Knowledge (XXG)

Hypacrosaurus

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called homeothermy) and therefore there should be little variation in the oxygen-isotope ratio when measured in different bones. Alternatively, the oxygen-isotope ratio differs considerably when measured throughout the body of an organism with an ectothermic (cold-blooded) physiology. Oxygen-isotope ratios calculated for
2261:—an actual life assemblage of animals. The cause of death in a volcanic ashfall is suffocation by the ash and by the gases released from volcanic eruptions. The preservation of this diverse group of dinosaurs provides researchers with a growth series, which is a sequence of growth stages from juvenile to adult. 2256:
in Montana was interpreted as a group of dinosaurs that was killed by a volcanic ashfall. This assemblage is considered autochthonous, meaning that the remains are thought to have been buried on or near the same spot where the individuals died. The variety of ages in this group supports that this was
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Examining the oxygen-isotope ratio from the bones from different parts of an extinct animal's body should indicate which thermoregulation mode an animal used during its lifetime. An endothermic (warm-blooded) animal should maintain a very similar body temperature throughout its entire body (which is
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cells tested positive to possible chemical markers of DNA, in a similar way to the emu cells, suggesting the potential preservation of this molecule. The team concluded that the find was not a product of fossil contamination, and DNA may last much longer than previously assumed.
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most likely had social functions, such as a visual signal allowing individuals to identify sex or species, and providing a resonating chamber for making noises. The crest and its associated nasal passages have also figured in the debate about
3567:; Barrett, Paul M.; Coria, Rodolfo A.; Le Loueff, Jean; Xu Xing; Zhao Xijin; Sahni, Ashok; Gomani, Elizabeth M.P.; Noto, Christopher N. (2004). "Dinosaur distribution". In Weishampel, David B.; Dodson, Peter; Osmólska Halszka (eds.). 3275:
Barrick, R. E., Showers, W. J. and Fischer, A. G. 1996. Comparison of thermoregulation of four ornithischian dinosaurs and a varanid lizard from the Cretaceous Two Medicine Formation: Evidence from oxygen isotopes. Palaios 11:
3078:"Testing Size–frequency Distributions As a Method of Ontogenetic Aging: A Life-history Assessment of Hadrosaurid Dinosaurs from the Dinosaur Park Formation of Alberta, Canada, with Implications for Hadrosaurid Paleoecology" 739:, but is more pointed along its top, not as tall, wider side to side, and has a small bony point at the rear. Unlike other lambeosaurines, the passages for the airways do not form an S-curve in the crest (at least not in 2075:
of 20 by 18.5 cm (7.9 by 7.3 in), with embryos 60 cm (24 in) long. Hatchlings were around 1.7 m (5.6 ft) long. Studies of lines of growth (i.e. lines of von Ebner) in the teeth of embryonic
2124:; rapidly growing hypacrosaurs would have had a better chance to reach a size large enough to be of defensive value, and beginning reproduction at an early age would also have been advantageous to a prey animal. 2238:
specimen inflicted by a bite from the teeth of a tyrannosaurid indicated that this, and other hadrosaurids were either preyed upon or scavenged by large theropod dinosaurs during the Late Cretaceous period.
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Behrensmeyer, A. K. 1991. Terrestrial vertebrate accumulations. In Allison P. A. and Briggs D. E. G. (Eds), Taphonomy: Releasing the Data Locked in the Fossil Record. New York: Plenum Press.
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indicates that this animal may have reached reproductive maturity at the age of 2 to 3 years, and reached full size at about 10 to 12 years old. The circumference of the
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Rogers, Raymond R. (1990). "Taphonomy of three dinosaur bone beds in the Upper Cretaceous Two Medicine Formation of northwestern Montana: evidence for drought-related mortality".
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that contained hundreds of teeth, only a relative handful of which were in use at any time. Plant material would have been cropped by its broad beak, and held in the jaws by a
735:, are 5 to 7 times the height of the body of their respective vertebrae in the back, which would have given it a tall back in profile. The skull's hollow crest is like that of 6200: 3698: 6250: 3242: 431:, it had a tall, hollow rounded crest, although not as large and straight. It is known from the remains of two species that spanned 75 to 67 million years ago, in the 539:
During this period, the remains of small hollow-crested duckbills were described as their own genera and species. The first of these that figure into the history of
6136: 3142:"Osteological and taxonomic reassessments of Sahaliyania elunchunorum (Dinosauria, Hadrosauridae) from the Upper Cretaceous Yuliangzi Formation, northeast China" 2088:. Young and embryonic individuals had deep skulls with only slight expansion in the bones that would one day form the crest. Growth was faster than that of an 631:, is known from 5 to 10 articulated skulls with some associated skeletal remains, from juvenile to adult individuals found in the Horseshoe Canyon Formation. 6235: 950:
and its close relatives was recovered in a 2022 phylogenetic analysis by Xing Hai and colleagues. Unlike other modern analyses, they found the genus
6240: 6123: 5657: 704:, and across the border in Alberta. These represent "the largest collection of baby skeletal material of any single species of hadrosaur known". 4742: 6215: 5652: 3338: 3729: 2614: 4756: 2514:
were also present. This formation was more distant from the Western Interior Seaway, higher and drier, with a more terrestrial influence.
6220: 2854:; Weishampel, David B.; Forster, Catherine A. (2004). "Hadrosauridae". In Weishampel, David B.; Dodson, Peter; Osmólska Halszka (eds.). 2396:, and are distinct from those in the formations above and below. The Horseshoe Canyon Formation is interpreted as having a significant 3127:
A revision of the Hadrosauridae (Reptilia:Ornithischia) and their evolution during the Campanian and Maastrichtian. Ph.D. dissertation
499: 3444:"First Evidence of Dinosaurian Secondary Cartilage in the Post-Hatching Skull of Hypacrosaurus stebingeri (Dinosauria, Ornithischia)" 3623: 3584: 3263: 2912: 2871: 2703: 530:). Brown described these remains, in combination with other postcranial bones, in 1913 as a new genus that he considered to be like 5639: 2000:(warm-blooded animals) who could lose a great deal of water while breathing because they breathe more often than comparably sized 6230: 6225: 830: 6018: 3692: 3192:
Ruben, J. A.; Hillenius, W.J.; Geist, N.R.; Leitch, A.; Jones, T.D.; Currie, P.J.; Horner, J.R.; Espe, G., III (August 1996).
6041: 6028: 3288:"Dinosaur incubation periods directly determined from growth-line counts in embryonic teeth show reptilian-grade development" 2539: 1329: 547:, based on a skull and assorted limb bones, vertebrae, and pelvic bones from the Horseshoe Canyon Formation. Not long after, 6245: 6033: 2903:(Ornithischia, Lambeosauridae) from Montana and Alberta". In Carpenter, Kenneth; Hirsch, Karl F.; Horner John R. (eds.). 2896: 2851: 2112:
at postulated reproductive maturity was about 40% that of its circumference at full size. The postulated growth rate of
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and function in smell. Respiratory turbinates function to prevent water loss through evaporation and are found only in
6210: 5663: 3057:(1981). "The nasal cavity of lambeosaurine hadrosaurids (Reptilia:Ornithischia): comparative anatomy and homologies". 1563: 601:
specimens studied by Dodson, and for good reason: it was much more like a species that would not be named until 1994,
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Martin, A.J. (2006). Introduction to the Study of Dinosaurs. Second Edition. Oxford, Blackwell Publishing. 560 pp.
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was named for a variety of remains, including hatchlings with associated eggs and nests, found near the top of the
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was not warm-blooded, which was based on the absence of nasal turbinates (see Crest functions subsection, above).
1946: 843:, and has been recognized as such since the description of its skull. Within the Lambeosaurinae, it is closest to 3722: 1643: 480: 2170:
analyses on skull and limb bones of nestling individuals of the specimen MOR 548, a large nesting ground in the
917:, this was rejected in a later, more comprehensive reanalysis of lambeosaurines, which found the two species of 3829: 2097: 1744: 1696: 3005: 3503:"Evidence of proteins, chromosomes and chemical markers of DNA in exceptionally preserved dinosaur cartilage" 3129:. Washington, D.C.: Graduate School of Arts and Sciences of The George Washington University. pp. 1–272. 743:). The animal is estimated to have been around 9.1 meters (30 feet) long, and to have weighed up to 4.0  6205: 2401: 1966: 1660: 1401: 1384: 701: 635:
is known from an unknown but substantial number of individuals, with an age range of embryos to adults. The
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was a homeotherm, and likely was endothermic. This is in contrast to the Ruben et al. (1996) finding that
697: 552: 1938:-like organ. Its feeding range would have extended from the ground to about 4 m (13 ft) above. 6162: 5998: 4554: 3856: 3236: 2140: 2159: 613: 5418: 3715: 3648: 3455: 3302: 3208: 3089: 2951: 2646: 388: 124: 5646: 3564: 3501:
Bailleul, A. M.; Zheng, W.; Horner, J. R.; Hall, B. K.; Holliday, C. M.; Schweitzer, M. H. (2020).
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Xing, Hai; Gu, Wei; Hai, Shulin; Yu, Tingxiang; Han, Dong; Zhang, Yuguang; Zhang, Shujun (2022).
3107: 3032: 3009: 2967: 2899:; Currie, Phillip J. (1994). "Embryonic and neonatal morphology and ontogeny of a new species of 2820: 2722: 2662: 2348: 352: 131: 2057: 751:). As with most duckbills, its skeleton is otherwise not particularly remarkable, although some 551:
and Nelda Wright identified an American Museum of Natural History skeleton (AMNH 5461) from the
6167: 3615: 3576: 2863: 2190:-like structures were found. Several of these structures were preserved in the final stages of 506:
skeleton consisting of several vertebrae and a partial pelvis (AMNH 5204), came from along the
6149: 6141: 5777: 5516: 5437: 5397: 4281: 4167: 4113: 4084: 3619: 3580: 3534: 3483: 3424: 3330: 3259: 2908: 2867: 2699: 2610: 2458: 2393: 2357: 2008:. Ruben and others in 1996 concluded that respiratory turbinates were probably not present in 112: 6177: 6154: 5918: 5911: 5721: 5714: 5243: 5154: 4992: 4893: 4879: 4593: 4561: 4196: 4105: 4019: 3954: 3656: 3568: 3524: 3514: 3473: 3463: 3414: 3406: 3375: 3320: 3310: 3216: 3153: 3097: 3001: 2992:, a crested hadrosaurid dinosaur (Ornithischia) from the Dinosaur Park Formation, Alberta". 2959: 2855: 2812: 2687: 2654: 2581: 2531: 2343: 2214: 1931: 693: 527: 2759:, a new genus and species of trachodont dinosaur from the Edmonton Cretaceous of Alberta". 6003: 5798: 5749: 5612: 5543: 5446: 5266: 4985: 4971: 4950: 4568: 4441: 4208: 3976: 3814: 3702: 2630: 2384: 2380: 2337: 2325: 2269: 2081: 1314: 728: 557: 432: 30: 2688: 2565: 3652: 3459: 3306: 3212: 3178:
Chinsamy, Anusuya; and Hillenius, Willem J. (2004). "Physiology of nonavian dinosaurs".
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of other established lambeosaurines. Although he was mostly concerned with the earlier,
6013: 6008: 5946: 5883: 5756: 5605: 5568: 5550: 5495: 5127: 4943: 4936: 4865: 4787: 4774: 4702: 4690: 4659: 4634: 4498: 4420: 4381: 4367: 4244: 4155: 4077: 4055: 4048: 3789: 3608: 3529: 3502: 3478: 3443: 3419: 3394: 3325: 2752: 2683: 2498: 2486: 2442: 2437: 2310: 2306: 2297: 2199: 2183: 1166: 1096: 724: 507: 368: 255: 213: 3193: 2932: 536:. No skull was known at this time, but two skulls were soon discovered and described. 443:, United States, and is the latest hollow-crested duckbill known from good remains in 6194: 5953: 5897: 5876: 5847: 5763: 5700: 5598: 5591: 5577: 5487: 5467: 5460: 5357: 5300: 5284: 5259: 5232: 5225: 5218: 5189: 5140: 5102: 5067: 5048: 5020: 4964: 4627: 4620: 4575: 4505: 4374: 4233: 4218: 4180: 4091: 3984: 3947: 3940: 3569: 3165: 3111: 2856: 2633:; Burns, M. E.; Sissons, R. L. (2009). "A redescription of the ankylosaurid dinosaur 2492: 2446: 2388: 2367: 2331: 2286: 2072: 1628: 1441: 1360: 1354: 1303: 1230: 1197: 1180: 1074: 1026: 897: 657: 523: 452: 444: 423: 406: 404:(meaning "near the highest lizard" , because it was almost but not quite as large as 268: 242: 70: 3228: 3013: 2971: 2666: 5861: 5837: 5825: 5791: 5784: 5735: 5728: 5534: 5478: 5453: 5425: 5314: 5116: 5081: 5027: 4957: 4929: 4815: 4808: 4648: 4613: 4540: 4533: 4455: 4396: 4351: 4253: 4127: 4120: 4069: 4033: 3991: 3841: 3780: 3603: 3395:"Relative growth rates of predator and prey dinosaurs reflect effects of predation" 3379: 2800: 2605:
Creisler, Benjamin S. (2007). "Deciphering duckbills". In Carpenter Kenneth (ed.).
2561: 2376: 2361: 2315: 2301: 2195: 2167: 2016: 2010: 1997: 1524: 1250: 1106: 1002: 887:, are regarded as the "helmeted" or "hooded" branch of the lambeosaurines, and the 784: 775: 628: 566: 495: 297: 288: 197: 3220: 3157: 811: 712: 3684: 3468: 6108: 6050: 5939: 5904: 5742: 5584: 5557: 5509: 5502: 5335: 5321: 5293: 5273: 5252: 5196: 5175: 5168: 5161: 5095: 5088: 5041: 5006: 4978: 4914: 4907: 4676: 4526: 4512: 4484: 4462: 4404: 4267: 4098: 4062: 4026: 3922: 3909: 3868: 3738: 2937:(Dinosauria; Ornithopoda): anatomy and systematic position within Hadrosauridae" 2803:(1975). "Taxonomic implications of relative growth in lambeosaurine dinosaurs". 2510: 2504: 2474: 2452: 2405: 2392:. The dinosaurs from this formation are sometimes known as Edmontonian, after a 2292: 2258: 2187: 2179: 2148: 2101: 1923: 1900: 1727: 1721: 1510: 1281: 1264: 1137: 1057: 952: 856: 840: 744: 731:
and the form of its crest. The neural spines, which project from the top of the
593:. This idea has become accepted, although not formally tested. The Two Medicine 532: 415: 376: 341: 229: 45: 800: 5992: 5967: 5960: 5932: 5925: 5816: 5805: 5691: 5390: 5370: 5328: 5203: 5055: 4922: 4872: 4837: 4830: 4801: 4641: 4519: 4491: 4360: 4333: 4308: 4148: 4141: 4134: 4012: 3366:
as hypothesized from lines of arrested growth and whole femur circumference".
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excluding other known hadrosaur species may be incorrect, as noted in Suzuki
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would turn out to be composed of juvenile individuals of the contemporaneous
5974: 5890: 5869: 5707: 5404: 5342: 4851: 4823: 4683: 4547: 4477: 4448: 4274: 4260: 4003: 3315: 2609:. Bloomington and Indianapolis: Indiana University Press. pp. 185–210. 2425: 2147:
MOR 548 supraoccipital with microscopic magnification showing the preserved
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they form is sometimes informally designated Lambeosaurini. Although Suzuki
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growth, for several years, based on the amount of bone growth seen between
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Bailleul, A. M.; Hall, B. K.; Horner, J. R. (2012). Dodson, Peter (ed.).
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Cooper, Lisa N.; Lee, Andrew H.; Taper, Mark L.; Horner, John R. (2008).
2735:
Canada Department of Mines Geological Survey Bulletin (Geological Series)
2658: 2372: 2206: 1985: 1120: 978: 732: 487: 418: 181: 168: 85: 80: 65: 60: 50: 2988:
Evans, David C.; Reisz, Robert R. (2007). "Anatomy and relationships of
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long incubation times, with a minimum incubation time of 171.4 days for
6128: 5147: 4582: 3668: 3102: 3077: 2824: 2637:
Parks, 1924 (Ornithischia: Ankylosauria) and a revision of the genus".
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outpaces those of tyrannosaurids (predators of hypacrosaurs) such as
2093: 1993: 752: 158: 6070: 3660: 2816: 2162:
and colleagues reported cartilage traces on a hatchling specimen of
3575:(2nd ed.). Berkeley: University of California Press. pp.  3286:
Erickson, G.M.; Zelenitsky, D.K.; Kay, D.I.; Norrell, M.A. (2017).
2862:(2nd ed.). Berkeley: University of California Press. pp.  2028:, thus there was no evidence that those animals were warm-blooded. 4858: 4388: 2397: 2268: 2139: 2109: 2056: 1945: 1935: 1927: 1919: 922: 888: 821: 810: 799: 723:
is most easily distinguished from other hollow-crested duckbills (
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that covered the midsection of North America through much of the
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Suzuki, Daisuke; Weishampel, David B.; Minoura, Nachio (2004).
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shared the Horseshoe Canyon Formation with fellow hadrosaurids
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showed that it was more likely that the "cheneosaurs" were the
3707: 2423:, was also populated by another well-known nesting hadrosaur, 2218: 2203: 2152: 447:. It was an obscure genus until the discovery in the 1990s of 2435:, which is also known from nesting traces. The tyrannosaurid 2907:. Cambridge: Cambridge University Press. pp. 312–336. 2694:. Jefferson, North Carolina: McFarland & Co. pp.  2041:
suggesting that the ratios varied little, indicating that
3362:
Cooper, Lisa N.; Horner, John R. (1999). "Growth rate of
3006:
10.1671/0272-4634(2007)27[373:AAROLM]2.0.CO;2
3194:"The metabolic status of some Late Cretaceous dinosaurs" 3076:
Wosik, M.; Chiba, K.; Therrien, F.; Evans, D.C. (2020).
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details are distinctive. Like other duckbills, it was a
2128:
has been found in the skull of a hatchling specimen of
681:. This was rejected by Evans and Reisz (2007), though. 2607:
Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs
2104:
in trees). Research by Lisa Cooper and colleagues on
1922:. Its skull permitted a grinding motion analogous to 871:, and Michael K. Brett-Surman (1989) suggesting that 3254: 3252: 960:, indicating it could be a potential third species. 565:
were accepted as valid genera until the 1970s, when
327: 304: 6083: 5984: 5835: 5815: 5690: 5567: 5533: 5477: 5436: 5380: 5356: 5283: 5242: 5126: 5065: 4785: 4701: 4658: 4592: 4418: 4350: 4306: 4243: 4207: 4166: 4002: 3965: 3920: 3867: 3840: 3813: 2419:The slightly older Two Medicine Formation, home to 774:, are not differentiated in the typical method, of 716:
Size comparison between the two species and a human
3614:. Princeton: Princeton University Press. pp.  3607: 3043:. Geological Society of America. pp. 206–208. 2574:Bulletin of the American Museum of Natural History 2221:chondrocytes, and these tied up to DNA fragments. 1980:that come in two types, with two functions. Nasal 782:was described as transitional between the earlier 2198:. Bailleul and colleagues isolated some of these 518:, Canada, from rocks of what is now known as the 2234:The discovery of tooth marks in the fibula of a 3295:Proceedings of the National Academy of Sciences 2151:-like structures, in comparison to those of an 2004:(cold-blooded animals) to support their higher 879:are the same genus. These genera, particularly 3696:in The Natural History Museum's Dino Directory 3039:. Geological Society of America Special Paper 3723: 2729:, with a description of the type specimen of 8: 3559: 3557: 3241:: CS1 maint: multiple names: authors list ( 1930:were continually replacing and packed into 6201:Late Cretaceous dinosaurs of North America 6071: 5687: 5676: 5362: 5132: 5073: 4793: 4782: 4769: 4598: 4426: 4318: 4314: 4172: 3932: 3928: 3917: 3904: 3846: 3819: 3810: 3746: 3730: 3716: 3708: 2570:, from the Edmonton Cretaceous of Alberta" 2416:may have preferred to stay more landward. 2194:, with some preserving putative traces of 2186:bone, and upon microscopic magnification, 597:, meanwhile, was not quite like the other 111: 20: 3528: 3518: 3477: 3467: 3418: 3324: 3314: 3101: 2926: 2924: 6251:Multispecific non-avian dinosaur genera 3610:The Horned Dinosaurs: A Natural History 3598: 3596: 3037:Hadrosaurian Dinosaurs of North America 3027: 3025: 3023: 2983: 2981: 2891: 2889: 2887: 2885: 2883: 2846: 2844: 2842: 2840: 2838: 2836: 2834: 2717: 2715: 2678: 2676: 2550: 3234: 2556: 2554: 2252:The large, monospecific assemblage of 7: 3688:, from the Canadian Museum of Nature 1969:, specifically in discussions about 1984:turbinates are found in all living 1712: 1688: 1635: 1623: 1555: 1531: 1502: 1495: 1471: 1447: 1376: 1345: 1321: 1309: 1256: 1246: 1222: 1172: 1162: 1112: 1102: 1049: 1042: 1018: 994: 970: 963: 901:found a close relationship between 3399:Proceedings of the Royal Society B 3368:Journal of Vertebrate Paleontology 3146:Journal of Vertebrate Paleontology 2994:Journal of Vertebrate Paleontology 2944:Journal of Vertebrate Paleontology 2639:Journal of Vertebrate Paleontology 2371:, a variety of poorly known small 946:illustrating the relationships of 500:American Museum of Natural History 14: 6236:Campanian genus first appearances 3125:Brett-Surman, Michael K. (1989). 2400:influence, due to an encroaching 5662: 5656: 5651: 5645: 5638: 4755: 4748: 4741: 2524: 831:Philip J. Currie Dinosaur Museum 135: 6241:Maastrichtian genus extinctions 6019:List of informal dinosaur names 3344:from the original on 2017-01-10 6029:Timeline of hadrosaur research 3380:10.1080/02724634.1999.10011202 2781:(1920). "Canadian dinosaurs". 2540:Timeline of hadrosaur research 1330:Parasaurolophus cyrtocristatus 1: 6216:Fossil taxa described in 1913 3221:10.1126/science.273.5279.1204 3158:10.1080/02724634.2021.2085111 1976:Turbinates are thin bones or 939:being the closest relatives. 3469:10.1371/journal.pone.0036112 2566:"A new trachodont dinosaur, 2429:, as well as the troodontid 669:was more closely related to 555:of Montana as a specimen of 3035:; Wright, Nelda E. (1942). 2990:Lambeosaurus magnicristatus 2690:Dinosaurs: The Encyclopedia 2202:in order to be tested with 1564:Lambeosaurus magnicristatus 6267: 6221:Taxa named by Barnum Brown 2935:Nipponosaurus sachaliensis 2635:Dyoplosaurus acutosquameus 1540:Lambeosaurus clavinitialis 520:Horseshoe Canyon Formation 494:were collected in 1910 by 5686: 5675: 5636: 5365: 5135: 5076: 4796: 4781: 4768: 4739: 4601: 4429: 4321: 4317: 4175: 3935: 3931: 3916: 3903: 3849: 3822: 3809: 3745: 2686:(1997). "Hypacrosaurus". 2178:, and the results showed 1741: 1717: 1710: 1693: 1686: 1657: 1644:Corythosaurus intermedius 1640: 1633: 1621: 1577: 1560: 1553: 1536: 1529: 1507: 1500: 1493: 1476: 1469: 1452: 1445: 1398: 1381: 1374: 1350: 1343: 1326: 1319: 1307: 1278: 1261: 1254: 1244: 1227: 1220: 1194: 1177: 1170: 1160: 1134: 1117: 1110: 1100: 1071: 1054: 1047: 1040: 1023: 1016: 999: 992: 975: 968: 766:. The two known species, 502:. The remains, a partial 481:Canadian Museum of Nature 421:similar in appearance to 358: 351: 324: 319: 303: 296: 132:Scientific classification 130: 119: 110: 23: 16:Extinct genus of reptiles 3364:Hypacrosaurus stebingeri 2905:Dinosaur Eggs and Babies 2757:Cheneosaurus tolmanensis 2723:Gilmore, Charles Whitney 2254:Hypacrosaurus stebingeri 2098:lines of arrested growth 2069:Hypacrosaurus stebingeri 1745:Hypacrosaurus stebingeri 1697:Hypacrosaurus altispinus 907:Hypacrosaurus stebingeri 895:s 2004 redescription of 863:is transitional between 686:Hypacrosaurus stebingeri 655:s 2004 redescription of 591:Hypacrosaurus altispinus 545:Cheneosaurus tolmanensis 309:Hypacrosaurus altispinus 6231:Paleontology in Montana 6226:Paleontology in Alberta 3507:National Science Review 3316:10.1073/pnas.1613716114 3059:Journal of Paleontology 2779:Matthew, William Diller 2490:, and horned dinosaurs 2402:Western Interior Seaway 2071:laid roughly spherical 1661:Corythosaurus casuarius 1402:Parasaurolophus walkeri 1385:Parasaurolophus tubicen 859:(1994) suggesting that 702:Glacier County, Montana 575:Dinosaur Park Formation 3411:10.1098/rspb.2008.0912 2725:(1924). "On the genus 2278: 2172:Two Medicine Formation 2155: 2065: 1957: 1918:, eating a variety of 833: 819: 808: 776:unique characteristics 717: 698:Two Medicine Formation 621: 553:Two Medicine Formation 483: 6163:Paleobiology Database 5999:Basturs Poble bonebed 2761:The Ottawa Naturalist 2272: 2166:. The team performed 2143: 2060: 1949: 825: 814: 803: 715: 663:phylogenetic analysis 616: 474: 467:Discovery and history 6246:Ornithischian genera 6021:(contains hadrosaur 5419:Probrachylophosaurus 3565:Weishampel, David B. 3055:Weishampel, David B. 2659:10.1671/039.029.0405 2462:, armored dinosaurs 2217:. They also exposed 1960:The hollow crest of 839:was a lambeosaurine 804:Life restoration of 585:, he suggested that 125:Royal Tyrrell Museum 3653:1990Palai...5..394R 3460:2012PLoSO...736112B 3405:(1651): 2609–2615. 3307:2017PNAS..114..540E 3213:1996Sci...273.1204R 3207:(5279): 1204–1207. 3094:2020Pbio...46..379W 3033:Lull, Richard Swann 2956:2004JVPal..24..145S 2731:Lambeosaurus lambei 2651:2009JVPal..29.1117A 2323:, horned dinosaurs 2182:cartilage within a 2126:Secondary cartilage 1967:dinosaur endothermy 1581:Lambeosaurus lambei 524:early Maastrichtian 6211:Maastrichtian life 5771:Ornithopodichnites 5382:Brachylophosaurini 3967:Hypsilophodontidae 3701:2007-07-13 at the 3520:10.1093/nsr/nwz206 3103:10.1017/pab.2020.2 2805:Systematic Zoology 2753:Lambe, Lawrence M. 2349:pachycephalosaurid 2279: 2156: 2092:and comparable to 2066: 1958: 1907:would have been a 909:, indicating that 834: 820: 809: 718: 622: 561:. These and other 549:Richard Swann Lull 484: 475:Skeletal mount of 6188: 6187: 6150:Open Tree of Life 6077:Taxon identifiers 6068: 6067: 6064: 6063: 6060: 6059: 5778:Shiraminesauropus 5671: 5670: 5634: 5633: 5630: 5629: 5626: 5625: 5622: 5621: 5517:Naashoibitosaurus 5438:Austrokritosauria 5398:Brachylophosaurus 5352: 5351: 4764: 4763: 4737: 4736: 4733: 4732: 4729: 4728: 4725: 4724: 4721: 4720: 4414: 4413: 4302: 4301: 4298: 4297: 4282:Transylvanosaurus 4168:Rhabdodontomorpha 4114:Notohypsilophodon 4085:Macrogryphosaurus 3899: 3898: 3895: 3894: 3891: 3890: 3887: 3886: 3374:(3, Suppl): 35A. 2616:978-0-253-34817-3 2478:, hypsilophodont 2459:Saurornitholestes 2450:, dromaeosaurids 2160:Alida M. Bailleul 2082:plesiomorphically 1891: 1890: 1882: 1881: 1873: 1872: 1864: 1863: 1855: 1854: 1846: 1845: 1837: 1836: 1828: 1827: 1819: 1818: 1810: 1809: 1801: 1800: 1792: 1791: 1783: 1782: 1774: 1773: 1765: 1764: 1756: 1755: 1672: 1671: 1610: 1609: 1601: 1600: 1592: 1591: 1431: 1430: 1422: 1421: 1413: 1412: 1293: 1292: 1209: 1208: 1149: 1148: 1086: 1085: 397: 396: 391: 372: 345: 292: 6258: 6181: 6180: 6171: 6170: 6158: 6157: 6145: 6144: 6132: 6131: 6119: 6118: 6117: 6104: 6103: 6102: 6072: 6045:mummy S.M. R4036 5919:Microhadrosaurus 5912:Mandschurosaurus 5722:Camptosaurichnus 5715:Bonaparteichnium 5688: 5681:Related articles 5677: 5666: 5660: 5655: 5650: 5649: 5642: 5363: 5244:Parasaurolophini 5155:Angulomastacator 5133: 5074: 4993:Shuangmiaosaurus 4894:Huehuecanauhtlus 4880:Gongpoquansaurus 4794: 4783: 4770: 4759: 4753: 4752: 4745: 4599: 4594:Hadrosauriformes 4562:Ratchasimasaurus 4427: 4319: 4315: 4197:Muttaburrasaurus 4173: 4106:Muttaburrasaurus 4020:Atlascopcosaurus 3955:Thescelosauridae 3933: 3929: 3918: 3905: 3847: 3820: 3811: 3804: 3803: 3747: 3732: 3725: 3718: 3709: 3673: 3672: 3636: 3630: 3629: 3613: 3600: 3591: 3590: 3574: 3561: 3552: 3549: 3543: 3542: 3532: 3522: 3498: 3492: 3491: 3481: 3471: 3439: 3433: 3432: 3422: 3390: 3384: 3383: 3359: 3353: 3352: 3350: 3349: 3343: 3328: 3318: 3292: 3283: 3277: 3273: 3267: 3256: 3247: 3246: 3240: 3232: 3198: 3189: 3183: 3176: 3170: 3169: 3137: 3131: 3130: 3122: 3116: 3115: 3105: 3073: 3067: 3066: 3051: 3045: 3044: 3029: 3018: 3017: 2985: 2976: 2975: 2964:10.1671/A1034-11 2941: 2928: 2919: 2918: 2893: 2878: 2877: 2861: 2848: 2829: 2828: 2797: 2791: 2790: 2775: 2769: 2768: 2749: 2743: 2742: 2719: 2710: 2709: 2693: 2680: 2671: 2670: 2645:(4): 1117–1135. 2627: 2621: 2620: 2602: 2596: 2595: 2593: 2592: 2558: 2534: 2532:Dinosaurs portal 2529: 2528: 2527: 2344:Pachyrhinosaurus 2053:Nests and growth 2032:Thermoregulation 1971:nasal turbinates 1932:dental batteries 1713: 1689: 1636: 1624: 1556: 1532: 1503: 1496: 1472: 1448: 1377: 1346: 1322: 1310: 1257: 1247: 1223: 1173: 1163: 1113: 1103: 1050: 1043: 1019: 995: 971: 964: 694:Upper Cretaceous 684:The new species 528:Upper Cretaceous 382: 367: 336: 329: 306: 287: 280: 267: 254: 241: 228: 212: 196: 140: 139: 115: 105: 42: 29:Temporal range: 21: 6266: 6265: 6261: 6260: 6259: 6257: 6256: 6255: 6191: 6190: 6189: 6184: 6176: 6174: 6166: 6161: 6153: 6148: 6140: 6135: 6127: 6122: 6113: 6112: 6107: 6098: 6097: 6092: 6079: 6069: 6056: 6037:mummy AMNH 5060 6004:Dakota (fossil) 5980: 5831: 5811: 5799:Tetrapodosaurus 5750:Hadrosaurichnus 5682: 5667: 5644: 5618: 5613:Shantungosaurus 5563: 5544:Augustynolophus 5529: 5473: 5447:Bonapartesaurus 5432: 5376: 5348: 5279: 5267:Parasaurolophus 5238: 5122: 5070: 5061: 4986:Qianjiangsaurus 4972:Probactrosaurus 4951:Penelopognathus 4790: 4777: 4760: 4747: 4717: 4697: 4654: 4588: 4569:Riabininohadros 4442:Bayannurosaurus 4423: 4410: 4346: 4311: 4294: 4239: 4209:Tenontosauridae 4203: 4162: 3998: 3977:Gideonmantellia 3961: 3925: 3912: 3883: 3863: 3836: 3830:Avemetatarsalia 3815:Avemetatarsalia 3805: 3752: 3751: 3741: 3736: 3703:Wayback Machine 3681: 3676: 3661:10.2307/3514834 3638: 3637: 3633: 3626: 3602: 3601: 3594: 3587: 3563: 3562: 3555: 3550: 3546: 3500: 3499: 3495: 3441: 3440: 3436: 3392: 3391: 3387: 3361: 3360: 3356: 3347: 3345: 3341: 3290: 3285: 3284: 3280: 3274: 3270: 3257: 3250: 3233: 3196: 3191: 3190: 3186: 3182:, 2nd. 643-659. 3177: 3173: 3152:(6): e2085111. 3139: 3138: 3134: 3124: 3123: 3119: 3075: 3074: 3070: 3065:(5): 1046–1057. 3053: 3052: 3048: 3031: 3030: 3021: 2987: 2986: 2979: 2939: 2930: 2929: 2922: 2915: 2897:Horner, John R. 2895: 2894: 2881: 2874: 2852:Horner, John R. 2850: 2849: 2832: 2817:10.2307/2412696 2799: 2798: 2794: 2783:Natural History 2777: 2776: 2772: 2751: 2750: 2746: 2721: 2720: 2713: 2706: 2684:Glut, Donald F. 2682: 2681: 2674: 2629: 2628: 2624: 2617: 2604: 2603: 2599: 2590: 2588: 2580:(20): 395–406. 2560: 2559: 2552: 2548: 2530: 2525: 2523: 2520: 2394:land mammal age 2338:Arrhinoceratops 2326:Montanoceratops 2273:Restoration of 2267: 2250: 2245: 2232: 2138: 2055: 2034: 1944: 1942:Crest functions 1897: 1892: 1883: 1874: 1865: 1856: 1847: 1838: 1829: 1820: 1811: 1802: 1793: 1784: 1775: 1766: 1757: 1673: 1611: 1602: 1593: 1432: 1423: 1414: 1315:Parasaurolophus 1294: 1210: 1150: 1087: 798: 710: 611: 599:Procheneosaurus 595:Procheneosaurus 558:Procheneosaurus 469: 433:Late Cretaceous 383:Prieto-Márquez 381: 366: 335: 315: 312: 286: 278: 265: 252: 239: 226: 210: 194: 134: 106: 104: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 53: 48: 37: 36: 34: 31:Late Cretaceous 27: 17: 12: 11: 5: 6264: 6262: 6254: 6253: 6248: 6243: 6238: 6233: 6228: 6223: 6218: 6213: 6208: 6206:Lambeosaurines 6203: 6193: 6192: 6186: 6185: 6183: 6182: 6172: 6159: 6146: 6133: 6120: 6105: 6089: 6087: 6081: 6080: 6075: 6066: 6065: 6062: 6061: 6058: 6057: 6055: 6054: 6047: 6039: 6031: 6026: 6016: 6014:Hypsilophodont 6011: 6009:Hadrosaur diet 6006: 6001: 5996: 5988: 5986: 5985:Other articles 5982: 5981: 5979: 5978: 5971: 5964: 5957: 5950: 5947:Stephanosaurus 5943: 5936: 5929: 5922: 5915: 5908: 5901: 5894: 5887: 5884:Fulgurotherium 5880: 5873: 5866: 5858: 5851: 5843: 5841: 5833: 5832: 5830: 5829: 5821: 5819: 5813: 5812: 5810: 5809: 5802: 5795: 5788: 5781: 5774: 5767: 5760: 5757:Iguanodontipus 5753: 5746: 5739: 5732: 5725: 5718: 5711: 5704: 5696: 5694: 5684: 5683: 5680: 5673: 5672: 5669: 5668: 5637: 5635: 5632: 5631: 5628: 5627: 5624: 5623: 5620: 5619: 5617: 5616: 5609: 5606:Laiyangosaurus 5602: 5595: 5588: 5581: 5573: 5571: 5569:Edmontosaurini 5565: 5564: 5562: 5561: 5554: 5551:Prosaurolophus 5547: 5539: 5537: 5531: 5530: 5528: 5527: 5520: 5513: 5506: 5499: 5496:Coahuilasaurus 5492: 5483: 5481: 5475: 5474: 5472: 5471: 5464: 5457: 5450: 5442: 5440: 5434: 5433: 5431: 5430: 5422: 5415: 5408: 5401: 5394: 5386: 5384: 5378: 5377: 5375: 5374: 5366: 5360: 5354: 5353: 5350: 5349: 5347: 5346: 5339: 5332: 5325: 5318: 5311: 5304: 5297: 5289: 5287: 5281: 5280: 5278: 5277: 5270: 5263: 5256: 5248: 5246: 5240: 5239: 5237: 5236: 5229: 5222: 5215: 5208: 5200: 5193: 5186: 5179: 5172: 5165: 5158: 5151: 5144: 5136: 5130: 5128:Lambeosaurinae 5124: 5123: 5121: 5120: 5113: 5106: 5099: 5092: 5085: 5077: 5071: 5066: 5063: 5062: 5060: 5059: 5052: 5045: 5038: 5031: 5024: 5017: 5010: 5003: 4996: 4989: 4982: 4975: 4968: 4961: 4954: 4947: 4944:Nanyangosaurus 4940: 4937:Nanningosaurus 4933: 4926: 4919: 4911: 4904: 4897: 4890: 4883: 4876: 4869: 4866:Gilmoreosaurus 4862: 4855: 4848: 4841: 4834: 4827: 4820: 4812: 4805: 4797: 4791: 4788:Hadrosauroidea 4786: 4779: 4778: 4775:Hadrosauroidea 4773: 4766: 4765: 4762: 4761: 4740: 4738: 4735: 4734: 4731: 4730: 4727: 4726: 4723: 4722: 4719: 4718: 4716: 4715: 4714: 4713: 4707: 4705: 4703:Hadrosauroidea 4699: 4698: 4696: 4695: 4691:Mantellisaurus 4687: 4680: 4673: 4664: 4662: 4660:Iguanodontidae 4656: 4655: 4653: 4652: 4645: 4638: 4635:Mantellisaurus 4631: 4624: 4617: 4610: 4602: 4596: 4590: 4589: 4587: 4586: 4579: 4572: 4565: 4558: 4551: 4544: 4537: 4530: 4523: 4516: 4509: 4502: 4499:Iguanacolossus 4495: 4488: 4481: 4474: 4466: 4459: 4452: 4445: 4438: 4430: 4424: 4421:Ankylopollexia 4419: 4416: 4415: 4412: 4411: 4409: 4408: 4401: 4393: 4385: 4382:Eousdryosaurus 4378: 4371: 4368:Dysalotosaurus 4364: 4356: 4354: 4348: 4347: 4345: 4344: 4337: 4330: 4322: 4312: 4307: 4304: 4303: 4300: 4299: 4296: 4295: 4293: 4292: 4285: 4278: 4271: 4264: 4257: 4249: 4247: 4245:Rhabdodontidae 4241: 4240: 4238: 4237: 4230: 4223: 4214: 4212: 4205: 4204: 4202: 4201: 4193: 4185: 4176: 4170: 4164: 4163: 4161: 4160: 4156:Weewarrasaurus 4152: 4145: 4138: 4131: 4124: 4117: 4110: 4102: 4095: 4088: 4081: 4078:Leaellynasaura 4074: 4066: 4059: 4056:Gasparinisaura 4052: 4049:Galleonosaurus 4045: 4037: 4030: 4023: 4016: 4008: 4006: 4000: 3999: 3997: 3996: 3988: 3981: 3972: 3970: 3963: 3962: 3960: 3959: 3951: 3944: 3936: 3926: 3921: 3914: 3913: 3908: 3901: 3900: 3897: 3896: 3893: 3892: 3889: 3888: 3885: 3884: 3882: 3881: 3880: 3879: 3873: 3871: 3865: 3864: 3862: 3861: 3860: 3859: 3850: 3844: 3838: 3837: 3835: 3834: 3833: 3832: 3823: 3817: 3807: 3806: 3802: 3801: 3792: 3790:Neornithischia 3783: 3774: 3765: 3759: 3750: 3743: 3742: 3737: 3735: 3734: 3727: 3720: 3712: 3706: 3705: 3690: 3680: 3679:External links 3677: 3675: 3674: 3647:(5): 394–413. 3631: 3624: 3592: 3585: 3571:The Dinosauria 3553: 3544: 3513:(4): 815−822. 3493: 3434: 3385: 3354: 3301:(3): 540–545. 3278: 3268: 3248: 3184: 3180:The Dinosauria 3171: 3132: 3117: 3088:(3): 379–404. 3068: 3046: 3019: 3000:(2): 373–393. 2977: 2950:(1): 145–164. 2920: 2913: 2879: 2872: 2858:The Dinosauria 2830: 2792: 2770: 2767:(10): 117–123. 2744: 2727:Stephanosaurus 2711: 2704: 2672: 2622: 2615: 2597: 2549: 2547: 2544: 2543: 2542: 2536: 2535: 2519: 2516: 2499:Brachyceratops 2487:Prosaurolophus 2438:Daspletosaurus 2381:dromaeosaurids 2358:ostrich-mimics 2311:Anodontosaurus 2298:hypsilophodont 2277:in environment 2266: 2263: 2249: 2246: 2244: 2241: 2231: 2230:Paleopathology 2228: 2196:celular nuclei 2184:supraoccipital 2174:attributed to 2137: 2134: 2100:(analogous to 2054: 2051: 2033: 2030: 1943: 1940: 1896: 1893: 1889: 1888: 1885: 1884: 1880: 1879: 1876: 1875: 1871: 1870: 1867: 1866: 1862: 1861: 1858: 1857: 1853: 1852: 1849: 1848: 1844: 1843: 1840: 1839: 1835: 1834: 1831: 1830: 1826: 1825: 1822: 1821: 1817: 1816: 1813: 1812: 1808: 1807: 1804: 1803: 1799: 1798: 1795: 1794: 1790: 1789: 1786: 1785: 1781: 1780: 1777: 1776: 1772: 1771: 1768: 1767: 1763: 1762: 1759: 1758: 1754: 1753: 1750: 1749: 1740: 1737: 1736: 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1167:Tsintaosaurini 1161: 1159: 1156: 1155: 1152: 1151: 1147: 1146: 1143: 1142: 1133: 1130: 1129: 1126: 1125: 1116: 1111: 1109: 1101: 1099: 1097:Lambeosaurinae 1093: 1092: 1089: 1088: 1084: 1083: 1080: 1079: 1070: 1067: 1066: 1063: 1062: 1053: 1048: 1046: 1041: 1039: 1036: 1035: 1032: 1031: 1022: 1017: 1015: 1012: 1011: 1008: 1007: 998: 993: 991: 988: 987: 984: 983: 974: 969: 967: 962: 942:The following 797: 796:Classification 794: 727:) by its tall 725:lambeosaurines 709: 706: 690:late Campanian 620:holotype skull 610: 607: 508:Red Deer River 468: 465: 439:, Canada, and 395: 394: 393: 392: 373: 356: 355: 349: 348: 347: 346: 322: 321: 320:Other species 317: 316: 313: 301: 300: 294: 293: 276: 272: 271: 263: 259: 258: 256:Lambeosaurinae 250: 246: 245: 237: 233: 232: 224: 217: 216: 214:Neornithischia 208: 201: 200: 192: 185: 184: 179: 172: 171: 166: 162: 161: 156: 152: 151: 146: 142: 141: 128: 127: 123:fossil at the 117: 116: 108: 107: 99: 94: 89: 84: 79: 74: 69: 64: 59: 54: 49: 44: 43: 28: 15: 13: 10: 9: 6: 4: 3: 2: 6263: 6252: 6249: 6247: 6244: 6242: 6239: 6237: 6234: 6232: 6229: 6227: 6224: 6222: 6219: 6217: 6214: 6212: 6209: 6207: 6204: 6202: 6199: 6198: 6196: 6179: 6173: 6169: 6164: 6160: 6156: 6151: 6147: 6143: 6138: 6134: 6130: 6125: 6121: 6116: 6115:Hypacrosaurus 6110: 6106: 6101: 6095: 6091: 6090: 6088: 6086: 6085:Hypacrosaurus 6082: 6078: 6073: 6053: 6052: 6048: 6046: 6044: 6043:Edmontosaurus 6040: 6038: 6036: 6035:Edmontosaurus 6032: 6030: 6027: 6024: 6020: 6017: 6015: 6012: 6010: 6007: 6005: 6002: 6000: 5997: 5995: 5994: 5990: 5989: 5987: 5983: 5977: 5976: 5972: 5970: 5969: 5965: 5963: 5962: 5958: 5956: 5955: 5954:Syngonosaurus 5951: 5949: 5948: 5944: 5942: 5941: 5937: 5935: 5934: 5930: 5928: 5927: 5923: 5921: 5920: 5916: 5914: 5913: 5909: 5907: 5906: 5902: 5900: 5899: 5898:Koutalisaurus 5895: 5893: 5892: 5888: 5886: 5885: 5881: 5879: 5878: 5877:Eucercosaurus 5874: 5872: 5871: 5867: 5864: 5863: 5859: 5857: 5856: 5852: 5850: 5849: 5848:Arstanosaurus 5845: 5844: 5842: 5840: 5839: 5834: 5828: 5827: 5823: 5822: 5820: 5818: 5814: 5808: 5807: 5803: 5801: 5800: 5796: 5794: 5793: 5789: 5787: 5786: 5782: 5780: 5779: 5775: 5773: 5772: 5768: 5766: 5765: 5764:Orcauichnites 5761: 5759: 5758: 5754: 5752: 5751: 5747: 5745: 5744: 5740: 5738: 5737: 5733: 5731: 5730: 5726: 5724: 5723: 5719: 5717: 5716: 5712: 5710: 5709: 5705: 5703: 5702: 5701:Amblydactylus 5698: 5697: 5695: 5693: 5689: 5685: 5678: 5674: 5665: 5661: 5659: 5654: 5648: 5641: 5615: 5614: 5610: 5608: 5607: 5603: 5601: 5600: 5599:Kundurosaurus 5596: 5594: 5593: 5592:Kerberosaurus 5589: 5587: 5586: 5582: 5580: 5579: 5578:Edmontosaurus 5575: 5574: 5572: 5570: 5566: 5560: 5559: 5555: 5553: 5552: 5548: 5546: 5545: 5541: 5540: 5538: 5536: 5532: 5526: 5525: 5521: 5519: 5518: 5514: 5512: 5511: 5507: 5505: 5504: 5500: 5498: 5497: 5493: 5490: 5489: 5488:Anasazisaurus 5485: 5484: 5482: 5480: 5476: 5470: 5469: 5468:Secernosaurus 5465: 5463: 5462: 5461:Kelumapusaura 5458: 5456: 5455: 5451: 5449: 5448: 5444: 5443: 5441: 5439: 5435: 5428: 5427: 5423: 5421: 5420: 5416: 5414: 5413: 5409: 5407: 5406: 5402: 5400: 5399: 5395: 5393: 5392: 5388: 5387: 5385: 5383: 5379: 5373: 5372: 5368: 5367: 5364: 5361: 5359: 5358:Saurolophinae 5355: 5345: 5344: 5340: 5338: 5337: 5333: 5331: 5330: 5326: 5324: 5323: 5319: 5317: 5316: 5312: 5310: 5309: 5308:Hypacrosaurus 5305: 5303: 5302: 5301:Corythosaurus 5298: 5296: 5295: 5291: 5290: 5288: 5286: 5285:Lambeosaurini 5282: 5276: 5275: 5271: 5269: 5268: 5264: 5262: 5261: 5260:Charonosaurus 5257: 5255: 5254: 5250: 5249: 5247: 5245: 5241: 5235: 5234: 5233:Tsintaosaurus 5230: 5228: 5227: 5226:Pararhabdodon 5223: 5221: 5220: 5219:Nipponosaurus 5216: 5214: 5213: 5209: 5206: 5205: 5201: 5199: 5198: 5194: 5192: 5191: 5190:Jaxartosaurus 5187: 5185: 5184: 5180: 5178: 5177: 5173: 5171: 5170: 5166: 5164: 5163: 5159: 5157: 5156: 5152: 5150: 5149: 5145: 5143: 5142: 5141:Adynomosaurus 5138: 5137: 5134: 5131: 5129: 5125: 5119: 5118: 5114: 5112: 5111: 5107: 5105: 5104: 5103:Lapampasaurus 5100: 5098: 5097: 5093: 5091: 5090: 5086: 5084: 5083: 5079: 5078: 5075: 5072: 5069: 5068:Hadrosauridae 5064: 5058: 5057: 5053: 5051: 5050: 5049:Zhanghenglong 5046: 5044: 5043: 5039: 5037: 5036: 5032: 5030: 5029: 5025: 5023: 5022: 5021:Telmatosaurus 5018: 5016: 5015: 5011: 5009: 5008: 5004: 5002: 5001: 4997: 4995: 4994: 4990: 4988: 4987: 4983: 4981: 4980: 4976: 4974: 4973: 4969: 4967: 4966: 4965:Portellsaurus 4962: 4960: 4959: 4955: 4953: 4952: 4948: 4946: 4945: 4941: 4939: 4938: 4934: 4932: 4931: 4927: 4925: 4924: 4920: 4917: 4916: 4912: 4910: 4909: 4905: 4903: 4902: 4898: 4896: 4895: 4891: 4889: 4888: 4884: 4882: 4881: 4877: 4875: 4874: 4870: 4868: 4867: 4863: 4861: 4860: 4856: 4854: 4853: 4849: 4847: 4846: 4842: 4840: 4839: 4835: 4833: 4832: 4828: 4826: 4825: 4821: 4818: 4817: 4813: 4811: 4810: 4806: 4804: 4803: 4799: 4798: 4795: 4792: 4789: 4784: 4780: 4776: 4771: 4767: 4758: 4754: 4751: 4744: 4711: 4710: 4709: 4708: 4706: 4704: 4700: 4693: 4692: 4688: 4686: 4685: 4681: 4679: 4678: 4674: 4671: 4670: 4666: 4665: 4663: 4661: 4657: 4651: 4650: 4646: 4644: 4643: 4639: 4637: 4636: 4632: 4630: 4629: 4628:Jinzhousaurus 4625: 4623: 4622: 4621:Hypselospinus 4618: 4616: 4615: 4611: 4609: 4608: 4604: 4603: 4600: 4597: 4595: 4591: 4585: 4584: 4580: 4578: 4577: 4576:Theiophytalia 4573: 4571: 4570: 4566: 4564: 4563: 4559: 4557: 4556: 4552: 4550: 4549: 4545: 4543: 4542: 4538: 4536: 4535: 4531: 4529: 4528: 4524: 4522: 4521: 4517: 4515: 4514: 4510: 4508: 4507: 4506:Lanzhousaurus 4503: 4501: 4500: 4496: 4494: 4493: 4489: 4487: 4486: 4482: 4480: 4479: 4475: 4472: 4471: 4467: 4465: 4464: 4460: 4458: 4457: 4453: 4451: 4450: 4446: 4444: 4443: 4439: 4437: 4436: 4432: 4431: 4428: 4425: 4422: 4417: 4407: 4406: 4402: 4399: 4398: 4394: 4391: 4390: 4386: 4384: 4383: 4379: 4377: 4376: 4375:Elrhazosaurus 4372: 4370: 4369: 4365: 4363: 4362: 4358: 4357: 4355: 4353: 4349: 4343: 4342: 4338: 4336: 4335: 4331: 4329: 4328: 4324: 4323: 4320: 4316: 4313: 4310: 4305: 4291: 4290: 4286: 4284: 4283: 4279: 4277: 4276: 4272: 4270: 4269: 4265: 4263: 4262: 4258: 4256: 4255: 4251: 4250: 4248: 4246: 4242: 4236: 4235: 4234:Tenontosaurus 4231: 4229: 4228: 4224: 4221: 4220: 4219:Convolosaurus 4216: 4215: 4213: 4210: 4206: 4199: 4198: 4194: 4191: 4190: 4186: 4183: 4182: 4181:Ampelognathus 4178: 4177: 4174: 4171: 4169: 4165: 4158: 4157: 4153: 4151: 4150: 4146: 4144: 4143: 4139: 4137: 4136: 4132: 4130: 4129: 4125: 4123: 4122: 4118: 4116: 4115: 4111: 4108: 4107: 4103: 4101: 4100: 4096: 4094: 4093: 4092:Mahuidacursor 4089: 4087: 4086: 4082: 4080: 4079: 4075: 4072: 4071: 4067: 4065: 4064: 4060: 4058: 4057: 4053: 4051: 4050: 4046: 4043: 4042: 4038: 4036: 4035: 4031: 4029: 4028: 4024: 4022: 4021: 4017: 4015: 4014: 4010: 4009: 4007: 4005: 4001: 3994: 3993: 3989: 3987: 3986: 3985:Hypsilophodon 3982: 3979: 3978: 3974: 3973: 3971: 3968: 3964: 3957: 3956: 3952: 3950: 3949: 3948:Callovosaurus 3945: 3943: 3942: 3941:Burianosaurus 3938: 3937: 3934: 3930: 3927: 3924: 3919: 3915: 3911: 3906: 3902: 3877: 3876: 3875: 3874: 3872: 3870: 3866: 3858: 3854: 3853: 3852: 3851: 3848: 3845: 3843: 3839: 3831: 3827: 3826: 3825: 3824: 3821: 3818: 3816: 3812: 3808: 3800: 3796: 3793: 3791: 3787: 3784: 3782: 3778: 3775: 3773: 3769: 3766: 3764: 3760: 3758: 3754: 3753: 3748: 3744: 3740: 3733: 3728: 3726: 3721: 3719: 3714: 3713: 3710: 3704: 3700: 3697: 3695: 3694:Hypacrosaurus 3691: 3689: 3687: 3686:Hypacrosaurus 3683: 3682: 3678: 3670: 3666: 3662: 3658: 3654: 3650: 3646: 3642: 3635: 3632: 3627: 3625:0-691-05900-4 3621: 3617: 3612: 3611: 3605: 3604:Dodson, Peter 3599: 3597: 3593: 3588: 3586:0-520-24209-2 3582: 3578: 3573: 3572: 3566: 3560: 3558: 3554: 3548: 3545: 3540: 3536: 3531: 3526: 3521: 3516: 3512: 3508: 3504: 3497: 3494: 3489: 3485: 3480: 3475: 3470: 3465: 3461: 3457: 3454:(4): e36112. 3453: 3449: 3445: 3438: 3435: 3430: 3426: 3421: 3416: 3412: 3408: 3404: 3400: 3396: 3389: 3386: 3381: 3377: 3373: 3369: 3365: 3358: 3355: 3340: 3336: 3332: 3327: 3322: 3317: 3312: 3308: 3304: 3300: 3296: 3289: 3282: 3279: 3272: 3269: 3265: 3264:1-4051-3413-5 3261: 3255: 3253: 3249: 3244: 3238: 3230: 3226: 3222: 3218: 3214: 3210: 3206: 3202: 3195: 3188: 3185: 3181: 3175: 3172: 3167: 3163: 3159: 3155: 3151: 3147: 3143: 3136: 3133: 3128: 3121: 3118: 3113: 3109: 3104: 3099: 3095: 3091: 3087: 3083: 3079: 3072: 3069: 3064: 3060: 3056: 3050: 3047: 3042: 3038: 3034: 3028: 3026: 3024: 3020: 3015: 3011: 3007: 3003: 2999: 2995: 2991: 2984: 2982: 2978: 2973: 2969: 2965: 2961: 2957: 2953: 2949: 2945: 2938: 2936: 2927: 2925: 2921: 2916: 2914:0-521-56723-8 2910: 2906: 2902: 2901:Hypacrosaurus 2898: 2892: 2890: 2888: 2886: 2884: 2880: 2875: 2873:0-520-24209-2 2869: 2865: 2860: 2859: 2853: 2847: 2845: 2843: 2841: 2839: 2837: 2835: 2831: 2826: 2822: 2818: 2814: 2810: 2806: 2802: 2801:Dodson, Peter 2796: 2793: 2788: 2784: 2780: 2774: 2771: 2766: 2762: 2758: 2754: 2748: 2745: 2740: 2736: 2732: 2728: 2724: 2718: 2716: 2712: 2707: 2705:0-89950-917-7 2701: 2697: 2692: 2691: 2685: 2679: 2677: 2673: 2668: 2664: 2660: 2656: 2652: 2648: 2644: 2640: 2636: 2632: 2626: 2623: 2618: 2612: 2608: 2601: 2598: 2587: 2583: 2579: 2575: 2571: 2569: 2568:Hypacrosaurus 2563: 2562:Brown, Barnum 2557: 2555: 2551: 2545: 2541: 2538: 2537: 2533: 2522: 2517: 2515: 2513: 2512: 2507: 2506: 2501: 2500: 2495: 2494: 2493:Achelousaurus 2489: 2488: 2483: 2482: 2477: 2476: 2471: 2470: 2465: 2461: 2460: 2455: 2454: 2449: 2448: 2447:Chirostenotes 2444: 2440: 2439: 2434: 2433: 2428: 2427: 2422: 2421:H. stebingeri 2417: 2415: 2414:H. altispinus 2411: 2407: 2403: 2399: 2395: 2391: 2390: 2389:Albertosaurus 2386: 2385:tyrannosaurid 2382: 2378: 2374: 2370: 2369: 2368:Struthiomimus 2364: 2363: 2359: 2355: 2354: 2350: 2346: 2345: 2340: 2339: 2334: 2333: 2332:Anchiceratops 2328: 2327: 2322: 2321: 2317: 2313: 2312: 2308: 2304: 2303: 2299: 2295: 2294: 2289: 2288: 2287:Edmontosaurus 2283: 2282:H. altispinus 2276: 2275:H. altispinus 2271: 2264: 2262: 2260: 2255: 2247: 2242: 2240: 2237: 2236:Hypacrosaurus 2229: 2227: 2224: 2223:H. stebingeri 2220: 2216: 2212: 2208: 2205: 2201: 2197: 2193: 2189: 2185: 2181: 2177: 2176:H. stebingeri 2173: 2169: 2165: 2164:H. stebingeri 2161: 2154: 2150: 2146: 2145:Hypacrosaurus 2142: 2135: 2133: 2131: 2130:H. stebingeri 2127: 2123: 2122:Tyrannosaurus 2119: 2118:Albertosaurus 2115: 2114:H. stebingeri 2111: 2107: 2106:H. stebingeri 2103: 2099: 2095: 2091: 2087: 2086:H. stebingeri 2083: 2079: 2078:H. stebingeri 2074: 2070: 2063: 2062:H. stebingeri 2059: 2052: 2050: 2048: 2047:Hypacrosaurus 2044: 2043:Hypacrosaurus 2040: 2039:Hypacrosaurus 2031: 2029: 2027: 2023: 2022:Hypacrosaurus 2019: 2018: 2013: 2012: 2007: 2003: 1999: 1995: 1991: 1987: 1983: 1979: 1974: 1972: 1968: 1963: 1962:Hypacrosaurus 1956: 1952: 1951:H. altispinus 1948: 1941: 1939: 1937: 1933: 1929: 1925: 1921: 1917: 1914: 1910: 1906: 1905:Hypacrosaurus 1902: 1894: 1887: 1886: 1878: 1877: 1869: 1868: 1860: 1859: 1851: 1850: 1842: 1841: 1833: 1832: 1824: 1823: 1815: 1814: 1806: 1805: 1797: 1796: 1788: 1787: 1779: 1778: 1770: 1769: 1761: 1760: 1752: 1751: 1748: 1747: 1746: 1739: 1738: 1735: 1734: 1731: 1730: 1729: 1724: 1723: 1715: 1714: 1708: 1707: 1704: 1703: 1700: 1699: 1698: 1691: 1690: 1684: 1683:Hypacrosaurus 1681: 1680: 1677: 1676: 1668: 1667: 1664: 1663: 1662: 1655: 1654: 1651: 1650: 1647: 1646: 1645: 1638: 1637: 1631: 1630: 1629:Corythosaurus 1626: 1625: 1619: 1618: 1615: 1614: 1606: 1605: 1597: 1596: 1588: 1587: 1584: 1583: 1582: 1575: 1574: 1571: 1570: 1567: 1566: 1565: 1558: 1557: 1551: 1550: 1547: 1546: 1543: 1542: 1541: 1534: 1533: 1527: 1526: 1522: 1521: 1518: 1517: 1514: 1513: 1512: 1505: 1504: 1498: 1497: 1491: 1490: 1487: 1486: 1483: 1482: 1481: 1474: 1473: 1467: 1466: 1463: 1462: 1459: 1458: 1457: 1450: 1449: 1443: 1442:Lambeosaurini 1440: 1439: 1436: 1435: 1427: 1426: 1418: 1417: 1409: 1408: 1405: 1404: 1403: 1396: 1395: 1392: 1391: 1388: 1387: 1386: 1379: 1378: 1372: 1371: 1368: 1367: 1364: 1363: 1362: 1357: 1356: 1355:Charonosaurus 1348: 1347: 1341: 1340: 1337: 1336: 1333: 1332: 1331: 1324: 1323: 1317: 1316: 1312: 1311: 1305: 1304:Corythosauria 1302: 1301: 1298: 1297: 1289: 1288: 1285: 1284: 1283: 1276: 1275: 1272: 1271: 1268: 1267: 1266: 1259: 1258: 1252: 1249: 1248: 1242: 1241: 1238: 1237: 1234: 1233: 1232: 1231:Jaxartosaurus 1225: 1224: 1218: 1217: 1214: 1213: 1205: 1204: 1201: 1200: 1199: 1198:Tsintaosaurus 1192: 1191: 1188: 1187: 1184: 1183: 1182: 1181:Pararhabdodon 1175: 1174: 1168: 1165: 1164: 1158: 1157: 1154: 1153: 1145: 1144: 1141: 1140: 1139: 1132: 1131: 1128: 1127: 1124: 1123: 1122: 1115: 1114: 1108: 1105: 1104: 1098: 1095: 1094: 1091: 1090: 1082: 1081: 1078: 1077: 1076: 1075:Edmontosaurus 1069: 1068: 1065: 1064: 1061: 1060: 1059: 1052: 1051: 1045: 1044: 1038: 1037: 1034: 1033: 1030: 1029: 1028: 1027:Telmatosaurus 1021: 1020: 1014: 1013: 1010: 1009: 1006: 1005: 1004: 997: 996: 990: 989: 986: 985: 982: 981: 980: 973: 972: 966: 965: 961: 959: 958:Hypacrosaurus 956:to be within 955: 954: 949: 945: 940: 938: 937: 932: 931:Corythosaurus 928: 927:Nipponosaurus 924: 920: 919:Hypacrosaurus 916: 912: 911:Hypacrosaurus 908: 904: 903:Nipponosaurus 900: 899: 898:Nipponosaurus 894: 890: 886: 885:Hypacrosaurus 882: 881:Corythosaurus 878: 877:Corythosaurus 874: 873:Hypacrosaurus 870: 869:H. altispinus 866: 862: 861:H. stebingeri 858: 854: 850: 849:Corythosaurus 846: 842: 838: 837:Hypacrosaurus 832: 829:mount at the 828: 827:Hypacrosaurus 824: 818: 817:H. altispinus 813: 807: 806:H. altispinus 802: 795: 793: 791: 790:Hypacrosaurus 787: 786: 781: 780:H. stebingeri 777: 773: 772:H. stebingeri 769: 768:H. altispinus 765: 762: 758: 754: 750: 746: 742: 741:H. altispinus 738: 737:Corythosaurus 734: 730: 729:neural spines 726: 722: 721:Hypacrosaurus 714: 707: 705: 703: 699: 695: 691: 687: 682: 680: 679:H. altispinus 676: 675:H. stebingeri 672: 671:H. altispinus 668: 667:Nipponosaurus 664: 660: 659: 658:Nipponosaurus 654: 650: 649:natural group 646: 645:H. stebingeri 642: 641:H. altispinus 638: 634: 633:H. stebingeri 630: 626: 625:H. altispinus 619: 618:H. stebingeri 615: 608: 606: 604: 603:H. stebingeri 600: 596: 592: 588: 584: 580: 579:Corythosaurus 576: 572: 568: 564: 560: 559: 554: 550: 546: 542: 541:Hypacrosaurus 537: 535: 534: 529: 525: 521: 517: 513: 509: 505: 501: 497: 493: 492:Hypacrosaurus 489: 482: 478: 477:H. altispinus 473: 466: 464: 462: 461:H. stebingeri 459:belonging to 458: 454: 450: 446: 445:North America 442: 438: 434: 430: 429:Corythosaurus 426: 425: 424:Corythosaurus 420: 417: 413: 409: 408: 407:Tyrannosaurus 403: 402: 401:Hypacrosaurus 390: 386: 379: 378: 374: 370: 365: 364: 360: 359: 357: 354: 350: 343: 339: 334: 333: 332:H. stebingeri 326: 325: 323: 318: 311: 310: 302: 299: 295: 290: 285: 284: 283:Hypacrosaurus 277: 274: 273: 270: 269:Lambeosaurini 264: 261: 260: 257: 251: 248: 247: 244: 243:Hadrosauridae 238: 235: 234: 231: 225: 222: 219: 218: 215: 209: 206: 203: 202: 199: 193: 190: 187: 186: 183: 180: 177: 174: 173: 170: 167: 164: 163: 160: 157: 154: 153: 150: 147: 144: 143: 138: 133: 129: 126: 122: 121:H. altispinus 118: 114: 109: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 52: 47: 41: 32: 26: 25:Hypacrosaurus 22: 19: 6084: 6049: 6042: 6034: 6022: 5991: 5973: 5966: 5959: 5952: 5945: 5938: 5931: 5924: 5917: 5910: 5903: 5896: 5889: 5882: 5875: 5868: 5862:Claorhynchus 5860: 5853: 5846: 5838:Nomina dubia 5836: 5826:Guegoolithus 5824: 5804: 5797: 5792:Staurichnium 5790: 5785:Sousaichnium 5783: 5776: 5769: 5762: 5755: 5748: 5741: 5736:Dakotasaurus 5734: 5729:Caririchnium 5727: 5720: 5713: 5706: 5699: 5643: 5611: 5604: 5597: 5590: 5583: 5576: 5556: 5549: 5542: 5535:Saurolophini 5522: 5515: 5508: 5501: 5494: 5486: 5479:Kritosaurini 5466: 5459: 5454:Huallasaurus 5452: 5445: 5426:Wulagasaurus 5424: 5417: 5410: 5403: 5396: 5389: 5369: 5341: 5334: 5327: 5320: 5315:Lambeosaurus 5313: 5307: 5306: 5299: 5292: 5272: 5265: 5258: 5251: 5231: 5224: 5217: 5210: 5202: 5195: 5188: 5181: 5174: 5167: 5160: 5153: 5146: 5139: 5117:Yamatosaurus 5115: 5108: 5101: 5094: 5087: 5082:Aquilarhinus 5080: 5054: 5047: 5040: 5033: 5028:Tethyshadros 5026: 5019: 5012: 5005: 4998: 4991: 4984: 4977: 4970: 4963: 4958:Plesiohadros 4956: 4949: 4942: 4935: 4930:Lophorhothon 4928: 4921: 4913: 4906: 4899: 4892: 4885: 4878: 4871: 4864: 4857: 4850: 4843: 4836: 4829: 4822: 4816:Batyrosaurus 4814: 4809:Bactrosaurus 4807: 4800: 4746: 4689: 4682: 4675: 4667: 4649:Ouranosaurus 4647: 4640: 4633: 4626: 4619: 4614:Brighstoneus 4612: 4605: 4581: 4574: 4567: 4560: 4553: 4546: 4541:Osmakasaurus 4539: 4534:Oblitosaurus 4532: 4525: 4518: 4511: 4504: 4497: 4490: 4483: 4476: 4468: 4461: 4456:Camptosaurus 4454: 4447: 4440: 4433: 4403: 4397:Kangnasaurus 4395: 4387: 4380: 4373: 4366: 4359: 4352:Dryosauridae 4339: 4332: 4325: 4287: 4280: 4273: 4266: 4259: 4254:Matheronodon 4252: 4232: 4225: 4217: 4195: 4187: 4179: 4154: 4147: 4140: 4133: 4128:Sektensaurus 4126: 4121:Qantassaurus 4119: 4112: 4104: 4097: 4090: 4083: 4076: 4070:Kangnasaurus 4068: 4061: 4054: 4047: 4039: 4034:Diluvicursor 4032: 4025: 4018: 4011: 3992:Vectidromeus 3990: 3983: 3975: 3953: 3946: 3939: 3857:Ornithischia 3842:Ornithischia 3794: 3785: 3781:Ornithischia 3776: 3767: 3693: 3685: 3644: 3640: 3634: 3609: 3570: 3547: 3510: 3506: 3496: 3451: 3447: 3437: 3402: 3398: 3388: 3371: 3367: 3363: 3357: 3346:. Retrieved 3298: 3294: 3281: 3271: 3237:cite journal 3204: 3200: 3187: 3179: 3174: 3149: 3145: 3135: 3126: 3120: 3085: 3082:Paleobiology 3081: 3071: 3062: 3058: 3049: 3040: 3036: 2997: 2993: 2989: 2947: 2943: 2934: 2904: 2900: 2857: 2811:(1): 37–54. 2808: 2804: 2795: 2786: 2782: 2773: 2764: 2760: 2756: 2755:(1917). "On 2747: 2741:(43): 29–48. 2738: 2734: 2730: 2726: 2689: 2642: 2638: 2634: 2631:Arbour, V.M. 2625: 2606: 2600: 2589:. Retrieved 2577: 2573: 2567: 2509: 2503: 2497: 2491: 2485: 2484:, hadrosaur 2479: 2473: 2467: 2463: 2457: 2451: 2445: 2443:caenagnathid 2436: 2430: 2424: 2420: 2418: 2413: 2387: 2366: 2362:Ornithomimus 2360: 2351: 2342: 2336: 2330: 2324: 2318: 2309: 2307:ankylosaurid 2302:Parksosaurus 2300: 2291: 2285: 2281: 2280: 2274: 2253: 2251: 2243:Paleoecology 2235: 2233: 2222: 2175: 2168:histological 2163: 2157: 2144: 2129: 2121: 2117: 2113: 2105: 2102:growth rings 2085: 2077: 2068: 2067: 2061: 2046: 2042: 2038: 2035: 2021: 2017:Ornithomimus 2015: 2011:Nanotyrannus 2009: 1975: 1961: 1959: 1950: 1904: 1898: 1895:Paleobiology 1743: 1742: 1726: 1720: 1718: 1695: 1694: 1682: 1659: 1658: 1642: 1641: 1627: 1579: 1578: 1562: 1561: 1538: 1537: 1525:Lambeosaurus 1523: 1509: 1508: 1478: 1477: 1454: 1453: 1400: 1399: 1383: 1382: 1359: 1353: 1351: 1328: 1327: 1313: 1280: 1279: 1263: 1262: 1251:Arenysaurini 1229: 1228: 1196: 1195: 1179: 1178: 1136: 1135: 1119: 1118: 1107:Aralosaurini 1073: 1072: 1056: 1055: 1025: 1024: 1003:Bactrosaurus 1001: 1000: 977: 976: 957: 951: 948:Lambeosaurus 947: 941: 934: 930: 926: 918: 915:paraphyletic 910: 906: 902: 896: 892: 884: 880: 876: 872: 868: 865:Lambeosaurus 864: 860: 848: 845:Lambeosaurus 844: 836: 835: 826: 816: 815:Hindfoot of 805: 789: 785:Lambeosaurus 783: 779: 771: 767: 740: 736: 720: 719: 685: 683: 678: 674: 670: 666: 656: 652: 644: 640: 632: 629:type species 624: 623: 617: 602: 598: 594: 590: 587:Cheneosaurus 586: 583:Lambeosaurus 582: 578: 567:Peter Dodson 556: 544: 540: 538: 531: 512:Tolman Ferry 496:Barnum Brown 491: 488:type remains 485: 476: 460: 428: 422: 405: 400: 399: 398: 384: 375: 363:Cheneosaurus 362: 361: 331: 330: 308: 307: 298:Type species 282: 281: 220: 204: 198:Ornithischia 188: 175: 120: 24: 18: 6109:Wikispecies 6051:Willinakaqe 6023:nomina nuda 5940:Siluosaurus 5905:Loncosaurus 5743:Dinehichnus 5585:Kamuysaurus 5558:Saurolophus 5510:Kritosaurus 5503:Gryposaurus 5336:Sahaliyania 5322:Magnapaulia 5294:Amurosaurus 5274:Tlatolophus 5253:Adelolophus 5197:Kazaklambia 5176:Blasisaurus 5169:Arenysaurus 5162:Aralosaurus 5096:Hadrosaurus 5089:Eotrachodon 5042:Yunganglong 5007:Sirindhorna 4979:Protohadros 4915:Koshisaurus 4908:Jintasaurus 4677:Comptonatus 4527:Napaisaurus 4513:Lurdusaurus 4485:Fukuisaurus 4463:Cedrorestes 4405:Valdosaurus 4268:Pareisactus 4099:Morrosaurus 4063:Isasicursor 4027:Chakisaurus 3923:Ornithopoda 3910:Ornithopoda 3869:Ornithopoda 3739:Ornithopoda 2789:(5): 1–162. 2511:Rubeosaurus 2505:Einiosaurus 2475:Scolosaurus 2453:Bambiraptor 2406:shallow sea 2377:troodontids 2293:Saurolophus 2265:Environment 2259:biocoenosis 2188:chondrocyte 2149:chondrocyte 2026:CT scanning 1913:quadrupedal 1901:hadrosaurid 1722:Magnapaulia 1511:Amurosaurus 1361:jiayinensis 1282:Arenysaurus 1265:Blasisaurus 1138:Aralosaurus 1058:Gryposaurus 953:Magnapaulia 857:Phil Currie 853:Jack Horner 841:hadrosaurid 761:quadrupedal 708:Description 665:found that 533:Saurolophus 504:postcranial 377:Magnapaulia 314:Brown, 1913 249:Subfamily: 230:Ornithopoda 38:75–67  6195:Categories 5993:Arkharavia 5968:Tichosteus 5961:Thespesius 5933:Pteropelyx 5926:Orthomerus 5806:Wintonopus 5391:Acristavus 5371:Barsboldia 5329:Olorotitan 5204:Latirhinus 5056:Zuoyunlong 4923:Levnesovia 4873:Gobihadros 4838:Datonglong 4831:Claosaurus 4802:Altirhinus 4712:see below↓ 4642:Morelladon 4520:Magnamanus 4492:Hippodraco 4361:Dryosaurus 4334:Hesperonyx 4309:Dryomorpha 4149:Trinisaura 4142:Tietasaura 4135:Talenkauen 4013:Anabisetia 3878:see below↓ 3772:Dinosauria 3348:2017-01-04 2733:, Parks". 2591:2007-05-02 2546:References 2481:Orodromeus 2469:Oohkotokia 2464:Edmontonia 2410:Cretaceous 2383:, and the 2375:including 2353:Stegoceras 2320:Edmontonia 2316:nodosaurid 2110:thigh bone 2006:metabolism 2002:ectotherms 1998:endotherms 1978:cartilages 1926:, and its 1728:laticaudus 1456:Olorotitan 936:Olorotitan 921:to form a 788:and later 747:(4.4  637:hypothesis 457:hatchlings 182:Dinosauria 5975:Trachodon 5891:Glishades 5870:Diclonius 5708:Anomoepus 5692:Ichnotaxa 5405:Maiasaura 5343:Velafrons 4852:Equijubus 4824:Choyrodon 4684:Iguanodon 4548:Planicoxa 4478:Dakotadon 4449:Calvarius 4275:Rhabdodon 4261:Mochlodon 4004:Elasmaria 3755:Kingdom: 3166:250463301 3112:221666530 2586:2246/1416 2426:Maiasaura 2373:theropods 2248:Taphonomy 2209:: stains 2180:calcified 2158:In 2020, 2090:alligator 2080:suggests 2024:based on 1996:, modern 1986:tetrapods 1982:olfactory 1916:herbivore 1480:Velafrons 944:cladogram 764:herbivore 733:vertebrae 571:juveniles 155:Kingdom: 149:Eukaryota 6094:Wikidata 5855:Cionodon 5524:Rhinorex 5412:Ornatops 5212:Minqaria 5183:Canardia 5110:Malefica 5035:Xuwulong 5000:Siamodon 4901:Jeyawati 4887:Gonkoken 4845:Eolambia 4669:Barilium 4470:Cumnoria 4435:Barilium 4341:Owenodon 4327:Draconyx 4289:Zalmoxes 4189:Fostoria 4041:Fostoria 3799:Cerapoda 3763:Chordata 3761:Phylum: 3757:Animalia 3699:Archived 3606:(1996). 3539:34692099 3488:22558351 3448:PLOS ONE 3429:18682367 3339:Archived 3335:28049837 3276:295–305. 3229:84693210 3014:86070917 2972:83841024 2667:85665879 2564:(1913). 2518:See also 2207:staining 1121:Canardia 979:Xuwulong 925:without 661:; their 498:for the 419:dinosaur 416:duckbill 410:) was a 353:Synonyms 236:Family: 169:Chordata 165:Phylum: 159:Animalia 145:Domain: 6178:4199661 6155:4128527 6142:1026975 6129:4823230 6100:Q311563 5148:Ajnabia 4583:Uteodon 3669:3514834 3649:Bibcode 3641:PALAIOS 3530:8289162 3479:3340333 3456:Bibcode 3420:2605812 3326:5255600 3303:Bibcode 3209:Bibcode 3201:Science 3090:Bibcode 2952:Bibcode 2825:2412696 2647:Bibcode 2432:Troodon 2192:mitosis 1994:mammals 1953:skull, 1924:chewing 1909:bipedal 929:, with 913:may be 893:et al.' 851:, with 757:bipedal 677:was to 653:et al.' 647:form a 609:Species 577:genera 516:Alberta 441:Montana 437:Alberta 427:. Like 275:Genus: 262:Tribe: 6175:uBio: 5817:Ootaxa 5014:Tanius 4607:Bolong 3667:  3622:  3583:  3579:–606. 3537:  3527:  3486:  3476:  3427:  3417:  3333:  3323:  3262:  3227:  3164:  3110:  3012:  2970:  2911:  2870:  2866:–463. 2823:  2702:  2698:–482. 2665:  2613:  2508:, and 2472:, and 2404:, the 2398:marine 2341:, and 2094:ratite 1920:plants 753:pelvic 745:tonnes 627:, the 455:, and 385:et al. 371:, 1917 344:, 1994 342:Currie 340:& 338:Horner 291:, 1913 6168:56405 6137:IRMNG 4859:Fylax 4389:Iyuku 3795:Clade 3786:Clade 3777:Clade 3768:Clade 3665:JSTOR 3616:14–15 3342:(PDF) 3291:(PDF) 3225:S2CID 3197:(PDF) 3162:S2CID 3108:S2CID 3010:S2CID 2968:S2CID 2940:(PDF) 2821:JSTOR 2663:S2CID 2200:cells 2136:Cells 1990:birds 1936:cheek 1928:teeth 1899:As a 923:clade 889:clade 778:, as 673:than 639:that 510:near 449:nests 412:genus 369:Lambe 289:Brown 221:Clade 205:Clade 189:Clade 176:Clade 6124:GBIF 4555:Proa 4227:Iani 3855:see 3828:see 3620:ISBN 3581:ISBN 3535:PMID 3484:PMID 3425:PMID 3331:PMID 3260:ISBN 3243:link 2909:ISBN 2868:ISBN 2700:ISBN 2611:ISBN 2456:and 2379:and 2365:and 2290:and 2213:and 2211:DAPI 2120:and 2073:eggs 2064:nest 1992:and 1955:AMNH 933:and 905:and 883:and 875:and 867:and 855:and 847:and 770:and 749:tons 643:and 581:and 563:taxa 543:was 486:The 453:eggs 389:2012 46:PreꞒ 3657:doi 3577:517 3525:PMC 3515:doi 3474:PMC 3464:doi 3415:PMC 3407:doi 3403:275 3376:doi 3321:PMC 3311:doi 3299:114 3217:doi 3205:273 3154:doi 3098:doi 3002:doi 2960:doi 2864:438 2813:doi 2696:478 2655:doi 2582:hdl 2219:emu 2204:DNA 2153:emu 2020:or 700:in 490:of 479:at 435:of 414:of 6197:: 6165:: 6152:: 6139:: 6126:: 6111:: 6096:: 3797:: 3788:: 3779:: 3770:: 3663:. 3655:. 3643:. 3618:. 3595:^ 3556:^ 3533:. 3523:. 3509:. 3505:. 3482:. 3472:. 3462:. 3450:. 3446:. 3423:. 3413:. 3401:. 3397:. 3372:19 3370:. 3337:. 3329:. 3319:. 3309:. 3297:. 3293:. 3251:^ 3239:}} 3235:{{ 3223:. 3215:. 3203:. 3199:. 3160:. 3150:41 3148:. 3144:. 3106:. 3096:. 3086:46 3084:. 3080:. 3063:55 3061:. 3041:40 3022:^ 3008:. 2998:27 2996:. 2980:^ 2966:. 2958:. 2948:24 2946:. 2942:. 2923:^ 2882:^ 2833:^ 2819:. 2809:24 2807:. 2787:20 2785:. 2765:30 2763:. 2739:38 2737:. 2714:^ 2675:^ 2661:. 2653:. 2643:29 2641:. 2578:32 2576:. 2572:. 2553:^ 2502:, 2496:, 2466:, 2441:, 2412:. 2356:, 2347:, 2335:, 2329:, 2314:, 2305:, 2296:, 2257:a 2215:PI 2132:. 2014:, 1973:. 1903:, 1725:" 1358:" 792:. 696:) 605:. 526:, 514:, 463:. 451:, 387:, 380:? 223:: 207:: 191:: 178:: 96:Pg 40:Ma 6025:) 5865:? 5491:? 5429:? 5207:? 4918:? 4819:? 4694:? 4672:? 4473:? 4400:? 4392:? 4222:? 4211:? 4200:? 4192:? 4184:? 4159:? 4109:? 4073:? 4044:? 3995:? 3980:? 3969:? 3958:? 3731:e 3724:t 3717:v 3671:. 3659:: 3651:: 3645:5 3628:. 3589:. 3541:. 3517:: 3511:7 3490:. 3466:: 3458:: 3452:7 3431:. 3409:: 3382:. 3378:: 3351:. 3313:: 3305:: 3266:. 3245:) 3231:. 3219:: 3211:: 3168:. 3156:: 3114:. 3100:: 3092:: 3016:. 3004:: 2974:. 2962:: 2954:: 2933:" 2917:. 2876:. 2827:. 2815:: 2708:. 2669:. 2657:: 2649:: 2619:. 2594:. 2584:: 1911:/ 1719:" 1352:" 759:/ 692:( 522:( 328:† 305:† 279:† 266:† 253:† 240:† 227:† 211:† 195:† 101:N 91:K 86:J 81:T 76:P 71:C 66:D 61:S 56:O 51:Ꞓ 35:~ 33:,

Index

Late Cretaceous
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Royal Tyrrell Museum
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Dinosauria
Ornithischia
Neornithischia
Ornithopoda
Hadrosauridae
Lambeosaurinae
Lambeosaurini
Hypacrosaurus
Brown

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