: 956: 953: 951: 948: 947: 943: 941: 938: 932: 924: 922: 918: 916: 912: 905: 904: 899: 896:described by 895: 891: 886: 881: 873: 871: 868: 866: 862: 858: 854: 849: 847: 841: 834: 832: 829: 825: 821: 817: 812: 810: 802: 800: 798: 792: 784: 782: 777: 769: 767: 765: 764:hymenopterans 757: 755: 753: 752:Albertus Seba 749: 747: 738: 736: 734: 730: 726: 722: 714: 712: 707:Paralectotype 706: 704: 702: 698: 693: 691: 686: 683: 682:Homo sapiens. 679: 675: 674: 669: 668:Carl Linnaeus 664: 662: 658: 650: 648: 643: 635: 633: 627: 625: 623: 614: 612: 610: 604: 596: 594: 592: 586: 578: 576: 574: 568: 566: 558: 555:type locality 548: 542: 541: 535: 528: 523: 519: 516: 515:nominal genus 512: 508: 505: 501: 497: 493: 489: 486: 482: 478: 477: 472: 468: 467: 463: 461: 459: 449: 443: 442: 437: 432: 425: 423: 421: 413: 410: 407: 404: 401: 398: 395: 394: 393: 391: 384:(article 10). 383: 379: 375: 373:(article 10). 372: 368: 364: 361: 357: 353: 352: 351: 348: 346: 342: 338: 334: 330: 326: 318: 313: 309: 305: 301: 300: 299: 297: 293: 289: 285: 281: 273: 270: 264: 262: 255: 253: 249: 241: 239: 237: 233: 232: 227: 223: 219: 214: 212: 208: 204: 200: 197: 193: 189: 184: 181: 177: 173: 169: 165: 160: 158: 154: 149: 145: 143: 139: 135: 131: 127: 123: 119: 118: 113: 105: 101: 100:Carl Linnaeus 97: 91: 84:Type specimen 83: 81: 79: 75: 70: 68: 64: 61:to which the 60: 56: 52: 48: 41: 40: 34: 30: 19: 1735: 1716:. Retrieved 1712: 1703: 1691:. Retrieved 1671: 1667: 1654: 1642: 1630: 1618: 1605: 1591: 1582: 1578: 1572: 1562: 1555: 1538: 1534: 1530: 1524: 1512:. Retrieved 1503: 1496: 1471: 1467: 1461: 1444: 1440: 1430: 1405: 1401: 1395: 1378: 1374: 1364: 1356:the original 1351: 1311: 1293:www.iczn.org 1292: 1283: 1250: 1246: 1239: 1219: 1199: 1195: 1185: 1176: 1166: 1155: 1148: 1136:. Retrieved 1126: 1079: 1073: 1065: 1053:. Retrieved 1015: 1011: 960:Nomen dubium 934: 919: 908: 901: 893: 880:Type species 874:Type species 869: 856: 850: 842: 838: 823: 813: 806: 794: 780: 761: 744: 742: 732: 728: 724: 718: 710: 701:Homo sapiens 700: 694: 690:Homo sapiens 689: 685: 681: 678:Homo sapiens 677: 673:Homo sapiens 671: 665: 654: 645: 631: 618: 606: 590: 588: 572: 569: 560: 552: 549: 545: 538: 492:type species 474: 455: 439: 419: 417: 389: 387: 349: 344: 340: 336: 332: 322: 316: 291: 287: 283: 277: 267: 257: 247: 245: 229: 217: 215: 198: 185: 179: 171: 161: 157:illustration 150: 146: 121: 115: 109: 104:Homo sapiens 103: 71: 50: 44: 37: 29: 1447:: 109–114. 1408:(1): 4–22. 1381:: 109–114. 1138:17 November 1074:Marleyimyia 1055:17 November 966:Nomen nudum 890:common toad 828:color morph 715:Hapantotype 697:Edward Cope 522:type series 464:Definitions 317:Vienna Code 261:spĂ©cifique. 196:systematics 126:taxonomists 1768:Categories 1754:(archived) 1693:13 January 989:References 931:Type genus 925:Type genus 861:generitype 855:, cotype, 820:bushshrike 797:kleptotype 795:The term " 791:Kleptotype 785:Kleptotype 758:Ergatotype 721:Protistans 666:Formally, 511:type genus 426:In zoology 308:microscope 1736:ICZN Code 1688:1175-5334 1453:0097-3157 1422:1063-5157 1387:0097-3157 1100:1313-2970 977:Pathotype 971:Genetypes 962:(zoology) 950:Archetype 894:Bufo bufo 739:Iconotype 651:Lectotype 563:type host 329:dandelion 296:herbarium 274:In botany 231:PhyloCode 228:. In the 134:herbarium 1746:Fishbase 1718:18 April 1514:18 April 1275:82003952 1118:26487819 944:See also 915:genetics 898:Linnaeus 857:topotype 853:allotype 770:Hypotype 754:(1734). 657:syntypes 615:Allotype 603:Paratype 597:Paratype 591:holotype 585:Holotype 579:Holotype 573:holotype 504:subgenus 485:skullcap 319:, 2006). 192:families 176:holotype 122:specimen 59:organism 55:specimen 1668:Zootaxa 1488:1220790 1267:3283335 1157:congrès 1109:4607853 1081:ZooKeys 1032:2434993 865:isotype 846:neotype 642:Syntype 636:Syntype 628:Neotype 551:as its 513:is the 494:is the 471:species 371:species 226:zoology 47:biology 1686:  1486:  1451:  1420:  1385:  1319:  1273:  1265:  1227:  1116:  1106:  1098:  1030:  937:family 448:dorsal 382:genera 378:family 222:botany 199:cannot 188:Genera 130:museum 1760:(PDF) 1664:(PDF) 1508:(PDF) 1484:JSTOR 1468:Taxon 1271:S2CID 1263:JSTOR 1028:JSTOR 500:genus 367:genus 288:typus 269:type. 252:taxon 67:taxon 1720:2024 1695:2011 1684:ISSN 1672:1513 1516:2024 1449:ISSN 1418:ISSN 1383:ISSN 1317:ISBN 1225:ISBN 1140:2015 1114:PMID 1096:ISSN 1057:2015 903:Bufo 888:The 727:and 622:ICZN 520:The 388:The 360:taxa 284:type 282:, a 224:and 218:type 207:taxa 190:and 140:and 51:type 49:, a 1676:doi 1543:doi 1476:doi 1445:149 1410:doi 1379:149 1255:doi 1104:PMC 1086:doi 1020:doi 913:in 818:(a 502:or 456:In 446:1) 420:ICN 390:ICN 278:In 132:or 74:set 45:In 1770:: 1738:: 1711:. 1682:. 1670:. 1666:. 1581:. 1539:26 1537:. 1482:. 1472:33 1470:. 1443:. 1439:. 1416:. 1404:. 1377:. 1373:. 1350:. 1331:^ 1301:^ 1291:. 1269:. 1261:. 1251:78 1249:. 1208:^ 1200:25 1198:. 1194:. 1175:. 1112:. 1102:. 1094:. 1078:. 1039:^ 1026:, 1014:, 1010:, 996:^ 892:, 863:, 859:, 692:. 624:. 611:. 509:A 490:A 438:, 434:A 376:A 365:A 290:, 238:. 144:. 1722:. 1697:. 1678:: 1585:. 1583:3 1549:. 1545:: 1518:. 1490:. 1478:: 1455:. 1424:. 1412:: 1406:8 1389:. 1325:. 1295:. 1277:. 1257:: 1233:. 1142:. 1120:. 1088:: 1059:. 1022:: 1016:8 778:. 748:, 684:" 506:. 444:: 286:( 106:. 92:. 20:)

Index

Botanical type

Marocaster coronatus
biology
specimen
organism
scientific name
taxon
set
species description
type specimen (mineralogy)

Carl Linnaeus
International Code of Zoological Nomenclature
International Code of Nomenclature for algae, fungi, and plants
taxonomists
museum
herbarium
scientific nomenclature
alpha taxonomy
nomenclature code
illustration
Natural History Museum
spotted harrier
holotype
Genera
families
systematics
name-bearing type
taxa

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