573:
118:
144:
2223:
812:
2526:
469:
838:
of eurypterids (Adelophthalmoidea, Pterygotioidea and the waeringopteroids), as inferred by Tetlie and
Poschmann in 2008, based on the results of a 2008 analysis specifically pertaining to the Adelophthalmoidea and a preceding 2004 analysis.
706:
was recognized as sufficiently different to represent its own genus due to the anterior position of its eyes, much more exaggerated than in the other adelophthalmid genera. The first part of the generic name is composed of the word
1735:
756:
is considered to be the most basal adelophthalmid genus due to the position of its eyes. These are placed closer to the margin than to the ocelli, in contrast to any other adelopthalmid. In fact, the carapace of
560:
with respect to the rest of adelophthalmids, with the eyes closer to the margin than to the ocelli, suggesting that the eyes migrated towards a central position from the basal genera to
510:(limbs). The prosoma was rather broad, with a parabolic (approximately U-shaped) outline. It was 19.5 millimetres (0.77 in) long and 22 mm (0.87 in) wide. Although the
538:(simple eyes) were located in the posterior half of the prosoma and consisted of two circular spots. The posterior part was smooth, but the previous part had a distinctive
534:
were reniform (bean-shaped) and with a thin and light-colored test (external "shield") on its surface. They measured 4.5 mm (0.18 in) in length. The
490:
was a small eurypterid. The total size of the only known specimen is estimated at only 8 centimetres (3.1 inches), far from the largest adelophthalmids like
2576:
2655:
2302:
2292:
1814:
2640:
1780:
2254:
592:
572:
2269:
645:
species, essentially the shape of the prosoma and the form or position of the eyes. He also suggested a close relationship between
831:
below presents the inferred phylogenetic positions of most of the genera included in the three most derived superfamilies of the
2513:
2297:
2239:
1485:
641:, named after Patten, who died months after the meeting. Størmer noticed several differences between his new species and other
2650:
2287:
2277:
2231:
808:
of a group formed by
Pterygotioidea and Adelophthalmoidea, but until more fossil material is found this can not be proven.
2645:
2249:
2244:
2282:
2259:
515:
143:
2635:
535:
1751:
2630:
607:
408:
117:
1915:
1807:
1480:
1506:"Cope's rule and Romer's theory: patterns of diversity and gigantism in eurypterids and Palaeozoic vertebrates"
800:
trait (a trait present in a common ancestor). These characteristics have led some authors to question whether
2222:
2544:
611:
530:
shield of the prosoma). The prosoma surface was gently arched, with the central portion rather level. The
433:
genus that was distinguished from the rest of adelophthalmids by the position near the head margin of the
2602:
739:
553:
430:
354:
2529:
1800:
1621:. Vol. 37. Publications of the Geological Institution of the University of Tartu. pp. 1–8.
1553:
1385:
366:
539:
446:
719:, a suffix commonly used in eurypterids which means "wing" (and therefore, translated as "wing of
1763:
1703:
1676:
1592:
627:
271:
138:
2607:
1781:"Eurypterid-Associated Biota of the Rootsikula Horizon, Saaremaa, Estonia: Rootsikula, Estonia"
2589:
2118:
2051:
2043:
1755:
1622:
1545:
1527:
949:
856:
519:
2594:
1657:"Phylogeny and palaeoecology of the Adelophthalmoidea (Arthropoda; Chelicerata; Eurypterida)"
2458:
2172:
2164:
2154:
2146:
2126:
2097:
2013:
1962:
1747:
1668:
1584:
1535:
1519:
1068:
1044:
1011:
699:
678:
543:
527:
442:
373:
342:
339:
224:
211:
1721:
2487:
2333:
2190:
2131:
2087:
1990:
1977:
1510:
1464:
1392:
1172:
774:
769:
380:
623:
1694:
Kjellesvig-Waering, Erik N. (1961). "The
Silurian Eurypterida of the Welsh Borderland".
2403:
2361:
2319:
2182:
2109:
2069:
2061:
1985:
1736:"Pterygotids (Chelicerata; Eurypterida) from the Silurian Vernon Formation of New York"
1540:
1425:
1134:
1085:
894:
846:
832:
514:
part of the prosoma is not preserved, the fossil mold indicates a pointed portion. The
502:
is a little known eurypterid, with only one specimen collected that only preserves the
492:
480:
811:
2624:
2410:
2382:
2375:
2347:
2326:
2079:
2018:
2008:
2000:
1954:
1944:
1936:
1892:
1856:
1767:
1656:
1568:
1445:
1285:
531:
507:
454:
434:
350:
68:
1680:
1596:
1505:
750:, but it was considered different enough to represent a new separate genus in 2008.
2368:
2200:
2136:
2092:
1967:
1389:
1220:
1148:
911:
870:
805:
797:
785:
763:
695:
656:
However, in 1961, the
American paleontologist Erik Norman Kjellesvig-Waering split
636:
619:
377:
358:
250:
30:
622:). During a meeting between Patten and the Norwegian paleontologist and geologist
468:
2567:
1616:
2465:
2442:
2340:
2034:
1927:
1865:
1847:
1430:
994:
185:
43:
1573:(Eurypterida, Chelicerata) from the Devonian of the South Minusinsk Depression"
2431:
2417:
2396:
2389:
2354:
2195:
1883:
1874:
1823:
1672:
1626:
1588:
1434:
1419:
1413:
1268:
1244:
835:
820:
702:
by the
Norwegian Odd Erik Tetlie and German Markus Poschmann paleontologists,
483:
327:
198:
88:
53:
1759:
1531:
2508:
2492:
2482:
1841:
1468:
1452:
828:
661:
604:
600:
542:. This ornamentation extended until the end of the eyes and was composed of
416:
412:
331:
175:
155:
93:
37:
1549:
1523:
780:, the eyes were separated from the margin by the marginal rim, whereas in
2561:
2451:
2424:
1456:
1448:
1395:
of the
Rootsikula Formation of Saaremaa, Estonia. This formation was the
1382:
1196:
615:
588:
523:
511:
457:
438:
384:
370:
126:
83:
78:
63:
58:
48:
1455:(the physical characteristics of the rocks) of the zone was composed of
788:
genera. In addition, the triangular anterior carapace margin present in
2581:
1707:
1396:
631:
503:
404:
388:
98:
73:
630:
of eurypterids, among which he recognized a carapace of a new unknown
403:, a traditional suffix for the eurypterid genera, meaning "wing". The
1835:
1460:
712:
596:
396:
165:
2538:
682:) for small reniform intramarginal (within the margin) eyes. Since
1400:
810:
735:
571:
557:
467:
346:
323:
1437:
2542:
1796:
1752:
10.1666/pleo0022-3360(2007)081[0725:PEFTSV]2.0.CO;2
734:
1792:
587:
is only known from one single specimen (and is therefore the
1618:
A new eurypterid from the
Saaremaa- (Oesel-) beds in Estonia
784:
they were completely marginal, a feature present in all
518:
border was slightly convex. A narrow rim surrounded the
1724:. In World Spider Catalog. Natural History Museum Bern.
449:. With an estimated length of 8 cm (3.1 in),
1722:
A summary list of fossil spiders and their relatives
634:. He described it in 1934 as belonging to the genus
2551:
2501:
2475:
2441:
2311:
2268:
2230:
2181:
2163:
2145:
2117:
2106:
2078:
2060:
2042:
2031:
1999:
1976:
1953:
1935:
1924:
1908:
1901:
761:and other basal genera belonging to superfamilies,
746:was originally described as a species of the genus
649:and other species, especially the Scottish species
672:) for species with large ovoid marginal eyes and
1720:Dunlop, J. A., Penney, D. & Jekel, D. 2018.
1655:Erik Tetlie, O; Poschmann, Markus (2008-06-01).
460:quiet community along other eurypterid species.
441:was parabolic (approximately U-shaped) and with
437:, different from the rest of its relatives. Its
1567:Shpinev, Evgeniy S.; Filimonov, A. N. (2018).
1808:
1504:Lamsdell, James C.; Braddy, Simon J. (2009).
686:fulfilled this condition, it was referred to
357:of eurypterids. One fossil of the single and
8:
1734:Ciurca, Samuel J.; Tetlie, O. Erik (2007).
1440:have also been found. The habitat in which
804:really represents an adelophthalmid or the
2539:
2114:
2039:
1932:
1905:
1815:
1801:
1793:
715:name for Saaremaa, and the second part by
116:
20:
1539:
610:during his extensive explorations at the
496:of 32 cm (12.6 in) in length.
419:who discovered the only known fossil of
133:based on Tetlie and Poschmann's of 2008
1496:
694:). It was not until 2008 when during a
698:review of the entire Adelophthalmidae
453:was a small eurypterid. It lived in a
595:) which preserves the carapace. This
308:Kjellesvig-Waering & Leutze, 1966
7:
1650:
1648:
1646:
1644:
1642:
1640:
1638:
1636:
1610:
1608:
1606:
1661:Journal of Systematic Palaeontology
1260:
1236:
1212:
1188:
1164:
1140:
1060:
1036:
986:
979:
955:
945:
886:
862:
852:
842:
626:in 1932, Størmer observed Patten's
1429:. Fossil remains of indeterminate
593:American Museum of Natural History
14:
546:lines developed as deep furrows.
2525:
2524:
2221:
796:, indicating that it might be a
773:, were almost identical. In the
142:
2514:Timeline of eurypterid research
1486:Timeline of eurypterid research
576:Restoration of the carapace of
353:("advanced") Adelophthalmoidea
599:was collected by the American
330:, an extinct group of aquatic
1:
2656:Fossil taxa described in 2008
1414:Eurypterus tetragonophthalmus
1377:The single known specimen of
338:is classified as part of the
1444:lived is considered to be a
618:, Estonia (then part of the
591:, AMNH 32720, housed at the
244:Tetlie & Poschmann, 2008
1403:of eurypterids, among them
815:Restoration of the related
391:. The genus is named after
16:Extinct genus of arthropods
2672:
2641:Silurian animals of Europe
580:based on Størmer's of 1934
493:Adelophthalmus khakassicus
2522:
2219:
1916:List of eurypterid genera
1830:
1785:The Paleobiology Database
1673:10.1017/S1477201907002416
1589:10.1134/S0031030118130129
1481:List of eurypterid genera
1282:
1265:
1258:
1241:
1234:
1217:
1210:
1193:
1186:
1169:
1162:
1145:
1138:
1082:
1065:
1058:
1041:
1034:
1008:
991:
984:
977:
960:
953:
943:
908:
891:
884:
867:
860:
850:
445:deep furrows forming the
365:, has been discovered in
277:
270:
256:
249:
139:Scientific classification
137:
124:
115:
23:
1381:has been recovered from
305:Nanahughmilleria patteni
300:Kjellesvig-Waering, 1961
1740:Journal of Paleontology
1696:Journal of Paleontology
1577:Paleontological Journal
1426:Strobilopterus laticeps
399:name for Saaremaa, and
1615:Størmer, Leif (1934).
1524:10.1098/rsbl.2009.0700
1405:Erettopterus laticauda
824:
581:
476:
2651:Eurypterids of Europe
2603:Paleobiology Database
1554:Supplemental material
1451:quiet community. The
814:
738:("group") within the
578:Eysyslopterus patteni
575:
471:
260:Eysyslopterus patteni
2646:Silurian eurypterids
612:Rootsikula Formation
281:Hughmilleria patteni
823:of Adelophthalmidae
742:Adelophthalmoidea.
568:History of research
472:Size comparison of
125:Restoration of the
2636:Fossils of Estonia
2044:Onychopterelloidea
1420:Mixopterus simonsi
825:
628:private collection
582:
477:
429:is a little-known
2631:Adelophthalmoidea
2618:
2617:
2590:Open Tree of Life
2545:Taxon identifiers
2536:
2535:
2217:
2216:
2213:
2212:
2209:
2208:
2165:Adelophthalmoidea
2147:Waeringopteroidea
2119:Carcinosomatoidea
2052:Onychopterellidae
2027:
2026:
1583:(13): 1553–1560.
1569:"A New Record of
1369:
1368:
1360:
1359:
1351:
1350:
1342:
1341:
1333:
1332:
1324:
1323:
1315:
1314:
1306:
1305:
1297:
1296:
1124:
1123:
1115:
1114:
1106:
1105:
1097:
1096:
1023:
1022:
950:Adelophthalmoidea
932:
931:
923:
922:
857:Waeringopteroidea
506:("head") with no
315:
314:
309:
301:
285:
245:
212:Adelophthalmoidea
36:422.9–418.7
2663:
2611:
2610:
2598:
2597:
2585:
2584:
2572:
2571:
2570:
2540:
2528:
2527:
2502:Related articles
2459:Merostomichnites
2225:
2173:Adelophthalmidae
2155:Waeringopteridae
2127:Carcinosomatidae
2115:
2098:Strobilopteridae
2040:
2014:Hibbertopteridae
1963:Parastylonuridae
1933:
1906:
1817:
1810:
1803:
1794:
1788:
1778:
1772:
1771:
1731:
1725:
1718:
1712:
1711:
1691:
1685:
1684:
1652:
1631:
1630:
1612:
1601:
1600:
1564:
1558:
1557:
1543:
1501:
1261:
1237:
1213:
1189:
1165:
1141:
1069:Parahughmilleria
1061:
1045:Nanahughmilleria
1037:
1012:Pittsfordipterus
987:
980:
956:
946:
887:
863:
853:
843:
792:was shared with
692:Nanahughmilleria
679:Nanahughmilleria
343:Adelophthalmidae
307:
299:
293:Nanahughmilleria
283:
243:
236:
225:Adelophthalmidae
223:
210:
197:
147:
146:
120:
110:
40:
29:Temporal range:
21:
2671:
2670:
2666:
2665:
2664:
2662:
2661:
2660:
2621:
2620:
2619:
2614:
2606:
2601:
2593:
2588:
2580:
2575:
2566:
2565:
2560:
2547:
2537:
2532:
2518:
2497:
2488:Chasmataspidida
2471:
2437:
2334:Campylocephalus
2307:
2264:
2226:
2205:
2191:Hughmilleriidae
2177:
2159:
2141:
2132:Megalograptidae
2108:
2102:
2088:Dolichopteridae
2074:
2062:Moselopteroidea
2056:
2033:
2023:
2009:Drepanopteridae
1995:
1991:Hardieopteridae
1978:Kokomopteroidea
1972:
1949:
1926:
1920:
1897:
1826:
1821:
1791:
1779:
1775:
1733:
1732:
1728:
1719:
1715:
1693:
1692:
1688:
1654:
1653:
1634:
1614:
1613:
1604:
1566:
1565:
1561:
1511:Biology Letters
1503:
1502:
1498:
1494:
1477:
1375:
1370:
1361:
1352:
1343:
1334:
1325:
1316:
1307:
1298:
1173:Herefordopterus
1125:
1116:
1107:
1098:
1024:
933:
924:
782:Herefordopterus
775:waeringopteroid
770:Herefordopterus
729:
570:
522:margins of the
479:Like the other
466:
266:
263:
242:
234:
221:
208:
195:
141:
111:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
35:
34:
27:
17:
12:
11:
5:
2669:
2667:
2659:
2658:
2653:
2648:
2643:
2638:
2633:
2623:
2622:
2616:
2615:
2613:
2612:
2599:
2586:
2573:
2557:
2555:
2549:
2548:
2543:
2534:
2533:
2523:
2520:
2519:
2517:
2516:
2511:
2505:
2503:
2499:
2498:
2496:
2495:
2490:
2485:
2479:
2477:
2476:Related groups
2473:
2472:
2470:
2469:
2462:
2455:
2447:
2445:
2439:
2438:
2436:
2435:
2428:
2421:
2414:
2407:
2404:Onychopterella
2400:
2393:
2386:
2379:
2372:
2365:
2362:Hibbertopterus
2358:
2351:
2344:
2337:
2330:
2323:
2320:Adelophthalmus
2315:
2313:
2312:Notable genera
2309:
2308:
2306:
2305:
2300:
2295:
2290:
2285:
2280:
2274:
2272:
2266:
2265:
2263:
2262:
2257:
2252:
2247:
2242:
2236:
2234:
2228:
2227:
2220:
2218:
2215:
2214:
2211:
2210:
2207:
2206:
2204:
2203:
2198:
2193:
2187:
2185:
2183:Pterygotioidea
2179:
2178:
2176:
2175:
2169:
2167:
2161:
2160:
2158:
2157:
2151:
2149:
2143:
2142:
2140:
2139:
2134:
2129:
2123:
2121:
2112:
2110:Diploperculata
2104:
2103:
2101:
2100:
2095:
2090:
2084:
2082:
2076:
2075:
2073:
2072:
2070:Moselopteridae
2066:
2064:
2058:
2057:
2055:
2054:
2048:
2046:
2037:
2029:
2028:
2025:
2024:
2022:
2021:
2016:
2011:
2005:
2003:
1997:
1996:
1994:
1993:
1988:
1986:Kokomopteridae
1982:
1980:
1974:
1973:
1971:
1970:
1965:
1959:
1957:
1951:
1950:
1948:
1947:
1941:
1939:
1937:Rhenopteroidea
1930:
1922:
1921:
1919:
1918:
1912:
1910:
1903:
1899:
1898:
1896:
1895:
1886:
1877:
1868:
1859:
1850:
1844:
1838:
1831:
1828:
1827:
1822:
1820:
1819:
1812:
1805:
1797:
1790:
1789:
1773:
1746:(4): 725–736.
1726:
1713:
1702:(4): 789–835.
1686:
1667:(2): 237–249.
1632:
1602:
1571:Adelophthalmus
1559:
1518:(2): 265–269.
1495:
1493:
1490:
1489:
1488:
1483:
1476:
1473:
1374:
1371:
1367:
1366:
1363:
1362:
1358:
1357:
1354:
1353:
1349:
1348:
1345:
1344:
1340:
1339:
1336:
1335:
1331:
1330:
1327:
1326:
1322:
1321:
1318:
1317:
1313:
1312:
1309:
1308:
1304:
1303:
1300:
1299:
1295:
1294:
1291:
1290:
1281:
1278:
1277:
1274:
1273:
1264:
1259:
1257:
1254:
1253:
1250:
1249:
1240:
1235:
1233:
1230:
1229:
1226:
1225:
1216:
1211:
1209:
1206:
1205:
1202:
1201:
1192:
1187:
1185:
1182:
1181:
1178:
1177:
1168:
1163:
1161:
1158:
1157:
1154:
1153:
1144:
1139:
1137:
1135:Pterygotioidea
1131:
1130:
1127:
1126:
1122:
1121:
1118:
1117:
1113:
1112:
1109:
1108:
1104:
1103:
1100:
1099:
1095:
1094:
1091:
1090:
1086:Adelophthalmus
1081:
1078:
1077:
1074:
1073:
1064:
1059:
1057:
1054:
1053:
1050:
1049:
1040:
1035:
1033:
1030:
1029:
1026:
1025:
1021:
1020:
1017:
1016:
1007:
1004:
1003:
1000:
999:
990:
985:
983:
978:
976:
973:
972:
969:
968:
959:
954:
952:
944:
942:
939:
938:
935:
934:
930:
929:
926:
925:
921:
920:
917:
916:
907:
904:
903:
900:
899:
895:Waeringopterus
890:
885:
883:
880:
879:
876:
875:
866:
861:
859:
851:
849:
847:Diploperculata
841:
833:Diploperculata
817:Adelophthalmus
728:
727:Classification
725:
608:William Patten
569:
566:
562:Adelophthalmus
556:("primitive")
481:adelophthalmid
465:
462:
411:, an American
409:William Patten
313:
312:
311:
310:
302:
286:
275:
274:
268:
267:
264:
254:
253:
247:
246:
232:
228:
227:
219:
215:
214:
206:
202:
201:
193:
189:
188:
183:
179:
178:
173:
169:
168:
163:
159:
158:
153:
149:
148:
135:
134:
122:
121:
113:
112:
104:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
41:
28:
15:
13:
10:
9:
6:
4:
3:
2:
2668:
2657:
2654:
2652:
2649:
2647:
2644:
2642:
2639:
2637:
2634:
2632:
2629:
2628:
2626:
2609:
2604:
2600:
2596:
2591:
2587:
2583:
2578:
2574:
2569:
2563:
2559:
2558:
2556:
2554:
2553:Eysyslopterus
2550:
2546:
2541:
2531:
2521:
2515:
2512:
2510:
2507:
2506:
2504:
2500:
2494:
2491:
2489:
2486:
2484:
2481:
2480:
2478:
2474:
2468:
2467:
2463:
2461:
2460:
2456:
2454:
2453:
2449:
2448:
2446:
2444:
2440:
2434:
2433:
2429:
2427:
2426:
2422:
2420:
2419:
2415:
2413:
2412:
2411:Pentecopterus
2408:
2406:
2405:
2401:
2399:
2398:
2394:
2392:
2391:
2387:
2385:
2384:
2383:Megalograptus
2380:
2378:
2377:
2376:Jaekelopterus
2373:
2371:
2370:
2366:
2364:
2363:
2359:
2357:
2356:
2352:
2350:
2349:
2348:Drepanopterus
2345:
2343:
2342:
2338:
2336:
2335:
2331:
2329:
2328:
2327:Brachyopterus
2324:
2322:
2321:
2317:
2316:
2314:
2310:
2304:
2303:South America
2301:
2299:
2296:
2294:
2293:North America
2291:
2289:
2286:
2284:
2281:
2279:
2276:
2275:
2273:
2271:
2267:
2261:
2258:
2256:
2255:Carboniferous
2253:
2251:
2248:
2246:
2243:
2241:
2238:
2237:
2235:
2233:
2232:Geochronology
2229:
2224:
2202:
2199:
2197:
2194:
2192:
2189:
2188:
2186:
2184:
2180:
2174:
2171:
2170:
2168:
2166:
2162:
2156:
2153:
2152:
2150:
2148:
2144:
2138:
2135:
2133:
2130:
2128:
2125:
2124:
2122:
2120:
2116:
2113:
2111:
2105:
2099:
2096:
2094:
2091:
2089:
2086:
2085:
2083:
2081:
2080:Eurypteroidea
2077:
2071:
2068:
2067:
2065:
2063:
2059:
2053:
2050:
2049:
2047:
2045:
2041:
2038:
2036:
2030:
2020:
2019:Mycteroptidae
2017:
2015:
2012:
2010:
2007:
2006:
2004:
2002:
2001:Mycteropoidea
1998:
1992:
1989:
1987:
1984:
1983:
1981:
1979:
1975:
1969:
1966:
1964:
1961:
1960:
1958:
1956:
1955:Stylonuroidea
1952:
1946:
1945:Rhenopteridae
1943:
1942:
1940:
1938:
1934:
1931:
1929:
1923:
1917:
1914:
1913:
1911:
1907:
1904:
1900:
1894:
1893:Sclerophorata
1890:
1887:
1885:
1881:
1878:
1876:
1872:
1869:
1867:
1863:
1860:
1858:
1857:Euchelicerata
1854:
1851:
1849:
1845:
1843:
1839:
1837:
1833:
1832:
1829:
1825:
1818:
1813:
1811:
1806:
1804:
1799:
1798:
1795:
1786:
1782:
1777:
1774:
1769:
1765:
1761:
1757:
1753:
1749:
1745:
1741:
1737:
1730:
1727:
1723:
1717:
1714:
1709:
1705:
1701:
1697:
1690:
1687:
1682:
1678:
1674:
1670:
1666:
1662:
1658:
1651:
1649:
1647:
1645:
1643:
1641:
1639:
1637:
1633:
1628:
1624:
1620:
1619:
1611:
1609:
1607:
1603:
1598:
1594:
1590:
1586:
1582:
1578:
1574:
1572:
1563:
1560:
1555:
1551:
1547:
1542:
1537:
1533:
1529:
1525:
1521:
1517:
1513:
1512:
1507:
1500:
1497:
1491:
1487:
1484:
1482:
1479:
1478:
1474:
1472:
1470:
1466:
1462:
1458:
1454:
1450:
1447:
1443:
1442:Eysyslopterus
1439:
1436:
1432:
1428:
1427:
1422:
1421:
1416:
1415:
1410:
1409:E. osiliensis
1406:
1402:
1398:
1394:
1391:
1387:
1384:
1380:
1379:Eysyslopterus
1372:
1365:
1364:
1356:
1355:
1347:
1346:
1338:
1337:
1329:
1328:
1320:
1319:
1311:
1310:
1302:
1301:
1293:
1292:
1289:
1288:
1287:
1286:Jaekelopterus
1280:
1279:
1276:
1275:
1272:
1271:
1270:
1263:
1262:
1256:
1255:
1252:
1251:
1248:
1247:
1246:
1239:
1238:
1232:
1231:
1228:
1227:
1224:
1223:
1222:
1215:
1214:
1208:
1207:
1204:
1203:
1200:
1199:
1198:
1191:
1190:
1184:
1183:
1180:
1179:
1176:
1175:
1174:
1167:
1166:
1160:
1159:
1156:
1155:
1152:
1151:
1150:
1143:
1142:
1136:
1133:
1132:
1129:
1128:
1120:
1119:
1111:
1110:
1102:
1101:
1093:
1092:
1089:
1088:
1087:
1080:
1079:
1076:
1075:
1072:
1071:
1070:
1063:
1062:
1056:
1055:
1052:
1051:
1048:
1047:
1046:
1039:
1038:
1032:
1031:
1028:
1027:
1019:
1018:
1015:
1014:
1013:
1006:
1005:
1002:
1001:
998:
997:
996:
989:
988:
982:
981:
975:
974:
971:
970:
967:
966:
965:
964:Eysyslopterus
958:
957:
951:
948:
947:
941:
940:
937:
936:
928:
927:
919:
918:
915:
914:
913:
906:
905:
902:
901:
898:
897:
896:
889:
888:
882:
881:
878:
877:
874:
873:
872:
865:
864:
858:
855:
854:
848:
845:
844:
840:
837:
834:
830:
822:
818:
813:
809:
807:
803:
802:Eysyslopterus
799:
798:plesiomorphic
795:
791:
790:Eysyslopterus
787:
783:
779:
776:
772:
771:
766:
765:
760:
759:Eysyslopterus
755:
754:Eysyslopterus
751:
749:
745:
741:
737:
733:
732:Eysyslopterus
726:
724:
722:
718:
714:
710:
705:
701:
697:
693:
689:
685:
681:
680:
675:
671:
667:
663:
659:
654:
652:
651:H. lanceolata
648:
644:
640:
638:
633:
629:
625:
621:
617:
613:
609:
606:
602:
598:
594:
590:
586:
585:Eysyslopterus
579:
574:
567:
565:
563:
559:
555:
551:
550:Eysyslopterus
547:
545:
541:
540:ornamentation
537:
533:
529:
525:
521:
517:
513:
509:
505:
501:
500:Eysyslopterus
497:
495:
494:
489:
488:Eysyslopterus
485:
482:
475:
470:
463:
461:
459:
456:
452:
451:Eysyslopterus
448:
447:ornamentation
444:
440:
436:
432:
428:
427:Eysyslopterus
424:
422:
421:Eysyslopterus
418:
414:
410:
406:
402:
398:
394:
390:
386:
382:
379:
375:
372:
368:
364:
360:
356:
352:
348:
344:
341:
337:
336:Eysyslopterus
333:
329:
325:
321:
320:
319:Eysyslopterus
306:
303:
298:
294:
290:
287:
284:Størmer, 1934
282:
279:
278:
276:
273:
269:
265:Størmer, 1934
262:
261:
255:
252:
248:
241:
240:
239:Eysyslopterus
233:
230:
229:
226:
220:
217:
216:
213:
207:
205:Superfamily:
204:
203:
200:
194:
191:
190:
187:
184:
181:
180:
177:
174:
171:
170:
167:
164:
161:
160:
157:
154:
151:
150:
145:
140:
136:
132:
128:
123:
119:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
26:
25:Eysyslopterus
22:
19:
2552:
2464:
2457:
2450:
2430:
2423:
2416:
2409:
2402:
2395:
2388:
2381:
2374:
2369:Hughmilleria
2367:
2360:
2353:
2346:
2339:
2332:
2325:
2318:
2201:Pterygotidae
2137:Mixopteridae
2093:Eurypteridae
1968:Stylonuridae
1888:
1879:
1870:
1861:
1852:
1784:
1776:
1743:
1739:
1729:
1716:
1699:
1695:
1689:
1664:
1660:
1617:
1580:
1576:
1570:
1562:
1515:
1509:
1499:
1441:
1431:osteostracid
1424:
1418:
1412:
1408:
1404:
1378:
1376:
1373:Paleoecology
1284:
1283:
1267:
1266:
1243:
1242:
1221:Erettopterus
1219:
1218:
1195:
1194:
1171:
1170:
1149:Hughmilleria
1147:
1146:
1084:
1083:
1067:
1066:
1043:
1042:
1010:
1009:
993:
992:
963:
962:
961:
912:Grossopterus
910:
909:
893:
892:
871:Orcanopterus
869:
868:
826:
816:
806:sister taxon
801:
794:Hughmilleria
793:
789:
786:pterygotioid
781:
778:Orcanopterus
777:
768:
764:Orcanopterus
762:
758:
753:
752:
748:Hughmilleria
747:
743:
731:
730:
720:
716:
708:
703:
696:phylogenetic
691:
687:
683:
677:
673:
670:Hughmilleria
669:
665:
658:Hughmilleria
657:
655:
650:
646:
643:Hughmilleria
642:
637:Hughmilleria
635:
624:Leif Størmer
620:Soviet Union
584:
583:
577:
561:
549:
548:
499:
498:
491:
487:
478:
473:
450:
426:
425:
420:
407:name honors
400:
392:
369:of the Late
362:
359:type species
335:
318:
317:
316:
304:
296:
292:
289:Hughmilleria
288:
280:
259:
258:
251:Type species
238:
237:
130:
24:
18:
2466:Palmichnium
2443:Ichnogenera
2341:Carcinosoma
2035:Eurypterina
1928:Stylonurina
1866:Prosomapoda
1848:Chelicerata
1846:Subphylum:
1824:Eurypterids
1435:thelodontid
995:Bassipterus
740:superfamily
484:eurypterids
464:Description
355:superfamily
349:within the
345:, the only
199:Eurypterida
186:Chelicerata
182:Subphylum:
2625:Categories
2432:Stylonurus
2418:Pterygotus
2397:Mixopterus
2390:Megarachne
2355:Eurypterus
2240:Ordovician
2196:Slimonidae
2107:Infraorder
1884:Dekatriata
1875:Planaterga
1842:Arthropoda
1627:1006783631
1492:References
1269:Acutiramus
1245:Pterygotus
836:infraorder
821:type genus
744:E. patteni
704:N. patteni
684:H. patteni
647:H. patteni
544:transverse
508:appendages
474:E. patteni
443:transverse
363:E. patteni
332:arthropods
328:eurypterid
176:Arthropoda
131:E. patteni
2568:Q48850505
2509:Metastoma
2493:Xiphosura
2483:Arachnida
2270:Geography
1834:Kingdom:
1768:140668235
1760:0022-3360
1532:1744-9561
1469:limestone
1453:lithology
1446:nearshore
829:cladogram
662:subgenera
605:zoologist
601:biologist
516:posterior
455:nearshore
417:zoologist
413:biologist
162:Kingdom:
156:Eukaryota
2562:Wikidata
2530:Category
2452:Arcuites
2425:Slimonia
2250:Devonian
2245:Silurian
2032:Suborder
1925:Suborder
1902:Taxonomy
1840:Phylum:
1836:Animalia
1681:59488956
1597:91741388
1550:19828493
1475:See also
1465:dolomite
1457:mudcrack
1449:lagoonal
1386:deposits
1383:Silurian
1197:Slimonia
616:Saaremaa
589:holotype
524:carapace
512:anterior
458:lagoonal
439:carapace
385:Saaremaa
371:Silurian
367:deposits
272:Synonyms
218:Family:
172:Phylum:
166:Animalia
152:Domain:
127:carapace
2595:7690487
2582:8512215
2298:Oceania
2260:Permian
1708:1301214
1541:2865068
1461:burrows
1397:habitat
1388:of the
721:Eysysla
717:opterus
709:Eysysla
660:in two
639:patteni
632:species
520:lateral
504:prosoma
405:species
401:opterus
393:Eysysla
389:Estonia
351:derived
297:patteni
231:Genus:
192:Order:
105:↓
2608:259141
2288:Europe
2278:Africa
1909:Genera
1766:
1758:
1706:
1679:
1625:
1595:
1548:
1538:
1530:
1438:fishes
1390:Ludlow
713:Viking
711:, the
700:family
597:fossil
552:was a
536:ocelli
528:dorsal
397:Viking
395:, the
378:Ludlow
374:period
340:family
31:Ludlow
1889:Clade
1880:Clade
1871:Clade
1862:Clade
1853:Clade
1764:S2CID
1704:JSTOR
1677:S2CID
1593:S2CID
1401:fauna
1399:of a
1393:epoch
736:clade
558:genus
554:basal
431:basal
383:) in
381:epoch
347:clade
324:genus
322:is a
2577:GBIF
2283:Asia
1756:ISSN
1623:OCLC
1546:PMID
1528:ISSN
1467:and
1459:and
1433:and
1423:and
827:The
767:and
723:").
603:and
532:eyes
435:eyes
415:and
44:PreꞒ
1748:doi
1669:doi
1585:doi
1536:PMC
1520:doi
1463:of
614:in
326:of
129:of
2627::
2605::
2592::
2579::
2564::
1891::
1882::
1873::
1864::
1855::
1783:.
1762:.
1754:.
1744:81
1742:.
1738:.
1700:35
1698:.
1675:.
1663:.
1659:.
1635:^
1605:^
1591:.
1581:52
1579:.
1575:.
1552:.
1544:.
1534:.
1526:.
1514:.
1508:.
1471:.
1417:,
1411:,
1407:,
819:,
688:H.
674:H.
666:H.
664:,
653:.
564:.
486:,
423:.
387:,
361:,
334:.
295:)
94:Pg
38:Ma
33:,
1816:e
1809:t
1802:v
1787:.
1770:.
1750::
1710:.
1683:.
1671::
1665:6
1629:.
1599:.
1587::
1556:.
1522::
1516:6
690:(
676:(
668:(
526:(
376:(
291:(
257:†
235:†
222:†
209:†
196:†
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:Ꞓ
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