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Carpediemonas

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substrate, while the anterior flagellum beats less rapidly and in a slow sweeping motion. Further studies by Simpson and Patterson (1999) go into greater detail about the flagella and describe the flagellar apparatus as having a third, barren basal body. Supporting the dorsal side of the cell is a microtubular fan with a microtubular root at the anterior end. On the ventral side, microtubules extending from different flagellar roots support the ventral groove. The anterior flagellum has a "9+2" axoneme. Simpson and Patterson described that in addition to the "9+2" axoneme, the posterior flagellum also has "three radiating lamellae of electron-dense material which form the central components of vanes". The first lamella arises from after the flagellar insertion and is directed ventrally. The second lamella originates opposite from the first lamella. The third lamella supports the third vane, which is located more distally and lies perpendicular to the other two vanes or lamellae. All three lamella have striations when viewed in a longitudinal section and these striations are perpendicular to the "9+2" axoneme.
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all subunits of the Ndc80 complex involved in chromosome segregation. ORC and Cdc6 are proteins in charge of replication initiation and licensing in eukaryotes, and their absence appears to indicate the existence of a non-standard and as-yet undescribed mechanism to start replication. The absence of Ndc80 complex proteins also suggest that a non-standard mechanism could be in place for chromosome attachment to microtubules for chromosome segregation.
463:. In short: replication is proposed to proceed by a Dmc1-dependent homologous recombination mechanism that does not require origins of replication and that is mediated by RNA:DNA hybrids. This hypothesis still needs to be experimentally proven. Sexual or parasexual reproduction have not been directly observed in Metamonada. However, the study confirms the conservation of key meiotic proteins in the group with the bonus finding that 449:
are even further reduced. These genomes lack the DNA replication proteins of the origin recognition complex (ORC), Cdc6, some components of the GINS complex and some subunits of polymerases delta and epsilon, as well as most structural kinetochore subunits, a microtubule plus-end tracking complex and
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also has no mitochondria, which is typical of metamonads. Instead, it has hydrogenosomes, likely derived from anaerobic mitochondria. It also contains a single Golgi dictyosome, located anteriorly, dorsally, and to the left of the flagellar apparatus. The endoplasmic reticulum in this genus is mainly
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species have homologs from the tmcB family and sperm-specific channel subunits, the latter only reported previously only in Opisthokonta and three other protists. The presence of such proteins means that further investigations are required to understand if sex occurs, and if these proteins actually
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has a longitudinal depression that spans almost the entire ventral side of the cell. It often has two unequal flagella inserting to the anterior side of the ventral groove, but may sometimes have three or four flagella. The acronematic posterior flagellum is used in feeding and to attach to
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Salas-Leiva, Dayana E.; Tromer, Eelco C.; Curtis, Bruce A.; Jerlström-Hultqvist, Jon; Kolisko, Martin; Yi, Zhenzhen; Salas-Leiva, Joan S.; Gallot-Lavallée, Lucie; Williams S., Williams S.; Kops, Geert J. P. L.; Archibald, John M.; Simpson, Alastair G. B.; Roger, Andrew J. (2021).
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and contain a longitudinal groove. However, they did not have any evidence that the non-dividing organisms had more than two flagella. The species also contained a pouch with threads that may be difficult to discern from flagella. Ekebom et al. (1996) then renamed the organism as
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reproduces, it is obvious that they do replicate their DNA and researchers have proposed a hypothesis on how replication starts. This hypothesis uses elements of processes observed in other species but takes into account the specific protein complements found in
755:"The excavate protozoan phyla Metamonada Grasse emend. (Anaeromonadea, Parabasalia, Carpediemonas, Eopharyngia) and Loukozoa emend. (Jakobea, Malawimonas): their evolutionary affinities and new higher taxa" 227:
can be found in anaerobic intertidal sediment, where it feeds on bacteria. A feature of this species is the presence of a feeding groove, a characteristic of the excavates. Like most other metamonads,
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Yubuki, Naoji; Simpson, Alastair G.B.; Leander, Brian S. (2013). "Comprehensive Ultrastructure of Kipferlia bialata Provides Evidence for Character Evolution within the Fornicata (Excavata)".
421:-like organisms (CLOs) have allowed for the better understanding of the evolution of anaerobic excavates by studying their cytoskeletal traits and modified mitochondria. An example of a 239:
that are used to produce ATP. This organism has two flagella: a posterior one used for feeding on the substrate, and an anterior one that moves in a slower sweeping motion.
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Simpson, Alastair G.B.; Patterson, David J. (1999). "The ultrastructure of Carpediemonas membranifera (Eukaryota) with reference to the "excavate hypothesis"".
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found near the periphery of the cell. Around the cytoplasm, food vacuoles containing bacterial contents can be found. Also, three centrioles are present in
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Hamann, Emmo; Tegetmeyer, Halina E; Riedel, Dietmar; Littmann, Sten; Ahmerkamp, Soeren; Chen, Jianwei; Hach, Philipp F; Strous, Marc (2017-02-17).
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has revealed extensive loss of the DNA replication and segregation protein complements within Metamonada and has highlighted that the genomes of
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when it was found from substrate samples in the Great Barrier Reef and classified it as a metamonad. Additionally, a metabolic relationship of
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Ekebom, J.; Patterson, D.J.; Vørs, N. (1996). "Heterotrophic Flagellates from Coral Reef Sediments (Great Barrier Reef, Australia)".
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J. Ekebom; D. J. Patterson; N. Vørs (1996). "Heterotrophic flagellates from coral reef sediments (Great Barrier Reef, Australia)".
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contains a single ovate nucleus, located anteriorly in the cell. The nucleolus can also be found subcentrally within the nucleus.
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is the first known eukaryote to possess such drastically altered DNA replication and segregation machineries. Since
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is classified as an excavate because it has the characteristic feeding groove of the group. Within the excavates,
4212: 819:"Genomic analysis finds no evidence of canonical eukaryotic DNA processing complexes in a free-living protist" 251:
is a member of the metamonads, it is unusual in the sense that it is free-living and has three basal bodies.
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for incompletely digested organic material and the oxidation of various biomolecules. On the other hand,
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was found to release biomolecules that have been predigested. Prokaryotic communities would rely on
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was first discovered by Larsen and Patterson (1990) who identified it as a previously unidentified
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as organisms with a size of approximately 5 μm long (with a range of 4–7.5 μm).
17: 698:"Syntrophic linkage between predatory Carpediemonas and specific prokaryotic populations" 834: 3699: 3610: 3320: 3275: 3152: 3140: 3109: 2629: 2609: 2534: 2235: 2073: 2063: 2027: 2002: 1691: 1482: 1383: 1305: 1114: 946: 851: 818: 730: 697: 303: 676: 584: 540: 504: 4201: 3375: 3266: 3183: 2798: 2738: 2589: 2562: 2444: 2109: 1908: 1900: 1730: 1651: 1591: 1502: 1476: 1205: 236: 232: 3579: 3366: 3079: 2776: 2733: 2728: 2379: 2354: 2142: 1969: 1949: 1889: 1793: 1766: 1713: 1671: 1638: 1413: 1191: 1130: 425:-like organism that was used to study the evolutionary history within excavates is 366:
intertidal sediments, where it feeds on bacteria. It can be found co-existing with
294: 160: 4142: 4060: 3952: 3899: 3870: 3801: 622: 247:-like organisms are used in researching the evolution within excavates. Although 4151: 4069: 3981: 3961: 3879: 3810: 3595: 3462: 3257: 3195: 3074: 2771: 2754: 2710: 2701: 2359: 2033: 1959: 1811: 1775: 1755: 1703: 1677: 1567: 1515: 1488: 1470: 1443: 1210: 1079: 1068: 842: 3525: 3190: 2763: 2691: 2646: 2455: 2114: 1938: 1929: 1914: 1782: 1719: 1578: 1539: 1534: 1420: 1337: 1232: 1144: 322: 277: 208: 780: 721: 630: 592: 548: 3472: 3449: 3396: 3356: 3163: 3050: 3043: 2929: 2897: 2602: 2364: 2013: 1954: 1662: 1627: 1585: 1562: 1544: 1431: 1367: 1242: 1119: 1062: 1049: 901: 307: 79: 69: 59: 870: 860: 788: 739: 713: 638: 771: 754: 4136: 4054: 3946: 3864: 3795: 3403: 3283: 3243: 2892: 2597: 2404: 2346: 2274: 2191: 2020: 1982: 1883: 1860: 1407: 1402: 1394: 1324: 1199: 1074: 1057: 1035: 985: 216: 212: 4089: 3994: 3912: 3830: 2337: 2316: 2136: 1871: 1708: 1554: 1461: 1450: 1314: 1220: 1176: 1084: 803:"Microbe somehow survives without key proteins for replicating its DNA" 178: 1526: 1256: 1158: 1100: 3772: 2096: 1736: 275:, indicating a unicellular organism. The organism was named after 219:
eukaryote that was first discovered in substrate samples from the
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International Journal of Systematic and Evolutionary Microbiology
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participate during sex and what their role could be. In short,
172:(Larsen & Patterson 1990) Ekebom, Patterson & Vørs 1996 2160: 4126: 4044: 3936: 3854: 3785: 3652: 3603: 3594: 3518: 3493: 3471: 3447: 3374: 3365: 3354: 3319: 3312: 3305: 3274: 3265: 3256: 3241: 3151: 3120: 3067: 3060: 3035: 3014: 2963: 2937: 2928: 2850: 2824: 2797: 2762: 2753: 2709: 2700: 2689: 2588: 2552: 2545: 2524: 2503: 2496: 2471: 2464: 2453: 2442: 2412: 2403: 2396: 2345: 2336: 2329: 2314: 2095: 2072: 2049: 1999: 1968: 1928: 1899: 1870: 1858: 1765: 1729: 1690: 1661: 1637: 1624: 1553: 1525: 1501: 1460: 1430: 1393: 1380: 1350: 1323: 1303: 1241: 1219: 1173: 1143: 1128: 1097: 1048: 1033: 417:is assigned to the fornicates. In the fornicates, 302:. Larsen and Patterson treated this organism as a 433:DNA replication, chromosome segregation, and sex 2176: 886: 243:is assigned to the fornicates, where similar 8: 3773: 3600: 3477: 3455: 3444: 3371: 3362: 3316: 3309: 3271: 3262: 3253: 3249: 3064: 2934: 2803: 2759: 2706: 2697: 2549: 2500: 2468: 2461: 2450: 2409: 2400: 2342: 2333: 2326: 2322: 2183: 2169: 2161: 2046: 1876: 1867: 1855: 1762: 1726: 1696: 1643: 1634: 1621: 1507: 1436: 1390: 1377: 1360: 1347: 1320: 1300: 1296: 1225: 1183: 1140: 1136: 1107: 1045: 1041: 1030: 1026: 893: 879: 871: 306:, because it would occasionally have four 31: 850: 770: 729: 263:originates from three Latin roots, with 235:to produce energy. Instead, it contains 481: 2151:denotes groups with over 1,000 species 472:are intriguing microbial eukaryotes. 271:meaning "the day", and the suffix of 7: 604: 602: 566: 564: 562: 560: 558: 522: 520: 518: 516: 514: 343:relies on the prokaryotic organism, 677:"Taxonomy – It Came from the Pond" 154:Ekebom, Patterson & Vørs, 1996 25: 753:Cavalier-Smith, T. (2003-11-01). 372:in these anaerobic environments. 215:. This organism is a unicellular 573:European Journal of Protistology 46: 380:Ekebom et al. (1996) describes 1630:      1626:      1386:      1382:      1: 585:10.1016/s0932-4739(99)80044-3 541:10.1016/s0003-9365(96)80013-3 505:10.1016/S0003-9365(96)80013-3 623:10.1016/j.protis.2013.02.002 231:does not rely on an aerobic 211:, and belongs to the group 4229: 843:10.1038/s41467-021-26077-2 314:Carpediemonas membranifera 168:Carpediemonas membranifera 18:Carpediemonas membranifera 3480: 3458: 3443: 3252: 2806: 2325: 2198: 2127: 1879: 1854: 1699: 1646: 1620: 1510: 1439: 1376: 1363: 1346: 1299: 1295: 1228: 1186: 1139: 1110: 1044: 1029: 1025: 909: 529:Archiv für Protistenkunde 492:Archiv für Protistenkunde 325:communities was found in 300:Percolomonas membranifera 184: 177: 166: 159: 43:Scientific classification 41: 34: 1356:     1352:     714:10.1038/ismej.2016.197 298:and provided the name 823:Nature Communications 772:10.1099/ijs.0.02548-0 437:A recent study using 369:Cafeteria marsupialis 3693:Spirotrichonymphidae 3689:Spirotrichonymphida 439:comparative genomics 328:Carpediemonas frisia 3683:Hypotrichomonadidae 3679:Hypotrichomonadida 2028:Gunflint microbiota 1789:Klebsormidiophyceae 1573:Hyphochytriomycetes 835:2021NatCo..12.6003S 355:Habitat and ecology 345:Deltaproteobacteria 3740:Tritrichomonadidae 3730:Monocercomonadidae 3721:Tritrichomonadida 3627:Spirotrichosomidae 2886:Trypanoplasmatidae 2851:Metakinetoplastina 2831:Prokinetoplastida 2578:Psalteriomonadidae 2568:Paravahlkampfiidae 1799:Coleochaetophyceae 1179:    1175:    1103:    1099:    657:comenius.susqu.edu 349:hydrogen oxidizing 221:Great Barrier Reef 4195: 4194: 4167:Open Tree of Life 4016:Open Tree of Life 3886:Carpediemonadidae 3856:Carpediemonadidae 3779:Taxon identifiers 3770: 3769: 3766: 3765: 3762: 3761: 3758: 3757: 3754: 3753: 3750: 3749: 3735:Simplicimonadidae 3705:Honigbergiellidae 3590: 3589: 3508:Carpediemonadidae 3439: 3438: 3429:Streblomastigidae 3387:Paratrimastigidae 3383:Paratrimastigida 3350: 3349: 3346: 3345: 3342: 3341: 3301: 3300: 3297: 3296: 3237: 3236: 3233: 3232: 3229: 3228: 3225: 3224: 3221: 3220: 3201:Euglenamorphaceae 3174:Euglenomonadaceae 3170:Euglenomonadales 2924: 2923: 2920: 2919: 2825:Prokinetoplastina 2749: 2748: 2685: 2684: 2681: 2680: 2677: 2676: 2667:Stephanopogonidae 2535:Neovahlkampfiidae 2531:Neovahlkampfiida 2492: 2491: 2482:Pharyngomonadidae 2478:Pharyngomonadida 2472:Pharyngomonadidea 2438: 2437: 2434: 2433: 2430: 2429: 2392: 2391: 2388: 2387: 2158: 2157: 2123: 2122: 2059:Spironematellidae 2040:Meteora sporadica 1995: 1994: 1991: 1990: 1924: 1923: 1850: 1849: 1846: 1845: 1842: 1841: 1838: 1837: 1834: 1833: 1830: 1829: 1826: 1825: 1822: 1821: 1750:Prasinodermophyta 1686: 1685: 1616: 1615: 1612: 1611: 1608: 1607: 1604: 1603: 1600: 1599: 1497: 1496: 1333:Centroplasthelida 1291: 1290: 1287: 1286: 1283: 1282: 1279: 1278: 1275: 1274: 1271: 1270: 1267: 1266: 1169: 1168: 1093: 1092: 427:Kipferlia bialata 267:meaning "seize", 200: 199: 155: 138:Carpediemonadidae 129: 113: 97: 27:Genus of protists 16:(Redirected from 4220: 4213:Metamonad genera 4188: 4187: 4175: 4174: 4162: 4161: 4160: 4147: 4146: 4145: 4119: 4118: 4106: 4105: 4093: 4092: 4080: 4079: 4078: 4065: 4064: 4063: 4037: 4036: 4024: 4023: 4011: 4010: 3998: 3997: 3985: 3984: 3972: 3971: 3970: 3968:Carpediemonadida 3957: 3956: 3955: 3938:Carpediemonadida 3929: 3928: 3916: 3915: 3903: 3902: 3890: 3889: 3888: 3875: 3874: 3873: 3847: 3846: 3834: 3833: 3821: 3820: 3819: 3806: 3805: 3804: 3774: 3632:Staurojoeninidae 3601: 3580:Retortamonadidae 3503:Carpediemonadida 3478: 3456: 3445: 3424:Saccinobaculidae 3372: 3363: 3317: 3310: 3272: 3263: 3254: 3250: 3206:Euglenocapsaceae 3090:Heteronemataceae 3086:Heteronematales 3065: 2935: 2910:Trypanosomatidae 2871:Rhynchomonadidae 2835:Ichthyobodonidae 2804: 2760: 2707: 2698: 2662:Percolomonadidae 2573:Plaesiobystridae 2550: 2501: 2469: 2462: 2451: 2422:Tsukubamonadidae 2410: 2401: 2343: 2334: 2327: 2323: 2227: 2207: 2185: 2178: 2171: 2162: 2105:Collodictyonidae 2051:Hemimastigophora 2047: 2006: 1877: 1868: 1864: 1856: 1805:Zygnematophyceae 1763: 1743: 1727: 1697: 1644: 1635: 1631: 1622: 1508: 1437: 1391: 1387: 1378: 1361: 1357: 1348: 1321: 1308: 1301: 1297: 1251:Choanoflagellata 1226: 1184: 1180: 1141: 1137: 1133: 1108: 1104: 1046: 1042: 1038: 1031: 1027: 938: 918: 895: 888: 881: 872: 865: 864: 854: 813: 807: 806: 799: 793: 792: 774: 765:(6): 1741–1758. 750: 744: 743: 733: 708:(5): 1205–1217. 702:The ISME Journal 693: 687: 686: 684: 683: 673: 667: 666: 664: 663: 653:"CARPEDIEMONADA" 649: 643: 642: 606: 597: 596: 568: 553: 552: 535:(3–4): 251–272. 524: 509: 508: 499:(3–4): 251–272. 486: 362:can be found in 285:History of study 153: 125: 122:Carpediemonadida 109: 93: 51: 50: 32: 21: 4228: 4227: 4223: 4222: 4221: 4219: 4218: 4217: 4198: 4197: 4196: 4191: 4183: 4178: 4170: 4165: 4158:Carpediemonadia 4156: 4155: 4150: 4141: 4140: 4135: 4128:Carpediemonadia 4122: 4114: 4109: 4101: 4096: 4088: 4083: 4076:Carpediemonadea 4074: 4073: 4068: 4059: 4058: 4053: 4046:Carpediemonadea 4040: 4032: 4027: 4019: 4014: 4006: 4001: 3993: 3988: 3980: 3975: 3966: 3965: 3960: 3951: 3950: 3945: 3932: 3924: 3919: 3911: 3906: 3898: 3893: 3884: 3883: 3878: 3869: 3868: 3863: 3850: 3842: 3837: 3829: 3824: 3815: 3814: 3809: 3800: 3799: 3794: 3781: 3771: 3746: 3715:Trichomonadidae 3710:Tricercomitidae 3664:Cristamonadida 3648: 3642:Trichonymphidae 3637:Teratonymphidae 3586: 3575:Chilomastigidae 3565:Retortamonadida 3514: 3495:Carpediemonadea 3489: 3467: 3452: 3435: 3359: 3338: 3329:Ancyromonadidae 3293: 3289:Malawimonadidae 3246: 3217: 3147: 3116: 3056: 3031: 3010: 3004:Sphenomonadidae 2994:Sphenomonadidae 2959: 2953:Sphenomonadidae 2948:Petalomonadidae 2944:Petalomonadida 2916: 2905:Trypanosomatida 2846: 2820: 2816:Trypanophididae 2811:Bordnamonadidae 2793: 2782:Eupelagonemidae 2745: 2694: 2673: 2647:Percolomonadida 2584: 2541: 2525:Neovahlkampfiea 2520: 2488: 2458: 2447: 2426: 2414:Tsukubamonadida 2384: 2319: 2310: 2309: 2225: 2203: 2194: 2189: 2159: 2154: 2119: 2091: 2082:Ancyromonadidae 2068: 2045: 2000: 1987: 1978:Malawimonadidae 1964: 1920: 1895: 1859: 1818: 1761: 1734: 1733: 1725: 1682: 1657: 1625: 1596: 1549: 1521: 1493: 1456: 1426: 1381: 1372: 1351: 1342: 1319: 1304: 1263: 1237: 1215: 1174: 1165: 1129: 1124: 1098: 1089: 1034: 1021: 1020: 936: 914: 905: 899: 869: 868: 815: 814: 810: 801: 800: 796: 752: 751: 747: 695: 694: 690: 681: 679: 675: 674: 670: 661: 659: 651: 650: 646: 608: 607: 600: 570: 569: 556: 526: 525: 512: 488: 487: 483: 478: 443:C. membranifera 435: 408: 378: 357: 287: 257: 193:C. membranifera 173: 170: 152: 124: 108: 106:Carpediemonadea 92: 90:Carpediemonadia 45: 28: 23: 22: 15: 12: 11: 5: 4226: 4224: 4216: 4215: 4210: 4200: 4199: 4193: 4192: 4190: 4189: 4176: 4163: 4148: 4132: 4130: 4124: 4123: 4121: 4120: 4107: 4094: 4081: 4066: 4050: 4048: 4042: 4041: 4039: 4038: 4025: 4012: 3999: 3986: 3973: 3958: 3942: 3940: 3934: 3933: 3931: 3930: 3917: 3904: 3891: 3876: 3860: 3858: 3852: 3851: 3849: 3848: 3835: 3822: 3807: 3791: 3789: 3783: 3782: 3777: 3768: 3767: 3764: 3763: 3760: 3759: 3756: 3755: 3752: 3751: 3748: 3747: 3745: 3744: 3743: 3742: 3737: 3732: 3727: 3725:Dientamoebidae 3719: 3718: 3717: 3712: 3707: 3700:Trichomonadida 3697: 3696: 3695: 3687: 3686: 3685: 3677: 3676: 3675: 3673:Devescovinidae 3670: 3662: 3660:Hexamastigidae 3656: 3654: 3650: 3649: 3647: 3646: 3645: 3644: 3639: 3634: 3629: 3624: 3622:Hoplonymphidae 3619: 3617:Burmanymphidae 3611:Trichonymphida 3607: 3605: 3598: 3592: 3591: 3588: 3587: 3585: 3584: 3583: 3582: 3577: 3572: 3570:Caviomonadidae 3562: 3561: 3560: 3550: 3549: 3548: 3543: 3541:Spironucleidae 3538: 3533: 3522: 3520: 3516: 3515: 3513: 3512: 3511: 3510: 3499: 3497: 3491: 3490: 3488: 3487: 3481: 3475: 3469: 3468: 3466: 3465: 3463:Anaeramoebidae 3459: 3453: 3448: 3441: 3440: 3437: 3436: 3434: 3433: 3432: 3431: 3426: 3421: 3419:Pyrsonymphidae 3416: 3414:Polymastigidae 3411: 3401: 3400: 3399: 3391: 3390: 3389: 3380: 3378: 3369: 3360: 3355: 3352: 3351: 3348: 3347: 3344: 3343: 3340: 3339: 3337: 3336: 3334:Planomonadidae 3331: 3325: 3323: 3321:Ancyromonadida 3314: 3307: 3303: 3302: 3299: 3298: 3295: 3294: 3292: 3291: 3286: 3280: 3278: 3276:Malawimonadida 3269: 3260: 3247: 3242: 3239: 3238: 3235: 3234: 3231: 3230: 3227: 3226: 3223: 3222: 3219: 3218: 3216: 3215: 3214: 3213: 3208: 3203: 3198: 3188: 3187: 3186: 3178: 3177: 3176: 3168: 3167: 3166: 3157: 3155: 3153:Euglenophyceae 3149: 3148: 3146: 3145: 3144: 3143: 3141:Peranemataceae 3135: 3134: 3133: 3124: 3122: 3118: 3117: 3115: 3114: 3113: 3112: 3110:Neometanemidae 3107: 3102: 3094: 3093: 3092: 3084: 3083: 3082: 3071: 3069: 3062: 3058: 3057: 3055: 3054: 3047: 3039: 3037: 3033: 3032: 3030: 3029: 3028: 3027: 3025:Entosiphonidae 3021:Entosiphonida 3018: 3016: 3012: 3011: 3009: 3008: 3007: 3006: 2998: 2997: 2996: 2990:Heterostavida 2988: 2987: 2986: 2984:Lentomonadidae 2981: 2976: 2967: 2965: 2961: 2960: 2958: 2957: 2956: 2955: 2950: 2941: 2939: 2932: 2926: 2925: 2922: 2921: 2918: 2917: 2915: 2914: 2913: 2912: 2902: 2901: 2900: 2890: 2889: 2888: 2883: 2875: 2874: 2873: 2868: 2863: 2854: 2852: 2848: 2847: 2845: 2844: 2843: 2842: 2837: 2828: 2826: 2822: 2821: 2819: 2818: 2813: 2807: 2801: 2795: 2794: 2792: 2791: 2790: 2789: 2784: 2779: 2768: 2766: 2757: 2751: 2750: 2747: 2746: 2744: 2743: 2742: 2741: 2736: 2726: 2725: 2724: 2715: 2713: 2704: 2695: 2690: 2687: 2686: 2683: 2682: 2679: 2678: 2675: 2674: 2672: 2671: 2670: 2669: 2664: 2659: 2654: 2644: 2643: 2642: 2634: 2633: 2632: 2630:Vahlkampfiidae 2624: 2623: 2622: 2617: 2607: 2606: 2605: 2594: 2592: 2586: 2585: 2583: 2582: 2581: 2580: 2575: 2570: 2565: 2556: 2554: 2547: 2543: 2542: 2540: 2539: 2538: 2537: 2528: 2526: 2522: 2521: 2519: 2518: 2517: 2516: 2507: 2505: 2498: 2494: 2493: 2490: 2489: 2487: 2486: 2485: 2484: 2475: 2473: 2466: 2465:Pharyngomonada 2459: 2454: 2448: 2443: 2440: 2439: 2436: 2435: 2432: 2431: 2428: 2427: 2425: 2424: 2418: 2416: 2407: 2398: 2394: 2393: 2390: 2389: 2386: 2385: 2383: 2382: 2377: 2372: 2367: 2362: 2357: 2351: 2349: 2340: 2331: 2320: 2315: 2312: 2311: 2308: 2307: 2306: 2305: 2304: 2303: 2300:Mesomycetozoea 2297: 2292: 2282: 2272: 2271: 2270: 2265: 2260: 2255: 2250: 2249: 2248: 2236:Diaphoretickes 2233: 2228: 2223: 2218: 2213: 2208: 2200: 2199: 2196: 2195: 2190: 2188: 2187: 2180: 2173: 2165: 2156: 2155: 2153: 2152: 2146: 2139: 2135:excluded from 2128: 2125: 2124: 2121: 2120: 2118: 2117: 2112: 2107: 2101: 2099: 2093: 2092: 2090: 2089: 2087:Planomonadidae 2084: 2078: 2076: 2074:Ancyromonadida 2070: 2069: 2067: 2066: 2064:Paramastigidae 2061: 2055: 2053: 2044: 2043: 2036: 2030: 2024: 2016: 2009: 2007: 2003:Incertae sedis 1997: 1996: 1993: 1992: 1989: 1988: 1986: 1985: 1980: 1974: 1972: 1966: 1965: 1963: 1962: 1957: 1952: 1947: 1942: 1934: 1932: 1926: 1925: 1922: 1921: 1919: 1918: 1911: 1905: 1903: 1897: 1896: 1894: 1893: 1886: 1880: 1874: 1865: 1852: 1851: 1848: 1847: 1844: 1843: 1840: 1839: 1836: 1835: 1832: 1831: 1828: 1827: 1824: 1823: 1820: 1819: 1817: 1816: 1808: 1801: 1796: 1791: 1786: 1779: 1771: 1769: 1760: 1759: 1752: 1746: 1744: 1724: 1723: 1716: 1711: 1706: 1700: 1694: 1692:Archaeplastida 1688: 1687: 1684: 1683: 1681: 1680: 1675: 1667: 1665: 1659: 1658: 1656: 1655: 1647: 1641: 1632: 1618: 1617: 1614: 1613: 1610: 1609: 1606: 1605: 1602: 1601: 1598: 1597: 1595: 1594: 1589: 1582: 1575: 1570: 1565: 1559: 1557: 1551: 1550: 1548: 1547: 1542: 1537: 1531: 1529: 1523: 1522: 1520: 1519: 1511: 1505: 1499: 1498: 1495: 1494: 1492: 1491: 1486: 1483:Dinoflagellata 1479: 1477:Chrompodellida 1474: 1466: 1464: 1458: 1457: 1455: 1454: 1447: 1440: 1434: 1428: 1427: 1425: 1424: 1417: 1410: 1405: 1399: 1397: 1388: 1374: 1373: 1371: 1370: 1364: 1358: 1344: 1343: 1341: 1340: 1335: 1329: 1327: 1318: 1317: 1311: 1309: 1306:Diaphoretickes 1293: 1292: 1289: 1288: 1285: 1284: 1281: 1280: 1277: 1276: 1273: 1272: 1269: 1268: 1265: 1264: 1262: 1261: 1253: 1247: 1245: 1239: 1238: 1236: 1235: 1229: 1223: 1217: 1216: 1214: 1213: 1208: 1203: 1195: 1187: 1181: 1171: 1170: 1167: 1166: 1164: 1163: 1155: 1149: 1147: 1134: 1126: 1125: 1123: 1122: 1117: 1111: 1105: 1095: 1094: 1091: 1090: 1088: 1087: 1082: 1077: 1072: 1065: 1060: 1054: 1052: 1039: 1023: 1022: 1019: 1018: 1017: 1016: 1015: 1014: 1011:Mesomycetozoea 1008: 1003: 993: 983: 982: 981: 976: 971: 966: 961: 960: 959: 947:Diaphoretickes 944: 939: 934: 929: 924: 919: 911: 910: 907: 906: 904:classification 900: 898: 897: 890: 883: 875: 867: 866: 808: 794: 745: 688: 668: 644: 617:(3): 423–439. 598: 579:(4): 353–370. 554: 510: 480: 479: 477: 474: 434: 431: 407: 404: 377: 374: 356: 353: 304:heterolobosean 286: 283: 256: 253: 237:hydrogenosomes 198: 197: 196: 195: 190: 182: 181: 175: 174: 171: 164: 163: 157: 156: 145: 141: 140: 135: 131: 130: 127:Cavalier-Smith 119: 115: 114: 111:Cavalier-Smith 103: 99: 98: 95:Cavalier-Smith 87: 83: 82: 77: 73: 72: 67: 63: 62: 57: 53: 52: 39: 38: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 4225: 4214: 4211: 4209: 4206: 4205: 4203: 4186: 4181: 4177: 4173: 4168: 4164: 4159: 4153: 4149: 4144: 4138: 4134: 4133: 4131: 4129: 4125: 4117: 4112: 4108: 4104: 4099: 4095: 4091: 4086: 4082: 4077: 4071: 4067: 4062: 4056: 4052: 4051: 4049: 4047: 4043: 4035: 4030: 4026: 4022: 4017: 4013: 4009: 4004: 4000: 3996: 3991: 3987: 3983: 3978: 3974: 3969: 3963: 3959: 3954: 3948: 3944: 3943: 3941: 3939: 3935: 3927: 3922: 3918: 3914: 3909: 3905: 3901: 3896: 3892: 3887: 3881: 3877: 3872: 3866: 3862: 3861: 3859: 3857: 3853: 3845: 3840: 3836: 3832: 3827: 3823: 3818: 3817:Carpediemonas 3812: 3808: 3803: 3797: 3793: 3792: 3790: 3788: 3787:Carpediemonas 3784: 3780: 3775: 3741: 3738: 3736: 3733: 3731: 3728: 3726: 3723: 3722: 3720: 3716: 3713: 3711: 3708: 3706: 3703: 3702: 3701: 3698: 3694: 3691: 3690: 3688: 3684: 3681: 3680: 3678: 3674: 3671: 3669: 3668:Calonymphidae 3666: 3665: 3663: 3661: 3658: 3657: 3655: 3653:Trichomonadea 3651: 3643: 3640: 3638: 3635: 3633: 3630: 3628: 3625: 3623: 3620: 3618: 3614: 3613: 3612: 3609: 3608: 3606: 3604:Trichonymphea 3602: 3599: 3597: 3593: 3581: 3578: 3576: 3573: 3571: 3568: 3567: 3566: 3563: 3559: 3556: 3555: 3554: 3551: 3547: 3544: 3542: 3539: 3537: 3534: 3532: 3529: 3528: 3527: 3526:Diplomonadida 3524: 3523: 3521: 3517: 3509: 3506: 3505: 3504: 3501: 3500: 3498: 3496: 3492: 3486: 3483: 3482: 3479: 3476: 3474: 3470: 3464: 3461: 3460: 3457: 3454: 3451: 3446: 3442: 3430: 3427: 3425: 3422: 3420: 3417: 3415: 3412: 3410: 3407: 3406: 3405: 3402: 3398: 3397:Trimastigidae 3395: 3394: 3393:Trimastigida 3392: 3388: 3385: 3384: 3382: 3381: 3379: 3377: 3376:Anaeromonadea 3373: 3370: 3368: 3364: 3361: 3358: 3353: 3335: 3332: 3330: 3327: 3326: 3324: 3322: 3318: 3315: 3311: 3308: 3304: 3290: 3287: 3285: 3282: 3281: 3279: 3277: 3273: 3270: 3268: 3267:Malawimonadea 3264: 3261: 3259: 3255: 3251: 3248: 3245: 3240: 3212: 3209: 3207: 3204: 3202: 3199: 3197: 3194: 3193: 3192: 3189: 3185: 3184:Eutreptiaceae 3182: 3181: 3180:Eutreptiales 3179: 3175: 3172: 3171: 3169: 3165: 3162: 3161: 3159: 3158: 3156: 3154: 3150: 3142: 3139: 3138: 3137:Peranematida 3136: 3132: 3131:Teloproctidae 3129: 3128: 3127:Acroglissida 3126: 3125: 3123: 3119: 3111: 3108: 3106: 3103: 3101: 3098: 3097: 3096:Natomonadida 3095: 3091: 3088: 3087: 3085: 3081: 3078: 3077: 3076: 3073: 3072: 3070: 3066: 3063: 3059: 3053: 3052: 3048: 3046: 3045: 3041: 3040: 3038: 3034: 3026: 3023: 3022: 3020: 3019: 3017: 3013: 3005: 3002: 3001: 2999: 2995: 2992: 2991: 2989: 2985: 2982: 2980: 2977: 2975: 2972: 2971: 2969: 2968: 2966: 2962: 2954: 2951: 2949: 2946: 2945: 2943: 2942: 2940: 2938:Petalomonadea 2936: 2933: 2931: 2927: 2911: 2908: 2907: 2906: 2903: 2899: 2896: 2895: 2894: 2891: 2887: 2884: 2882: 2881:Parabodonidae 2879: 2878: 2877:Parabodonida 2876: 2872: 2869: 2867: 2864: 2862: 2861:Allobodonidae 2859: 2858: 2856: 2855: 2853: 2849: 2841: 2840:Perkinselidae 2838: 2836: 2833: 2832: 2830: 2829: 2827: 2823: 2817: 2814: 2812: 2809: 2808: 2805: 2802: 2800: 2799:Kinetoplastea 2796: 2788: 2787:Hemistasiidae 2785: 2783: 2780: 2778: 2775: 2774: 2773: 2770: 2769: 2767: 2765: 2761: 2758: 2756: 2752: 2740: 2739:Postgaardidae 2737: 2735: 2732: 2731: 2730: 2727: 2723: 2720: 2719: 2717: 2716: 2714: 2712: 2708: 2705: 2703: 2699: 2696: 2693: 2688: 2668: 2665: 2663: 2660: 2658: 2655: 2653: 2650: 2649: 2648: 2645: 2641: 2638: 2637: 2635: 2631: 2628: 2627: 2625: 2621: 2618: 2616: 2613: 2612: 2611: 2608: 2604: 2601: 2600: 2599: 2596: 2595: 2593: 2591: 2590:Heterolobosea 2587: 2579: 2576: 2574: 2571: 2569: 2566: 2564: 2563:Gruberellidae 2561: 2560: 2559:Lyromonadida 2558: 2557: 2555: 2551: 2548: 2544: 2536: 2533: 2532: 2530: 2529: 2527: 2523: 2515: 2514:Selenaionidae 2512: 2511: 2510:Selenaionida 2509: 2508: 2506: 2502: 2499: 2495: 2483: 2480: 2479: 2477: 2476: 2474: 2470: 2467: 2463: 2460: 2457: 2452: 2449: 2446: 2445:Discicristata 2441: 2423: 2420: 2419: 2417: 2415: 2411: 2408: 2406: 2402: 2399: 2397:Tsukubamonada 2395: 2381: 2378: 2376: 2373: 2371: 2370:Moramonadidae 2368: 2366: 2363: 2361: 2358: 2356: 2353: 2352: 2350: 2348: 2344: 2341: 2339: 2335: 2332: 2328: 2324: 2321: 2318: 2313: 2301: 2298: 2296: 2293: 2291: 2288: 2287: 2286: 2283: 2281: 2278: 2277: 2276: 2273: 2269: 2266: 2264: 2263:Stramenopiles 2261: 2259: 2256: 2254: 2251: 2247: 2244: 2243: 2242: 2239: 2238: 2237: 2234: 2232: 2229: 2226:(major groups 2224: 2222: 2219: 2217: 2214: 2212: 2209: 2206: 2202: 2201: 2197: 2193: 2186: 2181: 2179: 2174: 2172: 2167: 2166: 2163: 2150: 2147: 2144: 2140: 2138: 2134: 2131:¹traditional 2130: 2129: 2126: 2116: 2113: 2111: 2110:Mantamonadida 2108: 2106: 2103: 2102: 2100: 2098: 2094: 2088: 2085: 2083: 2080: 2079: 2077: 2075: 2071: 2065: 2062: 2060: 2057: 2056: 2054: 2052: 2048: 2042: 2041: 2037: 2035: 2031: 2029: 2025: 2023: 2022: 2017: 2015: 2011: 2010: 2008: 2005: 2004: 1998: 1984: 1981: 1979: 1976: 1975: 1973: 1971: 1967: 1961: 1958: 1956: 1953: 1951: 1948: 1946: 1943: 1941: 1940: 1936: 1935: 1933: 1931: 1927: 1917: 1916: 1912: 1910: 1909:Heterolobosea 1907: 1906: 1904: 1902: 1901:Discicristata 1898: 1892: 1891: 1887: 1885: 1882: 1881: 1878: 1875: 1873: 1869: 1866: 1862: 1857: 1853: 1814: 1813: 1809: 1807: 1806: 1802: 1800: 1797: 1795: 1792: 1790: 1787: 1785: 1784: 1780: 1778: 1777: 1773: 1772: 1770: 1768: 1764: 1758: 1757: 1753: 1751: 1748: 1747: 1745: 1741: 1740: 1732: 1731:Viridiplantae 1728: 1722: 1721: 1717: 1715: 1712: 1710: 1707: 1705: 1702: 1701: 1698: 1695: 1693: 1689: 1679: 1676: 1674: 1673: 1669: 1668: 1666: 1664: 1660: 1654: 1653: 1652:Microheliella 1649: 1648: 1645: 1642: 1640: 1636: 1633: 1629: 1623: 1619: 1593: 1592:Actinophryida 1590: 1588: 1587: 1583: 1581: 1580: 1576: 1574: 1571: 1569: 1566: 1564: 1561: 1560: 1558: 1556: 1552: 1546: 1543: 1541: 1538: 1536: 1533: 1532: 1530: 1528: 1524: 1518: 1517: 1513: 1512: 1509: 1506: 1504: 1503:Stramenopiles 1500: 1490: 1487: 1485: 1484: 1480: 1478: 1475: 1473: 1472: 1468: 1467: 1465: 1463: 1459: 1453: 1452: 1448: 1445: 1442: 1441: 1438: 1435: 1433: 1429: 1423: 1422: 1418: 1416: 1415: 1411: 1409: 1406: 1404: 1401: 1400: 1398: 1396: 1392: 1389: 1385: 1379: 1375: 1369: 1366: 1365: 1362: 1359: 1355: 1349: 1345: 1339: 1336: 1334: 1331: 1330: 1328: 1326: 1322: 1316: 1313: 1312: 1310: 1307: 1302: 1298: 1294: 1259: 1258: 1254: 1252: 1249: 1248: 1246: 1244: 1240: 1234: 1231: 1230: 1227: 1224: 1222: 1218: 1212: 1209: 1207: 1206:Ichthyosporea 1204: 1202: 1201: 1196: 1194: 1193: 1189: 1188: 1185: 1182: 1178: 1172: 1161: 1160: 1156: 1154: 1153:Rotosphaerida 1151: 1150: 1148: 1146: 1142: 1138: 1135: 1132: 1127: 1121: 1118: 1116: 1115:Apusomonadida 1113: 1112: 1109: 1106: 1102: 1096: 1086: 1083: 1081: 1078: 1076: 1073: 1071: 1070: 1066: 1064: 1061: 1059: 1056: 1055: 1053: 1051: 1047: 1043: 1040: 1037: 1032: 1028: 1024: 1012: 1009: 1007: 1004: 1002: 999: 998: 997: 994: 992: 989: 988: 987: 984: 980: 977: 975: 974:Stramenopiles 972: 970: 967: 965: 962: 958: 955: 954: 953: 950: 949: 948: 945: 943: 940: 937:(major groups 935: 933: 930: 928: 925: 923: 920: 917: 913: 912: 908: 903: 896: 891: 889: 884: 882: 877: 876: 873: 862: 858: 853: 848: 844: 840: 836: 832: 828: 824: 820: 812: 809: 804: 798: 795: 790: 786: 782: 778: 773: 768: 764: 760: 756: 749: 746: 741: 737: 732: 727: 723: 719: 715: 711: 707: 703: 699: 692: 689: 678: 672: 669: 658: 654: 648: 645: 640: 636: 632: 628: 624: 620: 616: 612: 605: 603: 599: 594: 590: 586: 582: 578: 574: 567: 565: 563: 561: 559: 555: 550: 546: 542: 538: 534: 530: 523: 521: 519: 517: 515: 511: 506: 502: 498: 494: 493: 485: 482: 475: 473: 471: 470:Carpediemonas 466: 465:Carpediemonas 462: 461:Carpediemonas 457: 456:Carpediemonas 453: 452:Carpediemonas 448: 444: 440: 432: 430: 428: 424: 423:Carpediemonas 420: 419:Carpediemonas 416: 415:Carpediemonas 412: 411:Carpediemonas 405: 403: 401: 400:Carpediemonas 396: 395:Carpediemonas 392: 391:Carpediemonas 387: 386:Carpediemonas 383: 382:Carpediemonas 375: 373: 371: 370: 365: 361: 360:Carpediemonas 354: 352: 350: 346: 342: 338: 334: 330: 329: 324: 320: 319:Carpediemonas 316: 315: 309: 305: 301: 297: 296: 291: 290:Carpediemonas 284: 282: 280: 279: 274: 270: 266: 262: 261:Carpediemonas 254: 252: 250: 249:Carpediemonas 246: 245:Carpediemonas 242: 241:Carpediemonas 238: 234: 233:mitochondrion 230: 229:Carpediemonas 226: 225:Carpediemonas 222: 218: 214: 210: 206: 205: 204:Carpediemonas 194: 191: 189: 186: 185: 183: 180: 176: 169: 165: 162: 158: 151: 150: 149:Carpediemonas 146: 143: 142: 139: 136: 133: 132: 128: 123: 120: 117: 116: 112: 107: 104: 101: 100: 96: 91: 88: 85: 84: 81: 78: 75: 74: 71: 68: 65: 64: 61: 58: 55: 54: 49: 44: 40: 37: 36:Carpediemonas 33: 30: 19: 4127: 4045: 3937: 3855: 3786: 3507: 3485:Kipferliidae 3409:Oxymonadidae 3367:Anaeromonada 3313:Planomonadea 3080:Anisonemidae 3049: 3042: 3015:Entosiphonea 2979:Keelungiidae 2974:Decastavidae 2970:Decastavida 2866:Neobodonidae 2857:Neobodonida 2777:Diplonemidae 2734:Calkinsiidae 2729:Postgaardida 2722:Bihospitidae 2718:Bihospitida 2626:Tetramitida 2620:Naegleriidae 2615:Tulamoebidae 2546:Eutetramitia 2380:Stygiellidae 2355:Andaluciidae 2285:Opisthokonta 2148: 2143:paraphyletic 2038: 2019: 2001: 1970:Malawimonada 1950:Anaeromonada 1945:Anaeramoebae 1937: 1913: 1890:Tsukubamonas 1888: 1810: 1803: 1794:Charophyceae 1781: 1774: 1767:Streptophyta 1754: 1738: 1718: 1714:Rhodelphidia 1672:Palpitomonas 1670: 1650: 1639:Pancryptista 1584: 1577: 1514: 1481: 1469: 1449: 1419: 1414:Foraminifera 1412: 1255: 1198: 1192:Tunicaraptor 1190: 1157: 1131:Opisthokonta 1067: 996:Opisthokonta 826: 822: 811: 797: 762: 758: 748: 705: 701: 691: 680:. 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Index

Carpediemonas membranifera
Scientific classification
Edit this classification
Eukaryota
Metamonada
Fornicata
Carpediemonadia
Cavalier-Smith
Carpediemonadea
Cavalier-Smith
Carpediemonadida
Cavalier-Smith
Carpediemonadidae
Carpediemonas
Type species
Species
Metamonada
Excavata
flagellated
Great Barrier Reef
mitochondrion
hydrogenosomes
carpe diem
Percolomonas
heterolobosean
flagella
Carpediemonas membranifera
prokaryotic
Carpediemonas frisia
Deltaproteobacteria

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