Knowledge (XXG)

Fusulinida

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Thirteen superfamilies are presently recognised, based on taxa (families) included in the three superfamilies given in the Treatise. Three are based on families in the Parathuramminacea, 1964, and nine families in the Endothyracea, 1964. The Fusulinacea remains the same in both sources (Treatise 1964
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Leppig, Ursula; Forke, Holger C.; Montenari, Michael; Fohrer, Beate (2005). "A three- and two-dimensional documentation of structural elements in schwagerinids (superfamily Fusulinoidea) exemplified by silicified material from the Upper Carboniferous of the Carnic Alps (Austria/Italy): a comparison
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Later species of fusulinids grew to much larger size, with some forms reaching 5 cm in length; reportedly, some specimens reach up to 14 cm in length, making them among the largest foraminifera extant or extinct. Fusulinids are the earliest lineage of foraminifera thought to have evolved
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revealed that this supposed structure actually represented tests that had been extensively modified by diagenetic processes. Instead, living fusulinids had low-magnesium hyaline tests with spherical nanograins up to 100 nm across, similar to the tests of the
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Traditionally, fusulinid tests were considered to have been composed of very small, tightly-packed calcite crystals with no preferred orientation—a so-called microgranular structure. However. a 2017 study using
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The term "fusulinid" applies to any of the Fusulinida. The Fusulinida are fusulinids (sensu lato). However, the term "fusulinid" is often applied just to the fusiform Fusulinacea and not to the entire order.
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The term fusulinata has traditionally been used to refer to all palaeozoic foraminifera with multi-chambered tests. However, recent studies based on test microstructure have suggested that fusulinids may be
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A 2021 study further examined test microstructure and suggested instead that the forams examined in the 2017 study were not true fusulinids, but rather considered them their own group containing
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for determining ages and correlating Upper Mississippian to Permian strata. In some places fusulinaceans may be so abundant as to be a significant component of
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Stevens, Calvin H. (September 1995). "A Giant Permian Fusulinid from East-Central Alaska with Comparisons of All Giant Fusulinids in Western North America".
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and relatives. These authors considers that true fusulinids did in fact have microgranular tests. A third group consisting of forms related to
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calcite. Like all forams, they were single-celled organisms. In advanced forms the test wall was differentiated into two or more layers.
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Saraswati, Pratul Kumar; Srinivasan, M. S. (2016), Saraswati, Pratul Kumar; Srinivasan, M.S. (eds.), "Calcareous-Walled Microfossils",
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Fusulinid limestone, Upper Pennsylvanian; Elk County, Kansas. Field of view is 3.9 cm wide. Link at source to view of entire slab.
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The fusulinids are among the earliest calcareous-walled foraminifera; they appear in the fossil record during the
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Tappan, Helen; Loeblich, Alfred R. (1988). "Foraminiferal Evolution, Diversification, and Extinction".
329:. While the latter is true, a more supported projected timespan is from the Mid-Carboniferous period. 2212: 2129: 1901: 1724: 906: 567: 500: 264: 2004: 1913: 1907: 1714: 1485: 1360: 1202: 379: 272: 224: 2235: 1988: 1453: 1254: 228: 1947: 1857: 1810: 1719: 1370: 1365: 1146: 268: 232: 1077: 1055: 1043: 1018: 930: 922: 797: 731: 346:
and consist of at least three distinct lineages, and as such are in need of systematic revision.
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was also found to have a distinct test microstructure, and was suggested to be a third lineage.
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Dubicka, Zofia; Gajewska, Maria; Kozłowski, Wojciech; Mikhalevich, Valeria (29 June 2021).
2039: 1576: 856: 294: 910: 863: 571: 556:"Unlocking the biomineralization style and affinity of Paleozoic fusulinid foraminifera" 504: 418:. The group then gradually declined in diversity until finally going extinct during the 1994: 1805: 1709: 1438: 1314: 1174: 1130: 1033: 598: 555: 531: 488: 318: 252: 248: 165: 152: 2229: 1895: 1867: 1683: 1480: 1380: 1194: 1179: 934: 880: 801: 415: 365: 322: 306: 244: 195: 72: 2121: 2018: 1936: 1875: 1847: 1828: 1817: 1799: 1790: 1780: 1616: 1581: 1568: 1558: 1463: 1448: 1443: 1433: 1415: 1301: 1291: 1226: 1169: 1161: 440: 298: 236: 185: 817: 622: 2155: 2101: 1890: 1885: 1752: 1747: 1734: 1626: 1586: 1184: 691: 455: 240: 47: 580: 2027: 1953: 1742: 1701: 1691: 1673: 1621: 1596: 1591: 1423: 1393: 1347: 1329: 1319: 918: 793: 407: 92: 57: 2092: 727: 589: 522: 1880: 1842: 1545: 1355: 1272: 513: 444: 361: 343: 139: 97: 889: 607: 540: 411: 957: 2086: 2047: 2043: 1941: 1553: 1528: 1264: 1119: 1072: 988: 396: 87: 82: 67: 62: 52: 17: 2147: 2031: 1657: 951: 926: 735: 175: 102: 77: 2160: 489:"Test structure in some pioneer multichambered Paleozoic foraminifera" 2051: 749: 2063: 624:
The Carboniferous of the World: China, Korea, Japan & S.E. Asia
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Protista 2: Sarcodina Chiefly "Thecamoebians" and Foraminiferida
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Fossils of fusulinids have been found on all continents except
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Dubicka, Zofia; Gorzelak, Przemysław (9 November 2017).
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Payne, Jonathan; Jost, Adam; Wang, Steve (March 2013).
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Members, especially of the Fusulinacea, are excellent
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are traditionally considered to have been composed of
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sp. cross-section; Plattsmouth Chert; Red Oak, Iowa;
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(4 October 2013). 824:Treatise on Invertebrate Paleontology 7: 482: 480: 25: 327:Permian–Triassic extinction event 43:Lower Pennsylvanian–Upper Permian 881:10.1111/j.1558-5646.2012.01807.x 338:and Loeblich and Tappan, 1988). 126: 420:Permo-Triassic extinction event 317:, 1988, gives a range from the 621:Wagner, Robert Herman (1983). 1: 357:scanning electron microscopy 816:(1964). Moore, R.C. (ed.). 692:10.1007/978-3-319-14574-7_6 2277: 2256:Silurian first appearances 581:10.1038/s41598-017-15666-1 2013: 1774: 1667: 1650: 1522: 1410: 1342: 1310: 1285: 1220: 1112: 996: 919:10.1017/S0022336000035484 846:Geological Survey of Iran 794:10.1007/s10347-005-0014-4 222: 217: 123:Scientific classification 121: 112: 36: 750:"Fusulinids | GeoKansas" 899:Journal of Paleontology 716:Journal of Paleontology 514:10.1073/pnas.2100656118 955: 844:, E-Book published by 460: 286: 2261:Lopingian extinctions 2195:Paleobiology Database 946: 855:23 March 2012 at the 458: 284: 627:. IGME. p. 88. 387:Evolutionary history 2241:Foraminifera orders 2026:organization type: 911:1995JPal...69..805S 810:Loeblich, Alfred R. 572:2017NatSR...715218D 505:2021PNAS..11800656D 395:epoch of the early 257:Palaeotextulariacea 2246:Carboniferous life 1660:    1656:    956: 560:Scientific Reports 461: 287: 2223: 2222: 2182:Open Tree of Life 2070:Taxon identifiers 2061: 2060: 2044:heliozoan amoebae 2036:amoeboflagellates 2000:Labyrinthomyxidae 1974: 1973: 1970: 1969: 1966: 1965: 1962: 1961: 1919:Globigerinitoidea 1770: 1769: 1766: 1765: 1646: 1645: 1642: 1641: 1605: 1604: 1518: 1517: 1514: 1513: 1510: 1509: 1506: 1505: 1502: 1501: 1498: 1497: 1494: 1493: 1429:Phaeogymnocellida 1406: 1405: 1402: 1401: 1338: 1337: 1325:Paracercomonadida 1281: 1280: 1216: 1215: 833:978-0-8137-3003-5 701:978-3-319-14574-7 661:978-0-226-92537-0 634:978-84-300-9949-8 279: 278: 16:(Redirected from 2268: 2216: 2215: 2203: 2202: 2190: 2189: 2177: 2176: 2164: 2163: 2151: 2150: 2138: 2137: 2125: 2124: 2112: 2111: 2110: 2097: 2096: 2095: 2065: 1853:Silicoloculinida 1787: 1783: 1776: 1680: 1676: 1669: 1665: 1661: 1652: 1632:Plasmodiophorida 1542: 1535: 1531: 1524: 1412: 1389:Thaumatomonadida 1377: 1344: 1308: 1298: 1294: 1287: 1240: 1233: 1229: 1222: 1137: 1127: 1123: 1114: 1025: 1005: 982: 975: 968: 959: 938: 893: 883: 837: 805: 788:(1–4): 541–553. 765: 764: 762: 760: 754:geokansas.ku.edu 746: 740: 739: 711: 705: 704: 679: 673: 672: 670: 668: 645: 639: 638: 618: 612: 611: 601: 583: 551: 545: 544: 534: 516: 484: 350:Test composition 261:Parathuraminacea 210: 199: 131: 130: 117: 107: 44: 40:Temporal range: 34: 21: 2276: 2275: 2271: 2270: 2269: 2267: 2266: 2265: 2226: 2225: 2224: 2219: 2211: 2206: 2198: 2193: 2185: 2180: 2172: 2167: 2159: 2154: 2146: 2141: 2133: 2128: 2120: 2115: 2106: 2105: 2100: 2091: 2090: 2085: 2072: 2062: 2057: 2040:testate amoebae 2009: 1958: 1923: 1902:Globotruncanida 1862: 1823: 1779: 1762: 1729: 1725:Symphyacanthida 1696: 1672: 1655: 1638: 1601: 1577:Claustrosporida 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1944: 1939: 1933: 1931: 1929:Incertae sedis 1925: 1924: 1922: 1921: 1916: 1914:Guembelitriida 1910: 1908:Heterohelicida 1904: 1898: 1893: 1888: 1883: 1878: 1872: 1870: 1864: 1863: 1861: 1860: 1855: 1850: 1845: 1840: 1833: 1831: 1825: 1824: 1822: 1821: 1814: 1808: 1806:Xenophyophorea 1803: 1796: 1794: 1784: 1772: 1771: 1768: 1767: 1764: 1763: 1761: 1760: 1755: 1750: 1745: 1739: 1737: 1731: 1730: 1728: 1727: 1722: 1717: 1715:Chaunocanthida 1712: 1710:Arthracanthida 1706: 1704: 1698: 1697: 1695: 1694: 1688: 1686: 1677: 1662: 1648: 1647: 1644: 1643: 1640: 1639: 1637: 1636: 1635: 1634: 1629: 1619: 1613: 1611: 1607: 1606: 1603: 1602: 1600: 1599: 1594: 1589: 1584: 1579: 1573: 1571: 1565: 1564: 1562: 1561: 1556: 1550: 1548: 1539: 1532: 1520: 1519: 1516: 1515: 1512: 1511: 1508: 1507: 1504: 1503: 1500: 1499: 1496: 1495: 1492: 1491: 1489: 1488: 1486:Ventricleftida 1483: 1478: 1477: 1476: 1471: 1466: 1458: 1457: 1456: 1451: 1446: 1441: 1439:Phaeosphaerida 1436: 1431: 1420: 1418: 1408: 1407: 1404: 1403: 1400: 1399: 1397: 1396: 1391: 1385: 1383: 1374: 1373: 1368: 1363: 1361:Spongomonadida 1358: 1352: 1350: 1340: 1339: 1336: 1335: 1333: 1332: 1327: 1322: 1317: 1315:Glissomonadida 1311: 1305: 1295: 1283: 1282: 1279: 1278: 1276: 1275: 1269: 1267: 1261: 1260: 1258: 1257: 1252: 1246: 1244: 1237: 1230: 1218: 1217: 1214: 1213: 1211: 1210: 1205: 1203:Chlorarachnida 1199: 1197: 1191: 1190: 1188: 1187: 1182: 1177: 1175:Desmothoracida 1172: 1166: 1164: 1158: 1157: 1155: 1154: 1149: 1143: 1141: 1134: 1131:Reticulofilosa 1124: 1110: 1109: 1106: 1105: 1104: 1103: 1102: 1101: 1098:Mesomycetozoea 1095: 1090: 1080: 1070: 1069: 1068: 1063: 1058: 1053: 1048: 1047: 1046: 1034:Diaphoretickes 1031: 1026: 1021: 1016: 1011: 1006: 998: 997: 994: 993: 991:classification 987: 985: 984: 977: 970: 962: 940: 939: 905:(5): 805–812. 894: 874:(3): 816–827. 859: 838: 832: 806: 775: 773: 770: 767: 766: 741: 722:(5): 695–714. 706: 700: 674: 660: 640: 633: 613: 546: 475: 474: 472: 469: 468: 467: 452: 449: 436: 433: 427: 424: 388: 385: 380:Semitextularia 351: 348: 334: 331: 319:Lower Silurian 293:is an extinct 277: 276: 273:Tournayellacea 253:Nodosinellacea 249:Moravamminacea 225:Archaediscacea 220: 219: 218:Superfamilies 215: 214: 206: 202: 201: 193: 189: 188: 183: 179: 178: 173: 169: 168: 163: 156: 155: 153:Diaphoretickes 150: 143: 142: 137: 133: 132: 119: 118: 110: 109: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 51: 46: 45: 39: 28: 24: 14: 13: 10: 9: 6: 4: 3: 2: 2273: 2262: 2259: 2257: 2254: 2252: 2249: 2247: 2244: 2242: 2239: 2237: 2234: 2233: 2231: 2214: 2209: 2205: 2201: 2196: 2192: 2188: 2183: 2179: 2175: 2170: 2166: 2162: 2157: 2153: 2149: 2144: 2140: 2136: 2131: 2127: 2123: 2118: 2114: 2109: 2103: 2099: 2094: 2088: 2084: 2083: 2081: 2079: 2075: 2071: 2066: 2053: 2049: 2045: 2041: 2037: 2033: 2029: 2025: 2022: 2020: 2016: 2015: 2012: 2006: 2003: 2001: 1998: 1996: 1993: 1991: 1990: 1986: 1985: 1983: 1981: 1977: 1955: 1952: 1950: 1949: 1945: 1943: 1940: 1938: 1935: 1934: 1932: 1930: 1926: 1920: 1917: 1915: 1911: 1909: 1905: 1903: 1899: 1897: 1896:Globigerinida 1894: 1892: 1889: 1887: 1884: 1882: 1879: 1877: 1874: 1873: 1871: 1869: 1868:Globothalamea 1865: 1859: 1856: 1854: 1851: 1849: 1846: 1844: 1841: 1839: 1835: 1834: 1832: 1830: 1826: 1820: 1819: 1815: 1812: 1809: 1807: 1804: 1801: 1798: 1797: 1795: 1792: 1788: 1785: 1782: 1777: 1773: 1759: 1756: 1754: 1751: 1749: 1746: 1744: 1741: 1740: 1738: 1736: 1732: 1726: 1723: 1721: 1718: 1716: 1713: 1711: 1708: 1707: 1705: 1703: 1699: 1693: 1690: 1689: 1687: 1685: 1684:Sticholonchea 1681: 1678: 1675: 1670: 1666: 1663: 1659: 1653: 1649: 1633: 1630: 1628: 1625: 1624: 1623: 1620: 1618: 1615: 1614: 1612: 1608: 1598: 1595: 1593: 1590: 1588: 1585: 1583: 1580: 1578: 1575: 1574: 1572: 1570: 1566: 1560: 1557: 1555: 1552: 1551: 1549: 1547: 1543: 1540: 1536: 1533: 1530: 1525: 1521: 1487: 1484: 1482: 1481:Tectofilosida 1479: 1475: 1472: 1470: 1467: 1465: 1462: 1461: 1459: 1455: 1454:Phaeoconchida 1452: 1450: 1447: 1445: 1442: 1440: 1437: 1435: 1432: 1430: 1427: 1426: 1425: 1422: 1421: 1419: 1417: 1413: 1409: 1395: 1392: 1390: 1387: 1386: 1384: 1382: 1381:Silicofilosea 1378: 1372: 1369: 1367: 1364: 1362: 1359: 1357: 1354: 1353: 1351: 1349: 1345: 1341: 1331: 1330:Cercomonadida 1328: 1326: 1323: 1321: 1318: 1316: 1313: 1312: 1309: 1306: 1303: 1299: 1296: 1293: 1288: 1284: 1274: 1271: 1270: 1268: 1266: 1262: 1256: 1255:Metromonadida 1253: 1251: 1248: 1247: 1245: 1241: 1238: 1234: 1231: 1228: 1223: 1219: 1209: 1206: 1204: 1201: 1200: 1198: 1196: 1195:Chlorarachnea 1192: 1186: 1183: 1181: 1180:Leucodictyida 1178: 1176: 1173: 1171: 1168: 1167: 1165: 1163: 1159: 1153: 1150: 1148: 1145: 1144: 1142: 1138: 1135: 1132: 1128: 1125: 1121: 1115: 1111: 1099: 1096: 1094: 1091: 1089: 1086: 1085: 1084: 1081: 1079: 1076: 1075: 1074: 1071: 1067: 1064: 1062: 1061:Stramenopiles 1059: 1057: 1054: 1052: 1049: 1045: 1042: 1041: 1040: 1037: 1036: 1035: 1032: 1030: 1027: 1024:(major groups 1022: 1020: 1017: 1015: 1012: 1010: 1007: 1004: 1000: 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258: 254: 250: 246: 245:Geinitzinacea 242: 238: 234: 230: 229:Colaniellacea 226: 221: 216: 213: 207: 204: 203: 200: 196:Globothalamea 194: 191: 190: 187: 184: 181: 180: 177: 174: 171: 170: 167: 164: 161: 158: 157: 154: 151: 148: 145: 144: 141: 138: 135: 134: 129: 124: 120: 116: 111: 104: 99: 94: 89: 84: 79: 74: 69: 64: 59: 54: 49: 35: 32: 27: 19: 2251:Permian life 2077: 2019:paraphyletic 1989:Lapot gusevi 1987: 1979: 1948:Heterogromia 1946: 1937:Involutinida 1928: 1876:Textulariida 1858:Ammodiscidae 1848:Spirillinida 1837: 1829:Tubothalamea 1818:Reticulomyxa 1816: 1811:Astrorhizida 1800:Allogromiida 1791:Monothalamea 1781:Foraminifera 1735:Polycystinea 1720:Holocanthida 1617:Vampyrellida 1582:Haplosporida 1569:Ascetosporea 1559:Reticulosida 1464:Cryomonadida 1449:Phaeogromida 1444:Phaeocalpida 1434:Phaeocystida 1416:Thecofilosea 1371:Variglissida 1366:Marimonadida 1302:Sarcomonadea 1292:Ventrifilosa 1243:Metromonadea 1227:Monadofilosa 1170:Cryptofilida 1162:Granofilosea 1147:Aquavolonida 1083:Opisthokonta 947: 902: 898: 871: 867: 845: 841: 818: 785: 781: 757:. 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Retrieved 650: 643: 623: 616: 566:(1): 15218. 563: 559: 549: 496: 492: 438: 429: 405: 401: 393:Llandoverian 390: 378: 372: 370: 353: 344:polyphyletic 340: 336: 299:Foraminifera 290: 288: 269:Tetrataxacea 237:Endothyracea 233:Earlandiacea 223: 211: 186:Foraminifera 159: 146: 31: 26: 2156:iNaturalist 2102:Wikispecies 2052:fungus-like 2028:flagellates 1891:Carterinida 1886:Robertinida 1753:Spumellaria 1748:Nassellaria 1627:Phagomyxida 1610:Proteomyxia 1587:Mikrocytida 1185:Limnofilida 1140:Skiomonadea 435:Application 426:Terminology 297:within the 241:Fusulinacea 182:Subphylum: 2236:Fusulinida 2230:Categories 2108:Fusulinida 2078:Fusulinida 1954:Komokiacea 1838:Fusulinida 1743:Collodaria 1702:Acantharea 1692:Taxopodida 1674:Radiolaria 1622:Phytomyxea 1597:Paramyxida 1592:Paradinida 1424:Phaeodarea 1394:Euglyphida 1348:Imbricatea 1320:Katabiidae 1208:Minorisida 948:Triticites 667:10 January 471:References 408:Antarctica 291:Fusulinida 212:Fusulinida 37:Fusulinida 1881:Rotaliida 1843:Miliolida 1546:Gromiidea 1538:Marimyxia 1460:Eothecia 1356:Krakenida 1273:Helkesida 1250:Metopiida 1152:Tremulida 1078:Amoebozoa 1056:Alveolata 1044:Cryptista 1019:Eukaryota 935:132061868 868:Evolution 802:129848257 728:0022-3360 590:2045-2322 523:0027-8424 445:limestone 374:Nanicella 362:Rotaliida 198: (?) 140:Eukaryota 18:Fusulinid 2135:10568606 2087:Wikidata 2023:†extinct 1942:Lagenida 1758:Orodaria 1554:Gromiida 1529:Endomyxa 1474:Matazida 1469:Ebriacea 1265:Helkesea 1236:Eoglissa 1120:Cercozoa 1073:Amorphea 1051:Rhizaria 1039:Hacrobia 1029:Excavata 1014:Bacteria 989:Rhizaria 890:23461330 853:Archived 848:, 2005, 608:29123221 541:34155110 451:See also 397:Silurian 333:Taxonomy 311:Loeblich 172:Phylum: 136:Domain: 2187:5584883 2148:3235825 2093:Q134577 2032:amoebae 1658:Retaria 1088:Animals 1009:Archaea 952:Permian 927:1306346 907:Bibcode 736:1305391 599:5680253 568:Bibcode 532:8255957 501:Bibcode 321:to the 205:Order: 192:Class: 176:Retaria 2213:592502 2200:239725 2161:348316 1122:  1118:  1066:Plants 1003:Domain 933:  925:  888:  850:Online 830:  800:  782:Facies 759:16 May 734:  726:  698:  658:  631:  606:  596:  588:  539:  529:  521:  499:(26). 412:Visean 315:Tappan 2208:WoRMS 2174:12845 2169:IRMNG 2122:7ZJHF 1093:Fungi 931:S2CID 923:JSTOR 798:S2CID 732:JSTOR 303:tests 295:order 160:Clade 147:Clade 2143:GBIF 886:PMID 828:ISBN 761:2020 724:ISSN 696:ISBN 669:2023 656:ISBN 629:ISBN 604:PMID 586:ISSN 537:PMID 519:ISSN 313:and 289:The 48:Preęž’ 2130:EoL 2117:CoL 915:doi 876:doi 790:doi 688:doi 594:PMC 576:doi 527:PMC 509:doi 497:118 166:SAR 2232:: 2210:: 2197:: 2184:: 2171:: 2158:: 2145:: 2132:: 2119:: 2104:: 2089:: 2050:; 2046:; 2042:; 2038:; 2034:; 2030:; 929:. 921:. 913:. 903:69 901:. 884:. 872:67 870:. 866:. 822:. 812:; 796:. 786:51 784:. 752:. 730:. 720:62 718:. 694:, 602:. 592:. 584:. 574:. 562:. 558:. 535:. 525:. 517:. 507:. 495:. 491:. 479:^ 447:. 422:. 368:. 162:: 149:: 98:Pg 2054:. 2017:* 1912:† 1906:† 1900:† 1836:† 1813:* 1802:* 1793:* 1304:* 1133:* 1100:) 981:e 974:t 967:v 954:. 937:. 917:: 909:: 892:. 878:: 836:. 804:. 792:: 763:. 738:. 690:: 671:. 637:. 610:. 578:: 570:: 564:7 543:. 511:: 503:: 209:† 103:N 93:K 88:J 83:T 78:P 73:C 68:D 63:S 58:O 53:ęž’ 20:)

Index

Fusulinid
Preęž’
ęž’
O
S
D
C
P
T
J
K
Pg
N

Scientific classification
Edit this classification
Eukaryota
Diaphoretickes
SAR
Retaria
Foraminifera
Globothalamea (?)
Fusulinida
Archaediscacea
Colaniellacea
Earlandiacea
Endothyracea
Fusulinacea
Geinitzinacea
Moravamminacea

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