124:
253:
264:
specimens are known, ranging from about 1 to 4 cm in diameter; they are circular in outline and their eight arms, with closed ends, spiral either clockwise or counterclockwise. Ridges cut across both the inside and outside of the spiral arms. The arms of the
Australian individuals are longer
476:
Trying to relate organisms that are preserved in the two modes is immensely problematic, because they preserve such different parts of organisms; the imprints preserve a casts of the organism's outline, whereas the carbonaceous films display a shadow left by any resistant organic matter. This has
415:(Ediacara-type preservation), sandy casts in the beds of sandstone (Nama-type preservation), Avalon-type preservation characterized imprints on the top mudstone surface covered by volcanic ash, and in Khatyspyt-type preservation the fossils are found in finely laminated to medium-bedded, nodular
111:
308:
or not. If it is, it would be the earliest known fossil of an adult animal; and its anatomy is consistent with that expected from the earliest animals. However, it is not perfectly clear that it is an animal;
325:
In the
Ediacaran (Vendian) period there are two biotas of multicellular organisms; as now understood, these biota do not generally share members, as was considered until recently:
598:
Wang Yue; Wang
Xunlian; Huang Yuming (June 2008). "Megascopic Symmetrical Metazoans from the Ediacaran Doushantuo Formation in the Northeastern Guizhou, South China".
468:
and others. These fossils are of the form of flat organic carbonaceous films; it is shadow left by organic matter of the organism flesh (it is looks like of the
481:
is one of the few organisms which can convincingly be compared in both taphonomic modes. Such cases of preservation are known also for organisms related to the
411:
and others. These organisms mainly are negative and positive imprints on the base of sandstone beds with the "elephant skin" and tubercle texture diagnostic of
329:
Ediacara-type biota (it is in broad sense: Ediacara-, Nama-, Avalon-type ecological assemblage) it is community of the
Metazoa and problematic organisms:
265:
and more tightly coiled than those of the
Chinese, despite the Australian individuals not attaining as large a diameter; they are more often kinked.
811:
637:
695:
D. V. Grazhdankin; K. E. Nagovitsin; A.V. Maslov (2007). "Late
Vendian Miaohe-type Ecological Assemblage of the East European Platform".
281:. However, the discovery of the Chinese fossils, which have preserved organic matter, ruled out these interpretations, because the
638:"Macroscopic carbonaceous compressions in a terminal Proterozoic shale: A systematic reassessment of the Miaohe biota: South China"
469:
282:
865:
289:
also preserved in the
Doushantuo formation; the exact purpose still remains out on this until intermediate forms are found.
285:
displayed required relatively robust organic material to start with. Its spiral form has also led to comparison with the
304:, but it lacks sufficient physical characteristics to ascertain with any degree of certainty whether it is indeed an
123:
237:
875:
656:
524:
870:
779:
668:
837:
194:
751:
704:
572:
416:
241:
560:
561:"Eight-armed Ediacara fossil preserved in contrasting taphonomic windows from China and Australia"
460:
118:
842:
731:
824:
660:
615:
829:
759:
712:
652:
607:
580:
408:
755:
708:
576:
493:
301:
286:
229:
155:
611:
859:
412:
397:
355:
313:, the dominant constituent of the Doushantuo biota, cannot be ruled out, except that
68:
373:
278:
274:
559:
Maoyan Zhu; James G. Gehling; Shuhai Xiao; Yuanlong Zhao; Mary L. Droser (2008).
507:
391:
379:
361:
331:
43:
802:
716:
343:
297:
293:
88:
53:
664:
619:
498:
487:
403:
367:
233:
226:
135:
93:
37:
30:
252:
796:
83:
78:
63:
58:
48:
110:
816:
483:
432:
349:
98:
73:
584:
763:
502:
454:
385:
337:
305:
261:
145:
773:
512:
423:
310:
251:
777:
277:, and has also been considered to represent an agglutinating
657:
10.1666/0022-3360(2002)076<0347:MCCIAT>2.0.CO;2
232:
consisting of eight radial spiral arms, and known from two
690:
688:
240:
in
Australia, and as carbonaceous compressions from the
477:
made relating the two taphonomic types very difficult:
636:
Xiao, S.; Yuan, X.; Steiner, M.; Knoll, A.H. (2007).
292:
The organism bears a very superficial resemblance to
786:
631:
629:
491:-like fossils from the Doushantuo Formation, for
744:Geological Society, London, Special Publications
422:Miaohe-type biota characterizes assemblage of
8:
213:Tang, Yin, Bengtson, Liu, Wang and Gao, 2008
505:and Doushantuo Formation, and possible for
774:
600:Journal of China University of Geosciences
109:
20:
732:"Comparative taphonomy of Vendian genera
554:
552:
550:
548:
546:
544:
542:
540:
273:The organism was first interpreted as a
536:
7:
497:from the Lyamtsa Formation of the
14:
511:from Perevalok Formation of the
244:of China, where it is abundant.
122:
470:Burgess shale type preservation
283:Burgess shale type preservation
1:
612:10.1016/S1002-0705(08)60039-4
317:seems a little too complex.
16:Species of Ediacaran animal
892:
238:Ediacara type preservation
788:Eoandromeda octobrachiata
740:: taxonomy and lifestyle"
717:10.1134/S1028334X07080107
494:Beltanelloides sorichevae
300:and to some of the other
207:Eoandromeda octobrachiata
200:
193:
119:Scientific classification
117:
108:
23:
525:List of Ediacaran genera
645:Journal of Paleontology
321:Taphonomic significance
697:Doklady Earth Sciences
257:
866:Enigmatic animal taxa
838:Paleobiology Database
730:M. V. Leonov (2007).
417:bituminous limestones
255:
186:E. octobrachiata
242:Doushantuo formation
756:2007GSLSP.286..259L
709:2007DokES.417.1183G
577:2008Geo....36..867Z
508:Cyclomedusa davidi
258:
853:
852:
825:Open Tree of Life
780:Taxon identifiers
585:10.1130/G25203A.1
409:palaeopascichnids
218:
217:
883:
876:Fossils of China
846:
845:
833:
832:
820:
819:
807:
806:
805:
775:
768:
767:
764:10.1144/SP286.18
727:
721:
720:
703:(8): 1183β1187.
692:
683:
682:
680:
679:
673:
667:. Archived from
642:
633:
624:
623:
595:
589:
588:
556:
256:Life restoration
234:taphonomic modes
209:
205:
169:
127:
126:
113:
103:
40:
29:Temporal range:
21:
891:
890:
886:
885:
884:
882:
881:
880:
856:
855:
854:
849:
841:
836:
828:
823:
815:
810:
801:
800:
795:
782:
772:
771:
729:
728:
724:
694:
693:
686:
677:
675:
671:
640:
635:
634:
627:
597:
596:
592:
571:(11): 867β870.
558:
557:
538:
533:
521:
450:Liulingjitaenia
323:
271:
250:
236:: the standard
214:
211:
203:
202:
189:
167:
121:
104:
102:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
35:
34:
27:
17:
12:
11:
5:
889:
887:
879:
878:
873:
871:Ediacaran life
868:
858:
857:
851:
850:
848:
847:
834:
821:
808:
792:
790:
784:
783:
778:
770:
769:
750:(1): 259β267.
734:Beltanelloides
722:
684:
651:(2): 347β376.
625:
606:(3): 200β206.
590:
535:
534:
532:
529:
528:
527:
520:
517:
474:
473:
428:Beltanelloides
426:macrofossils:
420:
413:microbial mats
322:
319:
302:Ediacara biota
287:fossil embryos
270:
267:
249:
246:
216:
215:
212:
198:
197:
191:
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182:
180:
176:
175:
165:
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157:incertae sedis
153:
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143:
139:
138:
133:
129:
128:
115:
114:
106:
105:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
41:
28:
15:
13:
10:
9:
6:
4:
3:
2:
888:
877:
874:
872:
869:
867:
864:
863:
861:
844:
839:
835:
831:
826:
822:
818:
813:
809:
804:
798:
794:
793:
791:
789:
785:
781:
776:
765:
761:
757:
753:
749:
745:
741:
739:
735:
726:
723:
718:
714:
710:
706:
702:
698:
691:
689:
685:
674:on 2011-07-18
670:
666:
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658:
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650:
646:
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632:
630:
626:
621:
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582:
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566:
562:
555:
553:
551:
549:
547:
545:
543:
541:
537:
530:
526:
523:
522:
518:
516:
514:
513:Central Urals
510:
509:
504:
500:
496:
495:
490:
489:
485:
480:
471:
467:
463:
462:
457:
456:
451:
447:
443:
439:
435:
434:
429:
425:
421:
418:
414:
410:
406:
405:
400:
399:
398:Tribrachidium
394:
393:
388:
387:
382:
381:
376:
375:
370:
369:
364:
363:
358:
357:
356:Charniodiscus
352:
351:
346:
345:
340:
339:
334:
333:
328:
327:
326:
320:
318:
316:
312:
307:
303:
299:
295:
290:
288:
284:
280:
279:foraminiferan
276:
268:
266:
263:
254:
247:
245:
243:
239:
235:
231:
228:
224:
223:
210:
208:
199:
196:
195:Binomial name
192:
188:
187:
181:
178:
177:
174:
173:
166:
163:
162:
159:
158:
154:
151:
150:
147:
144:
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140:
137:
134:
131:
130:
125:
120:
116:
112:
107:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
36:580β550
32:
26:
22:
19:
787:
747:
743:
737:
733:
725:
700:
696:
676:. Retrieved
669:the original
648:
644:
603:
599:
593:
568:
564:
506:
492:
482:
478:
475:
465:
459:
453:
449:
445:
442:Jiuqunaoella
441:
437:
431:
427:
402:
396:
390:
384:
378:
374:Parvancorina
372:
366:
360:
354:
348:
342:
336:
330:
324:
314:
291:
275:trace fossil
272:
260:A few dozen
259:
221:
220:
219:
206:
201:
185:
184:
171:
170:
156:
24:
18:
479:Eoandromeda
438:Sinospongia
392:Fractofusus
380:Pteridinium
362:Cyclomedusa
332:Dickinsonia
315:Eoandromeda
298:ctenophores
294:echinoderms
222:Eoandromeda
172:Eoandromeda
25:Eoandromeda
860:Categories
678:2008-11-15
531:References
461:Calyptrina
344:Kimberella
248:Morphology
33:, approx.
665:0022-3360
620:1002-0705
499:White Sea
488:Albumares
404:Hiemalora
368:Ediacaria
227:Ediacaran
179:Species:
142:Kingdom:
136:Eukaryota
31:Ediacaran
803:Q5381582
797:Wikidata
519:See also
466:Cucullus
446:Grypania
269:Affinity
230:organism
152:Phylum:
146:Animalia
132:Domain:
830:7996718
817:4941137
752:Bibcode
738:Nemiana
705:Bibcode
573:Bibcode
565:Geology
484:Anfesta
433:Mezenia
350:Charnia
204:†
183:†
164:Genus:
843:148137
663:
618:
503:Russia
501:area,
455:Tawuia
386:Rangea
338:Yorgia
306:animal
262:fossil
225:is an
672:(PDF)
641:(PDF)
424:algal
311:algae
812:GBIF
736:and
661:ISSN
616:ISSN
44:Preκ
760:doi
748:286
713:doi
701:417
653:doi
608:doi
581:doi
862::
840::
827::
814::
799::
758:.
746:.
742:.
711:.
699:.
687:^
659:.
649:76
647:.
643:.
628:^
614:.
604:19
602:.
579:.
569:36
567:.
563:.
539:^
515:.
472:).
464:,
458:,
452:,
448:,
444:,
440:,
436:,
430:,
407:,
401:,
395:,
389:,
383:,
377:,
371:,
365:,
359:,
353:,
347:,
341:,
335:,
296:,
94:Pg
38:Ma
766:.
762::
754::
719:.
715::
707::
681:.
655::
622:.
610::
587:.
583::
575::
486:-
419:.
168:β
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:κ
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