492:
44:
88:
387:
409:
63:
1008:
482:
Due to its tiny size, this tortoise is heavily preyed on by crows, ostriches, jackals, baboons, dogs, and a wide range of other predators. Consequently, it spends most of its time hiding under rocks, foliage, and other cover. It lays a clutch of 2–4 eggs, which hatch 150–320 days later – usually on a
416:
The males are smaller than the females, and can be distinguished by their slightly longer tails and their distinctive heads. Males have larger heads, with a larger beak and a more pointy snout. The noses of dominant males also become bright orange or red in the breeding season.
405:, and unlike the other padlopers, it has only four toes on its front feet, as well as its hind feet. The average length is 110 mm, though females are larger than males. Their adult weight ranges from 140 to 300 g. It has a sharp, distinctly hooked beak.
397:
The common padloper is a small, rather flat tortoise. Its shell usually has varied colouration, ranging between olive-green and brown. The shields of the carapace are flat, with large raised areolae, and a thin black edging.
420:
In colour, males are frequently uniformly orange to light brown (compared to the deeper olive brown of the females). Males also have more lightly coloured bellies, though they do not exhibit the
875:
Schleicher, A. 2005. Die
Areolen-Flachschildkröte, Homopus areolatus (THUNBERG 1787). Beobachtungen und Erfahrungen: Lebensweise, Haltung, Nach- und Aufzucht. Reptilia (Münster) 10 (6): 36-40
872:
Loveridge, Arthur & Williams, Ernest E. 1957. Revision of the
African Tortoises and Turtles of the Suborder Cryptodira. Bulletin of the Museum of Comparative Zoology 115 (6): 163-557
857:
Fritz, Uwe and Olaf R.P. Bininda-Emonds 2007. When genes meet nomenclature: Tortoise phylogeny and the shifting generic concepts of
Testudo and Geochelone. Zoology 110 (4): 298-307
4381:
884:
Hughes, B. 1986. Longevity
Records of African Captive Amphibians and Reptiles: Part 1: Introduction and Species List 1 - Amphibians and Chelonians Jour. Herp. Ass. Afr. (32): 1-5
780:
863:
Greig, J.C., and P.D. Burdett. 1976. Patterns in the distributions of
Southern African terrestrial tortoises (Cryptodira: Testudinidae). Zool. Africana 11(2): 250–267.
4311:
510:
species is strictly illegal in South Africa and
Namibia. Any captive specimens are systematically registered in noncommercial studbooks, and any commercial sale of
4350:
881:
Thunberg, C. P. 1787. Beskrifning pé Trenne Skölpaddor Kongl. Vetenskaps
Academiens Nya Handlingar, För M_naderne 1787 (Julius, Augustus, September): 178-180
603:(2014). "Turtles of the World". In Rhodin, A.G.J.; Pritchard, P.C.H.; van Dijk, P.P.; Saumure, R.A.; Buhlmann, K.A.; Iverson, J.B.; Mittermeier, R.A. (eds.).
4417:
467:
At certain points, favourable conditions allow populations to survive further inland. In the
Western Cape, there are inland populations at Sutherland and
350:
is known by a wide range of common names. In its native region in southern Africa (and in much of the scientific community) it is usually known as the
4285:
869:
Lacepède, B. G. E. 1788. Histoire
Naturelle des Quadrupe des Ovipares et des Serpens. Vol.1. Imprimerie du Roi, Hôtel de Thou, Paris, xvii + 651 pp.
4324:
1253:
491:
1442:
1436:
909:
848:
Duméril, A. M.C., G. Bibron & A. Duméril 1854. Erpétologie générale ou
Histoire Naturelle complète des Reptiles. Vol. 9. Paris, XX + 440 S.
605:
Conservation Biology of Freshwater Turtles and Tortoises: A Compilation Project of the IUCN/SSC Tortoise and Freshwater Turtle Specialist Group
842:
Baard, E.H.W. 1996. Inter- and Intra-Individual Variation in the measurement of southern Padloper Homopus areolatus Afr. Herp News (25): 22-25
695:
2013:
522:
The species can adapt well to captivity – as its diet, while varied, is not highly specialized. It is, however, relatively rare as a pet.
854:
Fleck, J. & Fleck, S. 2001. Erfolgreiche Nachzucht von Chersina angulata und Homopus areolatus über mehrere Jahre. Elaphe 9 (3): 5-14
845:
Bour, R. 1980. Essai sur la taxinomie des Testudinidae actuels (Reptilia, Chelonii). Bull. Mus. natl. Hist. nat. Paris (4) 2 (2): 541-546
4432:
807:
738:
503:, traffic on roads, and increased frequency of wildfires. Another threat comes from introduced species, such as domestic dogs and pigs.
4427:
464:. The mild climate allows it to stay active throughout the year. It is generally intolerant of the arid conditions further inland.
1064:
670:
1295:
784:
43:
4046:
3129:
1424:
1418:
1290:
1049:
4329:
4173:
Turtles of the World 2017 Update: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status
1982:
turtles of the world 2017 update: Annotated checklist and atlas of taxonomy, synonymy, distribution, and conservation status
1678:
1493:
1663:
1653:
1315:
600:
4023:
4171:
1981:
4182:
2705:
2089:
1803:
1668:
1320:
1057:
4355:
2438:
1658:
1264:
902:
1673:
1430:
1081:
851:
Ernst, C.H. and Barbour, R.W. 1989. Turtles of the World. Smithsonian Institution Press, Washington D.C. - London
362:, due to the relatively large beak that the males of the species possess. Other names include the older name of
87:
3084:
2006:
1584:
1305:
1274:
1243:
1131:
860:
Gorseman, P. 1980. Opmerkingen over biotoop en voortplanting van Homopus areolatus. Lacerta 38 (10-11): 107-111
437:
1340:
1213:
3940:
1953:
1503:
1352:
1346:
1900:
1894:
1852:
651:
4199:
3912:
2712:
2670:
2310:
1912:
1858:
1552:
1534:
1310:
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1207:
1183:
1177:
1149:
1143:
1137:
1112:
1042:
4237:
1888:
1558:
1540:
2656:
2158:
1969:
1963:
1924:
1906:
1870:
1757:
1745:
1714:
1546:
1460:
1393:
1300:
1280:
1195:
1073:
895:
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202:
1948:
1930:
1630:
1564:
1528:
1228:
1223:
1882:
4272:
4246:
3898:
2564:
2188:
1918:
1465:
1201:
1189:
565:
1876:
1864:
1846:
3883:
2242:
1999:
1958:
1751:
1635:
1398:
1172:
500:
52:
1331:
3290:
3255:
3122:
2920:
2861:
2831:
1775:
1739:
1719:
1625:
1519:
1238:
238:
82:
2839:
2593:
4316:
4259:
3833:
2816:
2768:
2759:
2642:
2219:
1780:
1607:
1370:
1259:
1105:
1100:
878:
Schleicher, A. 2005. Homopus areolatus - The Parrot-Beaked Tortoise. Reptilia (GB) (43): 26-30
691:
804:
735:
4422:
4376:
4342:
3625:
3013:
2988:
2949:
2897:
2628:
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1826:
1696:
1388:
1268:
1024:
560:
402:
4363:
4251:
4073:
3721:
3450:
3418:
3055:
2964:
2740:
2326:
2071:
1940:
1808:
1498:
1488:
1483:
1284:
811:
742:
461:
457:
866:
Hoogmoed, M.S. 1980. Herpetologische waarnemingen in Ghana . Lacerta 38 (10-11): 112-116
621:
3968:
3958:
3855:
3705:
3545:
3247:
3192:
2934:
2869:
2409:
2400:
2369:
2277:
2166:
2080:
1795:
1575:
1508:
1167:
542:
468:
386:
4411:
4337:
4148:
4031:
3931:
3920:
3891:
3805:
3689:
3673:
3633:
3567:
3477:
3442:
3386:
3370:
3354:
3328:
3316:
3300:
3278:
3270:
3077:
2788:
2621:
2482:
2429:
2377:
1785:
1688:
1034:
967:
551:
159:
72:
67:
408:
4264:
4083:
4008:
3984:
3501:
3378:
3362:
3338:
3239:
3184:
3108:
3024:
2998:
2912:
2823:
2802:
2726:
2691:
2501:
2467:
2334:
2250:
2196:
1818:
1730:
1409:
1123:
973:
824:
445:
441:
421:
375:
336:
448:
Provinces. Here it favours coastal lowlands, especially along the south coast, in
4394:
4298:
4231:
4061:
4016:
3862:
3847:
3769:
3761:
3753:
3729:
3609:
3593:
3577:
3493:
3485:
3434:
3410:
3402:
3394:
3262:
3214:
3137:
3062:
2927:
2883:
2809:
2795:
2747:
2604:
2522:
2460:
2453:
2446:
2361:
2303:
2295:
2212:
2204:
2151:
2141:
1767:
1617:
1159:
1092:
479:
is relatively plentiful within its range, and hence the name "common padloper".
314:
17:
4140:
4053:
4039:
4000:
3777:
3697:
3665:
3649:
3617:
3555:
3528:
3509:
3426:
3231:
3151:
3144:
3092:
3069:
2979:
2846:
2578:
2385:
2318:
2235:
2227:
2130:
2121:
1452:
1380:
1016:
961:
955:
607:. Chelonian Research Monographs. Vol. 5 (7th ed.). pp. 329–479.
453:
149:
4222:
688:
The Completely Illustrated Atlas of Reptiles and Amphibians for the Terrarium
4132:
3992:
3976:
3905:
3869:
3819:
3812:
3737:
3713:
3657:
3601:
3308:
3100:
3048:
3006:
2719:
2677:
2585:
2571:
2557:
2550:
2543:
2536:
2508:
2353:
2181:
99:
4368:
471:. In the Eastern Cape, their range extends inland at one point, as far as
4216:
3947:
3876:
3796:
3785:
3681:
3346:
3223:
3115:
2957:
2942:
2904:
2853:
2779:
2684:
2663:
2649:
2635:
2491:
2418:
2287:
2266:
2258:
2040:
1837:
1706:
1475:
1362:
993:
937:
918:
433:
326:
169:
119:
712:
Reptiles and Amphibians in Captivity – Breeding, Longevity and Inventory
590:. Kongliga Vetenskaps Academiens Nya Handlingar, Stockholm (2)8:178–180.
4290:
4091:
3840:
3745:
3641:
3585:
3165:
3158:
3040:
2890:
2876:
2733:
2698:
2613:
2515:
2055:
2046:
1645:
1594:
949:
943:
331:
179:
129:
4303:
3826:
3469:
2061:
2034:
2022:
931:
449:
139:
109:
4193:
4389:
3033:
2100:
887:
655:
490:
407:
385:
2529:
4277:
4197:
1995:
891:
354:
due to its being by far the most commonly occurring of all the
686:
Fritz Jürgen Obst; Klaus Richter; Udo Jacob (1 January 1988).
536:
534:
1991:
412:
Orange-brown male, showing the larger, pointed head and beak
690:. Distributed in the United States by T.F.H. Publications.
637:
514:
tortoises is almost without exception strictly illegal.
358:("padloper") species. It is also commonly known as the
830:, World Chelonian Trust (retrieved August 20, 2013).
628:", World Chelonian Trust (retrieved August 20, 2013)
4206:
4123:
4116:
4109:
4071:
3957:
3930:
3795:
3565:
3543:
3536:
3527:
3460:
3326:
3288:
3213:
3206:
3175:
3023:
2987:
2978:
2778:
2757:
2603:
2490:
2481:
2428:
2408:
2399:
2344:
2276:
2140:
2129:
2120:
2088:
2079:
2070:
1939:
1835:
1817:
1794:
1766:
1728:
1705:
1687:
1644:
1616:
1593:
1573:
1517:
1474:
1451:
1407:
1379:
1361:
1329:
1222:
1158:
1122:
1091:
1033:
1015:
982:
681:
679:
566:10.2305/IUCN.UK.2018-2.RLTS.T170520A115656133.en
495:Injured beaked padloper, in an urban environment
757:. Southern Book Publishers: Johannesburg. 1988.
424:concavity that many other tortoise species do.
726:, Reptile & Amphibian Magazine, 1996-1997.
4170:Phylogenetic arrangement of turtles based on
2007:
1980:Phylogenetic arrangement of turtles based on
903:
768:Cape Tortoises: Their identification and care
370:. The latter name refers to its species name
8:
374:, and describes the raised rings around the
4194:
4120:
4113:
3540:
3533:
3210:
2984:
2487:
2405:
2137:
2126:
2094:
2085:
2076:
2014:
2000:
1992:
986:
910:
896:
888:
61:
42:
31:
647:
645:
564:
267:Lacepède 1788:166 (09:6) (nomen rejectum)
671:Homopus areolatus (Beaked Cape Tortoise)
736:IUCN Appendix 1. Regional Species Lists
616:
614:
530:
1443:Saddle-backed Rodrigues giant tortoise
1437:Saddle-backed Mauritius giant tortoise
666:
664:
475:. Unlike the other padloper species,
335:, indigenous to the southern part of
7:
4418:IUCN Red List least concern species
710:Slavens, F. and K. Slavens (1991).
638:Homopus Research Foundation website
588:Beskrifning på trenne sköld-paddor.
552:IUCN Red List of Threatened Species
541:Hofmeyr, M.D.; Keswick, T. (2018).
25:
770:. Cape Nature Conservation. 1994.
440:and is found specifically in the
1006:
392:"A Monograph of the Testudinata"
275:Daudin 1801:294 (junior homonym)
86:
755:The South African Tortoise Book
586:Thunberg, Carl Petter. (1787).
259:Thunberg 1788:206 (nomen nudum)
1425:Domed Rodrigues giant tortoise
1419:Domed Mauritius giant tortoise
1291:Santiago Island giant tortoise
401:Like its larger relative, the
1:
1244:Chatham Island giant tortoise
601:Turtle Taxonomy Working Group
499:The species is threatened by
1311:Volcán Darwin giant tortoise
1306:Volcán Alcedo giant tortoise
1296:Pinzón Island giant tortoise
4183:List of Testudines families
1804:Flat-backed spider tortoise
1679:Speke's hinge-back tortoise
1669:Lobatse hinge-back tortoise
1494:Goode's thornscrub tortoise
1321:Western Santa Cruz tortoise
1301:Sierra Negra giant tortoise
1249:Eastern Santa Cruz tortoise
814:(Retrieved March 17, 2010).
4449:
4433:Reptiles described in 1787
1664:Home's hinge-back tortoise
1659:Forest hinge-back tortoise
1654:Bell's hinge-back tortoise
1316:Volcán Wolf giant tortoise
1265:Hood Island giant tortoise
1254:Fernandina Island tortoise
1004:
724:The Herpetology Sourcebook
4166:
2097:
2029:
1978:
1674:Natal hinge-back tortoise
1239:Cerro Azul giant tortoise
1082:Aldabrachelys grandidieri
989:
926:
753:R. Boycott, O. Bourquin:
244:
237:
227:
223:
208:
201:
83:Scientific classification
81:
59:
50:
41:
34:
4428:Reptiles of South Africa
1585:Hesperotestudo percrassa
1341:Cheirogaster bacharidisi
1275:Santa Fe Island tortoise
1260:Floreana Island tortoise
1214:Chelonoidis sombrerensis
1132:African spurred tortoise
828:(Padloper Tortoise) Care
487:Threats and conservation
438:Republic of South Africa
428:Distribution and habitat
390:19th century plate from
297:Testudo areolata pallida
1504:Sonoran Desert tortoise
559:: e.T170520A115656133.
325:, is a tiny species of
232: Extant (resident)
1431:Réunion giant tortoise
1353:Cheirogaster schafferi
1347:Cheirogaster gymnesica
1184:Chelonoidis alburyorum
1178:Yellow-footed tortoise
1150:Centrochelys vulcanica
1144:Centrochelys burchardi
1138:Centrochelys atlantica
1113:Astrochelys rogerbouri
1043:Aldabra giant tortoise
652:Parrot-beaked tortoise
496:
413:
394:
360:parrot-beaked tortoise
323:parrot-beaked tortoise
1964:Spur-thighed tortoise
1901:Stylemys nebrascensis
1895:Stylemys karakolensis
1853:Stylemys calaverensis
1758:Megalochelys sondaari
1746:Megalochelys cautleyi
1715:Asian forest tortoise
1461:Burmese star tortoise
1281:Pinta Island tortoise
1074:Aldabrachelys abrupta
805:Membres.multimania.fr
494:
483:misty, overcast day.
411:
389:
289:Chersine tetradactyla
1954:Kleinmann's tortoise
1913:Stylemys oregonensis
1859:Stylemys canetotiana
1553:Hadrianus schucherti
1535:Hadrianus majusculus
1466:Indian star tortoise
1208:Chelonoidis sellovii
1202:Chelonoidis monensis
1190:Chelonoidis cubensis
364:beaked Cape tortoise
1959:Marginated tortoise
1889:Stylemys frizaciana
1752:Megalochelys margae
1636:Travancore tortoise
1559:Hadrianus utahensis
1541:Hadrianus octonaria
1229:Chelonoidis complex
1173:Red-footed tortoise
506:Trade in collected
501:habitat destruction
53:Conservation status
27:Species of tortoise
3291:Nanhsiungchelyidae
3256:Macrocephalochelys
1970:Testudo brevitesta
1949:Hermann's tortoise
1925:Stylemys uintensis
1907:Stylemys neglectus
1871:Stylemys conspecta
1776:Geometric tortoise
1740:Megalochelys atlas
1720:Impressed tortoise
1631:Forsten's tortoise
1626:Elongated tortoise
1547:Hadrianus robustus
1224:Galápagos tortoise
1196:Chelonoidis lutzae
810:2010-03-25 at the
781:"Biodiversity.org"
741:2010-07-16 at the
497:
414:
395:
368:areolated tortoise
251:Thunberg, 1787:180
4405:
4404:
4252:Homopus_areolatus
4238:Homopus areolatus
4208:Homopus areolatus
4200:Taxon identifiers
4191:
4190:
4162:
4161:
4158:
4157:
4105:
4104:
4101:
4100:
3523:
3522:
3519:
3518:
3202:
3201:
2974:
2973:
2477:
2476:
2395:
2394:
2116:
2115:
2112:
2111:
2108:
2107:
1989:
1988:
1984:. Key: †=extinct.
1931:Stylemys undabuna
1781:Serrated tortoise
1565:Hadrianus vialovi
1529:Hadrianus corsoni
1399:Speckled padloper
1371:Angulate tortoise
1106:Radiated tortoise
1101:Angonoka tortoise
1002:
1001:
697:978-0-86622-958-6
673:, ZipcodeZoo.com.
624:Homopus areolatus
622:Common Padloper (
545:Homopus areolatus
477:Homopus areolatus
348:Homopus areolatus
310:Homopus areolatus
306:
305:
300:
292:
284:
276:
268:
260:
252:
212:Homopus areolatus
194:H. areolatus
76:
36:Homopus areolatus
16:(Redirected from
4440:
4398:
4397:
4385:
4384:
4372:
4371:
4359:
4358:
4346:
4345:
4333:
4332:
4320:
4319:
4307:
4306:
4294:
4293:
4281:
4280:
4268:
4267:
4255:
4254:
4242:
4241:
4240:
4227:
4226:
4225:
4195:
4121:
4114:
3626:Chupacabrachelys
3541:
3534:
3211:
2989:Carettochelyidae
2985:
2488:
2406:
2138:
2127:
2095:
2086:
2077:
2016:
2009:
2002:
1993:
1883:Stylemys emiliae
1827:Leopard tortoise
1697:Pancake tortoise
1608:Greater padloper
1025:Russian tortoise
1010:
1009:
987:
912:
905:
898:
889:
831:
821:
815:
802:
796:
795:
793:
792:
783:. Archived from
777:
771:
764:
758:
751:
745:
733:
727:
721:
715:
708:
702:
701:
683:
674:
668:
659:
658:species gallery.
649:
640:
635:
629:
618:
609:
608:
597:
591:
584:
578:
577:
575:
573:
568:
538:
403:greater padloper
295:
287:
283:Hermann 1804:218
281:Testudo africana
279:
273:Testudo fasciata
271:
263:
255:
249:Testudo areolata
247:
231:
218:(Thunberg, 1787)
214:
91:
90:
70:
65:
64:
46:
32:
21:
4448:
4447:
4443:
4442:
4441:
4439:
4438:
4437:
4408:
4407:
4406:
4401:
4393:
4388:
4380:
4375:
4367:
4364:Observation.org
4362:
4354:
4349:
4341:
4336:
4328:
4323:
4315:
4310:
4302:
4297:
4289:
4284:
4276:
4271:
4263:
4258:
4250:
4245:
4236:
4235:
4230:
4221:
4220:
4215:
4202:
4192:
4187:
4154:
4097:
4074:Sahonachelyidae
4067:
3953:
3926:
3791:
3722:Phosphatochelys
3561:
3515:
3456:
3451:Terlinguachelys
3419:Pneumatoarthrus
3322:
3284:
3198:
3171:
3019:
2970:
2774:
2753:
2741:Siebenrockiella
2599:
2473:
2424:
2391:
2340:
2327:Mesodermochelys
2272:
2133:
2104:
2066:
2025:
2020:
1990:
1985:
1974:
1935:
1919:Stylemys pygmea
1831:
1813:
1809:Spider tortoise
1790:
1762:
1724:
1701:
1683:
1640:
1612:
1603:Common padloper
1589:
1569:
1513:
1499:Gopher tortoise
1489:Desert tortoise
1484:Bolson tortoise
1470:
1447:
1403:
1375:
1357:
1325:
1285:Lonesome George
1218:
1154:
1118:
1087:
1065:A. g. hololissa
1029:
1011:
1007:
998:
991:Species of the
978:
922:
916:
839:
837:Further reading
834:
822:
818:
812:Wayback Machine
803:
799:
790:
788:
779:
778:
774:
765:
761:
752:
748:
743:Wayback Machine
734:
730:
722:
718:
709:
705:
698:
685:
684:
677:
669:
662:
650:
643:
636:
632:
619:
612:
599:
598:
594:
585:
581:
571:
569:
540:
539:
532:
528:
520:
489:
462:valley bushveld
430:
384:
352:common padloper
345:
319:common padloper
265:Testudo miniata
233:
229:
219:
216:
210:
197:
85:
77:
66:
62:
55:
28:
23:
22:
18:Common padloper
15:
12:
11:
5:
4446:
4444:
4436:
4435:
4430:
4425:
4420:
4410:
4409:
4403:
4402:
4400:
4399:
4386:
4373:
4360:
4347:
4334:
4321:
4308:
4295:
4282:
4269:
4256:
4243:
4228:
4212:
4210:
4204:
4203:
4198:
4189:
4188:
4186:
4185:
4178:
4177:
4176:. † = extinct.
4167:
4164:
4163:
4160:
4159:
4156:
4155:
4153:
4152:
4144:
4136:
4127:
4125:
4118:
4111:
4107:
4106:
4103:
4102:
4099:
4098:
4096:
4095:
4087:
4078:
4076:
4069:
4068:
4066:
4065:
4057:
4050:
4043:
4035:
4027:
4020:
4012:
4004:
3996:
3988:
3980:
3972:
3969:Albertwoodemys
3963:
3961:
3959:Podocnemididae
3955:
3954:
3952:
3951:
3944:
3936:
3934:
3928:
3927:
3925:
3924:
3916:
3909:
3902:
3895:
3887:
3880:
3873:
3866:
3859:
3856:Lomalatachelys
3851:
3844:
3837:
3830:
3823:
3816:
3809:
3801:
3799:
3793:
3792:
3790:
3789:
3781:
3773:
3765:
3757:
3749:
3741:
3733:
3725:
3717:
3709:
3706:Labrostochelys
3701:
3693:
3685:
3677:
3669:
3661:
3653:
3645:
3637:
3629:
3621:
3613:
3605:
3597:
3589:
3581:
3572:
3570:
3563:
3562:
3560:
3559:
3550:
3548:
3546:Araripemydidae
3538:
3531:
3525:
3524:
3521:
3520:
3517:
3516:
3514:
3513:
3505:
3497:
3489:
3481:
3473:
3464:
3462:
3458:
3457:
3455:
3454:
3446:
3438:
3430:
3422:
3414:
3406:
3398:
3390:
3382:
3374:
3366:
3358:
3350:
3342:
3333:
3331:
3324:
3323:
3321:
3320:
3312:
3304:
3295:
3293:
3286:
3285:
3283:
3282:
3274:
3266:
3259:
3251:
3248:Emarginachelys
3243:
3235:
3227:
3219:
3217:
3208:
3204:
3203:
3200:
3199:
3197:
3196:
3193:Sinaspideretes
3188:
3179:
3177:
3173:
3172:
3170:
3169:
3162:
3155:
3148:
3141:
3133:
3126:
3119:
3112:
3104:
3096:
3088:
3081:
3073:
3066:
3059:
3052:
3044:
3037:
3029:
3027:
3021:
3020:
3018:
3017:
3010:
3002:
2993:
2991:
2982:
2976:
2975:
2972:
2971:
2969:
2968:
2961:
2953:
2946:
2938:
2931:
2924:
2916:
2908:
2901:
2894:
2887:
2880:
2873:
2870:Hesperotestudo
2865:
2857:
2850:
2843:
2835:
2827:
2820:
2813:
2806:
2799:
2792:
2784:
2782:
2776:
2775:
2773:
2772:
2764:
2762:
2760:Platysternidae
2755:
2754:
2752:
2751:
2744:
2737:
2730:
2723:
2716:
2709:
2702:
2695:
2688:
2681:
2674:
2667:
2660:
2653:
2646:
2639:
2632:
2625:
2617:
2609:
2607:
2601:
2600:
2598:
2597:
2589:
2582:
2575:
2568:
2561:
2554:
2547:
2540:
2533:
2526:
2519:
2512:
2505:
2496:
2494:
2485:
2479:
2478:
2475:
2474:
2472:
2471:
2464:
2457:
2450:
2442:
2434:
2432:
2426:
2425:
2423:
2422:
2414:
2412:
2410:Dermatemydidae
2403:
2401:Kinosternoidea
2397:
2396:
2393:
2392:
2390:
2389:
2381:
2373:
2370:Procolpochelys
2365:
2357:
2348:
2346:
2342:
2341:
2339:
2338:
2330:
2322:
2314:
2307:
2299:
2291:
2282:
2280:
2278:Dermochelyidae
2274:
2273:
2271:
2270:
2262:
2254:
2246:
2239:
2231:
2223:
2216:
2208:
2200:
2192:
2185:
2177:
2170:
2167:Carolinochelys
2162:
2155:
2146:
2144:
2135:
2124:
2118:
2117:
2114:
2113:
2110:
2109:
2106:
2105:
2098:
2092:
2083:
2074:
2068:
2067:
2065:
2064:
2058:
2049:
2043:
2037:
2030:
2027:
2026:
2021:
2019:
2018:
2011:
2004:
1996:
1987:
1986:
1979:
1976:
1975:
1973:
1972:
1966:
1961:
1956:
1951:
1945:
1943:
1937:
1936:
1934:
1933:
1927:
1921:
1915:
1909:
1903:
1897:
1891:
1885:
1879:
1877:Stylemys copei
1873:
1867:
1865:Stylemys capax
1861:
1855:
1849:
1847:Stylemys botti
1842:
1840:
1833:
1832:
1830:
1829:
1823:
1821:
1815:
1814:
1812:
1811:
1806:
1800:
1798:
1792:
1791:
1789:
1788:
1783:
1778:
1772:
1770:
1764:
1763:
1761:
1760:
1754:
1748:
1742:
1735:
1733:
1726:
1725:
1723:
1722:
1717:
1711:
1709:
1703:
1702:
1700:
1699:
1693:
1691:
1685:
1684:
1682:
1681:
1676:
1671:
1666:
1661:
1656:
1650:
1648:
1642:
1641:
1639:
1638:
1633:
1628:
1622:
1620:
1614:
1613:
1611:
1610:
1605:
1599:
1597:
1591:
1590:
1588:
1587:
1580:
1578:
1576:Hesperotestudo
1571:
1570:
1568:
1567:
1561:
1555:
1549:
1543:
1537:
1531:
1524:
1522:
1515:
1514:
1512:
1511:
1509:Texas tortoise
1506:
1501:
1496:
1491:
1486:
1480:
1478:
1472:
1471:
1469:
1468:
1463:
1457:
1455:
1449:
1448:
1446:
1445:
1439:
1433:
1427:
1421:
1414:
1412:
1405:
1404:
1402:
1401:
1396:
1394:Karoo padloper
1391:
1385:
1383:
1377:
1376:
1374:
1373:
1367:
1365:
1359:
1358:
1356:
1355:
1349:
1343:
1336:
1334:
1327:
1326:
1324:
1323:
1318:
1313:
1308:
1303:
1298:
1293:
1288:
1277:
1272:
1262:
1256:
1251:
1246:
1241:
1235:
1233:
1232:
1231:
1220:
1219:
1217:
1216:
1210:
1204:
1198:
1192:
1186:
1180:
1175:
1170:
1168:Chaco tortoise
1164:
1162:
1156:
1155:
1153:
1152:
1146:
1140:
1134:
1128:
1126:
1120:
1119:
1117:
1116:
1108:
1103:
1097:
1095:
1089:
1088:
1086:
1085:
1077:
1069:
1061:
1058:A. g. daudinii
1053:
1046:
1039:
1037:
1031:
1030:
1028:
1027:
1021:
1019:
1013:
1012:
1005:
1003:
1000:
999:
990:
984:
980:
979:
977:
976:
970:
964:
958:
952:
946:
940:
934:
927:
924:
923:
917:
915:
914:
907:
900:
892:
886:
885:
882:
879:
876:
873:
870:
867:
864:
861:
858:
855:
852:
849:
846:
843:
838:
835:
833:
832:
816:
797:
772:
766:E.H.W. Baard:
759:
746:
728:
716:
703:
696:
675:
660:
641:
630:
610:
592:
579:
529:
527:
524:
519:
516:
488:
485:
469:Nieuwoudtville
458:albany thicket
429:
426:
383:
380:
378:of its shell.
344:
341:
315:commonly known
304:
303:
302:
301:
293:
291:Merrem 1820:32
285:
277:
269:
261:
257:Testudo minuta
253:
242:
241:
235:
234:
228:
225:
224:
221:
220:
217:
206:
205:
199:
198:
191:
189:
185:
184:
177:
173:
172:
167:
163:
162:
157:
153:
152:
147:
143:
142:
137:
133:
132:
127:
123:
122:
117:
113:
112:
107:
103:
102:
97:
93:
92:
79:
78:
60:
57:
56:
51:
48:
47:
39:
38:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
4445:
4434:
4431:
4429:
4426:
4424:
4421:
4419:
4416:
4415:
4413:
4396:
4391:
4387:
4383:
4378:
4374:
4370:
4365:
4361:
4357:
4352:
4348:
4344:
4339:
4335:
4331:
4326:
4322:
4318:
4313:
4309:
4305:
4300:
4296:
4292:
4287:
4283:
4279:
4274:
4270:
4266:
4261:
4257:
4253:
4248:
4244:
4239:
4233:
4229:
4224:
4218:
4214:
4213:
4211:
4209:
4205:
4201:
4196:
4184:
4180:
4179:
4175:
4174:
4169:
4168:
4165:
4151:
4150:
4149:Tacuarembemys
4145:
4143:
4142:
4137:
4135:
4134:
4129:
4128:
4126:
4122:
4119:
4115:
4112:
4108:
4094:
4093:
4088:
4086:
4085:
4084:Sahonachelys
4080:
4079:
4077:
4075:
4070:
4064:
4063:
4058:
4056:
4055:
4051:
4049:
4048:
4047:Peltocephalus
4044:
4042:
4041:
4036:
4034:
4033:
4032:Lapparentemys
4028:
4026:
4025:
4021:
4019:
4018:
4013:
4011:
4010:
4005:
4003:
4002:
3997:
3995:
3994:
3989:
3987:
3986:
3981:
3979:
3978:
3973:
3971:
3970:
3965:
3964:
3962:
3960:
3956:
3950:
3949:
3945:
3943:
3942:
3938:
3937:
3935:
3933:
3932:Pelomedusidae
3929:
3923:
3922:
3921:Yaminuechelys
3917:
3915:
3914:
3910:
3908:
3907:
3903:
3901:
3900:
3896:
3894:
3893:
3892:Prochelidella
3888:
3886:
3885:
3881:
3879:
3878:
3874:
3872:
3871:
3867:
3865:
3864:
3860:
3858:
3857:
3852:
3850:
3849:
3845:
3843:
3842:
3838:
3836:
3835:
3831:
3829:
3828:
3824:
3822:
3821:
3817:
3815:
3814:
3810:
3808:
3807:
3806:Acanthochelys
3803:
3802:
3800:
3798:
3794:
3788:
3787:
3782:
3780:
3779:
3774:
3772:
3771:
3766:
3764:
3763:
3758:
3756:
3755:
3750:
3748:
3747:
3742:
3740:
3739:
3734:
3732:
3731:
3726:
3724:
3723:
3718:
3716:
3715:
3710:
3708:
3707:
3702:
3700:
3699:
3694:
3692:
3691:
3690:Kinkonychelys
3686:
3684:
3683:
3678:
3676:
3675:
3674:Itapecuruemys
3670:
3668:
3667:
3662:
3660:
3659:
3654:
3652:
3651:
3646:
3644:
3643:
3638:
3636:
3635:
3634:Eotaphrosphys
3630:
3628:
3627:
3622:
3620:
3619:
3614:
3612:
3611:
3606:
3604:
3603:
3598:
3596:
3595:
3590:
3588:
3587:
3582:
3580:
3579:
3574:
3573:
3571:
3569:
3568:Bothremydidae
3564:
3558:
3557:
3552:
3551:
3549:
3547:
3542:
3539:
3535:
3532:
3530:
3526:
3512:
3511:
3506:
3504:
3503:
3498:
3496:
3495:
3490:
3488:
3487:
3482:
3480:
3479:
3478:Argillochelys
3474:
3472:
3471:
3466:
3465:
3463:
3459:
3453:
3452:
3447:
3445:
3444:
3443:Santanachelys
3439:
3437:
3436:
3431:
3429:
3428:
3423:
3421:
3420:
3415:
3413:
3412:
3407:
3405:
3404:
3399:
3397:
3396:
3391:
3389:
3388:
3387:Desmatochelys
3383:
3381:
3380:
3375:
3373:
3372:
3371:Calcarichelys
3367:
3365:
3364:
3359:
3357:
3356:
3355:Atlantochelys
3351:
3349:
3348:
3343:
3341:
3340:
3335:
3334:
3332:
3330:
3329:Protostegidae
3325:
3319:
3318:
3317:Jiangxichelys
3313:
3311:
3310:
3305:
3303:
3302:
3301:Anomalochelys
3297:
3296:
3294:
3292:
3287:
3281:
3280:
3279:Protochelydra
3275:
3273:
3272:
3271:Planiplastron
3267:
3265:
3264:
3260:
3258:
3257:
3252:
3250:
3249:
3244:
3242:
3241:
3236:
3234:
3233:
3228:
3226:
3225:
3221:
3220:
3218:
3216:
3212:
3209:
3205:
3195:
3194:
3189:
3187:
3186:
3181:
3180:
3178:
3174:
3168:
3167:
3163:
3161:
3160:
3156:
3154:
3153:
3149:
3147:
3146:
3142:
3140:
3139:
3134:
3132:
3131:
3127:
3125:
3124:
3120:
3118:
3117:
3113:
3111:
3110:
3105:
3103:
3102:
3097:
3095:
3094:
3089:
3087:
3086:
3082:
3080:
3079:
3078:Drazinderetes
3074:
3072:
3071:
3067:
3065:
3064:
3060:
3058:
3057:
3053:
3051:
3050:
3045:
3043:
3042:
3038:
3036:
3035:
3031:
3030:
3028:
3026:
3022:
3016:
3015:
3014:Carettochelys
3011:
3009:
3008:
3003:
3001:
3000:
2995:
2994:
2992:
2990:
2986:
2983:
2981:
2977:
2967:
2966:
2962:
2960:
2959:
2954:
2952:
2951:
2947:
2945:
2944:
2939:
2937:
2936:
2932:
2930:
2929:
2925:
2923:
2922:
2917:
2915:
2914:
2909:
2907:
2906:
2902:
2900:
2899:
2898:Malacochersus
2895:
2893:
2892:
2888:
2886:
2885:
2881:
2879:
2878:
2874:
2872:
2871:
2866:
2864:
2863:
2858:
2856:
2855:
2851:
2849:
2848:
2844:
2842:
2841:
2836:
2834:
2833:
2828:
2826:
2825:
2821:
2819:
2818:
2814:
2812:
2811:
2807:
2805:
2804:
2800:
2798:
2797:
2793:
2791:
2790:
2789:Aldabrachelys
2786:
2785:
2783:
2781:
2777:
2771:
2770:
2766:
2765:
2763:
2761:
2756:
2750:
2749:
2745:
2743:
2742:
2738:
2736:
2735:
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2671:Leucocephalon
2668:
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2647:
2645:
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2640:
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2622:Banhxeochelys
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2134:(Sea turtles)
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1050:A. g. arnoldi
1047:
1044:
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1040:
1038:
1036:
1035:Aldabrachelys
1032:
1026:
1023:
1022:
1020:
1018:
1014:
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968:Testudinoidea
966:Superfamily:
965:
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787:on 2019-12-15
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366:, as well as
365:
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357:
353:
349:
342:
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338:
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329:of the genus
328:
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299:Gray 1831d:13
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203:Binomial name
200:
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160:Testudinoidea
158:
156:Superfamily:
155:
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141:
138:
135:
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128:
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94:
89:
84:
80:
74:
69:
68:Least Concern
58:
54:
49:
45:
40:
37:
33:
30:
19:
4207:
4172:
4147:
4139:
4131:
4090:
4082:
4060:
4052:
4045:
4038:
4030:
4024:Erymnochelys
4022:
4015:
4009:Cerrejonemys
4007:
3999:
3991:
3985:Brontochelys
3983:
3975:
3967:
3946:
3939:
3919:
3911:
3904:
3899:Pseudemydura
3897:
3890:
3882:
3875:
3868:
3861:
3854:
3846:
3839:
3832:
3825:
3818:
3811:
3804:
3784:
3776:
3768:
3760:
3752:
3744:
3736:
3728:
3720:
3712:
3704:
3696:
3688:
3680:
3672:
3664:
3656:
3648:
3640:
3632:
3624:
3616:
3608:
3600:
3592:
3584:
3576:
3554:
3508:
3502:Prionochelys
3500:
3492:
3484:
3476:
3468:
3449:
3441:
3433:
3425:
3417:
3409:
3401:
3393:
3385:
3379:Cratochelone
3377:
3369:
3363:Bouliachelys
3361:
3353:
3345:
3339:Alienochelys
3337:
3315:
3307:
3299:
3277:
3269:
3261:
3254:
3246:
3240:Chelydropsis
3238:
3230:
3222:
3191:
3185:Basilochelys
3183:
3164:
3157:
3150:
3143:
3136:
3128:
3121:
3114:
3109:Khunnuchelys
3107:
3099:
3091:
3083:
3076:
3068:
3061:
3054:
3047:
3039:
3032:
3025:Trionychidae
3012:
3005:
2999:Allaeochelys
2997:
2963:
2956:
2950:Stigmochelys
2948:
2941:
2933:
2926:
2919:
2913:Megalochelys
2911:
2903:
2896:
2889:
2882:
2875:
2868:
2860:
2852:
2845:
2838:
2830:
2824:Cylindraspis
2822:
2815:
2808:
2803:Centrochelys
2801:
2794:
2787:
2780:Testudinidae
2769:Platysternon
2767:
2748:Vijayachelys
2746:
2739:
2732:
2727:Rhinoclemmys
2725:
2718:
2711:
2704:
2697:
2692:Melanochelys
2690:
2683:
2676:
2669:
2662:
2655:
2648:
2641:
2634:
2627:
2620:
2612:
2592:
2584:
2577:
2570:
2563:
2556:
2549:
2542:
2535:
2528:
2521:
2514:
2507:
2502:Acherontemys
2500:
2468:Sternotherus
2466:
2459:
2452:
2445:
2437:
2417:
2384:
2376:
2368:
2360:
2352:
2335:Psephophorus
2333:
2325:
2317:
2309:
2302:
2294:
2286:
2265:
2257:
2251:Pacifichelys
2249:
2241:
2234:
2226:
2220:Lepidochelys
2218:
2211:
2203:
2197:Gigantatypus
2195:
2189:Eretmochelys
2187:
2180:
2172:
2165:
2157:
2150:
2131:Chelonioidea
2099:
2051:
1819:Stigmochelys
1731:Megalochelys
1602:
1410:Cylindraspis
1332:Cheirogaster
1124:Centrochelys
1111:
1080:
1072:
1063:
1056:
1048:
1045:(subspecies:
992:
974:Testudinidae
825:
823:Corton, M.,
819:
800:
789:. Retrieved
785:the original
775:
767:
762:
754:
749:
731:
723:
719:
711:
706:
687:
633:
623:
620:Loehr, V., "
604:
595:
587:
582:
570:. Retrieved
556:
550:
544:
521:
511:
507:
505:
498:
481:
476:
466:
446:Eastern Cape
442:Western Cape
431:
419:
415:
400:
396:
391:
371:
367:
363:
359:
355:
351:
347:
346:
337:South Africa
330:
322:
318:
309:
308:
307:
296:
288:
280:
272:
264:
256:
248:
211:
209:
193:
192:
180:
170:Testudinidae
35:
29:
4299:iNaturalist
4232:Wikispecies
4062:Stupendemys
4017:Cordichelys
3863:Mesoclemmys
3848:Hydromedusa
3770:Taphrosphys
3762:Sankuchemys
3754:Rhothonemys
3730:Polysternon
3610:Cearachelys
3594:Azabbaremys
3578:Araiochelys
3494:Ctenochelys
3486:Bashuchelys
3435:Rhinochelys
3411:Ocepechelon
3403:Notochelone
3395:Iserosaurus
3263:Macrochelys
3215:Chelydridae
3138:Palaeoamyda
3063:Cyclanorbis
2928:Psammobates
2921:Oligopherus
2884:Indotestudo
2832:Cymatholcus
2810:Chelonoidis
2796:Astrochelys
2605:Geoemydidae
2523:Deirochelys
2461:Staurotypus
2454:Kinosternon
2447:Hoplochelys
2362:Peritresius
2311:Dermochelys
2304:Cosmochelys
2296:Corsochelys
2213:Itilochelys
2205:Glarichelys
2152:Allopleuron
2142:Cheloniidae
2081:Superfamily
1768:Psammobates
1618:Indotestudo
1160:Chelonoidis
1093:Astrochelys
572:19 November
382:Description
372:"areolatus"
4412:Categories
4141:Caririemys
4054:Podocnemis
4040:Latentemys
4001:Carbonemys
3941:Pelomedusa
3778:Ummulisani
3698:Kurmademys
3666:Inaechelys
3650:Galianemys
3618:Chedighaii
3556:Araripemys
3529:Pleurodira
3510:Toxochelys
3427:Protostega
3232:Chelydrops
3152:Pelodiscus
3145:Pelochelys
3093:Gilmoremys
3070:Cycloderma
2980:Trionychia
2847:Geochelone
2840:Floridemys
2706:Notochelys
2594:Wilburemys
2565:Malaclemys
2419:Dermatemys
2386:Puppigerus
2319:Eosphargis
2236:Miocaretta
2228:Mexichelys
2122:Cryptodira
2062:Testudines
2023:Testudines
1453:Geochelone
1381:Chersobius
1017:Agrionemys
962:Cryptodira
960:Suborder:
956:Testudines
948:Subclass:
791:2013-06-23
526:References
454:strandveld
150:Cryptodira
146:Suborder:
140:Testudines
4382:areolatus
4181:See also
4133:Caribemys
3993:Caninemys
3977:Bauruemys
3906:Rheodytes
3870:Myuchelys
3813:Chelodina
3738:Puentemys
3714:Nigeremys
3658:Ilatardia
3602:Bothremys
3309:Basilemys
3123:Nilssonia
3101:Hutchemys
3049:Axestemys
3007:Anosteira
2862:Hadrianus
2720:Pangshura
2678:Malayemys
2643:Geoclemys
2586:Trachemys
2579:Terrapene
2572:Pseudemys
2558:Graptemys
2551:Glyptemys
2544:Emydoidea
2537:Actinemys
2509:Chrysemys
2354:Euclastes
2182:Eochelone
2033:Kingdom:
1520:Hadrianus
930:Kingdom:
518:Captivity
188:Species:
106:Kingdom:
100:Eukaryota
4390:Species+
4317:11141287
4217:Wikidata
3948:Pelusios
3913:Rhinemys
3884:Platemys
3877:Phrynops
3797:Chelidae
3786:Zolhafah
3682:Jainemys
3347:Archelon
3224:Chelydra
3116:Lissemys
2958:Stylemys
2943:Solitudo
2905:Manouria
2854:Gopherus
2817:Chersina
2685:Mauremys
2664:Heosemys
2657:Hardella
2650:Geoemyda
2636:Cyclemys
2492:Emydidae
2439:Claudius
2288:Arabemys
2267:Tasbacka
2259:Syllomus
2174:Chelonia
2072:Suborder
2056:Diapsida
2047:Reptilia
2041:Chordata
2039:Phylum:
2035:Animalia
1838:Stylemys
1707:Manouria
1476:Gopherus
1363:Chersina
994:tortoise
972:Family:
950:Anapsida
944:Reptilia
938:Chordata
936:Phylum:
932:Animalia
919:Tortoise
808:Archived
739:Archived
422:plastral
327:tortoise
239:Synonyms
166:Family:
130:Reptilia
120:Chordata
116:Phylum:
110:Animalia
96:Domain:
73:IUCN 3.1
4423:Homopus
4291:5220261
4223:Q642538
4092:Sokatra
3841:Emydura
3746:Rosasia
3642:Foxemys
3586:Arenila
3166:Trionyx
3159:Rafetus
3085:Dogania
3041:Apalone
2965:Testudo
2891:Kinixys
2877:Homopus
2734:Sacalia
2713:Orlitia
2699:Morenia
2614:Batagur
2516:Clemmys
2243:Natator
2159:Caretta
2060:Order:
2045:Class:
1941:Testudo
1646:Kinixys
1595:Homopus
954:Order:
942:Class:
826:Homopus
512:Homopus
508:Homopus
473:Cradock
436:to the
434:endemic
356:Homopus
332:Homopus
317:as the
181:Homopus
176:Genus:
136:Order:
126:Class:
71: (
4369:100018
4356:285996
4343:170520
4330:551783
4304:102930
4278:963554
4124:
4117:
4110:
3834:Elusor
3827:Elseya
3820:Chelus
3537:
3470:Adocus
3461:
3207:
3176:
3056:Chitra
2758:
2345:
2090:Family
997:family
983:Genera
921:family
694:
450:fynbos
432:It is
376:scutes
343:Naming
230:
4312:IRMNG
4265:3MHM5
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