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Eritherium

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1144: 1938: 248: 227: 1925: 25: 1932: 445:(M1-3)). The piece is about 6 inches long, 5 inches wide and just over 3 inches high. In addition, the fossils include 15 more objects including the skull bones (frontal and nasal bones), lower jaw fragments and teeth and the upper and lower jaw. It was about 20 cm tall at the shoulder and weighed about 5–6 kg. 464:, but their teeth are more specialised. The dentition of the mandible that was reconstructed (from two left fragments) made up the complete sequence of the original teeth of mammals: with three 495:
cusps on the occlusal surface-bearing structure). Between these bumps were approaches for forming transverse strips on the first two molars and on the rearmost molar, which is typical in
499:. The premolars had only one (lower jaw) or two (in the maxilla) cusps. The first incisor is relatively large and asymmetric and already showed signs of reduction. These facts link 996:(sharks and rays). This places the findings in a geological time ranging from 61.1 to 57.8 million years ago. The genus was named by Emmanuel Gheerbrant (2009). The name 2045: 2101: 1177: 2032: 1126:
Rodolphe Tabuce, Raphaël Sarr, Sylvain Adnet, Renaud Lebrun, Fabrice Lihoreau, Jeremy E. Martin, Bernard Sambou, Mustapha Thiam und Lionel Hautier:
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fossils were found in the "lower bone bed" of the phosphate layer. Other fossils from this layer include earliest evidence of the carnivorous
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was relatively far forward in the skull. Most other early Paenungulata had an orbit significantly shifted to the rear position of the skull.
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between the canine tooth on the front and back teeth. This primitive mammalian dentition is unique among Proboscideans.
61: 1163: 42: 247: 561:) which together form one of the most complete evolutionary sequences of early mammalian radiations following the 68: 35: 397:. It lived about 60 million years ago. It was first named by Emmanuel Gheerbrant in 2009 and the type species is 1924: 75: 1987: 593:
support the hypothesis that paenungulates originated in Africa and diversified rapidly in the Paleocene.
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honors the residents of the village of Ouled Azzouz, near where most of the fossils were discovered.
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comes from the Greek words "eρυ" (eri: old) and "θηρίον" (therion: animal), while the species name
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Filling a gap in the proboscidean fossil record: a new genus from the Lutetian of Senegal.
853: 511: 434: 226: 1049: 1034:"Paleocene emergence of elephant relatives and the rapid radiation of African ungulates" 82: 1890: 1722: 1598: 1554: 1463: 1433: 1377: 1323: 1068: 1033: 989: 869: 652: 539: 430: 2080: 1970: 1880: 1860: 1813: 1797: 1783: 1776: 1736: 1715: 1708: 1701: 1680: 1654: 1591: 1386: 1347: 1313: 1262: 993: 786: 702: 545: 477: 177: 1824: 1729: 1647: 1619: 1524: 1453: 1393: 1363: 1303: 1272: 969: 829: 752: 677: 586: 578: 557: 551: 526: 492: 469: 457: 453: 1155: 1131: 421:(specimen number MNHN PM69) is now in the Musée d'histoire naturelle - Guimet in 1965: 1870: 1752: 1687: 1670: 1633: 1612: 1605: 1503: 1356: 1241: 1226: 1187: 973: 734: 611: 574: 382: 299: 152: 24: 2010: 1038:
Proceedings of the National Academy of Sciences of the United States of America
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suggest that the closest relatives of the proboscid lineage are the manatees (
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with other early Proboscideans. Another primitive characteristic is the short
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The reconstruction of the upper portion of the skull showed that the
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phosphate basin in Morocco. These are 10 to 20 km south of
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shared similarities in the structure of their teeth with other
18: 1093:"Shoulder height, body mass and shape of proboscideans" 1896: 1886: 1876: 1866: 1856: 1846: 1794: 1747: 1665: 1565: 1535: 1484: 1469: 1459: 1449: 1439: 1429: 1374: 1344: 1329: 1319: 1309: 1299: 1288: 1278: 1268: 1258: 1994: 1823: 1793: 1746: 1664: 1564: 1534: 1483: 1414: 1373: 1343: 1239: 49:. Unsourced material may be challenged and removed. 405:is the oldest, smallest and most primitive known 537:to all other primitive proboscideans (including 984:) two early proboscideans were described. The 1171: 8: 1027: 1025: 1023: 1021: 1019: 1017: 1982: 1422: 1251: 1247: 1236: 1178: 1164: 1156: 525:is the oldest known representative of the 225: 129: 1111: 1067: 1057: 109:Learn how and when to remove this message 1013: 480:. The tooth row was closed and had no 7: 47:adding citations to reliable sources 2102:Prehistoric monotypic mammal genera 845: 802: 777: 725: 718: 693: 668: 643: 618: 606: 14: 1936: 1930: 1923: 1142: 589:. The great age and location of 460:or early representatives of the 246: 23: 1130:Journal of Paleontology, 2019, 34:needs additional citations for 992:, and a diverse assortment of 1: 2087:Fossil taxa described in 2009 1100:Acta Palaeontologica Polonica 2112:Prehistoric placental genera 1032:Emmanuel Gheerbrant (2009). 437:, each of the two posterior 2107:Paleocene mammals of Africa 2130: 599:in cladogram after Tabuce 487:The molars were generally 120: 1945: 1920: 1425: 1254: 1250: 1235: 1194: 866: 850: 843: 825: 807: 800: 782: 775: 748: 730: 723: 716: 698: 691: 673: 666: 648: 641: 623: 616: 349: 342: 243:Scientific classification 241: 233: 224: 132: 429:(with approaches of the 121:Not to be confused with 16:Extinct genus of mammals 2092:Paleocene proboscideans 1132:doi:10.1017/jpa.2019.98 1091:Larramendi, A. (2016). 1059:10.1073/pnas.0900251106 1113:10.4202/app.00136.2014 585:) they form the group 2059:Paleobiology Database 441:(P3 and 4) and three 399:Eritherium azzouzorum 356:Eritherium azzouzorum 236:Eritherium azzouzorum 2097:Transitional fossils 1951:Plesielephantiformes 1186:Genera of the order 956:History of discovery 563:K/T extinction event 456:such as the extinct 43:improve this article 1539:Choerolophodontidae 1150:Paleontology portal 1050:2009PNAS..10610717G 1044:(26): 10717–10721. 567:Cladistic analyses 435:maxillary branches 387:Ouled Abdoun basin 335:E. azzouzorum 2074: 2073: 1988:Taxon identifiers 1979: 1978: 1918: 1917: 1914: 1913: 1910: 1909: 1901:Stegotetrabelodon 1770:Paratetralophodon 1410: 1409: 976:, where in 1996 ( 952: 951: 943: 942: 934: 933: 925: 924: 916: 915: 907: 906: 898: 897: 889: 888: 880: 879: 764: 763: 581:and the hyraxes ( 507:of the mandible. 491:(i.e. with small 367: 366: 323: 119: 118: 111: 93: 2119: 2067: 2066: 2054: 2053: 2041: 2040: 2028: 2027: 2015: 2014: 2013: 1983: 1940: 1934: 1928: 1927: 1898: 1888: 1878: 1868: 1858: 1848: 1796: 1749: 1667: 1627:Progomphotherium 1567: 1537: 1486: 1471: 1461: 1451: 1441: 1431: 1423: 1376: 1346: 1331: 1321: 1311: 1301: 1290: 1280: 1270: 1260: 1252: 1248: 1237: 1180: 1173: 1166: 1157: 1152: 1147: 1146: 1145: 1134: 1124: 1118: 1117: 1115: 1097: 1088: 1082: 1081: 1071: 1061: 1029: 960:The findings of 846: 803: 778: 726: 719: 694: 669: 644: 619: 607: 577:. Together with 425:and includes an 362:Gheerbrant, 2009 358: 354: 318: 311: 251: 250: 229: 219: 149: 138:Temporal range: 130: 114: 107: 103: 100: 94: 92: 51: 27: 19: 2129: 2128: 2122: 2121: 2120: 2118: 2117: 2116: 2077: 2076: 2075: 2070: 2062: 2057: 2049: 2044: 2036: 2031: 2023: 2018: 2009: 2008: 2003: 1990: 1980: 1975: 1961:Elephantimorpha 1956:Numidotheriidae 1941: 1922: 1906: 1819: 1789: 1751: 1750:"Tetralophodont 1742: 1669: 1660: 1569:Amebelodontidae 1560: 1530: 1479: 1419: 1417:Elephantiformes 1406: 1401:Prodeinotherium 1369: 1339: 1244: 1231: 1190: 1184: 1148: 1143: 1141: 1138: 1137: 1125: 1121: 1095: 1090: 1089: 1085: 1031: 1030: 1015: 1010: 958: 953: 944: 935: 926: 917: 908: 899: 890: 881: 854:Elephantiformes 765: 520: 497:lophodont teeth 415: 363: 360: 352: 351: 338: 317: 309: 245: 220: 218: 217: 216: 215: 210: 205: 200: 195: 190: 185: 180: 175: 170: 165: 160: 155: 144: 143: 136: 126: 115: 104: 98: 95: 52: 50: 40: 28: 17: 12: 11: 5: 2127: 2126: 2123: 2115: 2114: 2109: 2104: 2099: 2094: 2089: 2079: 2078: 2072: 2071: 2069: 2068: 2055: 2042: 2029: 2016: 2000: 1998: 1992: 1991: 1986: 1977: 1976: 1974: 1973: 1968: 1963: 1958: 1953: 1946: 1943: 1942: 1921: 1919: 1916: 1915: 1912: 1911: 1908: 1907: 1905: 1904: 1894: 1891:Stegodibelodon 1884: 1874: 1864: 1854: 1844: 1837: 1829: 1827: 1821: 1820: 1818: 1817: 1810: 1802: 1800: 1791: 1790: 1788: 1787: 1780: 1773: 1766: 1758: 1756: 1744: 1743: 1741: 1740: 1733: 1726: 1723:Rhynchotherium 1719: 1712: 1705: 1698: 1691: 1684: 1676: 1674: 1662: 1661: 1659: 1658: 1651: 1644: 1637: 1630: 1623: 1616: 1609: 1602: 1599:Archaeobelodon 1595: 1588: 1581: 1573: 1571: 1562: 1561: 1559: 1558: 1555:Choerolophodon 1551: 1543: 1541: 1532: 1531: 1529: 1528: 1521: 1514: 1507: 1500: 1492: 1490: 1481: 1480: 1478: 1477: 1467: 1464:Palaeomastodon 1457: 1447: 1437: 1434:Dagbatitherium 1426: 1420: 1415: 1412: 1411: 1408: 1407: 1405: 1404: 1397: 1390: 1382: 1380: 1378:Deinotheriidae 1371: 1370: 1368: 1367: 1360: 1352: 1350: 1341: 1340: 1338: 1337: 1327: 1324:Phosphatherium 1317: 1307: 1297: 1286: 1276: 1266: 1255: 1245: 1240: 1233: 1232: 1230: 1229: 1220: 1214: 1208: 1202: 1195: 1192: 1191: 1185: 1183: 1182: 1175: 1168: 1160: 1154: 1153: 1136: 1135: 1119: 1083: 1012: 1011: 1009: 1006: 990:Hyaenodontidae 978:Phosphatherium 957: 954: 950: 949: 946: 945: 941: 940: 937: 936: 932: 931: 928: 927: 923: 922: 919: 918: 914: 913: 910: 909: 905: 904: 901: 900: 896: 895: 892: 891: 887: 886: 883: 882: 878: 877: 874: 873: 870:Deinotheriidae 865: 862: 861: 858: 857: 849: 844: 842: 839: 838: 835: 834: 824: 821: 820: 817: 816: 806: 801: 799: 796: 795: 792: 791: 781: 776: 774: 771: 770: 767: 766: 762: 761: 758: 757: 747: 744: 743: 740: 739: 729: 724: 722: 717: 715: 712: 711: 708: 707: 697: 692: 690: 687: 686: 683: 682: 672: 667: 665: 662: 661: 658: 657: 653:Phosphatherium 647: 642: 640: 637: 636: 633: 632: 622: 617: 615: 605: 540:Phosphatherium 519: 516: 431:zygomatic bone 414: 411: 365: 364: 361: 347: 346: 340: 339: 331: 329: 325: 324: 307: 303: 302: 297: 293: 292: 287: 283: 282: 277: 273: 272: 267: 263: 262: 257: 253: 252: 239: 238: 231: 230: 222: 221: 213: 212: 211: 206: 201: 196: 191: 186: 181: 176: 171: 166: 161: 156: 151: 150: 137: 117: 116: 31: 29: 22: 15: 13: 10: 9: 6: 4: 3: 2: 2125: 2124: 2113: 2110: 2108: 2105: 2103: 2100: 2098: 2095: 2093: 2090: 2088: 2085: 2084: 2082: 2065: 2060: 2056: 2052: 2047: 2043: 2039: 2034: 2030: 2026: 2021: 2017: 2012: 2006: 2002: 2001: 1999: 1997: 1993: 1989: 1984: 1972: 1971:Elephantoidea 1969: 1967: 1964: 1962: 1959: 1957: 1954: 1952: 1948: 1947: 1944: 1939: 1935: 1933: 1926: 1903: 1902: 1895: 1893: 1892: 1885: 1883: 1882: 1881:Selenetherium 1875: 1873: 1872: 1865: 1863: 1862: 1861:Palaeoloxodon 1855: 1853: 1852: 1845: 1843: 1842: 1838: 1836: 1835: 1831: 1830: 1828: 1826: 1822: 1816: 1815: 1814:Stegolophodon 1811: 1809: 1808: 1804: 1803: 1801: 1799: 1798:Stegodontidae 1792: 1786: 1785: 1784:Tetralophodon 1781: 1779: 1778: 1777:Pediolophodon 1774: 1772: 1771: 1767: 1765: 1764: 1760: 1759: 1757: 1754: 1745: 1739: 1738: 1737:Stegomastodon 1734: 1732: 1731: 1727: 1725: 1724: 1720: 1718: 1717: 1716:Notiomastodon 1713: 1711: 1710: 1709:Gomphotherium 1706: 1704: 1703: 1702:Gnathabelodon 1699: 1697: 1696: 1692: 1690: 1689: 1685: 1683: 1682: 1681:Blancotherium 1678: 1677: 1675: 1672: 1668:"Trilophodont 1663: 1657: 1656: 1655:Torynobelodon 1652: 1650: 1649: 1645: 1643: 1642: 1638: 1636: 1635: 1631: 1629: 1628: 1624: 1622: 1621: 1617: 1615: 1614: 1610: 1608: 1607: 1603: 1601: 1600: 1596: 1594: 1593: 1592:Aphanobelodon 1589: 1587: 1586: 1582: 1580: 1579: 1575: 1574: 1572: 1570: 1563: 1557: 1556: 1552: 1550: 1549: 1548:Afrochoerodon 1545: 1544: 1542: 1540: 1533: 1527: 1526: 1522: 1520: 1519: 1515: 1513: 1512: 1508: 1506: 1505: 1501: 1499: 1498: 1494: 1493: 1491: 1489: 1482: 1476: 1475: 1468: 1466: 1465: 1458: 1456: 1455: 1448: 1446: 1445: 1438: 1436: 1435: 1428: 1427: 1424: 1421: 1418: 1413: 1403: 1402: 1398: 1396: 1395: 1391: 1389: 1388: 1387:Chilgatherium 1384: 1383: 1381: 1379: 1372: 1366: 1365: 1361: 1359: 1358: 1354: 1353: 1351: 1349: 1348:Barytheriidae 1342: 1336: 1335: 1328: 1326: 1325: 1318: 1316: 1315: 1314:Numidotherium 1308: 1306: 1305: 1298: 1295: 1294: 1287: 1285: 1284: 1277: 1275: 1274: 1267: 1265: 1264: 1263:Arcanotherium 1257: 1256: 1253: 1249: 1246: 1243: 1238: 1234: 1228: 1224: 1221: 1219: 1215: 1213: 1209: 1207: 1203: 1201: 1197: 1196: 1193: 1189: 1181: 1176: 1174: 1169: 1167: 1162: 1161: 1158: 1151: 1140: 1133: 1129: 1123: 1120: 1114: 1109: 1105: 1101: 1094: 1087: 1084: 1079: 1075: 1070: 1065: 1060: 1055: 1051: 1047: 1043: 1039: 1035: 1028: 1026: 1024: 1022: 1020: 1018: 1014: 1007: 1005: 1003: 999: 995: 994:elasmobranchs 991: 987: 983: 979: 975: 971: 967: 966:Sidi Chennane 963: 955: 948: 947: 939: 938: 930: 929: 921: 920: 912: 911: 903: 902: 894: 893: 885: 884: 876: 875: 872: 871: 864: 863: 860: 859: 856: 855: 848: 847: 841: 840: 837: 836: 833: 832: 831: 823: 822: 819: 818: 815: 814: 813: 805: 804: 798: 797: 794: 793: 790: 789: 788: 787:Arcanotherium 780: 779: 773: 772: 769: 768: 760: 759: 756: 755: 754: 746: 745: 742: 741: 738: 737: 736: 728: 727: 721: 720: 714: 713: 710: 709: 706: 705: 704: 703:Numidotherium 696: 695: 689: 688: 685: 684: 681: 680: 679: 671: 670: 664: 663: 660: 659: 656: 655: 654: 646: 645: 639: 638: 635: 634: 631: 630: 629: 621: 620: 613: 609: 608: 604: 602: 598: 594: 592: 588: 584: 580: 576: 572: 568: 564: 560: 559: 554: 553: 548: 547: 546:Numidotherium 542: 541: 536: 532: 528: 524: 517: 515: 513: 508: 506: 502: 498: 494: 490: 485: 483: 479: 475: 471: 467: 463: 459: 455: 451: 446: 444: 440: 436: 432: 428: 424: 420: 412: 410: 408: 404: 400: 396: 392: 388: 385:found in the 384: 380: 377: 373: 372: 359: 357: 348: 345: 344:Binomial name 341: 337: 336: 330: 327: 326: 322: 316: 315: 308: 305: 304: 301: 298: 295: 294: 291: 288: 285: 284: 281: 278: 275: 274: 271: 268: 265: 264: 261: 258: 255: 254: 249: 244: 240: 237: 232: 228: 223: 209: 204: 199: 194: 189: 184: 179: 174: 169: 164: 159: 154: 148: 141: 135: 131: 128: 124: 113: 110: 102: 91: 88: 84: 81: 77: 74: 70: 67: 63: 60: –  59: 55: 54:Find sources: 48: 44: 38: 37: 32:This article 30: 26: 21: 20: 1995: 1929: 1899: 1889: 1879: 1869: 1859: 1849: 1839: 1832: 1825:Elephantidae 1812: 1805: 1782: 1775: 1768: 1761: 1753:gomphotheres 1735: 1730:Sinomastodon 1728: 1721: 1714: 1707: 1700: 1693: 1686: 1679: 1671:gomphotheres 1653: 1648:Stenobelodon 1646: 1639: 1632: 1625: 1620:Platybelodon 1618: 1611: 1604: 1597: 1590: 1583: 1578:Afromastodon 1576: 1553: 1546: 1525:Zygolophodon 1523: 1516: 1509: 1502: 1495: 1472: 1462: 1454:Hemimastodon 1452: 1442: 1432: 1399: 1394:Deinotherium 1392: 1385: 1364:Omanitherium 1362: 1355: 1332: 1322: 1312: 1304:Moeritherium 1302: 1291: 1282: 1281: 1273:Daouitherium 1271: 1261: 1222: 1216:Superorder: 1127: 1122: 1103: 1099: 1086: 1041: 1037: 1001: 997: 985: 982:Daouitherium 981: 980:) and 2002 ( 977: 970:Ouled Abdoun 968:, quarry in 961: 959: 867: 851: 830:Moeritherium 828: 826: 810: 808: 785: 783: 753:Omanitherium 751: 749: 733: 731: 701: 699: 678:Daouitherium 676: 674: 651: 649: 627: 626: 624: 600: 596: 595: 590: 587:Paenungulata 579:Embrithopoda 558:Daouitherium 556: 552:Moeritherium 550: 544: 538: 530: 522: 521: 509: 500: 486: 458:Embrithopoda 454:Paenungulata 449: 447: 416: 402: 398: 370: 369: 368: 355: 350: 334: 333: 319:Gheerbrant, 313: 312: 235: 133: 127: 105: 99:January 2013 96: 86: 79: 72: 65: 58:"Eritherium" 53: 41:Please help 36:verification 33: 1966:Elephantida 1871:Primelephas 1688:Cuvieronius 1634:Protanancus 1613:Konobelodon 1606:Eurybelodon 1504:Losodokodon 1357:Barytherium 1293:Khamsaconus 1242:Proboscidea 1227:Tethytheria 1188:Proboscidea 974:Grand Daoui 962:Eritherium 735:Barytherium 612:Proboscidea 575:Desmostylia 531:Eritherium 527:Proboscidea 518:Systematics 501:Eritherium 450:Eritherium 448:Generally, 413:Description 403:Eritherium 383:Proboscidea 300:Proboscidea 2081:Categories 1996:Eritherium 1949:See also: 1641:Serbelodon 1585:Amebelodon 1518:Sinomammut 1488:Mammutidae 1283:Eritherium 1218:Afrotheria 1008:References 1002:azzouzorum 998:Eritherium 986:Eritherium 964:come from 628:Eritherium 597:Eritherium 591:Eritherium 583:Hyracoidea 523:Eritherium 512:eye socket 476:and three 409:relative. 371:Eritherium 314:Eritherium 134:Eritherium 69:newspapers 1851:Mammuthus 1841:Loxodonta 1695:Eubelodon 1497:Eozygodon 1198:Kingdom: 603:., 2019: 505:symphysis 474:premolars 439:premolars 427:upper jaw 391:Thanetian 381:of early 328:Species: 266:Kingdom: 260:Eukaryota 140:Selandian 2025:31992899 2011:Q1354868 2005:Wikidata 1807:Stegodon 1444:Eritreum 1334:Saloumia 1212:Mammalia 1206:Chordata 1204:Phylum: 1200:Animalia 1078:19549873 812:Saloumia 489:bunodont 482:diastema 466:incisors 462:manatees 433:and two 419:holotype 407:elephant 290:Mammalia 280:Chordata 276:Phylum: 270:Animalia 256:Domain: 145:60  123:Eritreum 2051:1472676 2038:8672689 1834:Elephas 1763:Anancus 1474:Phiomia 1210:Class: 1069:2705600 1046:Bibcode 614:  571:Sirenia 472:, four 395:Morocco 389:(early 376:extinct 353:† 332:† 306:Genus: 296:Order: 286:Class: 234:Jaw of 214:↓ 83:scholar 2064:157501 1511:Mammut 1076:  1066:  868:  852:  827:  809:  784:  750:  732:  700:  675:  650:  625:  610:  573:) and 493:enamel 478:molars 470:canine 468:, one 443:molars 393:age), 374:is an 85:  78:  71:  64:  56:  2046:IRMNG 1223:Clade 1096:(PDF) 601:et al 535:basal 379:genus 90:JSTOR 76:books 2033:GBIF 1074:PMID 555:and 423:Lyon 417:The 321:2009 153:PreꞒ 62:news 2020:EoL 1108:doi 1064:PMC 1054:doi 1042:106 565:. 533:is 45:by 2083:: 2061:: 2048:: 2035:: 2022:: 2007:: 1225:: 1106:. 1104:61 1102:. 1098:. 1072:. 1062:. 1052:. 1040:. 1036:. 1016:^ 549:, 543:, 401:. 203:Pg 147:Ma 142:, 1897:† 1887:† 1877:† 1867:† 1857:† 1847:† 1795:† 1755:" 1748:† 1673:" 1666:† 1566:† 1536:† 1485:† 1470:† 1460:† 1450:† 1440:† 1430:† 1375:† 1345:† 1330:† 1320:† 1310:† 1300:† 1296:? 1289:† 1279:† 1269:† 1259:† 1179:e 1172:t 1165:v 1116:. 1110:: 1080:. 1056:: 1048:: 310:† 208:N 198:K 193:J 188:T 183:P 178:C 173:D 168:S 163:O 158:Ꞓ 125:. 112:) 106:( 101:) 97:( 87:· 80:· 73:· 66:· 39:.

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Eritreum
Selandian
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Eukaryota

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