125:
102:
283:
These beetles are 1.4 to 12 millimeters long as adults. They generally have very elongated bodies, some over 4 times longer than wide. They may be cylindrical or somewhat flattened. They are yellow to black in color, some with various patterning and some with red spots. They are hairless to slightly
686:
Robertson, James A.; Ślipiński, Adam; Moulton, Matthew; Shockley, Floyd W.; Giorgi, Adriano; Lord, Nathan P.; Mckenna, Duane D.; Tomaszewska, Wioletta; Forrester, Juanita; Miller, Kelly B.; Whiting, Michael F.; Mchugh, Joseph V. (October 2015).
689:"Phylogeny and classification of Cucujoidea and the recognition of a new superfamily Coccinelloidea (Coleoptera: Cucujiformia): Systematics of Cucujoidea and Coccinelloidea"
336:, with members of the former subfamily Bothriderinae constituting the only remaining members of the family. Genera placed in the family in the new circumscription include:
2874:
2913:
676:
Coleoptera, Beetles. Volume 2. Morphology and
Systematics (Polyphaga partim). (Eds. R. G. Beutel, R. A. B. Leschen and J. F. Lawrence). W. DeGruyter, Berlin.
2672:
2848:
2887:
661:
639:
785:
727:
288:
have 9 to 11 segments and are usually club-shaped at the tips. Larvae are up to 18 millimeters long and are elongate in shape.
2892:
595:
728:"Alveoderes gen. nov., the earliest fossil of Bothrideridae from mid-Cretaceous Burmese amber (Coleoptera: Coccinelloidea)"
2900:
124:
3000:
2757:
2734:
2918:
2827:
221:
2840:
778:
2995:
2990:
2687:
2762:
2949:
509:
2967:
501:
2928:
2796:
1924:
325:
397:
389:
2143:
771:
658:
517:
437:
429:
2725:
708:
477:
381:
341:
329:
230:
119:
560:
549:
469:
421:
349:
2954:
525:
493:
461:
405:
321:
2936:
2783:
2412:
1829:
749:
635:
533:
373:
357:
301:
285:
2879:
453:
2941:
2509:
2288:
2283:
2206:
1964:
980:
739:
700:
101:
2198:
2151:
1996:
1980:
1656:
1416:
1385:
1300:
1218:
1109:
1077:
665:
688:
2835:
2822:
2601:
2552:
2407:
2214:
1882:
1866:
1805:
1700:
1671:
1599:
1563:
1547:
1463:
1408:
1313:
1210:
1059:
1021:
1001:
897:
253:
206:
2984:
2739:
2583:
2387:
2378:
2119:
2090:
2082:
2074:
2048:
2023:
1919:
1906:
1845:
1763:
1666:
1661:
1542:
1476:
1447:
1395:
1365:
1318:
1272:
1239:
1155:
1147:
1118:
956:
628:
564:
309:
59:
712:
2560:
2484:
2230:
2222:
2177:
2161:
2111:
2032:
1988:
1940:
1858:
1771:
1640:
1627:
1607:
1568:
1468:
1455:
1426:
1360:
1348:
1244:
1201:
1184:
1176:
1126:
964:
934:
865:
854:
838:
305:
196:
744:
2788:
2905:
2861:
2719:
2646:
2617:
2591:
2578:
2518:
2476:
2460:
2452:
2444:
2428:
2420:
2356:
2314:
2306:
2275:
2190:
2169:
2135:
2127:
2040:
1972:
1956:
1948:
1911:
1898:
1874:
1792:
1784:
1734:
1726:
1684:
1648:
1619:
1591:
1586:
1502:
1497:
1489:
1439:
1375:
1370:
1284:
1249:
1234:
1226:
1142:
1134:
951:
445:
413:
365:
110:
34:
2573:
2534:
2526:
2468:
2331:
2293:
2246:
2238:
2058:
2006:
1890:
1821:
1692:
1676:
1632:
1573:
1537:
1517:
1481:
1390:
1380:
1254:
988:
972:
572:
568:
297:
272:
79:
44:
2710:
763:
753:
2809:
2638:
2630:
2622:
2609:
2565:
2364:
2339:
2267:
2095:
2066:
1750:
1581:
1555:
1507:
1421:
1323:
1292:
1264:
1097:
1072:
1067:
1048:
1013:
902:
885:
877:
826:
814:
485:
333:
265:
186:
156:
136:
84:
2770:
2749:
2704:
2542:
2492:
2436:
2395:
2344:
2001:
1932:
1813:
1525:
1331:
1305:
1163:
996:
943:
922:
832:
74:
69:
54:
49:
39:
2775:
2259:
2251:
2103:
1837:
1758:
1742:
704:
613:
541:
268:
89:
64:
2866:
1850:
1710:
1343:
820:
808:
795:
249:
176:
166:
146:
2853:
2681:
2814:
271:. In older literature, the family was often included in the family
2962:
296:
Most beetles in this family live under tree bark. The larvae are
2685:
1094:
1045:
919:
851:
767:
657:
2010. Slipiński, A. S., N. P. Lord and J. F. Lawrence, 10.28.
2801:
1166:(furniture beetles, death watch beetles, spider beetles)
630:
A field guide to the insects of
America north of Mexico
616:
Version 22 September 2009. Tree of Life Web
Project.
556:
320:
Genera that were assigned to the former subfamilies
2694:
2600:
2551:
2517:
2508:
2386:
2377:
2330:
2305:
2189:
2160:
2057:
2031:
2022:
1804:
1783:
1709:
1516:
1407:
1263:
1209:
1200:
1175:
1117:
1108:
933:
627:
2043:(jewel beetles, or metallic wood-boring beetles)
598:Integrated Taxonomic Information System (ITIS).
264:. They occur worldwide with most native to the
1766:(true weevils, bark beetles, ambrosia beetles)
779:
8:
2009:(ironclad beetles, cylindrical bark beetles)
275:(e.g., ), but is now considered unrelated.
2682:
2514:
2383:
2028:
1206:
1114:
1105:
1091:
1042:
930:
916:
848:
786:
772:
764:
100:
20:
2673:List of subgroups of the order Coleoptera
1753:(straight snout weevils, New York weevil)
743:
2180:(cicada beetle, cicada parasite beetles)
332:are now placed into the separate family
585:
2270:(false fireflies, long-lipped beetles)
653:
651:
614:Bothrideridae: Cocoon-forming beetles.
591:
589:
626:Donald Borror; Richard White (1970).
608:
606:
604:
7:
2968:396787A7-5C55-1B5D-FF1D-FC3FFEF6F14C
2929:a03387a2-8122-49cb-9193-9b7c583fcf98
2841:a81d0bc1-7f7f-4b30-aa2f-1ce573d72d87
2296:(false metallic wood-boring beetles)
726:Li, Yan-Da; Cai, Chen-Yang (2024).
300:of other insects, including other
14:
256:. They are known commonly as the
1907:Perimylopidae, or Promecheilidae
1492:(minute brown scavenger beetles)
123:
2122:(variegated mud-loving beetles)
284:hairy or scaly in texture. The
1:
1951:(narrow-waisted bark beetles)
1643:(parasitic flat bark beetles)
1594:(short-winged flower beetles)
1450:(lady beetles, or God's cows)
1187:(tooth-necked fungus beetles)
745:10.1590/s1984-4689.v41.e23084
659:Bothrideridae Erichson, 1845.
2463:(sand-loving scarab beetles)
1853:(minute tree-fungus beetles)
1679:(silvanid flat bark beetles)
2612:(primitive carrion beetles)
2568:(minute mud-loving beetles)
2431:(bumble bee scarab beetles)
2201:(soft-bodied plant beetles)
2172:(soft bodied plant beetles)
975:(predaceous diving beetles)
3017:
2447:(scavenger scarab beetles)
2439:(enigmatic scarab beetles)
1612:Lamingtonium binnaberrense
1326:(soft-wing flower beetles)
2670:
2586:(water scavenger beetles)
2319:Rhinorhipus tamborinensis
2098:(long-toed water beetles)
1991:(polypore fungus beetles)
1893:(palm and flower beetles)
1877:(tumbling flower beetles)
1824:(ant-like flower beetles)
1602:(lined flat bark beetles)
1576:(pleasing fungus beetles)
1471:(handsome fungus beetles)
1104:
1090:
1055:
1041:
1016:(burrowing water beetles)
929:
915:
861:
847:
803:
236:
229:
120:Scientific classification
118:
108:
99:
23:
2633:(feather-winged beetles)
1967:(false longhorn beetles)
1914:(jugular-horned beetles)
1861:(false darkling beetles)
1832:(cryptic fungus beetles)
1651:(shining flower beetles)
991:(crawling water beetles)
890:Sikhotealinia zhiltzovae
2114:(forest stream beetles)
1975:(synchroa bark beetles)
1935:(dead log bark beetles)
1927:(fire-coloured beetles)
1816:(ant-like leaf beetles)
1550:(silken fungus beetles)
1484:(well polished beetles)
1458:(minute fungus beetles)
1277:Acanthocnemus nigricans
2625:(round fungus beetles)
1959:(false flower beetles)
1943:(wedge-shaped beetles)
1901:(false blister beetle)
1885:(hairy fungus beetles)
1840:(conifer bark beetles)
1737:(leaf-rolling weevils)
1398:(bark-gnawing beetles)
1158:(wounded-tree beetles)
1024:(false ground beetles)
946:(trout-stream beetles)
258:cocoon-forming beetles
2950:Paleobiology Database
2537:(false clown beetles)
2359:(plate-thigh beetles)
2241:(false click beetles)
2146:(water-penny beetles)
1795:(ship-timber beetles)
1774:(pine flower weevils)
1622:(root-eating beetles)
1442:(minute bark beetles)
1353:Phloiophilus edwardsi
880:(reticulated beetles)
693:Systematic Entomology
2836:Fauna Europaea (new)
2415:(false stag beetles)
2400:Belohina inexpectata
2262:(net-winged beetles)
2225:(rare click beetles)
2138:(travertine beetles)
2130:(minute mud beetles)
1695:(dry-fungus beetles)
1530:Agapytho foveicollis
1287:(false skin beetles)
1150:(Jacobson's beetles)
870:Crowsoniella relicta
634:. Houghton Mifflin.
2413:Diphyllostomatidae
1745:(primitive weevils)
1558:(flat bark beetles)
1308:(checkered beetles)
1295:(fruitworm beetles)
983:(whirligig beetles)
732:Zoologia (Curitiba)
674:Handbook of Zoology
292:Biology and ecology
252:in the superfamily
3001:Polyphaga families
2278:(glowworm beetles)
1983:(darkling beetles)
1830:Archeocrypticidae
1687:(palmetto beetles)
1434:(dry bark beetles)
1221:(longhorn beetles)
705:10.1111/syen.12138
664:2015-12-22 at the
612:Lord, N. P. 2009.
302:woodboring beetles
2978:
2977:
2937:Open Tree of Life
2688:Taxon identifiers
2679:
2678:
2666:
2665:
2662:
2661:
2658:
2657:
2650:
2642:
2641:(carrion beetles)
2634:
2626:
2613:
2587:
2569:
2538:
2530:
2504:
2503:
2496:
2488:
2480:
2472:
2464:
2456:
2448:
2440:
2432:
2424:
2416:
2403:
2373:
2372:
2360:
2352:
2322:
2297:
2279:
2271:
2263:
2255:
2242:
2234:
2226:
2218:
2217:(soldier beetles)
2210:
2207:Brachypsectridae
2202:
2181:
2173:
2147:
2139:
2131:
2123:
2115:
2107:
2099:
2086:
2078:
2070:
2044:
2018:
2017:
2010:
1992:
1984:
1976:
1968:
1965:Stenotrachelidae
1960:
1952:
1944:
1936:
1928:
1915:
1902:
1894:
1886:
1878:
1870:
1869:(blister beetles)
1862:
1854:
1841:
1833:
1825:
1817:
1796:
1775:
1767:
1754:
1746:
1738:
1730:
1720:
1696:
1688:
1680:
1652:
1644:
1636:
1623:
1615:
1603:
1595:
1577:
1559:
1551:
1533:
1493:
1485:
1472:
1459:
1451:
1443:
1435:
1417:Akalyptoischiidae
1399:
1356:
1339:
1327:
1309:
1296:
1288:
1280:
1230:
1222:
1196:
1195:
1188:
1167:
1159:
1151:
1138:
1130:
1086:
1085:
1063:
1037:
1036:
1033:
1032:
1025:
1017:
1009:
992:
984:
976:
968:
960:
947:
911:
910:
893:
881:
873:
641:978-0-395-07436-7
510:Pseudobothrideres
243:
242:
225:
30:Cenomanian–Recent
16:Family of beetles
3008:
2971:
2970:
2958:
2957:
2945:
2944:
2932:
2931:
2922:
2921:
2909:
2908:
2906:NHMSYS0020151511
2896:
2895:
2883:
2882:
2870:
2869:
2857:
2856:
2844:
2843:
2831:
2830:
2818:
2817:
2805:
2804:
2792:
2791:
2779:
2778:
2766:
2765:
2753:
2752:
2743:
2742:
2730:
2729:
2728:
2715:
2714:
2713:
2683:
2648:
2640:
2632:
2624:
2611:
2585:
2567:
2536:
2528:
2515:
2510:Staphyliniformia
2494:
2486:
2478:
2470:
2462:
2454:
2446:
2438:
2430:
2422:
2414:
2397:
2384:
2358:
2349:Declinia relicta
2346:
2316:
2295:
2289:Sinopyrophoridae
2284:Rhagophthalmidae
2277:
2269:
2261:
2253:
2240:
2232:
2224:
2216:
2208:
2200:
2199:Artematopodidae
2179:
2171:
2145:
2137:
2129:
2121:
2113:
2106:(riffle beetles)
2105:
2097:
2085:(turtle beetles)
2084:
2076:
2068:
2042:
2029:
2008:
1990:
1982:
1974:
1966:
1958:
1950:
1942:
1934:
1926:
1913:
1900:
1892:
1884:
1876:
1868:
1860:
1852:
1839:
1831:
1823:
1815:
1794:
1773:
1765:
1752:
1744:
1736:
1729:(fungus weevils)
1728:
1719:
1718:
1714:
1701:Tasmosalpingidae
1694:
1686:
1678:
1650:
1642:
1634:
1621:
1609:
1601:
1593:
1575:
1557:
1549:
1527:
1491:
1483:
1470:
1457:
1449:
1441:
1433:
1397:
1350:
1333:
1325:
1307:
1294:
1286:
1274:
1273:Acanthocnemidae
1228:
1220:
1207:
1186:
1165:
1157:
1149:
1136:
1128:
1115:
1106:
1092:
1061:
1043:
1023:
1015:
1003:
990:
982:
974:
966:
959:(ground beetles)
958:
945:
931:
917:
887:
879:
867:
866:Crowsoniellidae
849:
788:
781:
774:
765:
758:
757:
747:
723:
717:
716:
683:
677:
655:
646:
645:
633:
623:
617:
610:
599:
593:
558:
502:Pseudantibothrus
262:dry bark beetles
220:
128:
127:
104:
94:
31:
27:Temporal range:
21:
3016:
3015:
3011:
3010:
3009:
3007:
3006:
3005:
2981:
2980:
2979:
2974:
2966:
2961:
2953:
2948:
2940:
2935:
2927:
2925:
2917:
2912:
2904:
2899:
2891:
2886:
2878:
2873:
2865:
2860:
2852:
2847:
2839:
2834:
2826:
2821:
2813:
2808:
2800:
2795:
2787:
2782:
2774:
2769:
2761:
2756:
2748:
2746:
2738:
2733:
2724:
2723:
2718:
2709:
2708:
2703:
2690:
2680:
2675:
2654:
2596:
2547:
2529:(clown beetles)
2500:
2471:(betsy beetles)
2379:Scarabaeiformia
2369:
2326:
2301:
2233:(click beetles)
2209:(Texas beetles)
2185:
2156:
2152:Ptilodactylidae
2077:(cedar beetles)
2053:
2014:
1997:Trictenotomidae
1883:Mycetophagidae
1800:
1779:
1716:
1715:
1713:
1705:
1657:Phloeostichidae
1608:Lamingtoniidae
1600:Laemophloeidae
1548:Cryptophagidae
1512:
1403:
1386:Thanerocleridae
1349:Phloiophilidae
1336:Metaxina ornata
1301:Chaetosomatidae
1259:
1192:
1171:
1129:(auger beetles)
1110:Bostrichiformia
1100:
1082:
1078:Torridincolidae
1062:(skiff beetles)
1060:Hydroscaphidae
1051:
1029:
1022:Trachypachidae
967:(tiger beetles)
925:
907:
857:
843:
799:
792:
762:
761:
725:
724:
720:
685:
684:
680:
666:Wayback Machine
656:
649:
642:
625:
624:
620:
611:
602:
594:
587:
582:
567:, Myanmar, mid-
326:Xylariophilinae
318:
298:ectoparasitoids
294:
281:
248:is a family of
219:
122:
95:
93:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
37:
29:
28:
25:
17:
12:
11:
5:
3014:
3012:
3004:
3003:
2998:
2996:Coccinelloidea
2993:
2983:
2982:
2976:
2975:
2973:
2972:
2959:
2946:
2933:
2923:
2910:
2897:
2884:
2871:
2858:
2845:
2832:
2823:Fauna Europaea
2819:
2806:
2793:
2780:
2767:
2754:
2744:
2731:
2716:
2700:
2698:
2692:
2691:
2686:
2677:
2676:
2671:
2668:
2667:
2664:
2663:
2660:
2659:
2656:
2655:
2653:
2652:
2649:(rove beetles)
2647:Staphylinidae
2644:
2636:
2628:
2620:
2615:
2606:
2604:
2602:Staphylinoidea
2598:
2597:
2595:
2594:
2589:
2584:Hydrophilidae
2581:
2576:
2571:
2563:
2557:
2555:
2553:Hydrophiloidea
2549:
2548:
2546:
2545:
2540:
2532:
2523:
2521:
2512:
2506:
2505:
2502:
2501:
2499:
2498:
2495:(hide beetles)
2490:
2482:
2479:(rain beetles)
2474:
2466:
2458:
2455:(stag beetles)
2450:
2442:
2434:
2426:
2418:
2410:
2408:Bolboceratidae
2405:
2392:
2390:
2381:
2375:
2374:
2371:
2370:
2368:
2367:
2362:
2354:
2342:
2336:
2334:
2328:
2327:
2325:
2324:
2315:Rhinorhipidae
2311:
2309:
2307:Rhinorhipoidea
2303:
2302:
2300:
2299:
2291:
2286:
2281:
2273:
2265:
2257:
2249:
2244:
2236:
2228:
2220:
2212:
2204:
2195:
2193:
2187:
2186:
2184:
2183:
2175:
2166:
2164:
2158:
2157:
2155:
2154:
2149:
2141:
2133:
2125:
2120:Heteroceridae
2117:
2109:
2101:
2093:
2088:
2083:Chelonariidae
2080:
2075:Callirhipidae
2072:
2069:(pill beetles)
2063:
2061:
2055:
2054:
2052:
2051:
2046:
2037:
2035:
2026:
2020:
2019:
2016:
2015:
2013:
2012:
2004:
1999:
1994:
1986:
1981:Tenebrionidae
1978:
1970:
1962:
1954:
1946:
1938:
1930:
1922:
1917:
1909:
1904:
1896:
1888:
1880:
1872:
1864:
1856:
1848:
1843:
1835:
1827:
1819:
1810:
1808:
1806:Tenebrionoidea
1802:
1801:
1799:
1798:
1789:
1787:
1781:
1780:
1778:
1777:
1769:
1764:Curculionidae
1761:
1756:
1748:
1740:
1732:
1723:
1721:
1711:Curculionoidea
1707:
1706:
1704:
1703:
1698:
1690:
1682:
1674:
1672:Protocucujidae
1669:
1664:
1659:
1654:
1646:
1638:
1630:
1625:
1617:
1605:
1597:
1589:
1584:
1579:
1571:
1566:
1564:Cybocephalidae
1561:
1553:
1545:
1540:
1535:
1522:
1520:
1514:
1513:
1511:
1510:
1505:
1500:
1495:
1487:
1479:
1474:
1466:
1464:Discolomatidae
1461:
1453:
1448:Coccinellidae
1445:
1437:
1432:Bothrideridae
1429:
1424:
1419:
1413:
1411:
1409:Coccinelloidea
1405:
1404:
1402:
1401:
1396:Trogossitidae
1393:
1388:
1383:
1378:
1373:
1368:
1363:
1358:
1346:
1341:
1329:
1321:
1316:
1314:Lophocateridae
1311:
1303:
1298:
1290:
1282:
1269:
1267:
1261:
1260:
1258:
1257:
1252:
1247:
1242:
1237:
1232:
1229:(leaf beetles)
1227:Chrysomelidae
1224:
1215:
1213:
1211:Chrysomeloidea
1204:
1198:
1197:
1194:
1193:
1191:
1190:
1181:
1179:
1173:
1172:
1170:
1169:
1161:
1156:Nosodendridae
1153:
1148:Jacobsoniidae
1145:
1140:
1137:(skin beetles)
1132:
1123:
1121:
1112:
1102:
1101:
1095:
1088:
1087:
1084:
1083:
1081:
1080:
1075:
1070:
1065:
1056:
1053:
1052:
1046:
1039:
1038:
1035:
1034:
1031:
1030:
1028:
1027:
1019:
1011:
1006:Meru phyllisae
999:
994:
986:
978:
970:
962:
954:
949:
940:
938:
927:
926:
920:
913:
912:
909:
908:
906:
905:
900:
898:Micromalthidae
895:
883:
875:
862:
859:
858:
852:
845:
844:
842:
841:
835:
829:
823:
817:
811:
804:
801:
800:
793:
791:
790:
783:
776:
768:
760:
759:
718:
699:(4): 745–778.
678:
647:
640:
618:
600:
596:Bothrideridae.
584:
583:
581:
578:
577:
576:
554:
546:
538:
530:
522:
514:
506:
498:
490:
482:
474:
466:
458:
450:
442:
434:
426:
418:
410:
402:
398:Cylindromicrus
394:
390:Craspedophilus
386:
378:
370:
362:
354:
346:
317:
314:
310:carpenter bees
293:
290:
280:
277:
254:Coccinelloidea
241:
240:
234:
233:
227:
226:
214:
210:
209:
207:Coccinelloidea
204:
200:
199:
194:
190:
189:
184:
180:
179:
174:
170:
169:
164:
160:
159:
154:
150:
149:
144:
140:
139:
134:
130:
129:
116:
115:
106:
105:
97:
96:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
33:
32:
26:
15:
13:
10:
9:
6:
4:
3:
2:
3013:
3002:
2999:
2997:
2994:
2992:
2991:Bothrideridae
2989:
2988:
2986:
2969:
2964:
2960:
2956:
2951:
2947:
2943:
2938:
2934:
2930:
2924:
2920:
2915:
2911:
2907:
2902:
2898:
2894:
2889:
2885:
2881:
2876:
2872:
2868:
2863:
2859:
2855:
2850:
2846:
2842:
2837:
2833:
2829:
2824:
2820:
2816:
2811:
2807:
2803:
2798:
2794:
2790:
2785:
2781:
2777:
2772:
2768:
2764:
2759:
2755:
2751:
2745:
2741:
2740:Bothrideridae
2736:
2732:
2727:
2726:Bothrideridae
2721:
2717:
2712:
2706:
2702:
2701:
2699:
2697:
2696:Bothrideridae
2693:
2689:
2684:
2674:
2669:
2651:
2645:
2643:
2637:
2635:
2629:
2627:
2621:
2619:
2616:
2614:
2608:
2607:
2605:
2603:
2599:
2593:
2590:
2588:
2582:
2580:
2577:
2575:
2572:
2570:
2564:
2562:
2559:
2558:
2556:
2554:
2550:
2544:
2541:
2539:
2535:Sphaeritidae
2533:
2531:
2525:
2524:
2522:
2520:
2516:
2513:
2511:
2507:
2497:
2491:
2489:
2485:Scarabaeidae
2483:
2481:
2475:
2473:
2467:
2465:
2459:
2457:
2451:
2449:
2443:
2441:
2435:
2433:
2427:
2425:
2423:(dor beetles)
2419:
2417:
2411:
2409:
2406:
2404:
2401:
2394:
2393:
2391:
2389:
2388:Scarabaeoidea
2385:
2382:
2380:
2376:
2366:
2363:
2361:
2355:
2353:
2350:
2343:
2341:
2338:
2337:
2335:
2333:
2329:
2323:
2320:
2313:
2312:
2310:
2308:
2304:
2298:
2292:
2290:
2287:
2285:
2282:
2280:
2274:
2272:
2266:
2264:
2258:
2256:
2250:
2248:
2245:
2243:
2237:
2235:
2229:
2227:
2223:Cerophytidae
2221:
2219:
2213:
2211:
2205:
2203:
2197:
2196:
2194:
2192:
2188:
2182:
2178:Rhipiceridae
2176:
2174:
2168:
2167:
2165:
2163:
2159:
2153:
2150:
2148:
2142:
2140:
2134:
2132:
2126:
2124:
2118:
2116:
2112:Eulichadidae
2110:
2108:
2102:
2100:
2094:
2092:
2091:Cneoglossidae
2089:
2087:
2081:
2079:
2073:
2071:
2065:
2064:
2062:
2060:
2056:
2050:
2049:Schizopodidae
2047:
2045:
2039:
2038:
2036:
2034:
2030:
2027:
2025:
2024:Elateriformia
2021:
2011:
2005:
2003:
2000:
1998:
1995:
1993:
1989:Tetratomidae
1987:
1985:
1979:
1977:
1971:
1969:
1963:
1961:
1955:
1953:
1947:
1945:
1941:Ripiphoridae
1939:
1937:
1931:
1929:
1925:Pyrochroidae
1923:
1921:
1920:Pterogeniidae
1918:
1916:
1910:
1908:
1905:
1903:
1897:
1895:
1889:
1887:
1881:
1879:
1873:
1871:
1865:
1863:
1859:Melandryidae
1857:
1855:
1849:
1847:
1846:Chalcodryidae
1844:
1842:
1836:
1834:
1828:
1826:
1820:
1818:
1812:
1811:
1809:
1807:
1803:
1797:
1791:
1790:
1788:
1786:
1782:
1776:
1772:Nemonychidae
1770:
1768:
1762:
1760:
1757:
1755:
1749:
1747:
1741:
1739:
1733:
1731:
1725:
1724:
1722:
1712:
1708:
1702:
1699:
1697:
1691:
1689:
1683:
1681:
1675:
1673:
1670:
1668:
1667:Propalticidae
1665:
1663:
1662:Priasilphidae
1660:
1658:
1655:
1653:
1647:
1645:
1641:Passandridae
1639:
1637:
1635:(sap beetles)
1631:
1629:
1626:
1624:
1618:
1616:
1613:
1606:
1604:
1598:
1596:
1590:
1588:
1585:
1583:
1580:
1578:
1572:
1570:
1567:
1565:
1562:
1560:
1554:
1552:
1546:
1544:
1543:Cavognathidae
1541:
1539:
1536:
1534:
1531:
1524:
1523:
1521:
1519:
1515:
1509:
1506:
1504:
1501:
1499:
1496:
1494:
1488:
1486:
1480:
1478:
1477:Eupsilobiidae
1475:
1473:
1469:Endomychidae
1467:
1465:
1462:
1460:
1456:Corylophidae
1454:
1452:
1446:
1444:
1438:
1436:
1430:
1428:
1425:
1423:
1420:
1418:
1415:
1414:
1412:
1410:
1406:
1400:
1394:
1392:
1389:
1387:
1384:
1382:
1379:
1377:
1374:
1372:
1371:Protopeltidae
1369:
1367:
1366:Prionoceridae
1364:
1362:
1359:
1357:
1354:
1347:
1345:
1342:
1340:
1337:
1330:
1328:
1322:
1320:
1319:Mauroniscidae
1317:
1315:
1312:
1310:
1304:
1302:
1299:
1297:
1291:
1289:
1283:
1281:
1278:
1271:
1270:
1268:
1266:
1262:
1256:
1253:
1251:
1248:
1246:
1243:
1241:
1240:Megalopodidae
1238:
1236:
1233:
1231:
1225:
1223:
1219:Cerambycidae
1217:
1216:
1214:
1212:
1208:
1205:
1203:
1199:
1189:
1185:Derodontidae
1183:
1182:
1180:
1178:
1177:Derodontoidea
1174:
1168:
1162:
1160:
1154:
1152:
1146:
1144:
1143:Endecatomidae
1141:
1139:
1133:
1131:
1127:Bostrichidae
1125:
1124:
1122:
1120:
1119:Bostrichoidea
1116:
1113:
1111:
1107:
1103:
1099:
1093:
1089:
1079:
1076:
1074:
1073:Sphaeriusidae
1071:
1069:
1066:
1064:
1058:
1057:
1054:
1050:
1044:
1040:
1026:
1020:
1018:
1012:
1010:
1007:
1000:
998:
995:
993:
987:
985:
979:
977:
971:
969:
965:Cicindelidae
963:
961:
955:
953:
950:
948:
942:
941:
939:
936:
932:
928:
924:
918:
914:
904:
901:
899:
896:
894:
891:
884:
882:
876:
874:
871:
864:
863:
860:
856:
850:
846:
840:
836:
834:
830:
828:
824:
822:
818:
816:
812:
810:
806:
805:
802:
797:
789:
784:
782:
777:
775:
770:
769:
766:
755:
751:
746:
741:
737:
733:
729:
722:
719:
714:
710:
706:
702:
698:
694:
690:
682:
679:
675:
671:
667:
663:
660:
654:
652:
648:
643:
637:
632:
631:
622:
619:
615:
609:
607:
605:
601:
597:
592:
590:
586:
579:
574:
570:
566:
565:Burmese amber
563:
562:
555:
552:
551:
547:
544:
543:
539:
536:
535:
531:
528:
527:
523:
520:
519:
518:Pseudososylus
515:
512:
511:
507:
504:
503:
499:
496:
495:
491:
488:
487:
483:
480:
479:
475:
472:
471:
467:
464:
463:
459:
456:
455:
451:
448:
447:
443:
440:
439:
435:
432:
431:
427:
424:
423:
419:
416:
415:
411:
408:
407:
403:
400:
399:
395:
392:
391:
387:
384:
383:
379:
376:
375:
371:
368:
367:
363:
360:
359:
355:
352:
351:
347:
344:
343:
339:
338:
337:
335:
331:
327:
323:
315:
313:
311:
307:
303:
299:
291:
289:
287:
278:
276:
274:
270:
267:
263:
259:
255:
251:
247:
246:Bothrideridae
239:
235:
232:
228:
223:
218:
217:Bothrideridae
215:
212:
211:
208:
205:
203:Superfamily:
202:
201:
198:
195:
192:
191:
188:
185:
182:
181:
178:
175:
172:
171:
168:
165:
162:
161:
158:
155:
152:
151:
148:
145:
142:
141:
138:
135:
132:
131:
126:
121:
117:
114:
112:
107:
103:
98:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
36:
24:Bothrideridae
22:
19:
2695:
2574:Helophoridae
2566:Georissidae
2561:Epimetopidae
2477:Pleocomidae
2461:Ochodaeidae
2445:Hybosoridae
2429:Glaphyridae
2421:Geotrupidae
2399:
2396:Belohinidae
2357:Eucinetidae
2348:
2345:Decliniidae
2318:
2276:Phengodidae
2215:Cantharidae
2170:Dascillidae
2162:Dascilloidea
2144:Psephenidae
2136:Lutrochidae
2128:Limnichidae
2041:Buprestidae
2033:Buprestoidea
1973:Synchroidae
1957:Scraptiidae
1949:Salpingidae
1912:Prostomidae
1899:Oedemeridae
1875:Mordellidae
1793:Lymexylidae
1785:Lymexyloidea
1735:Attelabidae
1727:Anthribidae
1685:Smicripidae
1649:Phalacridae
1633:Nitidulidae
1628:Myraboliidae
1620:Monotomidae
1611:
1592:Kateretidae
1569:Cyclaxyridae
1529:
1526:Agapythidae
1490:Latridiidae
1440:Cerylonidae
1431:
1427:Anamorphidae
1361:Phycosecidae
1352:
1335:
1332:Metaxinidae
1285:Biphyllidae
1276:
1245:Orsodacnidae
1202:Cucujiformia
1135:Dermestidae
1005:
944:Amphizoidae
889:
869:
855:Archostemata
839:Holometabola
837:Superorder:
831:Infraclass:
735:
731:
721:
696:
692:
681:
673:
669:
668:Pp. 411-422
629:
621:
559:
548:
540:
532:
524:
516:
508:
500:
492:
484:
476:
468:
460:
452:
444:
438:Leptoglyphus
436:
430:Erotylathris
428:
420:
412:
404:
396:
388:
380:
372:
364:
356:
348:
340:
319:
295:
282:
261:
257:
245:
244:
238:Anommatidae
237:
216:
197:Cucujiformia
193:Infraorder:
109:
18:
2862:iNaturalist
2720:Wikispecies
2618:Hydraenidae
2592:Spercheidae
2579:Hydrochidae
2543:Synteliidae
2527:Histeridae
2519:Histeroidea
2469:Passalidae
2437:Glaresidae
2294:Throscidae
2254:(fireflies)
2252:Lampyridae
2239:Eucnemidae
2231:Elateridae
2191:Elateroidea
2007:Zopheridae
1891:Mycteridae
1822:Anthicidae
1693:Sphindidae
1677:Silvanidae
1587:Hobartiidae
1574:Erotylidae
1503:Mycetaeidae
1498:Murmidiidae
1482:Euxestidae
1376:Rentoniidae
1250:Oxypeltidae
1235:Disteniidae
1068:Lepiceridae
997:Hygrobiidae
989:Haliplidae
973:Dytiscidae
952:Aspidytidae
478:Petalophora
446:Lithophorus
414:Deretaphrus
382:Cosmothroax
366:Bothrideres
342:Antibothrus
330:Anommatinae
279:Description
111:Bothrideres
2985:Categories
2639:Silphidae
2631:Ptiliidae
2623:Leiodidae
2610:Agyrtidae
2453:Lucanidae
2332:Scirtoidea
2268:Omethidae
2247:Jurasaidae
2096:Dryopidae
2067:Byrrhidae
2059:Byrrhoidea
1751:Brentidae
1556:Cucujidae
1538:Boganiidae
1518:Cucujoidea
1391:Thymalidae
1381:Rhadalidae
1324:Melyridae
1293:Byturidae
1255:Vesperidae
1164:Ptiniidae
1014:Noteridae
981:Gyrinidae
957:Carabidae
886:Jurodidae
878:Cupedidae
825:Subclass:
815:Arthropoda
796:Coleoptera
580:References
573:Cenomanian
569:Cretaceous
561:Alveoderes
553:Grouvelle.
550:Triboderus
489:Zimmermann
470:Patroderes
465:Ganglbauer
433:Motschulsk
422:Emmaglaeus
350:Acetoderes
306:wood wasps
273:Colydiidae
183:Suborder:
177:Coleoptera
157:Arthropoda
113:bipuncatus
2493:Trogidae
2487:(scarabs)
2365:Scirtidae
2340:Clambidae
1933:Pythidae
1867:Meloidae
1814:Aderidae
1717:(weevils)
1582:Helotidae
1508:Teredidae
1422:Alexiidae
1306:Cleridae
1265:Cleroidea
1098:Polyphaga
1096:Suborder
1049:Myxophaga
1047:Suborder
1002:Meruidae
921:Suborder
903:Ommatidae
853:Suborder
827:Pterygota
807:Kingdom:
754:1984-4689
526:Roplyctus
521:Grouvelle
513:Grouvelle
494:Machlotes
486:Prolyctus
473:Ślipiński
462:Ogmoderes
425:Fairmaire
406:Dastarcus
361:Grouvelle
334:Teredidae
322:Teredinae
266:Old World
187:Polyphaga
143:Kingdom:
137:Eukaryota
2771:BugGuide
2747:BioLib:
2705:Wikidata
2260:Lycidae
2104:Elmidae
2002:Ulodidae
1838:Boridae
1743:Belidae
1344:Peltidae
937:families
923:Adephaga
833:Neoptera
813:Phylum:
809:Animalia
798:families
713:55206626
662:Archived
545:Erichson
534:Shekarus
481:Westwood
374:Chinikus
358:Asosylus
316:Taxonomy
286:antennae
231:Synonyms
222:Erichson
213:Family:
153:Phylum:
147:Animalia
133:Domain:
2711:Q967786
1851:Ciidae
1759:Caridae
821:Insecta
819:Class:
794:Extant
571:(early
542:Sosylus
454:Mabomus
269:tropics
250:beetles
173:Order:
167:Insecta
163:Class:
2942:974222
2926:NZOR:
2919:196993
2893:678362
2880:115916
2867:173345
2815:1BOTRF
2776:159985
2763:165879
935:Extant
752:
711:
638:
417:Newman
409:Walker
393:Heinze
385:Kraatz
369:Dejean
328:, and
308:, and
224:, 1845
2963:Plazi
2955:70404
2875:IRMNG
2828:11056
2750:10637
709:S2CID
449:Sharp
441:Sharp
401:Sharp
345:Sharp
2914:NCBI
2888:ITIS
2854:7774
2849:GBIF
2810:EPPO
2802:7476
2758:BOLD
750:ISSN
636:ISBN
537:Pope
529:Pope
505:Pope
497:Horn
457:Pope
377:Pope
353:Pope
35:PreꞒ
2901:NBN
2797:EoL
2789:7BK
2784:CoL
2735:AFD
740:doi
701:doi
260:or
2987::
2965::
2952::
2939::
2916::
2903::
2890::
2877::
2864::
2851::
2838::
2825::
2812::
2799::
2786::
2773::
2760::
2737::
2722::
2707::
748:.
738:.
736:41
734:.
730:.
707:.
697:40
695:.
691:.
672::
670:In
650:^
603:^
588:^
324:,
312:.
304:,
85:Pg
2402:)
2398:(
2351:)
2347:(
2321:)
2317:(
1614:)
1610:(
1532:)
1528:(
1355:)
1351:(
1338:)
1334:(
1279:)
1275:(
1008:)
1004:(
892:)
888:(
872:)
868:(
787:e
780:t
773:v
756:.
742::
715:.
703::
644:.
575:)
557:†
90:N
80:K
75:J
70:T
65:P
60:C
55:D
50:S
45:O
40:Ꞓ
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