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Ophiocordyceps sinensis

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1340: 105: 1223: 632:. The perithecia are ordinally arranged and ovoid. The asci are cylindrical or slightly tapering at both ends, and may be straight or curved, with a capitate and hemispheroid apex, and may be two to four spored. Similarly, ascospores are hyaline, filiform, multiseptate at a length of 5–12 μm and subattenuated on both sides. Perithecial, ascus and ascospore characters in the fruiting bodies are the key identification characteristics of 80: 3043: 591: 53: 1123: 1195:
multihyphal structures that can remain dormant and then germinate to produce spores. After overwintering, the fungus ruptures the host body, forming the fruiting body, a sexual sporulating structure (a perithecial stroma) from the larval head that is connected to the sclerotia (dead larva) below ground and grows upward to emerge from the soil to complete the cycle.
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Winkler, D. 2005. Yartsa Gunbu – Cordyceps sinensis. Economy, Ecology & Ethno-mycology of a Fungus Endemic to the Tibetan Plateau. In: A. BOESI & F. CARDI (eds.). Wildlife and plants in traditional and modern Tibet: Conceptions, Exploitation and Conservation. Memorie della Società Italiana di
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at an altitude between 3,000 and 5,000 m (9,800 and 16,400 ft). The fungus is reported from the northern range of Nepal, Bhutan, and also from the northern states of India, apart from northern Yunnan, eastern Qinghai, eastern Tibet, western Sichuan, southwestern Gansu provinces. Fifty-seven
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Because of its high value, inter-village conflicts over access to its grassland habitats has become a headache for the local governing bodies and in several cases people were killed. In November 2011, a court in Nepal convicted 19 villagers over the murder of a group of farmers during a fight over
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The infected larvae tend to remain underground vertical to the soil surface with their heads up. After invading a host larva, the fungus ramifies throughout the host and eventually kills it. Gradually the host larvae become rigid because of the production of fungal sclerotia. Fungal sclerotia are
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has become the most important source of cash income. The fungi contributed 40% of the annual cash income to local households and 8.5% to the GDP in 2004. Prices have increased continuously, especially since the late 1990s. In 2008, one kilogram traded for US$ 3,000 (lowest quality) to over US$
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In 2004 the value of a kilogram of caterpillars was estimated at 30,000 to 60,000 Nepali rupees in Nepal, and about Rs 100,000 in India. In 2011, the value of a kilogram of caterpillars was estimated at 350,000 to 450,000 Nepali rupees in Nepal. A 2012 BBC article indicated that in north Indian
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as it is considered to be composed of both an animal and a vegetable. They are now cultivated on an industrial scale for their use in traditional Chinese medicine. However, no one has succeeded so far in rearing the fungus by infecting cultivated caterpillars; all products derived from cultured
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The stalk-like dark brown to black fruiting body (or mushroom) grows out of the head of the dead caterpillar and emerges from the soil in alpine meadows by early spring. During late summer, the fruiting body disperses spores. The caterpillars, which live underground feeding on roots, are most
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and in the Himalayas, between elevations of 3,000 and 5,000 metres (10,000 and 16,000 ft). The fungus germinates in the living larva, kills and mummifies it, and then a dark brown stalk-like fruiting body which is a few centimeters long emerges from the corpse and stands upright.
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in Nepal had only been legalised in 2001, and now demand is highest in countries such as China, Thailand, Vietnam, Korea and Japan. By 2002, the 'herb' was valued at R 105,000 ($ 1,435) per kilogram, allowing the government to charge a royalty of R 20,000 ($ 280) per kilogram.
617:. The stroma is the upper fungal part and is dark brown or black, but can be a yellow color when fresh, and longer than the caterpillar itself, usually 4–10 cm. It grows singly from the larval head, and is clavate, sublanceolate or fusiform, and distinct from the 1288:("New compilation of materia medica"). The ethno-mycological knowledge on caterpillar fungus among the Nepalese people is documented. The entire fungus-caterpillar combination is hand-collected for medicinal use. 533:
derivative. However, the fruiting bodies harvested in nature usually contain high amounts of arsenic and other heavy metals, so they are potentially toxic and sales have been strictly regulated by China's
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villages a single fungus was worth Rs 150 (about £2 or $ 3), which is more than the daily wage of a manual labourer. In 2012, a pound of top-quality yartsa had reached retail prices of $ 50,000.
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vulnerable to the fungus after shedding their skin, during late summer. In late autumn, chemicals on the skin of the caterpillar interact with the fungal spores and release the fungal
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Sung G. H.; et al. (2007). "A multi-gene phylogeny of Clavicipitaceae (Ascomycota, Fungi): identification of localized incongruence using a combinational bootstrap approach".
2881:"Criteria and indicator for assessing threat on Himalayan birch (B. utilis) at timberline ecotone of Nanda Devi Biosphere Reserve: A world heritage site, Western Himalaya, India" 650:
species, which produce either immersed or superficial perithecia perpendicular to stromal surface, and the ascospores at maturity are disarticulated into part spores. Generally
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as an endangered species in China. Additional research needs to be carried out in order to understand its morphology and growth habits for conservation and optimum utilization.
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Hsieh, C., et al., A Systematic Review of the Mysterious Caterpillar Fungus Ophiocordyceps sinensis in Dong-ChongXiaCao and Related Bioactive Ingredients. Vol. 3. 2013. 16–32.
1363:, about 900% between 1998 and 2008, an annual average of over 20% (after inflation). However, the value of large caterpillar fungus has increased more dramatically than small 3420: 842:(冬蟲夏草), meaning "winter worm, summer grass", which is a literal translation of the original Tibetan name. In traditional Chinese medicine, its name is often abbreviated as 2133:
Liu Z.; Liang Z.; Liu A.; Yao Y.; Hyde K. D.; Yu Z. (2002). "Molecular evidence for teleomorph-anamorph connections in Cordyceps based on ITS-5.8S rDNA sequences".
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where it grows. While it has been collected for centuries and is still common in such areas, current collection rates are much higher than in historical times.
772:. In colloquial Tibetan yartsa gunbu is often shortened to simply "bu" or "yartsa". The Tibetan name is transliterated in Nepali as यार्चागुन्बू, यार्चागुन्बा, 3425: 2730: 2110:
Hill H. Art. XXXVI: The Vegetable Caterpillar (Cordiceps robertsii). Transactions and Proceedings of the Royal Society of New Zealand. Vol 34, 1901;396–401
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apparently originated in Tibet and Nepal. So far the oldest known text documenting its use was written in the late 15th century by the Tibetan doctor
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grows at a comparatively low temperature, i.e., below 21 °C. Temperature requirements and growth rates are crucial factors that distinguish
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The fruiting bodies of the fungus are not yet cultivated commercially, but the mycelium form can be cultivated in vitro. Overharvesting and
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Martel J, Ko YF, Liau JC, Lee CS, Ojcius DM, Lai HC, Young JD (2017). "Myths and realities surrounding the mysterious caterpillar fungus".
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18,000 (best quality, largest larvae). The annual production on the Tibetan Plateau was estimated in 2009 at 80–175 tons. The Himalayan
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Zhu JS, Halpern GM, Jones K (1998). "The scientific rediscovery of an ancient Chinese herbal medicine: Cordyceps sinensis: part I".
3035: 3400: 1746:"Recent advances in Cordyceps sinensis polysaccharides: mycelial fermentation, isolation, structure, and bioactivities: a review" 1708: 2855: 2440: 3092: 273: 910:
The English term "vegetable caterpillar" is a misnomer, as no plant is involved. "Caterpillar fungus" is a preferred term.
3410: 1686: 287: 104: 2441:"Caterpillar Fungus (Ophiocordyceps sinensis) Production and Sustainability on the Tibetan Plateau and in the Himalayas" 568: 564: 1457:
and government forces fought for control of the lucrative export trade during the June–July harvest season. Collecting
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is recently also being harvested. The quality of the Bhutanese variety has been shown to be equal to the Tibetan one.
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Wu, Ding-Tao; Lv, Guang-Ping; Zheng, Jian; Li, Qian; Ma, Shuang-Cheng; Li, Shao-Ping; Zhao, Jing (December 2016).
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is not considered anamorph as it has widely distributed host range, and is not restricted only in high altitude.
3430: 3415: 895: 697: 646:) Kobayasi species produce whole ascospores and do not separate into part spores. This is different from other 28: 2704: 1709:"The World's Most Valuable Parasite Is in Trouble. And so are the livelihoods of the people who depend on it" 3435: 2580: 693: 478: 3047: 1627: 1246: 3405: 3103: 2566: 2266:
Xing X. K.; Guo S. X. (2008). "The Structure and Histochemistry of Sclerotia of Ophiocordyceps sinensis".
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is estimated to be around 10,000 billion VND/year. In the period 2017-2022, the production of cultivated
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in 2022 reached about 1,000 tons, an increase of five times compared to 2017. The selling price of fresh
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are synonyms. Evidence based on microcyclic conidiation from ascospores and molecular studies support
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The Global Circulation of Chinese Materia Medica, 1700-1949: A Microhistory of the Caterpillar Fungus
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The Global Circulation of Chinese Materia Medica, 1700-1949: A Microhistory of the Caterpillar Fungus
1837: 768:, "summer grass winter worm"). The name was first recorded in the 15th century by the Tibetan doctor 709: 679: 482: 166: 2571: 3246: 2344: 69: 3061: 3011: 2592: 2394: 2299: 2283: 2248: 2232: 2150: 2093: 1450: 970: 560: 465: 225: 99: 84: 2044:"Enumeratio Pyrenomycetum Hypocreaceorum hucusque cognitorum systemate carpologico dispositorum" 1545:
collectors near the treeline is reported to be depleting populations of tree species such as of
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is typically by growth of pure mycelia in liquid culture (in China) or on grains (in the West).
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has dark pigments and tough to pliant stromata, a typical characteristic feature of most of the
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Medicinal Plant Images Database (School of Chinese Medicine, Hong Kong Baptist University)
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generally live 15 cm (5.9 in) underground in alpine grass and shrub-lands on the
3065: 2992:. Medicine and Biomedical Sciences in Modern History. Palgrave Macmillan. pp. 1–294. 1470: 1360: 1141: 705: 701: 549: 486: 421: 146: 2481:
from high altitudes of the Indian Himalaya: Conservation and biotechnological priorities"
2043: 1261:("Instructions on a Myriad of Medicines"), where he describes its use as an aphrodisiac. 2950: 2627: 2074:. Medicine and Biomedical Sciences in Modern History. Palgrave Macmillan. pp. 1–3. 1892:"What is the Chinese caterpillar fungus Ophiocordyceps sinensis (Ophiocordycipitaceae)?" 1841: 1179:, and 14 others not correctly identified to genus) are recognized as potential hosts of 27:"Vegetable caterpillar" redirects here. For the other fungus with this common name, see 3331: 3176: 2969: 2928: 2652: 2611: 1996: 1967: 1860: 1829: 1770: 1745: 1273: 1207: 928: 751: 722: 671: 398: 176: 1040:
appeared only in one article and thus is discarded because of incomplete information.
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has very low similarity in RAPD polymorphism, hence it is not the anamorph. Likewise,
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Jiang, Y. Y., & Yao, Y. J. (n.d). Names related to Cordyceps sinensis anamorph.
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were identified only up to generic level, and it is dubious that they are anamorph.
3323: 3155: 2612:"Cordyceps collected from Bhutan, an appropriate alternative of Cordyceps sinensis" 1966:
Sung GH, Hywel-Jones NL, Sung JM, Luangsa-Ard JJ, Shrestha B, Spatafora JW (2007).
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Scienze Naturali e del Museo Civico di Storia Naturale di Milano, Vol. 33.1:69–85.
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In traditional Chinese medicine, it is regarded as having an excellent balance of
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is not considered anamorph as there is no valid published information. Similarly,
590: 52: 17: 3309: 2959: 2676:"NGÀNH NÔNG NGHIỆP ĐẨY MẠNH XUẤT KHẨU, GIẢI QUYẾT CÁC RÀO CẢN KỸ THUẬT PHÁT SINH" 2544: 1908: 1891: 3241: 3202: 3135: 3053: 2897: 2557:
Baral N, Heinen JT (2005). "The Maoist people's war and conservation in Nepal".
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ranges from 100-200 million VND/kg. Therefore, the economic value of cultivated
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the prized aphrodisiac fungus. Seven farmers were killed in the remote northern
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The fertile part of the stroma is the head. The head is granular because of the
621:(stalk). The stipe is slender, glabrous, and longitudinally furrowed or ridged. 522: 156: 3042: 2506: 1495:, and producers claim it may offer improved consistency. Artificial culture of 485:. It is mainly found in the meadows above 3,500 metres (11,500 ft) on the 2997: 2390: 2146: 2079: 1761: 1295: 1169: 1157: 1151: 1025: 959: 935: 629: 610: 545: 526: 510: 136: 3126: 2906: 2643: 2460: 2186: 3163: 1850: 1175: 1163: 1145: 1073: 900: 684: 643: 530: 116: 3070: 2978: 2661: 2588: 2295: 2244: 2005: 1987: 1949: 1869: 1814: 1779: 3030: 2194: 3357: 3303: 3228: 3120: 2937: 2929:"High Diversity of the Fungal Community Structure in Naturally-Occurring 1188: 506: 2287: 2236: 440: 3349: 3194: 1531: 1515: 1503: 1344: 1291: 1231: 1051:, have no valid information and thus are not considered as anamorph to 625: 3207: 2635: 3362: 3233: 2279: 1477: 1000: 989: 918:
Since the 1980s, 22 species in 13 genera have been attributed to the
861: 494: 126: 3097: 2377:) and the fungal commodification of the rural economy in Tibet AR". 821: 3168: 1338: 1320: 1290: 1235: 1221: 1121: 589: 502: 498: 490: 453: 449: 439: 245: 216: 1665:. New York, New York: M. Evans and Company, Inc. pp. 64–65. 2805:"Phát triển các mô hình trồng và chế biến dược liệu giá trị cao" 3101: 2319:"Demand for 'Himalayan Viagra' Fungus Heats Up, Maybe Too Much" 1587:(Berk.) G.H. Sung, J.M. Sung, Hywel-Jones & Spatafora 2007" 1256: 1250: 403: 389: 426: 2705:"Đông trùng hạ thảo, sâm Ngọc Linh.... không còn quá xa xỉ" 2357:
Journal of the International Association of Tibetan Studies
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the Japanese reading of the characters for the Chinese name
1830:"Host insect species of Ophiocordyceps sinensis: a review" 654:
species possess brightly colored and fleshy stromata, but
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Shrestha B.; Weimin Z.; Yongjie Z.; Xingzhong L. (2010).
879:. Strangely, sometimes in Chinese English-language texts 2414:"Ben cao cong xin" – "New compilation of materia medica" 1311:
are derived from mycelia grown on grains or in liquids.
219:) G.H.Sung, J.M.Sung, Hywel-Jones & Spatafora (2007) 2879:
Maletha, Ajay; Maikhuri, R. K.; Bargali, S. S. (2020).
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10.2990/1471-5457(2005)24[2:TMPWAC]2.0.CO;2
848:(蟲草 "insect plant"), a name that also applies to other 3059:
An Electronic Monograph of Cordyceps and Related Fungi
3058: 2545:'Himalayan viagra': Six men get life for Nepal murders 2345:"The mushrooming fungi market in Tibet exemplified by 1469:
is often perceived to threaten the environment of the
792: 836: 330: 63:(left) growing out of the head of a dead caterpillar 2927:
Zhang Y.; Zhang S.; Wang M.; Bai F.; Liu X. (2010).
2507:"The 'Viagra' transforming local economies in India" 2210:"Last Stand for the Body Snatcher of the Himalayas?" 1446:
in June 2009 after going to forage for Yarchagumba.
1298:, a compound isolated from the "caterpillar fungus". 3293: 3110: 1082:which had similar morphological characteristics to 420: 415: 397: 383: 378: 362: 357: 347: 342: 324: 317: 312: 300: 286: 272: 267: 258: 1359:is reported to have increased dramatically on the 670:The species was first described scientifically by 544:parasitizes the larvae of moths within the family 2856:"Việt Nam sản xuất cao từ nấm đông trùng hạ thảo" 2831:"Việt Nam sản xuất cao từ nấm đông trùng hạ thảo" 1687:"Caterpillar Fungus: The Viagra of the Himalayas" 1070:is considered the validly published anamorph of 1036:are discarded based on the molecular evidence. 481:(a fungus that grows on insects) in the family 2368: 2366: 1522:ranges from 10-20 million VND/kg, while dried 872: 1508:Ministry of Agriculture and Rural Development 1491:mycelium is an alternative to wild-harvested 998:and is not the anamorph. Several isolates of 292: 278: 8: 1226:Weighing the precious caterpillar fungus in 2885:Environmental and Sustainability Indicators 1961: 1959: 1885: 1883: 1881: 1879: 1538:has grown at an average rate of 40%/year.. 1110:as the anamorph of the caterpillar fungus, 525:. Caterpillar fungus contains the compound 3421:Fungi used in traditional Chinese medicine 3098: 3054:Yartsa Gunbu (Cordyceps sinensis) in Tibet 1702: 1700: 1651:《总局关于停止冬虫夏草用于保健食品试点工作的通知 -食药监食监三〔2016〕21号》 1024:is discarded as anamorph as it belongs to 922:(asexually reproducing mold-like form) of 708:, resulting in the naming of a new family 536:State Administration for Market Regulation 375: 309: 78: 51: 40: 2968: 2958: 2896: 2651: 2570: 2459: 1995: 1907: 1859: 1849: 1769: 1626:National Cancer Institute (2011-02-02). 1369: 1332:production might not exceed a few tons. 987:but there is lack of valid information. 1923: 1921: 1919: 1730: 1728: 1726: 1724: 1722: 1593:. International Mycological Association 1574: 1280:. In the 18th century it was listed in 1268:in traditional Chinese medicine was in 979:is only validly published in articles. 2683:Bộ Nông nghiệp và Phát triển nông thôn 2168: 2166: 2164: 1191:, which then infects the caterpillar. 255: 1930:Molecular Phylogenetics and Evolution 1506:, according to the statistics of the 1351:(冬虫夏草) among other local specialties. 763: 682:transferred the species to the genus 7: 3373:f0e86433-dfc9-4e3e-9d70-a54b96cc0367 3270:ef8ea3ee-8a0f-40ab-9191-3d1426fe8be4 2547:BBC News Asia, Retrieved 9 July 2012 1564:List of fungi by conservation status 1053:Ophiocordyceps sinensis. V. sinensis 609:consists of two parts, a fungal endo 563:, and its use has a long history in 3426:Taxa named by Miles Joseph Berkeley 790:. The transliteration in Bhutan is 548:, specifically genera found on the 2785:Tạp chí Doanh nghiệp và Thương mại 2534:, National Geographic, August 2012 2373:Winkler D. (2008). "Yartsa Gunbu ( 1828:Xiao-Liang W.; Yi-Jian Y. (2011). 1218:Use in traditional Asian medicines 578:have led to the classification of 25: 1241:The use of caterpillar fungus as 688:in 1878. The fungus was known as 586:Taxonomic history and systematics 3396:IUCN Red List vulnerable species 3077:Cordyceps sinensis (Berk.) Sacc. 3041: 3029: 2756:"Nâng tầm giá trị cây dược liệu" 2021:"On some entomogenous Sphaeriae" 1968:"Phylogenetic classification of 1610:Interview with Britt Bunyard on 1449:Its value gave it a role in the 983:possibly might be synonymous to 103: 2729:ONLINE, TUOI TRE (2023-04-02). 1972:and the clavicipitaceous fungi" 1744:Yan JK, Wang WQ, Wu JY (2014). 1510:, the production of cultivated 933:is the zygomycetous species of 899:, a species from south-eastern 696:analysis was used to amend the 517:which is valued in traditional 3093:National Geographic (magazine) 2559:Politics and the Life Sciences 1541:In India, fuelwood cutting by 1210:is suspected to be negatively 948:Cephalosporium dongchongxiacae 666:Developments in classification 404: 390: 331: 293: 279: 1: 2505:Jeffrey, Craig (2012-07-07). 2229:10.1126/science.322.5905.1182 1807:10.1016/j.tibtech.2017.06.011 1367:, regarded as lower quality. 1149:taxa from several genera (37 613:(within the caterpillar) and 493:and the Himalayan regions of 2960:10.1371/journal.pone.0015570 1909:10.1080/21501203.2010.536791 1434: 1424: 1414: 1411: 1403: 1400: 1392: 1252:Zur mkhar mnyam nyid rdo rje 914:Nomenclature of the anamorph 712:and the transfer of several 569:traditional Tibetan medicine 565:traditional Chinese medicine 2898:10.1016/j.indic.2020.100086 2781:"Đông trùng hạ thảo Hector" 1942:10.1016/j.ympev.2007.03.011 1257: 1251: 837: 822: 793: 594:Caterpillars with emerging 3452: 3073:information from Drugs.com 2317:KAI SCHULTZ (2016-06-26). 1628:"Definition of cordycepin" 1258:Man ngag bye ba ring bsrel 1206:from other similar fungi. 834:It is known in Chinese as 26: 2998:10.1007/978-3-031-24723-1 2709:Báo Người Lao Động Online 2543:Staff (14 November 2011) 2391:10.1007/s12231-008-9038-3 2147:10.1017/s0953756202006378 2080:10.1007/978-3-031-24723-1 1762:10.1016/j.jff.2013.11.024 873: 801:In India, it is known as 734:In Tibet, it is known as 473:), known colloquially as 434: 427: 374: 368: 308: 304:winter worm, summer grass 263: 231: 224: 205: 198: 100:Scientific classification 98: 76: 67: 59: 50: 43: 2461:10.1163/157342109X568829 2187:10.1089/acm.1998.4.3-289 2025:London Journal of Botany 1661:Halpern, Miller (2002). 975:are synonymous and only 896:Ophiocordyceps robertsii 274:Traditional Chinese 30:Ophiocordyceps robertsii 3401:Fungi described in 1843 3142:Ophiocordyceps sinensis 3112:Ophiocordyceps sinensis 3036:Ophiocordyceps sinensis 2931:Ophiocordyceps sinensis 2175:J Altern Complement Med 1851:10.3897/zookeys.127.802 1707:Yong, Ed (2018-10-22). 1585:Ophiocordyceps sinensis 1543:Ophiocordyceps sinensis 1512:Ophiocordyceps sinensis 1482:Ophiocordyceps sinensis 1467:Ophiocordyceps sinensis 1357:Ophiocordyceps sinensis 1266:Ophiocordyceps sinensis 1214:the mountain organism. 1127:Ophiocordyceps sinensis 981:Cephalosporium sinensis 765:[jɑ̀ːt͡sɑkỹ̀pu] 607:Ophiocordyceps sinensis 596:Ophiocordyceps sinensis 479:entomopathogenic fungus 461:Ophiocordyceps sinensis 288:Simplified Chinese 259:Ophiocordyceps sinensis 209:Ophiocordyceps sinensis 61:Ophiocordyceps sinensis 45:Ophiocordyceps sinensis 1988:10.3114/sim.2007.57.01 1412:429% (incl. inflation) 1401:467% (incl. inflation) 1352: 1299: 1272:’s 1694 compendium of 1238: 1136:prone to infection by 1129: 1118:Ecology and life cycle 1058:Metarhizium anisopliae 996:Paecilomyces farinosus 602:Morphological features 598: 456: 3089:Tibet’s Golden "Worm" 2809:Truyenhinhthanhhoa.vn 2532:Tibet’s Golden "Worm" 2042:Saccardo PA. (1878). 2019:Berkeley MJ. (1843). 1342: 1294: 1264:The first mention of 1247:Zurkhar Nyamnyi Dorje 1225: 1125: 1090:. This also confirms 1034:Sporothrix insectorum 1030:Paecilomyces sinensis 926:. Of the 22 species, 827:in Nepali, Hindi and 593: 443: 3411:Ophiocordycipitaceae 3038:at Wikimedia Commons 2439:Winkler, D. (2009). 2351:Tricholoma matsutake 2135:Mycological Research 1315:Economics and impact 770:Zurkhar Namnyi Dorje 748:dbyar rtswa dgun 'bu 710:Ophiocordycipitaceae 680:Pier Andrea Saccardo 483:Ophiocordycipitaceae 448:यार्सागुम्बा in the 167:Ophiocordycipitaceae 2951:2010PLoSO...515570Z 2628:2016NatSR...637668W 2343:Winkler D. (2008). 1976:Studies in Mycology 1842:2011ZooK..127...43W 1663:Medicinal Mushrooms 1632:NCI Drug Dictionary 1615:. BBC. Sep 9, 2023. 1067:Hirsutella sinensis 1045:sinense, P. hepiali 1022:Mortierella hepiali 559:is classified as a 391:Dbyar Rtswa Dgun'bu 70:Conservation status 3064:2014-05-17 at the 3048:Cordyceps sinensis 2616:Scientific Reports 2530:Finkel, M. (2012) 2479:Cordyceps sinensis 2475:Sharma S. (2004). 2375:Cordyceps sinensis 2347:Cordyceps sinensis 1451:Nepalese Civil War 1444:district of Manang 1353: 1349:Dōng chóng xià cǎo 1300: 1239: 1130: 1094:to be anamorph of 883:is referred to as 881:Cordyceps sinensis 838:dōng chóng xià cǎo 740:དབྱར་རྩྭ་དགུན་འབུ་ 716:species including 690:Cordyceps sinensis 599: 561:medicinal mushroom 475:caterpillar fungus 470:Cordyceps sinensis 457: 369:དབྱར་རྩྭ་དགུན་འབུ་ 332:dōng chóng xià cǎo 241:Cordyceps sinensis 18:Caterpillar fungus 3383: 3382: 3295:Sphaeria sinensis 3278:Open Tree of Life 3104:Taxon identifiers 3034:Media related to 3007:978-3-031-24722-4 2636:10.1038/srep37668 2089:978-3-031-24722-4 1836:(127): 12743–59. 1801:(11): 1017–1021. 1795:Trends Biotechnol 1672:978-0-87131-981-4 1478:Kingdom of Bhutan 1439: 1438: 1255:) ) in his text: 1198:The slow growing 1049:Scydalium hepiali 860:, it is known by 852:species, such as 692:until 2007, when 676:Sphaeria sinensis 444:Fruiting body of 438: 437: 411: 410: 338: 337: 319:Standard Mandarin 254: 253: 249: 238: 234:Sphaeria sinensis 93: 36:Species of fungus 16:(Redirected from 3443: 3376: 3375: 3366: 3365: 3353: 3352: 3340: 3339: 3327: 3326: 3314: 3313: 3312: 3286: 3285: 3273: 3272: 3263: 3262: 3250: 3249: 3237: 3236: 3224: 3223: 3211: 3210: 3198: 3197: 3185: 3184: 3172: 3171: 3159: 3158: 3146: 3145: 3144: 3131: 3130: 3129: 3099: 3085: 3082: 3046:Data related to 3045: 3033: 3019: 2982: 2972: 2962: 2911: 2910: 2900: 2876: 2870: 2869: 2867: 2866: 2852: 2846: 2845: 2843: 2842: 2826: 2820: 2819: 2817: 2816: 2801: 2795: 2794: 2792: 2791: 2776: 2770: 2769: 2767: 2766: 2752: 2746: 2745: 2743: 2742: 2726: 2720: 2719: 2717: 2716: 2700: 2694: 2693: 2691: 2690: 2680: 2672: 2666: 2665: 2655: 2607: 2601: 2600: 2574: 2554: 2548: 2541: 2535: 2528: 2522: 2521: 2519: 2517: 2502: 2496: 2495: 2485: 2472: 2466: 2465: 2463: 2445: 2436: 2430: 2424: 2418: 2417: 2409: 2403: 2402: 2370: 2361: 2360: 2340: 2334: 2333: 2331: 2329: 2314: 2308: 2307: 2280:10.3852/07-007R2 2263: 2257: 2256: 2214: 2208:Stone R (2008). 2205: 2199: 2198: 2170: 2159: 2158: 2141:(9): 1100–1108. 2130: 2124: 2117: 2111: 2108: 2102: 2101: 2065: 2059: 2058: 2048: 2039: 2033: 2032: 2016: 2010: 2009: 1999: 1963: 1954: 1953: 1936:(3): 1204–1223. 1925: 1914: 1913: 1911: 1887: 1874: 1873: 1863: 1853: 1825: 1819: 1818: 1790: 1784: 1783: 1773: 1741: 1735: 1732: 1717: 1716: 1704: 1695: 1694: 1683: 1677: 1676: 1658: 1652: 1649: 1643: 1642: 1640: 1638: 1623: 1617: 1616: 1608: 1602: 1601: 1599: 1598: 1579: 1547:Himalayan birch 1455:Nepalese Maoists 1435:125,000–500,000 1376:% Price Increase 1370: 1286:Ben cao cong xin 1260: 1254: 878: 876: 875: 840: 825: 796: 767: 741: 628:of the embedded 576:overexploitation 519:Chinese medicine 430: 429: 407: 406: 393: 392: 376: 370: 334: 333: 310: 296: 295: 282: 281: 256: 243: 236: 211: 191:O. sinensis 108: 107: 87: 82: 81: 55: 41: 21: 3451: 3450: 3446: 3445: 3444: 3442: 3441: 3440: 3431:Parasitic fungi 3416:Medicinal fungi 3386: 3385: 3384: 3379: 3371: 3369: 3361: 3356: 3348: 3343: 3335: 3330: 3322: 3317: 3308: 3307: 3302: 3289: 3281: 3276: 3268: 3266: 3258: 3253: 3245: 3240: 3232: 3227: 3219: 3214: 3206: 3201: 3193: 3188: 3180: 3175: 3167: 3162: 3154: 3149: 3140: 3139: 3134: 3125: 3124: 3119: 3106: 3083: 3080: 3066:Wayback Machine 3026: 3008: 2985: 2926: 2919: 2917:Further reading 2914: 2878: 2877: 2873: 2864: 2862: 2860:www.most.gov.vn 2854: 2853: 2849: 2840: 2838: 2837:(in Vietnamese) 2828: 2827: 2823: 2814: 2812: 2811:(in Vietnamese) 2803: 2802: 2798: 2789: 2787: 2778: 2777: 2773: 2764: 2762: 2754: 2753: 2749: 2740: 2738: 2737:(in Vietnamese) 2735:TUOI TRE ONLINE 2728: 2727: 2723: 2714: 2712: 2711:(in Vietnamese) 2702: 2701: 2697: 2688: 2686: 2678: 2674: 2673: 2669: 2609: 2608: 2604: 2572:10.1.1.474.3446 2556: 2555: 2551: 2542: 2538: 2529: 2525: 2515: 2513: 2504: 2503: 2499: 2488:Current Science 2483: 2474: 2473: 2469: 2443: 2438: 2437: 2433: 2429:, Devkota(2006) 2425: 2421: 2411: 2410: 2406: 2379:Economic Botany 2372: 2371: 2364: 2342: 2341: 2337: 2327: 2325: 2316: 2315: 2311: 2265: 2264: 2260: 2212: 2207: 2206: 2202: 2172: 2171: 2162: 2132: 2131: 2127: 2118: 2114: 2109: 2105: 2090: 2067: 2066: 2062: 2046: 2041: 2040: 2036: 2018: 2017: 2013: 1965: 1964: 1957: 1927: 1926: 1917: 1889: 1888: 1877: 1827: 1826: 1822: 1792: 1791: 1787: 1743: 1742: 1738: 1733: 1720: 1706: 1705: 1698: 1685: 1684: 1680: 1673: 1660: 1659: 1655: 1650: 1646: 1636: 1634: 1625: 1624: 1620: 1611: 1609: 1605: 1596: 1594: 1581: 1580: 1576: 1572: 1560: 1471:Tibetan Plateau 1465:The search for 1361:Tibetan Plateau 1317: 1278:Ben Cao Bei Yao 1220: 1142:Tibetan Plateau 1120: 1104:C. multiaxialis 1080:C. multiaxialis 994:is combined to 916: 889:, which is the 870: 794:yartsa guenboob 739: 732: 706:Clavicipitaceae 702:Cordycipitaceae 668: 604: 588: 550:Tibetan Plateau 513:and produces a 487:Tibetan Plateau 301:Literal meaning 239: 220: 213: 207: 194: 147:Sordariomycetes 102: 94: 83: 79: 72: 37: 34: 23: 22: 15: 12: 11: 5: 3449: 3447: 3439: 3438: 3436:Fungus species 3433: 3428: 3423: 3418: 3413: 3408: 3403: 3398: 3388: 3387: 3381: 3380: 3378: 3377: 3367: 3354: 3341: 3328: 3315: 3299: 3297: 3291: 3290: 3288: 3287: 3274: 3264: 3251: 3238: 3225: 3212: 3199: 3186: 3173: 3160: 3147: 3132: 3116: 3114: 3108: 3107: 3102: 3096: 3095: 3086: 3074: 3068: 3056: 3051: 3050:at Wikispecies 3039: 3025: 3024:External links 3022: 3021: 3020: 3006: 2986:Lu D. (2023). 2983: 2945:(12): e15570. 2924: 2918: 2915: 2913: 2912: 2871: 2847: 2821: 2796: 2771: 2760:lamdong.gov.vn 2747: 2721: 2695: 2667: 2602: 2549: 2536: 2523: 2497: 2467: 2454:(2): 291–316. 2448:Asian Medicine 2431: 2419: 2404: 2385:(3): 291–305. 2362: 2335: 2309: 2274:(4): 616–625. 2258: 2223:(5905): 1182. 2200: 2181:(3): 289–303. 2160: 2125: 2112: 2103: 2088: 2060: 2034: 2011: 1955: 1915: 1902:(4): 228–236. 1875: 1820: 1785: 1736: 1718: 1696: 1678: 1671: 1653: 1644: 1618: 1603: 1573: 1571: 1568: 1567: 1566: 1559: 1556: 1437: 1436: 1433: 1431: 1427: 1426: 1425:10,000–60,000 1423: 1421: 1417: 1416: 1413: 1410: 1406: 1405: 1402: 1399: 1395: 1394: 1391: 1389: 1385: 1384: 1377: 1374: 1330:Ophiocordyceps 1316: 1313: 1309:Ophiocordyceps 1274:materia medica 1219: 1216: 1208:Climate change 1119: 1116: 929:Cephalosporium 915: 912: 731: 728: 723:Ophiocordyceps 698:classification 672:Miles Berkeley 667: 664: 660:Ophiocordyceps 640:Ophiocordyceps 603: 600: 587: 584: 436: 435: 432: 431: 424: 418: 417: 413: 412: 409: 408: 401: 399:Tibetan Pinyin 395: 394: 387: 381: 380: 379:Transcriptions 372: 371: 366: 360: 359: 355: 354: 351: 345: 344: 340: 339: 336: 335: 328: 322: 321: 315: 314: 313:Transcriptions 306: 305: 302: 298: 297: 290: 284: 283: 276: 270: 269: 265: 264: 261: 260: 252: 251: 229: 228: 222: 221: 214: 203: 202: 196: 195: 188: 186: 182: 181: 178:Ophiocordyceps 174: 170: 169: 164: 160: 159: 154: 150: 149: 144: 140: 139: 134: 130: 129: 124: 120: 119: 114: 110: 109: 96: 95: 77: 74: 73: 68: 65: 64: 57: 56: 48: 47: 35: 24: 14: 13: 10: 9: 6: 4: 3: 2: 3448: 3437: 3434: 3432: 3429: 3427: 3424: 3422: 3419: 3417: 3414: 3412: 3409: 3407: 3406:Fungi of Asia 3404: 3402: 3399: 3397: 3394: 3393: 3391: 3374: 3368: 3364: 3359: 3355: 3351: 3346: 3342: 3338: 3333: 3329: 3325: 3320: 3316: 3311: 3305: 3301: 3300: 3298: 3296: 3292: 3284: 3279: 3275: 3271: 3265: 3261: 3256: 3252: 3248: 3243: 3239: 3235: 3230: 3226: 3222: 3217: 3213: 3209: 3204: 3200: 3196: 3191: 3187: 3183: 3178: 3174: 3170: 3165: 3161: 3157: 3152: 3148: 3143: 3137: 3133: 3128: 3122: 3118: 3117: 3115: 3113: 3109: 3105: 3100: 3094: 3090: 3087: 3078: 3075: 3072: 3069: 3067: 3063: 3060: 3057: 3055: 3052: 3049: 3044: 3040: 3037: 3032: 3028: 3027: 3023: 3017: 3013: 3009: 3003: 2999: 2995: 2991: 2990: 2984: 2980: 2976: 2971: 2966: 2961: 2956: 2952: 2948: 2944: 2940: 2939: 2934: 2932: 2925: 2921: 2920: 2916: 2908: 2904: 2899: 2894: 2890: 2886: 2882: 2875: 2872: 2861: 2857: 2851: 2848: 2836: 2835:vnexpress.net 2832: 2825: 2822: 2810: 2806: 2800: 2797: 2786: 2782: 2775: 2772: 2761: 2757: 2751: 2748: 2736: 2732: 2725: 2722: 2710: 2706: 2699: 2696: 2684: 2677: 2671: 2668: 2663: 2659: 2654: 2649: 2645: 2641: 2637: 2633: 2629: 2625: 2621: 2617: 2613: 2606: 2603: 2598: 2594: 2590: 2586: 2582: 2578: 2573: 2568: 2564: 2560: 2553: 2550: 2546: 2540: 2537: 2533: 2527: 2524: 2512: 2508: 2501: 2498: 2494:(12): 1614–9. 2493: 2489: 2482: 2480: 2471: 2468: 2462: 2457: 2453: 2449: 2442: 2435: 2432: 2428: 2423: 2420: 2416:(in Chinese). 2415: 2412:Wu Y (1757). 2408: 2405: 2400: 2396: 2392: 2388: 2384: 2380: 2376: 2369: 2367: 2363: 2358: 2354: 2352: 2348: 2339: 2336: 2324: 2320: 2313: 2310: 2305: 2301: 2297: 2293: 2289: 2285: 2281: 2277: 2273: 2269: 2262: 2259: 2254: 2250: 2246: 2242: 2238: 2234: 2230: 2226: 2222: 2218: 2211: 2204: 2201: 2196: 2192: 2188: 2184: 2180: 2176: 2169: 2167: 2165: 2161: 2156: 2152: 2148: 2144: 2140: 2136: 2129: 2126: 2122: 2116: 2113: 2107: 2104: 2099: 2095: 2091: 2085: 2081: 2077: 2073: 2072: 2068:Lu D (2023). 2064: 2061: 2057:(3): 277–325. 2056: 2052: 2045: 2038: 2035: 2030: 2026: 2022: 2015: 2012: 2007: 2003: 1998: 1993: 1989: 1985: 1981: 1977: 1973: 1971: 1962: 1960: 1956: 1951: 1947: 1943: 1939: 1935: 1931: 1924: 1922: 1920: 1916: 1910: 1905: 1901: 1897: 1893: 1886: 1884: 1882: 1880: 1876: 1871: 1867: 1862: 1857: 1852: 1847: 1843: 1839: 1835: 1831: 1824: 1821: 1816: 1812: 1808: 1804: 1800: 1796: 1789: 1786: 1781: 1777: 1772: 1767: 1763: 1759: 1755: 1751: 1750:J Funct Foods 1747: 1740: 1737: 1731: 1729: 1727: 1725: 1723: 1719: 1714: 1710: 1703: 1701: 1697: 1692: 1688: 1682: 1679: 1674: 1668: 1664: 1657: 1654: 1648: 1645: 1633: 1629: 1622: 1619: 1614: 1607: 1604: 1592: 1588: 1586: 1578: 1575: 1569: 1565: 1562: 1561: 1557: 1555: 1554: 1551: 1550: 1549:Betula utilis 1544: 1539: 1537: 1533: 1529: 1525: 1521: 1517: 1513: 1509: 1505: 1500: 1498: 1494: 1490: 1485: 1483: 1479: 1474: 1472: 1468: 1463: 1460: 1456: 1452: 1447: 1445: 1432: 1429: 1428: 1422: 1419: 1418: 1408: 1407: 1397: 1396: 1390: 1387: 1386: 1382: 1378: 1375: 1372: 1371: 1368: 1366: 1362: 1358: 1355:The price of 1350: 1346: 1341: 1337: 1333: 1331: 1326: 1322: 1314: 1312: 1310: 1305: 1297: 1293: 1289: 1287: 1283: 1279: 1275: 1271: 1267: 1262: 1259: 1253: 1248: 1244: 1243:folk medicine 1237: 1233: 1229: 1224: 1217: 1215: 1213: 1209: 1205: 1201: 1196: 1192: 1190: 1184: 1182: 1178: 1177: 1172: 1171: 1166: 1165: 1160: 1159: 1154: 1153: 1147: 1143: 1139: 1135: 1128: 1124: 1117: 1115: 1113: 1109: 1105: 1101: 1100:C. nepalensis 1098:and suggests 1097: 1093: 1089: 1085: 1081: 1077: 1075: 1072:O. sinensis, 1069: 1068: 1062: 1060: 1059: 1054: 1050: 1046: 1044: 1043:Tolypocladium 1039: 1035: 1031: 1027: 1023: 1019: 1017: 1012: 1010: 1006: 1002: 997: 993: 991: 986: 982: 978: 974: 972: 967: 963: 961: 956: 953: 949: 945: 943: 942:Chrysosporium 938: 937: 932: 930: 925: 921: 913: 911: 908: 906: 902: 898: 897: 892: 888: 887: 882: 869: 868: 863: 859: 855: 851: 847: 846: 841: 839: 832: 830: 826: 824: 823:ghaas fafoond 818: 817: 812: 811: 806: 805: 799: 797: 795: 789: 788: 783: 782: 777: 776: 771: 766: 761: 760:Lhasa dialect 757: 756:yarza g̈unbu' 753: 749: 745: 737: 729: 727: 725: 724: 720:to the genus 719: 715: 711: 707: 703: 699: 695: 691: 687: 686: 681: 677: 673: 665: 663: 661: 657: 653: 649: 645: 641: 637: 635: 631: 627: 622: 620: 616: 612: 608: 601: 597: 592: 585: 583: 581: 577: 572: 570: 566: 562: 558: 554: 551: 547: 543: 539: 537: 532: 528: 524: 520: 516: 515:fruiting body 512: 508: 504: 500: 496: 492: 488: 484: 480: 476: 472: 471: 467: 463: 462: 455: 451: 447: 442: 433: 425: 423: 419: 414: 402: 400: 396: 388: 386: 382: 377: 373: 367: 365: 361: 356: 352: 350: 346: 341: 329: 327: 323: 320: 316: 311: 307: 303: 299: 291: 289: 285: 277: 275: 271: 266: 262: 257: 250: 247: 242: 235: 230: 227: 223: 218: 212: 210: 204: 201: 200:Binomial name 197: 193: 192: 187: 184: 183: 180: 179: 175: 172: 171: 168: 165: 162: 161: 158: 155: 152: 151: 148: 145: 142: 141: 138: 135: 132: 131: 128: 125: 122: 121: 118: 115: 112: 111: 106: 101: 97: 91: 86: 75: 71: 66: 62: 58: 54: 49: 46: 42: 39: 32: 31: 19: 3294: 3111: 3091:August 2012 3084:(in Chinese) 3081:(in English) 2988: 2942: 2936: 2930: 2888: 2884: 2874: 2863:. 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Index

Caterpillar fungus
Ophiocordyceps robertsii

Conservation status
Vulnerable
IUCN 3.1
Scientific classification
Edit this classification
Eukaryota
Fungi
Ascomycota
Sordariomycetes
Hypocreales
Ophiocordycipitaceae
Ophiocordyceps
Binomial name
Berk.
Synonyms
Sacc.
Traditional Chinese
Simplified Chinese
Standard Mandarin
Hanyu Pinyin
Burmese
Tibetan
Wylie
Tibetan Pinyin
Nepali

Dolpo

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