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Carpediemonas

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substrate, while the anterior flagellum beats less rapidly and in a slow sweeping motion. Further studies by Simpson and Patterson (1999) go into greater detail about the flagella and describe the flagellar apparatus as having a third, barren basal body. Supporting the dorsal side of the cell is a microtubular fan with a microtubular root at the anterior end. On the ventral side, microtubules extending from different flagellar roots support the ventral groove. The anterior flagellum has a "9+2" axoneme. Simpson and Patterson described that in addition to the "9+2" axoneme, the posterior flagellum also has "three radiating lamellae of electron-dense material which form the central components of vanes". The first lamella arises from after the flagellar insertion and is directed ventrally. The second lamella originates opposite from the first lamella. The third lamella supports the third vane, which is located more distally and lies perpendicular to the other two vanes or lamellae. All three lamella have striations when viewed in a longitudinal section and these striations are perpendicular to the "9+2" axoneme.
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all subunits of the Ndc80 complex involved in chromosome segregation. ORC and Cdc6 are proteins in charge of replication initiation and licensing in eukaryotes, and their absence appears to indicate the existence of a non-standard and as-yet undescribed mechanism to start replication. The absence of Ndc80 complex proteins also suggest that a non-standard mechanism could be in place for chromosome attachment to microtubules for chromosome segregation.
452:. In short: replication is proposed to proceed by a Dmc1-dependent homologous recombination mechanism that does not require origins of replication and that is mediated by RNA:DNA hybrids. This hypothesis still needs to be experimentally proven. Sexual or parasexual reproduction have not been directly observed in Metamonada. However, the study confirms the conservation of key meiotic proteins in the group with the bonus finding that 438:
are even further reduced. These genomes lack the DNA replication proteins of the origin recognition complex (ORC), Cdc6, some components of the GINS complex and some subunits of polymerases delta and epsilon, as well as most structural kinetochore subunits, a microtubule plus-end tracking complex and
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also has no mitochondria, which is typical of metamonads. Instead, it has hydrogenosomes, likely derived from anaerobic mitochondria. It also contains a single Golgi dictyosome, located anteriorly, dorsally, and to the left of the flagellar apparatus. The endoplasmic reticulum in this genus is mainly
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species have homologs from the tmcB family and sperm-specific channel subunits, the latter only reported previously only in Opisthokonta and three other protists. The presence of such proteins means that further investigations are required to understand if sex occurs, and if these proteins actually
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has a longitudinal depression that spans almost the entire ventral side of the cell. It often has two unequal flagella inserting to the anterior side of the ventral groove, but may sometimes have three or four flagella. The acronematic posterior flagellum is used in feeding and to attach to
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Salas-Leiva, Dayana E.; Tromer, Eelco C.; Curtis, Bruce A.; Jerlström-Hultqvist, Jon; Kolisko, Martin; Yi, Zhenzhen; Salas-Leiva, Joan S.; Gallot-Lavallée, Lucie; Williams S., Williams S.; Kops, Geert J. P. L.; Archibald, John M.; Simpson, Alastair G. B.; Roger, Andrew J. (2021).
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and contain a longitudinal groove. However, they did not have any evidence that the non-dividing organisms had more than two flagella. The species also contained a pouch with threads that may be difficult to discern from flagella. Ekebom et al. (1996) then renamed the organism as
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reproduces, it is obvious that they do replicate their DNA and researchers have proposed a hypothesis on how replication starts. This hypothesis uses elements of processes observed in other species but takes into account the specific protein complements found in
744:"The excavate protozoan phyla Metamonada Grasse emend. (Anaeromonadea, Parabasalia, Carpediemonas, Eopharyngia) and Loukozoa emend. (Jakobea, Malawimonas): their evolutionary affinities and new higher taxa" 216:
can be found in anaerobic intertidal sediment, where it feeds on bacteria. A feature of this species is the presence of a feeding groove, a characteristic of the excavates. Like most other metamonads,
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Yubuki, Naoji; Simpson, Alastair G.B.; Leander, Brian S. (2013). "Comprehensive Ultrastructure of Kipferlia bialata Provides Evidence for Character Evolution within the Fornicata (Excavata)".
410:-like organisms (CLOs) have allowed for the better understanding of the evolution of anaerobic excavates by studying their cytoskeletal traits and modified mitochondria. An example of a 228:
that are used to produce ATP. This organism has two flagella: a posterior one used for feeding on the substrate, and an anterior one that moves in a slower sweeping motion.
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Simpson, Alastair G.B.; Patterson, David J. (1999). "The ultrastructure of Carpediemonas membranifera (Eukaryota) with reference to the "excavate hypothesis"".
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found near the periphery of the cell. Around the cytoplasm, food vacuoles containing bacterial contents can be found. Also, three centrioles are present in
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Hamann, Emmo; Tegetmeyer, Halina E; Riedel, Dietmar; Littmann, Sten; Ahmerkamp, Soeren; Chen, Jianwei; Hach, Philipp F; Strous, Marc (2017-02-17).
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has revealed extensive loss of the DNA replication and segregation protein complements within Metamonada and has highlighted that the genomes of
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when it was found from substrate samples in the Great Barrier Reef and classified it as a metamonad. Additionally, a metabolic relationship of
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Ekebom, J.; Patterson, D.J.; Vørs, N. (1996). "Heterotrophic Flagellates from Coral Reef Sediments (Great Barrier Reef, Australia)".
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J. Ekebom; D. J. Patterson; N. Vørs (1996). "Heterotrophic flagellates from coral reef sediments (Great Barrier Reef, Australia)".
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contains a single ovate nucleus, located anteriorly in the cell. The nucleolus can also be found subcentrally within the nucleus.
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is the first known eukaryote to possess such drastically altered DNA replication and segregation machineries. Since
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is classified as an excavate because it has the characteristic feeding groove of the group. Within the excavates,
4201: 808:"Genomic analysis finds no evidence of canonical eukaryotic DNA processing complexes in a free-living protist" 240:
is a member of the metamonads, it is unusual in the sense that it is free-living and has three basal bodies.
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for incompletely digested organic material and the oxidation of various biomolecules. On the other hand,
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was found to release biomolecules that have been predigested. Prokaryotic communities would rely on
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was first discovered by Larsen and Patterson (1990) who identified it as a previously unidentified
3473: 3397: 2967: 2962: 2854: 2710: 2608: 2603: 1933: 270:, meaning "to seize the day", in honour of the wife of one of the authors, who had recently died. 3805: 3534: 3524: 3093: 2640: 2363: 2268: 2246: 2234: 2209: 1787: 979: 957: 945: 920: 867: 352: 209: 31: 3558: 3541: 3519: 3088: 3199: 2628: 4155: 4004: 3965: 3883: 3832: 2655: 2278: 2241: 2229: 2219: 2157: 2121: 2047: 2028: 1738: 1321: 989: 952: 940: 930: 845: 773: 765: 724: 706: 623: 615: 577: 533: 4160: 4009: 3914: 2645: 3996: 3546: 3491: 2898: 2410: 2193: 2093: 2039: 1793: 1239: 904: 835: 827: 755: 714: 698: 607: 569: 525: 489: 4091: 3703: 3553: 3483: 3277: 2893: 2402: 1966: 1141: 373:
as organisms with a size of approximately 5 μm long (with a range of 4–7.5 μm).
687:"Syntrophic linkage between predatory Carpediemonas and specific prokaryotic populations" 823: 3688: 3599: 3309: 3264: 3141: 3129: 3098: 2618: 2598: 2523: 2224: 2062: 2052: 2016: 1991: 1680: 1471: 1372: 1294: 1103: 935: 840: 807: 719: 686: 292: 665: 573: 529: 493: 4190: 3364: 3255: 3172: 2787: 2727: 2578: 2551: 2433: 2098: 1897: 1889: 1719: 1640: 1580: 1491: 1465: 1194: 225: 221: 3568: 3355: 3068: 2765: 2722: 2717: 2368: 2343: 2131: 1958: 1938: 1878: 1782: 1755: 1702: 1660: 1627: 1402: 1180: 1119: 414:-like organism that was used to study the evolutionary history within excavates is 355:
intertidal sediments, where it feeds on bacteria. It can be found co-existing with
283: 149: 4131: 4049: 3941: 3888: 3859: 3790: 611: 236:-like organisms are used in researching the evolution within excavates. Although 4140: 4058: 3970: 3950: 3868: 3799: 3584: 3451: 3246: 3184: 3063: 2760: 2743: 2699: 2690: 2348: 2022: 1948: 1800: 1764: 1744: 1692: 1666: 1556: 1504: 1477: 1459: 1432: 1199: 1068: 1057: 831: 3514: 3179: 2752: 2680: 2635: 2444: 2103: 1927: 1918: 1903: 1771: 1708: 1567: 1528: 1523: 1409: 1326: 1221: 1133: 311: 266: 197: 769: 710: 619: 581: 537: 3461: 3438: 3385: 3345: 3152: 3039: 3032: 2918: 2886: 2591: 2353: 2002: 1943: 1651: 1616: 1574: 1551: 1533: 1420: 1356: 1231: 1108: 1051: 1038: 890: 296: 68: 58: 48: 859: 849: 777: 728: 702: 627: 760: 743: 4125: 4043: 3935: 3853: 3784: 3392: 3272: 3232: 2881: 2586: 2393: 2335: 2263: 2180: 2009: 1971: 1872: 1849: 1396: 1391: 1383: 1313: 1188: 1063: 1046: 1024: 974: 205: 201: 4078: 3983: 3901: 3819: 2326: 2305: 2125: 1860: 1697: 1543: 1450: 1439: 1303: 1209: 1165: 1073: 792:"Microbe somehow survives without key proteins for replicating its DNA" 167: 1515: 1245: 1147: 1089: 3761: 2085: 1725: 264:, indicating a unicellular organism. The organism was named after 208:
eukaryote that was first discovered in substrate samples from the
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International Journal of Systematic and Evolutionary Microbiology
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participate during sex and what their role could be. In short,
161:(Larsen & Patterson 1990) Ekebom, Patterson & Vørs 1996 2149: 4115: 4033: 3925: 3843: 3774: 3641: 3592: 3583: 3507: 3482: 3460: 3436: 3363: 3354: 3343: 3308: 3301: 3294: 3263: 3254: 3245: 3230: 3140: 3109: 3056: 3049: 3024: 3003: 2952: 2926: 2917: 2839: 2813: 2786: 2751: 2742: 2698: 2689: 2678: 2577: 2541: 2534: 2513: 2492: 2485: 2460: 2453: 2442: 2431: 2401: 2392: 2385: 2334: 2325: 2318: 2303: 2084: 2061: 2038: 1988: 1957: 1917: 1888: 1859: 1847: 1754: 1718: 1679: 1650: 1626: 1613: 1542: 1514: 1490: 1449: 1419: 1382: 1369: 1339: 1312: 1292: 1230: 1208: 1162: 1132: 1117: 1086: 1037: 1022: 406:is assigned to the fornicates. In the fornicates, 291:. Larsen and Patterson treated this organism as a 422:DNA replication, chromosome segregation, and sex 2165: 875: 232:is assigned to the fornicates, where similar 8: 3762: 3589: 3466: 3444: 3433: 3360: 3351: 3305: 3298: 3260: 3251: 3242: 3238: 3053: 2923: 2792: 2748: 2695: 2686: 2538: 2489: 2457: 2450: 2439: 2398: 2389: 2331: 2322: 2315: 2311: 2172: 2158: 2150: 2035: 1865: 1856: 1844: 1751: 1715: 1685: 1632: 1623: 1610: 1496: 1425: 1379: 1366: 1349: 1336: 1309: 1289: 1285: 1214: 1172: 1129: 1125: 1096: 1034: 1030: 1019: 1015: 882: 868: 860: 295:, because it would occasionally have four 20: 839: 759: 718: 252:originates from three Latin roots, with 224:to produce energy. Instead, it contains 470: 2140:denotes groups with over 1,000 species 461:are intriguing microbial eukaryotes. 260:meaning "the day", and the suffix of 7: 593: 591: 555: 553: 551: 549: 547: 511: 509: 507: 505: 503: 332:relies on the prokaryotic organism, 666:"Taxonomy – It Came from the Pond" 143:Ekebom, Patterson & Vørs, 1996 14: 742:Cavalier-Smith, T. (2003-11-01). 361:in these anaerobic environments. 204:. This organism is a unicellular 562:European Journal of Protistology 35: 369:Ekebom et al. (1996) describes 1619:      1615:      1375:      1371:      1: 574:10.1016/s0932-4739(99)80044-3 530:10.1016/s0003-9365(96)80013-3 494:10.1016/S0003-9365(96)80013-3 612:10.1016/j.protis.2013.02.002 220:does not rely on an aerobic 200:, and belongs to the group 4218: 832:10.1038/s41467-021-26077-2 303:Carpediemonas membranifera 157:Carpediemonas membranifera 3469: 3447: 3432: 3241: 2795: 2314: 2187: 2116: 1868: 1843: 1688: 1635: 1609: 1499: 1428: 1365: 1352: 1335: 1288: 1284: 1217: 1175: 1128: 1099: 1033: 1018: 1014: 898: 518:Archiv für Protistenkunde 481:Archiv für Protistenkunde 314:communities was found in 289:Percolomonas membranifera 173: 166: 155: 148: 32:Scientific classification 30: 23: 1345:     1341:     703:10.1038/ismej.2016.197 287:and provided the name 812:Nature Communications 761:10.1099/ijs.0.02548-0 426:A recent study using 358:Cafeteria marsupialis 3682:Spirotrichonymphidae 3678:Spirotrichonymphida 428:comparative genomics 317:Carpediemonas frisia 3672:Hypotrichomonadidae 3668:Hypotrichomonadida 2017:Gunflint microbiota 1778:Klebsormidiophyceae 1562:Hyphochytriomycetes 824:2021NatCo..12.6003S 344:Habitat and ecology 334:Deltaproteobacteria 3729:Tritrichomonadidae 3719:Monocercomonadidae 3710:Tritrichomonadida 3616:Spirotrichosomidae 2875:Trypanoplasmatidae 2840:Metakinetoplastina 2820:Prokinetoplastida 2567:Psalteriomonadidae 2557:Paravahlkampfiidae 1788:Coleochaetophyceae 1168:    1164:    1092:    1088:    646:comenius.susqu.edu 338:hydrogen oxidizing 210:Great Barrier Reef 4184: 4183: 4156:Open Tree of Life 4005:Open Tree of Life 3875:Carpediemonadidae 3845:Carpediemonadidae 3768:Taxon identifiers 3759: 3758: 3755: 3754: 3751: 3750: 3747: 3746: 3743: 3742: 3739: 3738: 3724:Simplicimonadidae 3694:Honigbergiellidae 3579: 3578: 3497:Carpediemonadidae 3428: 3427: 3418:Streblomastigidae 3376:Paratrimastigidae 3372:Paratrimastigida 3339: 3338: 3335: 3334: 3331: 3330: 3290: 3289: 3286: 3285: 3226: 3225: 3222: 3221: 3218: 3217: 3214: 3213: 3210: 3209: 3190:Euglenamorphaceae 3163:Euglenomonadaceae 3159:Euglenomonadales 2913: 2912: 2909: 2908: 2814:Prokinetoplastina 2738: 2737: 2674: 2673: 2670: 2669: 2666: 2665: 2656:Stephanopogonidae 2524:Neovahlkampfiidae 2520:Neovahlkampfiida 2481: 2480: 2471:Pharyngomonadidae 2467:Pharyngomonadida 2461:Pharyngomonadidea 2427: 2426: 2423: 2422: 2419: 2418: 2381: 2380: 2377: 2376: 2147: 2146: 2112: 2111: 2048:Spironematellidae 2029:Meteora sporadica 1984: 1983: 1980: 1979: 1913: 1912: 1839: 1838: 1835: 1834: 1831: 1830: 1827: 1826: 1823: 1822: 1819: 1818: 1815: 1814: 1811: 1810: 1739:Prasinodermophyta 1675: 1674: 1605: 1604: 1601: 1600: 1597: 1596: 1593: 1592: 1589: 1588: 1486: 1485: 1322:Centroplasthelida 1280: 1279: 1276: 1275: 1272: 1271: 1268: 1267: 1264: 1263: 1260: 1259: 1256: 1255: 1158: 1157: 1082: 1081: 416:Kipferlia bialata 256:meaning "seize", 189: 188: 144: 127:Carpediemonadidae 118: 102: 86: 16:Genus of protists 4209: 4202:Metamonad genera 4177: 4176: 4164: 4163: 4151: 4150: 4149: 4136: 4135: 4134: 4108: 4107: 4095: 4094: 4082: 4081: 4069: 4068: 4067: 4054: 4053: 4052: 4026: 4025: 4013: 4012: 4000: 3999: 3987: 3986: 3974: 3973: 3961: 3960: 3959: 3957:Carpediemonadida 3946: 3945: 3944: 3927:Carpediemonadida 3918: 3917: 3905: 3904: 3892: 3891: 3879: 3878: 3877: 3864: 3863: 3862: 3836: 3835: 3823: 3822: 3810: 3809: 3808: 3795: 3794: 3793: 3763: 3621:Staurojoeninidae 3590: 3569:Retortamonadidae 3492:Carpediemonadida 3467: 3445: 3434: 3413:Saccinobaculidae 3361: 3352: 3306: 3299: 3261: 3252: 3243: 3239: 3195:Euglenocapsaceae 3079:Heteronemataceae 3075:Heteronematales 3054: 2924: 2899:Trypanosomatidae 2860:Rhynchomonadidae 2824:Ichthyobodonidae 2793: 2749: 2696: 2687: 2651:Percolomonadidae 2562:Plaesiobystridae 2539: 2490: 2458: 2451: 2440: 2411:Tsukubamonadidae 2399: 2390: 2332: 2323: 2316: 2312: 2216: 2196: 2174: 2167: 2160: 2151: 2094:Collodictyonidae 2040:Hemimastigophora 2036: 1995: 1866: 1857: 1853: 1845: 1794:Zygnematophyceae 1752: 1732: 1716: 1686: 1633: 1624: 1620: 1611: 1497: 1426: 1380: 1376: 1367: 1350: 1346: 1337: 1310: 1297: 1290: 1286: 1240:Choanoflagellata 1215: 1173: 1169: 1130: 1126: 1122: 1097: 1093: 1035: 1031: 1027: 1020: 1016: 927: 907: 884: 877: 870: 861: 854: 853: 843: 802: 796: 795: 788: 782: 781: 763: 754:(6): 1741–1758. 739: 733: 732: 722: 697:(5): 1205–1217. 691:The ISME Journal 682: 676: 675: 673: 672: 662: 656: 655: 653: 652: 642:"CARPEDIEMONADA" 638: 632: 631: 595: 586: 585: 557: 542: 541: 524:(3–4): 251–272. 513: 498: 497: 488:(3–4): 251–272. 475: 351:can be found in 274:History of study 142: 114: 111:Carpediemonadida 98: 82: 40: 39: 21: 4217: 4216: 4212: 4211: 4210: 4208: 4207: 4206: 4187: 4186: 4185: 4180: 4172: 4167: 4159: 4154: 4147:Carpediemonadia 4145: 4144: 4139: 4130: 4129: 4124: 4117:Carpediemonadia 4111: 4103: 4098: 4090: 4085: 4077: 4072: 4065:Carpediemonadea 4063: 4062: 4057: 4048: 4047: 4042: 4035:Carpediemonadea 4029: 4021: 4016: 4008: 4003: 3995: 3990: 3982: 3977: 3969: 3964: 3955: 3954: 3949: 3940: 3939: 3934: 3921: 3913: 3908: 3900: 3895: 3887: 3882: 3873: 3872: 3867: 3858: 3857: 3852: 3839: 3831: 3826: 3818: 3813: 3804: 3803: 3798: 3789: 3788: 3783: 3770: 3760: 3735: 3704:Trichomonadidae 3699:Tricercomitidae 3653:Cristamonadida 3637: 3631:Trichonymphidae 3626:Teratonymphidae 3575: 3564:Chilomastigidae 3554:Retortamonadida 3503: 3484:Carpediemonadea 3478: 3456: 3441: 3424: 3348: 3327: 3318:Ancyromonadidae 3282: 3278:Malawimonadidae 3235: 3206: 3136: 3105: 3045: 3020: 2999: 2993:Sphenomonadidae 2983:Sphenomonadidae 2948: 2942:Sphenomonadidae 2937:Petalomonadidae 2933:Petalomonadida 2905: 2894:Trypanosomatida 2835: 2809: 2805:Trypanophididae 2800:Bordnamonadidae 2782: 2771:Eupelagonemidae 2734: 2683: 2662: 2636:Percolomonadida 2573: 2530: 2514:Neovahlkampfiea 2509: 2477: 2447: 2436: 2415: 2403:Tsukubamonadida 2373: 2308: 2299: 2298: 2214: 2192: 2183: 2178: 2148: 2143: 2108: 2080: 2071:Ancyromonadidae 2057: 2034: 1989: 1976: 1967:Malawimonadidae 1953: 1909: 1884: 1848: 1807: 1750: 1723: 1722: 1714: 1671: 1646: 1614: 1585: 1538: 1510: 1482: 1445: 1415: 1370: 1361: 1340: 1331: 1308: 1293: 1252: 1226: 1204: 1163: 1154: 1118: 1113: 1087: 1078: 1023: 1010: 1009: 925: 903: 894: 888: 858: 857: 804: 803: 799: 790: 789: 785: 741: 740: 736: 684: 683: 679: 670: 668: 664: 663: 659: 650: 648: 640: 639: 635: 597: 596: 589: 559: 558: 545: 515: 514: 501: 477: 476: 472: 467: 432:C. membranifera 424: 397: 367: 346: 276: 246: 182:C. membranifera 162: 159: 141: 113: 97: 95:Carpediemonadea 81: 79:Carpediemonadia 34: 17: 12: 11: 5: 4215: 4213: 4205: 4204: 4199: 4189: 4188: 4182: 4181: 4179: 4178: 4165: 4152: 4137: 4121: 4119: 4113: 4112: 4110: 4109: 4096: 4083: 4070: 4055: 4039: 4037: 4031: 4030: 4028: 4027: 4014: 4001: 3988: 3975: 3962: 3947: 3931: 3929: 3923: 3922: 3920: 3919: 3906: 3893: 3880: 3865: 3849: 3847: 3841: 3840: 3838: 3837: 3824: 3811: 3796: 3780: 3778: 3772: 3771: 3766: 3757: 3756: 3753: 3752: 3749: 3748: 3745: 3744: 3741: 3740: 3737: 3736: 3734: 3733: 3732: 3731: 3726: 3721: 3716: 3714:Dientamoebidae 3708: 3707: 3706: 3701: 3696: 3689:Trichomonadida 3686: 3685: 3684: 3676: 3675: 3674: 3666: 3665: 3664: 3662:Devescovinidae 3659: 3651: 3649:Hexamastigidae 3645: 3643: 3639: 3638: 3636: 3635: 3634: 3633: 3628: 3623: 3618: 3613: 3611:Hoplonymphidae 3608: 3606:Burmanymphidae 3600:Trichonymphida 3596: 3594: 3587: 3581: 3580: 3577: 3576: 3574: 3573: 3572: 3571: 3566: 3561: 3559:Caviomonadidae 3551: 3550: 3549: 3539: 3538: 3537: 3532: 3530:Spironucleidae 3527: 3522: 3511: 3509: 3505: 3504: 3502: 3501: 3500: 3499: 3488: 3486: 3480: 3479: 3477: 3476: 3470: 3464: 3458: 3457: 3455: 3454: 3452:Anaeramoebidae 3448: 3442: 3437: 3430: 3429: 3426: 3425: 3423: 3422: 3421: 3420: 3415: 3410: 3408:Pyrsonymphidae 3405: 3403:Polymastigidae 3400: 3390: 3389: 3388: 3380: 3379: 3378: 3369: 3367: 3358: 3349: 3344: 3341: 3340: 3337: 3336: 3333: 3332: 3329: 3328: 3326: 3325: 3323:Planomonadidae 3320: 3314: 3312: 3310:Ancyromonadida 3303: 3296: 3292: 3291: 3288: 3287: 3284: 3283: 3281: 3280: 3275: 3269: 3267: 3265:Malawimonadida 3258: 3249: 3236: 3231: 3228: 3227: 3224: 3223: 3220: 3219: 3216: 3215: 3212: 3211: 3208: 3207: 3205: 3204: 3203: 3202: 3197: 3192: 3187: 3177: 3176: 3175: 3167: 3166: 3165: 3157: 3156: 3155: 3146: 3144: 3142:Euglenophyceae 3138: 3137: 3135: 3134: 3133: 3132: 3130:Peranemataceae 3124: 3123: 3122: 3113: 3111: 3107: 3106: 3104: 3103: 3102: 3101: 3099:Neometanemidae 3096: 3091: 3083: 3082: 3081: 3073: 3072: 3071: 3060: 3058: 3051: 3047: 3046: 3044: 3043: 3036: 3028: 3026: 3022: 3021: 3019: 3018: 3017: 3016: 3014:Entosiphonidae 3010:Entosiphonida 3007: 3005: 3001: 3000: 2998: 2997: 2996: 2995: 2987: 2986: 2985: 2979:Heterostavida 2977: 2976: 2975: 2973:Lentomonadidae 2970: 2965: 2956: 2954: 2950: 2949: 2947: 2946: 2945: 2944: 2939: 2930: 2928: 2921: 2915: 2914: 2911: 2910: 2907: 2906: 2904: 2903: 2902: 2901: 2891: 2890: 2889: 2879: 2878: 2877: 2872: 2864: 2863: 2862: 2857: 2852: 2843: 2841: 2837: 2836: 2834: 2833: 2832: 2831: 2826: 2817: 2815: 2811: 2810: 2808: 2807: 2802: 2796: 2790: 2784: 2783: 2781: 2780: 2779: 2778: 2773: 2768: 2757: 2755: 2746: 2740: 2739: 2736: 2735: 2733: 2732: 2731: 2730: 2725: 2715: 2714: 2713: 2704: 2702: 2693: 2684: 2679: 2676: 2675: 2672: 2671: 2668: 2667: 2664: 2663: 2661: 2660: 2659: 2658: 2653: 2648: 2643: 2633: 2632: 2631: 2623: 2622: 2621: 2619:Vahlkampfiidae 2613: 2612: 2611: 2606: 2596: 2595: 2594: 2583: 2581: 2575: 2574: 2572: 2571: 2570: 2569: 2564: 2559: 2554: 2545: 2543: 2536: 2532: 2531: 2529: 2528: 2527: 2526: 2517: 2515: 2511: 2510: 2508: 2507: 2506: 2505: 2496: 2494: 2487: 2483: 2482: 2479: 2478: 2476: 2475: 2474: 2473: 2464: 2462: 2455: 2454:Pharyngomonada 2448: 2443: 2437: 2432: 2429: 2428: 2425: 2424: 2421: 2420: 2417: 2416: 2414: 2413: 2407: 2405: 2396: 2387: 2383: 2382: 2379: 2378: 2375: 2374: 2372: 2371: 2366: 2361: 2356: 2351: 2346: 2340: 2338: 2329: 2320: 2309: 2304: 2301: 2300: 2297: 2296: 2295: 2294: 2293: 2292: 2289:Mesomycetozoea 2286: 2281: 2271: 2261: 2260: 2259: 2254: 2249: 2244: 2239: 2238: 2237: 2225:Diaphoretickes 2222: 2217: 2212: 2207: 2202: 2197: 2189: 2188: 2185: 2184: 2179: 2177: 2176: 2169: 2162: 2154: 2145: 2144: 2142: 2141: 2135: 2128: 2124:excluded from 2117: 2114: 2113: 2110: 2109: 2107: 2106: 2101: 2096: 2090: 2088: 2082: 2081: 2079: 2078: 2076:Planomonadidae 2073: 2067: 2065: 2063:Ancyromonadida 2059: 2058: 2056: 2055: 2053:Paramastigidae 2050: 2044: 2042: 2033: 2032: 2025: 2019: 2013: 2005: 1998: 1996: 1992:Incertae sedis 1986: 1985: 1982: 1981: 1978: 1977: 1975: 1974: 1969: 1963: 1961: 1955: 1954: 1952: 1951: 1946: 1941: 1936: 1931: 1923: 1921: 1915: 1914: 1911: 1910: 1908: 1907: 1900: 1894: 1892: 1886: 1885: 1883: 1882: 1875: 1869: 1863: 1854: 1841: 1840: 1837: 1836: 1833: 1832: 1829: 1828: 1825: 1824: 1821: 1820: 1817: 1816: 1813: 1812: 1809: 1808: 1806: 1805: 1797: 1790: 1785: 1780: 1775: 1768: 1760: 1758: 1749: 1748: 1741: 1735: 1733: 1713: 1712: 1705: 1700: 1695: 1689: 1683: 1681:Archaeplastida 1677: 1676: 1673: 1672: 1670: 1669: 1664: 1656: 1654: 1648: 1647: 1645: 1644: 1636: 1630: 1621: 1607: 1606: 1603: 1602: 1599: 1598: 1595: 1594: 1591: 1590: 1587: 1586: 1584: 1583: 1578: 1571: 1564: 1559: 1554: 1548: 1546: 1540: 1539: 1537: 1536: 1531: 1526: 1520: 1518: 1512: 1511: 1509: 1508: 1500: 1494: 1488: 1487: 1484: 1483: 1481: 1480: 1475: 1472:Dinoflagellata 1468: 1466:Chrompodellida 1463: 1455: 1453: 1447: 1446: 1444: 1443: 1436: 1429: 1423: 1417: 1416: 1414: 1413: 1406: 1399: 1394: 1388: 1386: 1377: 1363: 1362: 1360: 1359: 1353: 1347: 1333: 1332: 1330: 1329: 1324: 1318: 1316: 1307: 1306: 1300: 1298: 1295:Diaphoretickes 1282: 1281: 1278: 1277: 1274: 1273: 1270: 1269: 1266: 1265: 1262: 1261: 1258: 1257: 1254: 1253: 1251: 1250: 1242: 1236: 1234: 1228: 1227: 1225: 1224: 1218: 1212: 1206: 1205: 1203: 1202: 1197: 1192: 1184: 1176: 1170: 1160: 1159: 1156: 1155: 1153: 1152: 1144: 1138: 1136: 1123: 1115: 1114: 1112: 1111: 1106: 1100: 1094: 1084: 1083: 1080: 1079: 1077: 1076: 1071: 1066: 1061: 1054: 1049: 1043: 1041: 1028: 1012: 1011: 1008: 1007: 1006: 1005: 1004: 1003: 1000:Mesomycetozoea 997: 992: 982: 972: 971: 970: 965: 960: 955: 950: 949: 948: 936:Diaphoretickes 933: 928: 923: 918: 913: 908: 900: 899: 896: 895: 893:classification 889: 887: 886: 879: 872: 864: 856: 855: 797: 783: 734: 677: 657: 633: 606:(3): 423–439. 587: 568:(4): 353–370. 543: 499: 469: 468: 466: 463: 423: 420: 396: 393: 366: 363: 345: 342: 293:heterolobosean 275: 272: 245: 242: 226:hydrogenosomes 187: 186: 185: 184: 179: 171: 170: 164: 163: 160: 153: 152: 146: 145: 134: 130: 129: 124: 120: 119: 116:Cavalier-Smith 108: 104: 103: 100:Cavalier-Smith 92: 88: 87: 84:Cavalier-Smith 76: 72: 71: 66: 62: 61: 56: 52: 51: 46: 42: 41: 28: 27: 15: 13: 10: 9: 6: 4: 3: 2: 4214: 4203: 4200: 4198: 4195: 4194: 4192: 4175: 4170: 4166: 4162: 4157: 4153: 4148: 4142: 4138: 4133: 4127: 4123: 4122: 4120: 4118: 4114: 4106: 4101: 4097: 4093: 4088: 4084: 4080: 4075: 4071: 4066: 4060: 4056: 4051: 4045: 4041: 4040: 4038: 4036: 4032: 4024: 4019: 4015: 4011: 4006: 4002: 3998: 3993: 3989: 3985: 3980: 3976: 3972: 3967: 3963: 3958: 3952: 3948: 3943: 3937: 3933: 3932: 3930: 3928: 3924: 3916: 3911: 3907: 3903: 3898: 3894: 3890: 3885: 3881: 3876: 3870: 3866: 3861: 3855: 3851: 3850: 3848: 3846: 3842: 3834: 3829: 3825: 3821: 3816: 3812: 3807: 3806:Carpediemonas 3801: 3797: 3792: 3786: 3782: 3781: 3779: 3777: 3776:Carpediemonas 3773: 3769: 3764: 3730: 3727: 3725: 3722: 3720: 3717: 3715: 3712: 3711: 3709: 3705: 3702: 3700: 3697: 3695: 3692: 3691: 3690: 3687: 3683: 3680: 3679: 3677: 3673: 3670: 3669: 3667: 3663: 3660: 3658: 3657:Calonymphidae 3655: 3654: 3652: 3650: 3647: 3646: 3644: 3642:Trichomonadea 3640: 3632: 3629: 3627: 3624: 3622: 3619: 3617: 3614: 3612: 3609: 3607: 3603: 3602: 3601: 3598: 3597: 3595: 3593:Trichonymphea 3591: 3588: 3586: 3582: 3570: 3567: 3565: 3562: 3560: 3557: 3556: 3555: 3552: 3548: 3545: 3544: 3543: 3540: 3536: 3533: 3531: 3528: 3526: 3523: 3521: 3518: 3517: 3516: 3515:Diplomonadida 3513: 3512: 3510: 3506: 3498: 3495: 3494: 3493: 3490: 3489: 3487: 3485: 3481: 3475: 3472: 3471: 3468: 3465: 3463: 3459: 3453: 3450: 3449: 3446: 3443: 3440: 3435: 3431: 3419: 3416: 3414: 3411: 3409: 3406: 3404: 3401: 3399: 3396: 3395: 3394: 3391: 3387: 3386:Trimastigidae 3384: 3383: 3382:Trimastigida 3381: 3377: 3374: 3373: 3371: 3370: 3368: 3366: 3365:Anaeromonadea 3362: 3359: 3357: 3353: 3350: 3347: 3342: 3324: 3321: 3319: 3316: 3315: 3313: 3311: 3307: 3304: 3300: 3297: 3293: 3279: 3276: 3274: 3271: 3270: 3268: 3266: 3262: 3259: 3257: 3256:Malawimonadea 3253: 3250: 3248: 3244: 3240: 3237: 3234: 3229: 3201: 3198: 3196: 3193: 3191: 3188: 3186: 3183: 3182: 3181: 3178: 3174: 3173:Eutreptiaceae 3171: 3170: 3169:Eutreptiales 3168: 3164: 3161: 3160: 3158: 3154: 3151: 3150: 3148: 3147: 3145: 3143: 3139: 3131: 3128: 3127: 3126:Peranematida 3125: 3121: 3120:Teloproctidae 3118: 3117: 3116:Acroglissida 3115: 3114: 3112: 3108: 3100: 3097: 3095: 3092: 3090: 3087: 3086: 3085:Natomonadida 3084: 3080: 3077: 3076: 3074: 3070: 3067: 3066: 3065: 3062: 3061: 3059: 3055: 3052: 3048: 3042: 3041: 3037: 3035: 3034: 3030: 3029: 3027: 3023: 3015: 3012: 3011: 3009: 3008: 3006: 3002: 2994: 2991: 2990: 2988: 2984: 2981: 2980: 2978: 2974: 2971: 2969: 2966: 2964: 2961: 2960: 2958: 2957: 2955: 2951: 2943: 2940: 2938: 2935: 2934: 2932: 2931: 2929: 2927:Petalomonadea 2925: 2922: 2920: 2916: 2900: 2897: 2896: 2895: 2892: 2888: 2885: 2884: 2883: 2880: 2876: 2873: 2871: 2870:Parabodonidae 2868: 2867: 2866:Parabodonida 2865: 2861: 2858: 2856: 2853: 2851: 2850:Allobodonidae 2848: 2847: 2845: 2844: 2842: 2838: 2830: 2829:Perkinselidae 2827: 2825: 2822: 2821: 2819: 2818: 2816: 2812: 2806: 2803: 2801: 2798: 2797: 2794: 2791: 2789: 2788:Kinetoplastea 2785: 2777: 2776:Hemistasiidae 2774: 2772: 2769: 2767: 2764: 2763: 2762: 2759: 2758: 2756: 2754: 2750: 2747: 2745: 2741: 2729: 2728:Postgaardidae 2726: 2724: 2721: 2720: 2719: 2716: 2712: 2709: 2708: 2706: 2705: 2703: 2701: 2697: 2694: 2692: 2688: 2685: 2682: 2677: 2657: 2654: 2652: 2649: 2647: 2644: 2642: 2639: 2638: 2637: 2634: 2630: 2627: 2626: 2624: 2620: 2617: 2616: 2614: 2610: 2607: 2605: 2602: 2601: 2600: 2597: 2593: 2590: 2589: 2588: 2585: 2584: 2582: 2580: 2579:Heterolobosea 2576: 2568: 2565: 2563: 2560: 2558: 2555: 2553: 2552:Gruberellidae 2550: 2549: 2548:Lyromonadida 2547: 2546: 2544: 2540: 2537: 2533: 2525: 2522: 2521: 2519: 2518: 2516: 2512: 2504: 2503:Selenaionidae 2501: 2500: 2499:Selenaionida 2498: 2497: 2495: 2491: 2488: 2484: 2472: 2469: 2468: 2466: 2465: 2463: 2459: 2456: 2452: 2449: 2446: 2441: 2438: 2435: 2434:Discicristata 2430: 2412: 2409: 2408: 2406: 2404: 2400: 2397: 2395: 2391: 2388: 2386:Tsukubamonada 2384: 2370: 2367: 2365: 2362: 2360: 2359:Moramonadidae 2357: 2355: 2352: 2350: 2347: 2345: 2342: 2341: 2339: 2337: 2333: 2330: 2328: 2324: 2321: 2317: 2313: 2310: 2307: 2302: 2290: 2287: 2285: 2282: 2280: 2277: 2276: 2275: 2272: 2270: 2267: 2266: 2265: 2262: 2258: 2255: 2253: 2252:Stramenopiles 2250: 2248: 2245: 2243: 2240: 2236: 2233: 2232: 2231: 2228: 2227: 2226: 2223: 2221: 2218: 2215:(major groups 2213: 2211: 2208: 2206: 2203: 2201: 2198: 2195: 2191: 2190: 2186: 2182: 2175: 2170: 2168: 2163: 2161: 2156: 2155: 2152: 2139: 2136: 2133: 2129: 2127: 2123: 2120:¹traditional 2119: 2118: 2115: 2105: 2102: 2100: 2099:Mantamonadida 2097: 2095: 2092: 2091: 2089: 2087: 2083: 2077: 2074: 2072: 2069: 2068: 2066: 2064: 2060: 2054: 2051: 2049: 2046: 2045: 2043: 2041: 2037: 2031: 2030: 2026: 2024: 2020: 2018: 2014: 2012: 2011: 2006: 2004: 2000: 1999: 1997: 1994: 1993: 1987: 1973: 1970: 1968: 1965: 1964: 1962: 1960: 1956: 1950: 1947: 1945: 1942: 1940: 1937: 1935: 1932: 1930: 1929: 1925: 1924: 1922: 1920: 1916: 1906: 1905: 1901: 1899: 1898:Heterolobosea 1896: 1895: 1893: 1891: 1890:Discicristata 1887: 1881: 1880: 1876: 1874: 1871: 1870: 1867: 1864: 1862: 1858: 1855: 1851: 1846: 1842: 1803: 1802: 1798: 1796: 1795: 1791: 1789: 1786: 1784: 1781: 1779: 1776: 1774: 1773: 1769: 1767: 1766: 1762: 1761: 1759: 1757: 1753: 1747: 1746: 1742: 1740: 1737: 1736: 1734: 1730: 1729: 1721: 1720:Viridiplantae 1717: 1711: 1710: 1706: 1704: 1701: 1699: 1696: 1694: 1691: 1690: 1687: 1684: 1682: 1678: 1668: 1665: 1663: 1662: 1658: 1657: 1655: 1653: 1649: 1643: 1642: 1641:Microheliella 1638: 1637: 1634: 1631: 1629: 1625: 1622: 1618: 1612: 1608: 1582: 1581:Actinophryida 1579: 1577: 1576: 1572: 1570: 1569: 1565: 1563: 1560: 1558: 1555: 1553: 1550: 1549: 1547: 1545: 1541: 1535: 1532: 1530: 1527: 1525: 1522: 1521: 1519: 1517: 1513: 1507: 1506: 1502: 1501: 1498: 1495: 1493: 1492:Stramenopiles 1489: 1479: 1476: 1474: 1473: 1469: 1467: 1464: 1462: 1461: 1457: 1456: 1454: 1452: 1448: 1442: 1441: 1437: 1434: 1431: 1430: 1427: 1424: 1422: 1418: 1412: 1411: 1407: 1405: 1404: 1400: 1398: 1395: 1393: 1390: 1389: 1387: 1385: 1381: 1378: 1374: 1368: 1364: 1358: 1355: 1354: 1351: 1348: 1344: 1338: 1334: 1328: 1325: 1323: 1320: 1319: 1317: 1315: 1311: 1305: 1302: 1301: 1299: 1296: 1291: 1287: 1283: 1248: 1247: 1243: 1241: 1238: 1237: 1235: 1233: 1229: 1223: 1220: 1219: 1216: 1213: 1211: 1207: 1201: 1198: 1196: 1195:Ichthyosporea 1193: 1191: 1190: 1185: 1183: 1182: 1178: 1177: 1174: 1171: 1167: 1161: 1150: 1149: 1145: 1143: 1142:Rotosphaerida 1140: 1139: 1137: 1135: 1131: 1127: 1124: 1121: 1116: 1110: 1107: 1105: 1104:Apusomonadida 1102: 1101: 1098: 1095: 1091: 1085: 1075: 1072: 1070: 1067: 1065: 1062: 1060: 1059: 1055: 1053: 1050: 1048: 1045: 1044: 1042: 1040: 1036: 1032: 1029: 1026: 1021: 1017: 1013: 1001: 998: 996: 993: 991: 988: 987: 986: 983: 981: 978: 977: 976: 973: 969: 966: 964: 963:Stramenopiles 961: 959: 956: 954: 951: 947: 944: 943: 942: 939: 938: 937: 934: 932: 929: 926:(major groups 924: 922: 919: 917: 914: 912: 909: 906: 902: 901: 897: 892: 885: 880: 878: 873: 871: 866: 865: 862: 851: 847: 842: 837: 833: 829: 825: 821: 817: 813: 809: 801: 798: 793: 787: 784: 779: 775: 771: 767: 762: 757: 753: 749: 745: 738: 735: 730: 726: 721: 716: 712: 708: 704: 700: 696: 692: 688: 681: 678: 667: 661: 658: 647: 643: 637: 634: 629: 625: 621: 617: 613: 609: 605: 601: 594: 592: 588: 583: 579: 575: 571: 567: 563: 556: 554: 552: 550: 548: 544: 539: 535: 531: 527: 523: 519: 512: 510: 508: 506: 504: 500: 495: 491: 487: 483: 482: 474: 471: 464: 462: 460: 459:Carpediemonas 455: 454:Carpediemonas 451: 450:Carpediemonas 446: 445:Carpediemonas 442: 441:Carpediemonas 437: 433: 429: 421: 419: 417: 413: 412:Carpediemonas 409: 408:Carpediemonas 405: 404:Carpediemonas 401: 400:Carpediemonas 394: 392: 390: 389:Carpediemonas 385: 384:Carpediemonas 381: 380:Carpediemonas 376: 375:Carpediemonas 372: 371:Carpediemonas 364: 362: 360: 359: 354: 350: 349:Carpediemonas 343: 341: 339: 335: 331: 327: 323: 319: 318: 313: 309: 308:Carpediemonas 305: 304: 298: 294: 290: 286: 285: 280: 279:Carpediemonas 273: 271: 269: 268: 263: 259: 255: 251: 250:Carpediemonas 243: 241: 239: 238:Carpediemonas 235: 234:Carpediemonas 231: 230:Carpediemonas 227: 223: 222:mitochondrion 219: 218:Carpediemonas 215: 214:Carpediemonas 211: 207: 203: 199: 195: 194: 193:Carpediemonas 183: 180: 178: 175: 174: 172: 169: 165: 158: 154: 151: 147: 140: 139: 138:Carpediemonas 135: 132: 131: 128: 125: 122: 121: 117: 112: 109: 106: 105: 101: 96: 93: 90: 89: 85: 80: 77: 74: 73: 70: 67: 64: 63: 60: 57: 54: 53: 50: 47: 44: 43: 38: 33: 29: 26: 25:Carpediemonas 22: 19: 4116: 4034: 3926: 3844: 3775: 3496: 3474:Kipferliidae 3398:Oxymonadidae 3356:Anaeromonada 3302:Planomonadea 3069:Anisonemidae 3038: 3031: 3004:Entosiphonea 2968:Keelungiidae 2963:Decastavidae 2959:Decastavida 2855:Neobodonidae 2846:Neobodonida 2766:Diplonemidae 2723:Calkinsiidae 2718:Postgaardida 2711:Bihospitidae 2707:Bihospitida 2615:Tetramitida 2609:Naegleriidae 2604:Tulamoebidae 2535:Eutetramitia 2369:Stygiellidae 2344:Andaluciidae 2274:Opisthokonta 2137: 2132:paraphyletic 2027: 2008: 1990: 1959:Malawimonada 1939:Anaeromonada 1934:Anaeramoebae 1926: 1902: 1879:Tsukubamonas 1877: 1799: 1792: 1783:Charophyceae 1770: 1763: 1756:Streptophyta 1743: 1727: 1707: 1703:Rhodelphidia 1661:Palpitomonas 1659: 1639: 1628:Pancryptista 1573: 1566: 1503: 1470: 1458: 1438: 1408: 1403:Foraminifera 1401: 1244: 1187: 1181:Tunicaraptor 1179: 1146: 1120:Opisthokonta 1056: 985:Opisthokonta 815: 811: 800: 786: 751: 747: 737: 694: 690: 680: 669:. 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Index

Scientific classification
Edit this classification
Eukaryota
Metamonada
Fornicata
Carpediemonadia
Cavalier-Smith
Carpediemonadea
Cavalier-Smith
Carpediemonadida
Cavalier-Smith
Carpediemonadidae
Carpediemonas
Type species
Species
Metamonada
Excavata
flagellated
Great Barrier Reef
mitochondrion
hydrogenosomes
carpe diem
Percolomonas
heterolobosean
flagella
Carpediemonas membranifera
prokaryotic
Carpediemonas frisia
Deltaproteobacteria
hydrogen oxidizing

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