Knowledge (XXG)

Primary nutritional groups

Source šŸ“

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microorganisms, however, are not limited to just one nutritional mode. For example, some algae live photoautotrophically in the light, but shift to chemoorganoheterotrophy in the dark. Even higher plants retained their ability to respire heterotrophically on starch at night which had been synthesised
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For example, plants are lithotrophs because they use water as their electron donor for the electron transport chain across the thylakoid membrane. Animals are organotrophs because they use organic compounds as electron donors to synthesize ATP (plants also do this, but this is not taken into
139:, divided in relation to the nutrition mode according to the sources of energy and carbon, needed for living, growth and reproduction. The sources of energy can be light or chemical compounds; the sources of carbon can be of organic or inorganic origin. 1043:, using organic molecules as energy, electron/hydrogen and carbon sources. Some bacteria are limited to only one nutritional group, whereas others are facultative and switch from one mode to the other, depending on the nutrient sources available. 305:
Organotrophic organisms are often also heterotrophic, using organic compounds as sources of both electrons and carbon. Similarly, lithotrophic organisms are often also autotrophic, using inorganic sources of electrons and
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for growth and development, and that obtains its energy from the decomposition of an organic compound. This group of organisms may be further subdivided according to what kind of organic substrate and compound they use.
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Some, usually unicellular, organisms can switch between different metabolic modes, for example between photoautotrophy, photoheterotrophy, and chemoheterotrophy in
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Illustrated Glossary of Protoctista: Vocabulary of the Algae, Apicomplexa, Ciliates, Foraminifera, Microspora, Water Molds, Slime Molds, and the Other Protoctists
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are rare because heterotrophy implies the availability of organic substrates, which can also serve as easy electron sources, making lithotrophy unnecessary.
2813: 53: 2883: 2420: 166:) do not refer to primary nutritional groups, but simply reflect the different use of possible electron acceptors in particular organisms, such as 2893: 2621: 1437: 1356: 1303: 2898: 3486: 2455: 100: 3086: 1655:
gen. nov., sp. nov., a thermophilic, microaerophilic, facultatively chemolithoheterotrophic bacterium from a deep-sea hydrothermal vent"
507:), fats and proteins. Chemoheterotrophs also obtain the carbon atoms that they need for cellular function from these organic compounds. 316:
The organic or inorganic substances (e.g., oxygen) used as electron acceptors needed in the catabolic processes of aerobic or anaerobic
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are uncommon since their organic source of electrons/hydrogens would provide an easy carbon source, resulting in heterotrophy.
162: 3176: 990:, since they use organic substances both as chemical energy sources and as electron/hydrogen donors and carbon sources. Some 817: 328:
account). Both use oxygen in respiration as electron acceptor, but this character is not used to define them as lithotrophs.
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Some lithotrophic bacteria can utilize diverse sources of electrons, depending on the availability of possible donors.
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Plants and some bacteria can alternate between phototrophy and chemotrophy, depending on the availability of light.
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All sorts of combinations may exist in nature, but some are more common than others. For example, most plants are
526:. The important differentiation amongst this group is that chemoorganotrophs oxidize only organic compounds while 46: 3891: 3813: 3568: 3384: 2996: 2888: 2746: 2731: 2726: 2117: 1286:
Kelly DP, Mason J, Wood A (1987). "Energy Metabolism in Chemolithotrophs". In van Verseveld HW, Duine JA (eds.).
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are examples of chemoorganoheterotrophs which obtain carbon and electrons or hydrogen from dead organic matter.
4104: 3911: 3632: 3573: 3479: 3374: 3369: 3339: 2606: 2488: 968:, due to their capability to use more resources than either photoautotrophic or organoheterotrophic organisms. 472: 3906: 929: 4005: 3754: 3749: 3735: 3710: 3218: 3081: 2991: 2859: 2741: 2711: 2568: 2533: 2253: 2220: 2195: 1048: 1649:
Miroshnichenko ML, L'Haridon S, Jeanthon C, Antipov AN, Kostrikina NA, Tindall BJ, et al. (May 2003).
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as their energy source and obtain electrons or hydrogen from the organic compounds, including sugars (i.e.
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are chemoheterotrophs (meaning they oxidize chemical compounds as a source of energy and carbon), as are
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Rippka R (March 1972). "Photoheterotrophy and chemoheterotrophy among unicellular blue-green algae".
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are examples of organisms that obtain carbon and electrons or hydrogen from living organic matter.
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Kellermann MY, Wegener G, Elvert M, Yoshinaga MY, Lin YS, Holler T, et al. (November 2012).
1039:, using organic molecules as both electron/hydrogen donors and carbon sources. Many bacteria are 531: 2913: 1856:"Carbon metabolic pathways in phototrophic bacteria and their broader evolutionary implications" 1536:
Gleizer S, Ben-Nissan R, Bar-On YM, Antonovsky N, Noor E, Zohar Y, et al. (November 2019).
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Yafremava LS, Wielgos M, Thomas S, Nasir A, Wang M, Mittenthal JE, Caetano-AnollƩs G (2013).
252:. Eventually, the energy is used for life processes such as moving, growth and reproduction. 3947: 3298: 3161: 3153: 3071: 2953: 2938: 2874: 2854: 2771: 2761: 2756: 2721: 2553: 2493: 2364: 2165: 2107: 2019: 1979: 1877: 1867: 1821: 1813: 1764: 1746: 1703: 1666: 1621: 1613: 1592:
Gassler T, Sauer M, Gasser B, Egermeier M, Troyer C, Causon T, et al. (December 2019).
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Morris, J. et al. (2019). "Biology: How Life Works", 3rd edition, W. H. Freeman.
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Proceedings of the National Academy of Sciences of the United States of America
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in anaerobic respiration, or various metabolic intermediates in fermentation.
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is converted from a heterotroph into an autotroph capable of growth on CO
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Table 1: Definitions of metabolic strategies to obtain carbon and energy
4015: 4000: 3379: 3186: 3056: 3051: 2678: 2626: 2286: 1931: 1707: 1107: 1052: 965: 782: 504: 249: 190: 174: 1538:"Conversion of Escherichia coli to Generate All Biomass Carbon from CO 957: 933: 790: 515: 511: 476: 167: 960:, use either light, inorganic or organic compounds for energy. Such 542:
The following table gives some examples for each nutritional group:
290:(electron donors) are needed by both phototrophs and chemotrophs in 1729:
Li, Meijie; Ning, Peng; Sun, Yi; Luo, Jie; Yang, Jianming (2022).
1130:, while the latter remained the source of electrons for the cells. 1019: 786: 716: 366: 291: 1659:
International Journal of Systematic and Evolutionary Microbiology
653:) perform anoxygenic photosynthesis and fix atmospheric carbon. 3468: 2651: 1904: 1106:
as the sole carbon source while using the one-carbon molecule
1081:
efforts enabled the transformation of the trophic mode of two
344:
organic compounds to obtain carbon for growth and development.
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Comparative Ecology of Microorganisms and Macroorganisms
1065:, using inorganic energy, electron, and carbon sources. 3697:(amino acidā†’pyruvate, acetyl CoA, or TCA intermediate) 376:
Classification of organisms based on their metabolism
374: 4072: 1321:"Nomenclature of nutritional types of microorganisms" 279:
use inorganic compounds as electron/hydrogen donors.
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Cold Spring Harbor Symposia on Quantitative Biology
60:. Unsourced material may be challenged and removed. 294:that transfer energy in the anabolic processes of 1319:Lwoff A, Van Niel CB, Ryan TF, Tatum EL (1946). 1432:. Jones & Bartlett Learning. pp. xxv. 1027:or sulfide as electron/hydrogen donors, and CO 984:as a carbon source. All animals and fungi are 3480: 2663: 1916: 1739:Frontiers in Bioengineering and Biotechnology 1453: 1451: 1449: 956:ā€“ another example ā€“ can grow with or without 8: 1587: 1585: 1531: 1529: 1085:from heterotrophy to chemoorganoautotrophy: 781:prokaryotes. Some eukaryotes (heterotrophic 434: 405: 392: 380: 3962: 3879: 3772: 3765: 3677: 3653: 3544: 3487: 3473: 3465: 2884:Latitudinal gradients in species diversity 2670: 2656: 2648: 1923: 1909: 1901: 1426:, McKhann HI, Olendzenski L, eds. (1993). 1122:yeast was genetically engineered to use CO 1881: 1871: 1854:Tang KH, Tang YJ, Blankenship RE (2011). 1825: 1768: 1750: 1670: 1625: 1565: 1509: 1499: 1399: 1389: 1263:"Todar's Online Textbook of Bacteriology" 1234: 120:Learn how and when to remove this message 2782:Predatorā€“prey (Lotkaā€“Volterra) equations 2421:Tritrophic interactions in plant defense 1021: 544: 4079: 2814:Random generalized Lotkaā€“Volterra model 1193: 259:Primary sources of reducing equivalents 2622:Herbivore adaptations to plant defense 1798:Applied and Environmental Microbiology 1207:Applied and Environmental Microbiology 226:and transform it into chemical energy. 444: 389: 7: 3597:Electron acceptors other than oxygen 2637:Predator avoidance in schooling fish 1731:"Characteristics and Application of 58:adding citations to reliable sources 3087:Intermediate disturbance hypothesis 2840:Ecological effects of biodiversity 310:as their inorganic carbon source. 25: 2176:Generalist and specialist species 995:phototrophically during the day. 499:which use the chemical energy in 324:are not taken into account here. 4082: 2899:Occupancyā€“abundance relationship 1288:Microbial Growth on C1 Compounds 1267:Nutrition and Growth of Bacteria 1126:as the carbon source instead of 818:anaerobic methanotrophic archaea 34: 3618:Substrate-level phosphorylation 2919:Relative abundance distribution 2632:Plant defense against herbivory 2499:Competitive exclusion principle 2211:Mesopredator release hypothesis 457: 452: 447: 437: 429: 424: 416: 408: 400: 397: 386: 383: 163:substrate-level phosphorylation 45:needs additional citations for 2504:Consumerā€“resource interactions 236:The freed energy is stored as 1: 4021:Reverse cholesterol transport 3350:Biological data visualization 3177:Environmental niche modelling 2904:Population viability analysis 964:organisms may dominate their 471:is one that requires organic 359:) as their source of carbon. 292:reduction-oxidation reactions 3673:(proteinā†’peptideā†’amino acid) 2835:Density-dependent inhibition 1735:as a Microbial Cell Factory" 1296:10.1007/978-94-009-3539-6_23 1290:. Springer. pp. 186ā€“7. 934: 824:, synthetically autotrophic 69:"Primary nutritional groups" 3304:Liebig's law of the minimum 3139:Resource selection function 2030:Metabolic theory of ecology 1110:as the source of electrons. 173:in aerobic respiration, or 4126: 4058:Phospagen system (ATP-PCr) 3528:Primary nutritional groups 3204:Niche apportionment models 2924:Relative species abundance 2128:Primary nutritional groups 2025:List of feeding behaviours 1733:Rhodopseudomonas palustris 1558:10.1016/j.cell.2019.11.009 1031:as carbon source, whereas 1017:, using light for energy, 1001:show a great diversity of 953:Rhodopseudomonas palustris 928: 920:when the source is water. 687:Purple non-sulfur bacteria 133:Primary nutritional groups 3920: 3892:Anoxygenic photosynthesis 3882: 3846: 3814:Pentose phosphate pathway 3809: 3792: 3775: 3569:Oxidative phosphorylation 3453: 3385:Ecosystem based fisheries 2997:Interspecific competition 2889:Minimum viable population 2747:Maximum sustainable yield 2732:Intraspecific competition 2727:Effective population size 2607:Anti-predator adaptations 2118:Photosynthetic efficiency 1792:Eiler A (December 2006). 1752:10.3389/fbioe.2022.897003 1618:10.1038/s41587-019-0363-0 1201:Eiler A (December 2006). 1100:evolved in the laboratory 1033:green non-sulfur bacteria 840: 744: 731: 657: 623:green non-sulfur bacteria 579: 570: 530:instead use oxidation of 419: 413: 332:Primary sources of carbon 268:use organic compounds as 233:release chemical energy. 213:Primary sources of energy 3912:Entner-Doudoroff pathway 3574:electron transport chain 3561:Pyruvate decarboxylation 3375:Ecological stoichiometry 3340:Alternative stable state 1873:10.3389/fmicb.2011.00165 1696:Archives of Microbiology 1391:10.3389/fgene.2013.00016 1351:. Springer. p. 68. 1041:chemoorganoheterotrophic 1037:photoorganoheterotrophic 987:chemoorganoheterotrophic 538:Primary metabolism table 466:chemoorganoheterotrophic 270:electron/hydrogen donors 4006:Sphingolipid metabolism 3907:DeLey-Doudoroff pathway 3755:carbohydrate catabolism 3750:Carbohydrate metabolism 3736:Purine nucleotide cycle 3219:Ontogenetic niche shift 3082:Ideal free distribution 2992:Ecological facilitation 2742:Malthusian growth model 2712:Consumer-resource model 2569:Paradox of the plankton 2534:Energy systems language 2254:Chemoorganoheterotrophy 2221:Optimal foraging theory 2196:Heterotrophic nutrition 1653:Oceanithermus profundus 1501:10.1073/pnas.1208795109 869:Oceanithermus profundus 534:as a source of energy. 286:or hydrogen atoms from 3975:Fatty acid degradation 3695:Amino acid degradation 3365:Ecological forecasting 3309:Marginal value theorem 3107:Landscape epidemiology 3042:Cross-boundary subsidy 2977:Biological interaction 2327:Microbial intelligence 2015:Green world hypothesis 1594:"The industrial yeast 1334:(5th ed.): 302ā€“3. 1096:genetically engineered 1068:Chemolithoheterotrophs 1011:purple sulfur bacteria 1003:nutritional categories 927:is from Ancient Greek 923:The common final part 495:Chemoorganotrophs are 372: 4011:Eicosanoid metabolism 3967:Fatty acid metabolism 3731:Pyrimidine metabolism 3590:Anaerobic respiration 3370:Ecological humanities 3269:Ecological energetics 3214:Niche differentiation 3077:Habitat fragmentation 2845:Ecological extinction 2792:Small population size 2544:Feed conversion ratio 2524:Ecological succession 2456:San Francisco Estuary 2370:Ecological efficiency 2312:Microbial cooperation 1672:10.1099/ijs.0.02367-0 1378:Frontiers in Genetics 1073:Photoorganoautotrophs 1015:photolithoautotrophic 978:photolithoautotrophic 370: 298:(in heterotrophs) or 151:anaerobic respiration 3985:Fatty acid synthesis 3690:Amino acid synthesis 3395:Evolutionary ecology 3360:Ecological footprint 3355:Ecological economics 3279:Ecological threshold 3274:Ecological indicator 3144:Sourceā€“sink dynamics 3097:Land change modeling 3092:Insular biogeography 2944:Species distribution 2683:Modelling ecosystems 2342:Microbial metabolism 2181:Intraguild predation 1970:Biogeochemical cycle 1936:Modelling ecosystems 1818:10.1128/AEM.01559-06 1606:Nature Biotechnology 1552:(6): 1255ā€“1263.e12. 1227:10.1128/AEM.01559-06 1189:Notes and references 1083:model microorganisms 1062:chemolithoautotrophs 1055:bacteria as well as 715:), some eukaryotes ( 288:reducing equivalents 54:improve this article 3549:Aerobic respiration 3445:Theoretical ecology 3420:Natural environment 3284:Ecosystem diversity 3254:Ecological collapse 3244:Bateman's principle 3199:Limiting similarity 3112:Landscape limnology 2934:Species homogeneity 2772:Population modeling 2767:Population dynamics 2584:Trophic state index 1810:2006ApEnM..72.7431E 1492:2012PNAS..10919321K 1345:Andrews JH (1991). 1219:2006ApEnM..72.7431E 532:inorganic compounds 426:Inorganic compounds 377: 145:aerobic respiration 4053:Ethanol metabolism 4001:Steroid metabolism 3726:Nucleotide salvage 3658:Protein metabolism 3456:Outline of ecology 3405:Industrial ecology 3400:Functional ecology 3264:Ecological deficit 3209:Niche construction 3172:Ecosystem engineer 2949:Speciesā€“area curve 2870:Introduced species 2685:: Other components 2617:Deimatic behaviour 2519:Ecological network 2451:North Pacific Gyre 2436:hydrothermal vents 2375:Ecological pyramid 2322:Microbial food web 2133:Primary production 2078:Foundation species 1708:10.1007/BF00424781 916:*Some authors use 375: 373: 27:Group of organisms 18:Chemoorganotrophic 4070: 4069: 4066: 4065: 4029: 4028: 3942: 3941: 3938: 3937: 3925:Xylose metabolism 3871: 3870: 3744: 3743: 3721:Purine metabolism 3666:Protein synthesis 3643: 3642: 3565:Citric acid cycle 3523:Metabolic network 3518:Metabolic pathway 3462: 3461: 3345:Balance of nature 3102:Landscape ecology 2987:Community ecology 2929:Species diversity 2865:Indicator species 2860:Gradient analysis 2737:Logistic function 2645: 2644: 2602:Animal coloration 2579:Trophic mutualism 2317:Microbial ecology 2108:Photoheterotrophs 2093:Myco-heterotrophy 2005:Ecosystem ecology 1990:Carrying capacity 1955:Abiotic component 1439:978-0-86720-081-2 1358:978-0-387-97439-2 1305:978-94-010-8082-8 1079:Synthetic biology 914: 913: 900:), some archaea ( 501:organic compounds 462: 461: 439:Organic compounds 410:Organic compounds 363:Energy and carbon 130: 129: 122: 104: 16:(Redirected from 4117: 4087: 4086: 4078: 4041:Metal metabolism 3963: 3948:Lipid metabolism 3880: 3773: 3766: 3678: 3654: 3580: 3545: 3489: 3482: 3475: 3466: 3162:Ecological niche 3134:selection theory 2954:Umbrella species 2939:Species richness 2875:Invasive species 2855:Flagship species 2762:Population cycle 2757:Overexploitation 2722:Ecological yield 2672: 2665: 2658: 2649: 2554:Mesotrophic soil 2494:Climax community 2426:Marine food webs 2365:Biomagnification 2166:Chemoorganotroph 2020:Keystone species 1980:Biotic component 1925: 1918: 1911: 1902: 1896: 1895: 1885: 1875: 1851: 1840: 1839: 1829: 1789: 1783: 1782: 1772: 1754: 1726: 1720: 1719: 1691: 1685: 1684: 1674: 1665:(Pt 3): 747ā€“52. 1646: 1640: 1639: 1629: 1589: 1580: 1579: 1569: 1533: 1524: 1523: 1513: 1503: 1471: 1465: 1455: 1444: 1443: 1420: 1414: 1413: 1403: 1393: 1369: 1363: 1362: 1342: 1336: 1335: 1325: 1316: 1310: 1309: 1283: 1277: 1276: 1274: 1273: 1261:Todar K (2009). 1258: 1252: 1251: 1238: 1198: 1091:Escherichia coli 1045:Sulfur-oxidizing 1026: 937: 932: 891: 888: 885: 864: 861: 858: 827:Escherichia coli 813: 810: 807: 767: 764: 761: 717:eukaryotic algae 708: 705: 702: 683: 680: 677: 646: 643: 640: 606: 603: 600: 545: 528:chemolithotrophs 378: 238:potential energy 222:absorb light in 204: 203: 202: 188: 187: 186: 125: 118: 114: 111: 105: 103: 62: 38: 30: 21: 4125: 4124: 4120: 4119: 4118: 4116: 4115: 4114: 4105:Trophic ecology 4095: 4094: 4093: 4081: 4073: 4071: 4062: 4046:Iron metabolism 4025: 3989: 3950: 3934: 3916: 3902:Carbon fixation 3867: 3842: 3805: 3788: 3784:Gluconeogenesis 3757: 3752: 3740: 3712: 3705: 3676: 3649: 3639: 3600: 3584: 3572: 3539: 3532: 3506: 3493: 3463: 3458: 3449: 3435:Systems ecology 3323: 3294:Extinction debt 3259:Ecological debt 3249:Bioluminescence 3230: 3223: 3192:marine habitats 3167:Ecological trap 3148: 3028: 3021: 2964: 2958: 2914:Rapoport's rule 2909:Priority effect 2850:Endemic species 2818: 2777:Population size 2693: 2686: 2676: 2646: 2641: 2594: 2588: 2574:Trophic cascade 2484:Bioaccumulation 2467: 2394: 2351: 2273: 2240: 2137: 2049: 2010:Ecosystem model 1943: 1929: 1899: 1860:Front Microbiol 1853: 1852: 1843: 1791: 1790: 1786: 1728: 1727: 1723: 1693: 1692: 1688: 1648: 1647: 1643: 1601: 1596:Pichia pastoris 1591: 1590: 1583: 1541: 1535: 1534: 1527: 1486:(47): 19321ā€“6. 1473: 1472: 1468: 1456: 1447: 1440: 1422: 1421: 1417: 1371: 1370: 1366: 1359: 1344: 1343: 1339: 1323: 1318: 1317: 1313: 1306: 1285: 1284: 1280: 1271: 1269: 1260: 1259: 1255: 1200: 1199: 1195: 1191: 1186: 1137: 1125: 1119:Pichia pastoris 1105: 1030: 1023: 1018: 1005:. For example, 983: 974: 944: 903:Methanobacteria 894:Some bacteria ( 889: 886: 883: 877: 867:Some bacteria ( 862: 859: 856: 850: 842: 833:Pichia pastoris 811: 808: 805: 799: 765: 762: 759: 753: 746: 737: 735: 733: 711:Some bacteria ( 706: 703: 700: 694: 681: 678: 675: 669: 659: 644: 641: 638: 630: 609:Some bacteria: 604: 601: 598: 590: 581: 572: 556: 554: 549: 540: 365: 358: 345: 334: 309: 273: 261: 227: 215: 201: 198: 197: 196: 194: 185: 182: 181: 180: 178: 171: 126: 115: 109: 106: 63: 61: 51: 39: 28: 23: 22: 15: 12: 11: 5: 4123: 4121: 4113: 4112: 4107: 4097: 4096: 4092: 4091: 4068: 4067: 4064: 4063: 4061: 4060: 4055: 4050: 4049: 4048: 4037: 4035: 4031: 4030: 4027: 4026: 4024: 4023: 4018: 4013: 4008: 4003: 3997: 3995: 3991: 3990: 3988: 3987: 3982: 3979:Beta oxidation 3971: 3969: 3960: 3944: 3943: 3940: 3939: 3936: 3935: 3933: 3932: 3927: 3921: 3918: 3917: 3915: 3914: 3909: 3904: 3899: 3897:Chemosynthesis 3894: 3889: 3887:Photosynthesis 3883: 3877: 3873: 3872: 3869: 3868: 3866: 3865: 3864: 3863: 3858: 3847: 3844: 3843: 3841: 3840: 3839: 3838: 3836:Leloir pathway 3828: 3827: 3826: 3824:Polyol pathway 3816: 3810: 3807: 3806: 3804: 3803: 3797:Glycogenolysis 3793: 3790: 3789: 3787: 3786: 3776: 3770: 3763: 3746: 3745: 3742: 3741: 3739: 3738: 3733: 3728: 3723: 3717: 3715: 3707: 3706: 3704: 3703: 3698: 3692: 3686: 3684: 3675: 3674: 3668: 3662: 3660: 3651: 3645: 3644: 3641: 3640: 3638: 3637: 3636: 3635: 3630: 3625: 3610: 3608: 3602: 3601: 3599: 3598: 3594: 3592: 3586: 3585: 3583: 3582: 3553: 3551: 3542: 3534: 3533: 3531: 3530: 3525: 3520: 3514: 3512: 3508: 3507: 3494: 3492: 3491: 3484: 3477: 3469: 3460: 3459: 3454: 3451: 3450: 3448: 3447: 3442: 3437: 3432: 3427: 3422: 3417: 3415:Microecosystem 3412: 3407: 3402: 3397: 3392: 3387: 3382: 3377: 3372: 3367: 3362: 3357: 3352: 3347: 3342: 3337: 3331: 3329: 3325: 3324: 3322: 3321: 3316: 3314:Thorson's rule 3311: 3306: 3301: 3296: 3291: 3286: 3281: 3276: 3271: 3266: 3261: 3256: 3251: 3246: 3241: 3239:Assembly rules 3235: 3233: 3225: 3224: 3222: 3221: 3216: 3211: 3206: 3201: 3196: 3195: 3194: 3184: 3179: 3174: 3169: 3164: 3158: 3156: 3150: 3149: 3147: 3146: 3141: 3136: 3124: 3122:Patch dynamics 3119: 3117:Metapopulation 3114: 3109: 3104: 3099: 3094: 3089: 3084: 3079: 3074: 3069: 3064: 3059: 3054: 3049: 3044: 3039: 3033: 3031: 3023: 3022: 3020: 3019: 3014: 3012:Storage effect 3009: 3004: 2999: 2994: 2989: 2984: 2979: 2974: 2968: 2966: 2960: 2959: 2957: 2956: 2951: 2946: 2941: 2936: 2931: 2926: 2921: 2916: 2911: 2906: 2901: 2896: 2894:Neutral theory 2891: 2886: 2881: 2879:Native species 2872: 2867: 2862: 2857: 2852: 2847: 2842: 2837: 2832: 2826: 2824: 2820: 2819: 2817: 2816: 2811: 2810: 2809: 2804: 2794: 2789: 2784: 2779: 2774: 2769: 2764: 2759: 2754: 2752:Overpopulation 2749: 2744: 2739: 2734: 2729: 2724: 2719: 2714: 2709: 2704: 2698: 2696: 2688: 2687: 2677: 2675: 2674: 2667: 2660: 2652: 2643: 2642: 2640: 2639: 2634: 2629: 2624: 2619: 2614: 2609: 2604: 2598: 2596: 2590: 2589: 2587: 2586: 2581: 2576: 2571: 2566: 2561: 2559:Nutrient cycle 2556: 2551: 2549:Feeding frenzy 2546: 2541: 2536: 2531: 2529:Energy quality 2526: 2521: 2516: 2511: 2506: 2501: 2496: 2491: 2489:Cascade effect 2486: 2481: 2475: 2473: 2469: 2468: 2466: 2465: 2464: 2463: 2458: 2453: 2448: 2443: 2438: 2433: 2423: 2418: 2413: 2408: 2402: 2400: 2396: 2395: 2393: 2392: 2387: 2382: 2377: 2372: 2367: 2361: 2359: 2353: 2352: 2350: 2349: 2344: 2339: 2334: 2332:Microbial loop 2329: 2324: 2319: 2314: 2309: 2304: 2299: 2297:Lithoautotroph 2294: 2289: 2283: 2281: 2279:Microorganisms 2275: 2274: 2272: 2271: 2266: 2261: 2256: 2250: 2248: 2242: 2241: 2239: 2238: 2236:Prey switching 2233: 2228: 2223: 2218: 2213: 2208: 2203: 2198: 2193: 2188: 2183: 2178: 2173: 2168: 2163: 2158: 2153: 2147: 2145: 2139: 2138: 2136: 2135: 2130: 2125: 2120: 2115: 2113:Photosynthesis 2110: 2105: 2100: 2095: 2090: 2085: 2080: 2075: 2070: 2068:Chemosynthesis 2065: 2059: 2057: 2051: 2050: 2048: 2047: 2042: 2037: 2032: 2027: 2022: 2017: 2012: 2007: 2002: 1997: 1992: 1987: 1982: 1977: 1972: 1967: 1962: 1960:Abiotic stress 1957: 1951: 1949: 1945: 1944: 1930: 1928: 1927: 1920: 1913: 1905: 1898: 1897: 1841: 1804:(12): 7431ā€“7. 1784: 1721: 1686: 1641: 1599: 1581: 1539: 1525: 1466: 1463:978-1319017637 1445: 1438: 1415: 1364: 1357: 1337: 1311: 1304: 1278: 1253: 1213:(12): 7431ā€“7. 1192: 1190: 1187: 1185: 1184: 1179: 1174: 1169: 1164: 1159: 1154: 1149: 1147:Chemosynthesis 1144: 1138: 1136: 1133: 1132: 1131: 1123: 1115:methylotrophic 1111: 1103: 1028: 981: 973: 970: 943: 940: 912: 911: 908:Chemosynthesis 892: 881: 876:Carbon dioxide 873: 872: 865: 854: 847: 838: 837: 822:Chemosynthesis 816:Some archaea ( 814: 803: 798:Carbon dioxide 795: 794: 768: 757: 750: 743: 729: 728: 725:Photosynthesis 709: 698: 693:Carbon dioxide 690: 689: 684: 673: 666: 655: 654: 649:Some archaea ( 647: 636: 629:Carbon dioxide 626: 625: 618:Heliobacterium 607: 596: 587: 578: 568: 567: 564: 561: 560:Carbon source 558: 551: 539: 536: 460: 459: 456: 454:Carbon dioxide 450: 449: 446: 443: 436: 432: 431: 428: 422: 421: 418: 415: 412: 407: 406:Electron donor 403: 402: 399: 395: 394: 391: 388: 385: 382: 364: 361: 356: 353:carbon dioxide 333: 330: 307: 260: 257: 224:photoreceptors 214: 211: 199: 183: 169: 135:are groups of 128: 127: 110:September 2010 42: 40: 33: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 4122: 4111: 4108: 4106: 4103: 4102: 4100: 4090: 4085: 4080: 4076: 4059: 4056: 4054: 4051: 4047: 4044: 4043: 4042: 4039: 4038: 4036: 4032: 4022: 4019: 4017: 4014: 4012: 4009: 4007: 4004: 4002: 3999: 3998: 3996: 3992: 3986: 3983: 3980: 3976: 3973: 3972: 3970: 3968: 3964: 3961: 3958: 3954: 3949: 3945: 3931: 3930:Radiotrophism 3928: 3926: 3923: 3922: 3919: 3913: 3910: 3908: 3905: 3903: 3900: 3898: 3895: 3893: 3890: 3888: 3885: 3884: 3881: 3878: 3874: 3862: 3859: 3857: 3854: 3853: 3852: 3851:Glycosylation 3849: 3848: 3845: 3837: 3834: 3833: 3832: 3829: 3825: 3822: 3821: 3820: 3817: 3815: 3812: 3811: 3808: 3802: 3798: 3795: 3794: 3791: 3785: 3781: 3778: 3777: 3774: 3771: 3767: 3764: 3761: 3756: 3751: 3747: 3737: 3734: 3732: 3729: 3727: 3724: 3722: 3719: 3718: 3716: 3714: 3708: 3702: 3699: 3696: 3693: 3691: 3688: 3687: 3685: 3683: 3679: 3672: 3669: 3667: 3664: 3663: 3661: 3659: 3655: 3652: 3646: 3634: 3631: 3629: 3626: 3624: 3621: 3620: 3619: 3615: 3612: 3611: 3609: 3607: 3603: 3596: 3595: 3593: 3591: 3587: 3579: 3575: 3570: 3566: 3562: 3558: 3555: 3554: 3552: 3550: 3546: 3543: 3541: 3535: 3529: 3526: 3524: 3521: 3519: 3516: 3515: 3513: 3509: 3505: 3501: 3497: 3490: 3485: 3483: 3478: 3476: 3471: 3470: 3467: 3457: 3452: 3446: 3443: 3441: 3440:Urban ecology 3438: 3436: 3433: 3431: 3428: 3426: 3423: 3421: 3418: 3416: 3413: 3411: 3408: 3406: 3403: 3401: 3398: 3396: 3393: 3391: 3388: 3386: 3383: 3381: 3378: 3376: 3373: 3371: 3368: 3366: 3363: 3361: 3358: 3356: 3353: 3351: 3348: 3346: 3343: 3341: 3338: 3336: 3333: 3332: 3330: 3326: 3320: 3317: 3315: 3312: 3310: 3307: 3305: 3302: 3300: 3299:Kleiber's law 3297: 3295: 3292: 3290: 3287: 3285: 3282: 3280: 3277: 3275: 3272: 3270: 3267: 3265: 3262: 3260: 3257: 3255: 3252: 3250: 3247: 3245: 3242: 3240: 3237: 3236: 3234: 3232: 3226: 3220: 3217: 3215: 3212: 3210: 3207: 3205: 3202: 3200: 3197: 3193: 3190: 3189: 3188: 3185: 3183: 3180: 3178: 3175: 3173: 3170: 3168: 3165: 3163: 3160: 3159: 3157: 3155: 3151: 3145: 3142: 3140: 3137: 3135: 3133: 3129: 3125: 3123: 3120: 3118: 3115: 3113: 3110: 3108: 3105: 3103: 3100: 3098: 3095: 3093: 3090: 3088: 3085: 3083: 3080: 3078: 3075: 3073: 3072:Foster's rule 3070: 3068: 3065: 3063: 3060: 3058: 3055: 3053: 3050: 3048: 3045: 3043: 3040: 3038: 3035: 3034: 3032: 3030: 3024: 3018: 3015: 3013: 3010: 3008: 3005: 3003: 3000: 2998: 2995: 2993: 2990: 2988: 2985: 2983: 2980: 2978: 2975: 2973: 2970: 2969: 2967: 2961: 2955: 2952: 2950: 2947: 2945: 2942: 2940: 2937: 2935: 2932: 2930: 2927: 2925: 2922: 2920: 2917: 2915: 2912: 2910: 2907: 2905: 2902: 2900: 2897: 2895: 2892: 2890: 2887: 2885: 2882: 2880: 2876: 2873: 2871: 2868: 2866: 2863: 2861: 2858: 2856: 2853: 2851: 2848: 2846: 2843: 2841: 2838: 2836: 2833: 2831: 2828: 2827: 2825: 2821: 2815: 2812: 2808: 2805: 2803: 2800: 2799: 2798: 2795: 2793: 2790: 2788: 2785: 2783: 2780: 2778: 2775: 2773: 2770: 2768: 2765: 2763: 2760: 2758: 2755: 2753: 2750: 2748: 2745: 2743: 2740: 2738: 2735: 2733: 2730: 2728: 2725: 2723: 2720: 2718: 2715: 2713: 2710: 2708: 2705: 2703: 2700: 2699: 2697: 2695: 2689: 2684: 2680: 2673: 2668: 2666: 2661: 2659: 2654: 2653: 2650: 2638: 2635: 2633: 2630: 2628: 2625: 2623: 2620: 2618: 2615: 2613: 2610: 2608: 2605: 2603: 2600: 2599: 2597: 2591: 2585: 2582: 2580: 2577: 2575: 2572: 2570: 2567: 2565: 2562: 2560: 2557: 2555: 2552: 2550: 2547: 2545: 2542: 2540: 2537: 2535: 2532: 2530: 2527: 2525: 2522: 2520: 2517: 2515: 2512: 2510: 2507: 2505: 2502: 2500: 2497: 2495: 2492: 2490: 2487: 2485: 2482: 2480: 2477: 2476: 2474: 2470: 2462: 2459: 2457: 2454: 2452: 2449: 2447: 2444: 2442: 2439: 2437: 2434: 2432: 2429: 2428: 2427: 2424: 2422: 2419: 2417: 2414: 2412: 2409: 2407: 2404: 2403: 2401: 2397: 2391: 2390:Trophic level 2388: 2386: 2383: 2381: 2378: 2376: 2373: 2371: 2368: 2366: 2363: 2362: 2360: 2358: 2354: 2348: 2347:Phage ecology 2345: 2343: 2340: 2338: 2337:Microbial mat 2335: 2333: 2330: 2328: 2325: 2323: 2320: 2318: 2315: 2313: 2310: 2308: 2305: 2303: 2300: 2298: 2295: 2293: 2292:Bacteriophage 2290: 2288: 2285: 2284: 2282: 2280: 2276: 2270: 2267: 2265: 2262: 2260: 2259:Decomposition 2257: 2255: 2252: 2251: 2249: 2247: 2243: 2237: 2234: 2232: 2229: 2227: 2224: 2222: 2219: 2217: 2214: 2212: 2209: 2207: 2206:Mesopredators 2204: 2202: 2199: 2197: 2194: 2192: 2189: 2187: 2184: 2182: 2179: 2177: 2174: 2172: 2169: 2167: 2164: 2162: 2159: 2157: 2154: 2152: 2151:Apex predator 2149: 2148: 2146: 2144: 2140: 2134: 2131: 2129: 2126: 2124: 2121: 2119: 2116: 2114: 2111: 2109: 2106: 2104: 2101: 2099: 2096: 2094: 2091: 2089: 2086: 2084: 2081: 2079: 2076: 2074: 2071: 2069: 2066: 2064: 2061: 2060: 2058: 2056: 2052: 2046: 2043: 2041: 2038: 2036: 2033: 2031: 2028: 2026: 2023: 2021: 2018: 2016: 2013: 2011: 2008: 2006: 2003: 2001: 1998: 1996: 1993: 1991: 1988: 1986: 1985:Biotic stress 1983: 1981: 1978: 1976: 1973: 1971: 1968: 1966: 1963: 1961: 1958: 1956: 1953: 1952: 1950: 1946: 1941: 1937: 1933: 1926: 1921: 1919: 1914: 1912: 1907: 1906: 1903: 1893: 1889: 1884: 1879: 1874: 1869: 1865: 1861: 1857: 1850: 1848: 1846: 1842: 1837: 1833: 1828: 1823: 1819: 1815: 1811: 1807: 1803: 1799: 1795: 1788: 1785: 1780: 1776: 1771: 1766: 1762: 1758: 1753: 1748: 1744: 1740: 1736: 1734: 1725: 1722: 1717: 1713: 1709: 1705: 1701: 1697: 1690: 1687: 1682: 1678: 1673: 1668: 1664: 1660: 1656: 1654: 1645: 1642: 1637: 1633: 1628: 1623: 1619: 1615: 1611: 1607: 1603: 1597: 1588: 1586: 1582: 1577: 1573: 1568: 1563: 1559: 1555: 1551: 1547: 1543: 1532: 1530: 1526: 1521: 1517: 1512: 1507: 1502: 1497: 1493: 1489: 1485: 1481: 1477: 1470: 1467: 1464: 1460: 1454: 1452: 1450: 1446: 1441: 1435: 1431: 1430: 1425: 1419: 1416: 1411: 1407: 1402: 1397: 1392: 1387: 1383: 1379: 1375: 1368: 1365: 1360: 1354: 1350: 1349: 1341: 1338: 1333: 1329: 1322: 1315: 1312: 1307: 1301: 1297: 1293: 1289: 1282: 1279: 1268: 1264: 1257: 1254: 1250: 1246: 1242: 1237: 1232: 1228: 1224: 1220: 1216: 1212: 1208: 1204: 1197: 1194: 1188: 1183: 1180: 1178: 1177:Organotrophic 1175: 1173: 1170: 1168: 1165: 1163: 1160: 1158: 1157:Heterotrophic 1155: 1153: 1150: 1148: 1145: 1143: 1140: 1139: 1134: 1129: 1121: 1120: 1116: 1112: 1109: 1101: 1097: 1093: 1092: 1088: 1087: 1086: 1084: 1080: 1076: 1074: 1070: 1069: 1064: 1063: 1058: 1054: 1050: 1046: 1042: 1038: 1034: 1025: 1016: 1012: 1008: 1007:cyanobacteria 1004: 1000: 996: 993: 989: 988: 979: 971: 969: 967: 963: 959: 955: 954: 949: 948:Chroococcales 941: 939: 938:"nutrition". 936: 931: 926: 921: 919: 909: 905: 904: 899: 898: 893: 882: 880: 875: 874: 870: 866: 855: 853: 848: 845: 839: 835: 834: 830:bacteria and 829: 828: 823: 819: 815: 804: 802: 797: 796: 792: 788: 784: 780: 776: 772: 769: 758: 756: 751: 749: 742: 741: 730: 726: 722: 718: 714: 713:cyanobacteria 710: 699: 697: 692: 691: 688: 685: 674: 672: 667: 664: 663: 656: 652: 648: 637: 635: 634: 628: 627: 624: 620: 619: 614: 613: 608: 597: 595: 594: 588: 586: 585: 577: 576: 569: 565: 562: 559: 552: 547: 546: 543: 537: 535: 533: 529: 525: 521: 517: 513: 508: 506: 502: 498: 493: 491: 487: 483: 478: 474: 470: 467: 455: 451: 442: 440: 435:Carbon source 433: 427: 423: 411: 404: 396: 381:Energy source 379: 371:Yellow fungus 369: 362: 360: 354: 350: 349: 343: 340: 339: 331: 329: 325: 323: 319: 314: 311: 303: 301: 297: 296:ATP synthesis 293: 289: 285: 280: 278: 277: 271: 267: 266: 258: 256: 253: 251: 247: 246:carbohydrates 243: 239: 234: 232: 231: 225: 221: 220: 212: 210: 208: 192: 176: 172: 165: 164: 159: 158: 153: 152: 147: 146: 140: 138: 134: 124: 121: 113: 102: 99: 95: 92: 88: 85: 81: 78: 74: 71: ā€“  70: 66: 65:Find sources: 59: 55: 49: 48: 43:This article 41: 37: 32: 31: 19: 3831:Galactolysis 3801:Glycogenesis 3606:Fermentation 3578:ATP synthase 3527: 3425:Regime shift 3410:Macroecology 3131: 3127: 3067:Edge effects 3037:Biogeography 2982:Commensalism 2830:Biodiversity 2707:Allee effect 2446:kelp forests 2399:Example webs 2264:Detritivores 2127: 2103:Organotrophs 2083:Kinetotrophs 2035:Productivity 1863: 1859: 1801: 1797: 1787: 1742: 1738: 1732: 1724: 1702:(1): 93ā€“98. 1699: 1695: 1689: 1662: 1658: 1652: 1644: 1612:(2): 210ā€“6. 1609: 1605: 1595: 1549: 1545: 1483: 1479: 1469: 1428: 1418: 1381: 1377: 1367: 1347: 1340: 1331: 1327: 1314: 1287: 1281: 1270:. Retrieved 1266: 1256: 1248: 1210: 1206: 1196: 1182:Phototrophic 1162:Lithotrophic 1152:Chemotrophic 1117: 1089: 1077: 1072: 1066: 1061: 1040: 1036: 1014: 997: 986: 977: 975: 951: 947: 945: 924: 922: 917: 915: 901: 895: 878: 868: 852:-heterotroph 851: 843: 831: 825: 800: 755:-heterotroph 754: 747: 738: 695: 671:-heterotroph 670: 660: 631: 617: 610: 593:-heterotroph 591: 582: 573: 541: 509: 494: 465: 463: 441: 347: 338:Heterotrophs 337: 335: 326: 322:fermentation 315: 312: 304: 300:biosynthesis 281: 275: 265:Organotrophs 264: 262: 254: 235: 229: 218: 216: 161: 157:fermentation 155: 149: 143: 141: 132: 131: 116: 107: 97: 90: 83: 76: 64: 52:Please help 47:verification 44: 3957:lipogenesis 3819:Fructolysis 3633:Lactic acid 3062:Disturbance 2965:interaction 2787:Recruitment 2717:Depensation 2509:Copiotrophs 2380:Energy flow 2302:Lithotrophy 2246:Decomposers 2226:Planktivore 2201:Insectivore 2191:Heterotroph 2156:Bacterivore 2123:Phototrophs 2073:Chemotrophs 2045:Restoration 1995:Competition 1172:Mixotrophic 1142:Autotrophic 1057:methanogens 999:Prokaryotes 962:mixotrophic 897:Nitrobacter 863:heterotroph 779:saprophytic 766:heterotroph 721:land plants 682:heterotroph 651:Haloarchaea 612:Rhodobacter 605:heterotroph 522:, and some 482:Decomposers 475:to get its 318:respiration 276:Lithotrophs 230:Chemotrophs 219:Phototrophs 4110:Physiology 4099:Categories 3780:Glycolysis 3713:metabolism 3711:Nucleotide 3701:Urea cycle 3682:Amino acid 3671:Catabolism 3614:Glycolysis 3557:Glycolysis 3540:metabolism 3500:catabolism 3496:Metabolism 3430:Sexecology 3007:Parasitism 2972:Antibiosis 2807:Resistance 2802:Resilience 2692:Population 2612:Camouflage 2564:Oligotroph 2479:Ascendency 2441:intertidal 2431:cold seeps 2385:Food chain 2186:Herbivores 2161:Carnivores 2088:Mixotrophs 2063:Autotrophs 1942:components 1745:: 897003. 1424:Margulis L 1272:2014-04-19 1167:Metabolism 992:eukaryotic 942:Mixotrophs 879:-autotroph 801:-autotroph 696:-autotroph 633:-autotroph 490:carnivores 486:Herbivores 473:substrates 348:Autotrophs 342:metabolize 142:The terms 80:newspapers 3953:lipolysis 3760:anabolism 3504:anabolism 3335:Allometry 3289:Emergence 3017:Symbiosis 3002:Mutualism 2797:Stability 2702:Abundance 2514:Dominance 2472:Processes 2461:tide pool 2357:Food webs 2231:Predation 2216:Omnivores 2143:Consumers 2098:Mycotroph 2055:Producers 2000:Ecosystem 1965:Behaviour 1761:2296-4185 1102:to use CO 1098:and then 1009:and many 890:autotroph 841:Inorganic 812:autotroph 775:parasitic 771:Predatory 736:Compounds 707:autotroph 658:Inorganic 645:autotroph 571:Sun Light 566:Examples 553:Electron/ 497:organisms 398:Molecules 284:electrons 137:organisms 3876:Nonhuman 3861:O-linked 3856:N-linked 3648:Specific 3390:Endolith 3319:Xerosere 3231:networks 3047:Ecocline 2593:Defense, 2269:Detritus 2171:Foraging 2040:Resource 1892:21866228 1836:17028233 1779:35646843 1681:12807196 1636:31844294 1576:31778652 1520:23129626 1410:23443991 1245:17028233 1135:See also 1128:methanol 972:Examples 783:protists 748:-organo- 734:Chemical 732:Breaking 584:-organo- 524:bacteria 520:protozoa 469:organism 448:hetero- 250:proteins 207:fumarate 4089:Biology 4016:Ketosis 3628:Ethanol 3511:General 3380:Ecopath 3187:Habitat 3057:Ecotype 3052:Ecotone 3029:ecology 3027:Spatial 2963:Species 2823:Species 2694:ecology 2679:Ecology 2627:Mimicry 2595:counter 2539:f-ratio 2287:Archaea 1975:Biomass 1948:General 1940:Trophic 1932:Ecology 1883:3149686 1866:: 165. 1827:1694265 1806:Bibcode 1770:9133744 1627:7008030 1567:6904909 1511:3511159 1488:Bibcode 1401:3580334 1236:1694265 1215:Bibcode 1108:formate 1053:anammox 1035:can be 1013:can be 966:habitat 918:-hydro- 849:Organic 844:-litho- 836:yeast. 791:animals 752:Organic 745:Organic 668:Organic 662:-litho- 621:, some 589:Organic 580:Organic 550:source 512:animals 505:glucose 430:litho- 420:  417:organo- 401:chemo- 393:-troph 191:sulfate 175:nitrate 94:scholar 4075:Portal 3538:Energy 2411:Rivers 2307:Marine 1890:  1880:  1834:  1824:  1777:  1767:  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Index

Chemoorganotrophic

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"Primary nutritional groups"
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JSTOR
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organisms
aerobic respiration
anaerobic respiration
fermentation
substrate-level phosphorylation
O2
nitrate
sulfate
fumarate
Phototrophs
photoreceptors
Chemotrophs
potential energy
ATP
carbohydrates
proteins
Organotrophs
electron/hydrogen donors

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