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Halisaurus

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Takuya Konishi, Michael W. Caldwell, Tomohiro Nishimura, Kazuhiko Sakurai & Kyo Tanoue (2015) A new halisaurine mosasaur (Squamata: Halisaurinae) from Japan: the first record in the western Pacific realm and the first documented insights into binocular vision in mosasaurs. Journal of Systematic
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Bardet, N., Pereda Suberbiola, X., Iarochene, M., Bouya, B. & Amaghzaz, M. 2005. A new species of Halisaurus from the Late Cretaceous phosphates of Morocco, and the phylogenetical relationships of the Halisaurinae (Squamata: Mosasauridae). Zoological Journal of the Linnean Society, 143,
1447:, among them the shape of its external nares (V-shaped anteriorly and U-shaped posteriorly), the shape of its quadrate (which has an oval vertical stapedial notch) and the presence of anterior ridges on the frontal. Its pterygoid bone also preserves twelve teeth (three more than in 570:
but is more symmetrically heart-shaped in anterior aspect and appears slightly inflated in later profile with a convex anteroventral outline that is continuous with the anteroexternal margin of the articular surface, as in the genus
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Lingham-Soliar, T. 1996. The first description of Halisaurus (Reptilia Mosasauridae) from Europe, from the Upper Cretaceous of Belgium. Bulletin de l'Institut royal des Sciences naturelles de Belgique, Sciences de la Terre, 66,
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Caldwell, Michael W.; Bell, Gorden L. Jr. (1995-09-14). "Halisaurus sp. (Mosasauridae) from the Upper Cretaceous (?Santonian) of east-central Peru, and the taxonomic utility of mosasaur cervical vertebrae".
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was a relatively poor swimmer is relatively surprising considering its small size, since other small and medium-sized mosasaurs were mostly adept swimmers. The description of
1542: 1428:. The type specimen, MNHN PMC 14, is an incomplete skeleton that includes a disarticulated skull and 27 associated articulated vertebrae from the Grand Daoui area near 2590: 2641: 2486: 437:. The genus remains a key taxon in mosasaur systematics due to its unique set of features and as the most complete representative of its subfamily, the 429:
Since its description, more complete remains have been uncovered from fossil deposits throughout the world with particularly complete remains found in
935:. Apart from the named species, fragmentary remains have been referred to the genus from across the world. Though the designation of these remains as 2577: 2636: 1781: 1694:
Shaker, A. A.; Longrich, N. R.; Strougo, A.; Asan, A.; Bardet, N.; Mousa, M. K.; Tantawy, A. A.; Abu El-Kheir, G. A. (2023). "A new species of
394:, already revealed a relatively unique combination of features and prompted a new genus to be described. Its name is a portmanteau of the 2631: 1593:"Mosasauroid phylogeny under multiple phylogenetic methods provides new insights on the evolution of aquatic adaptations in the group" 1310:
in 1869. The species name means "flat-spined". Referred specimens include the type specimen YPM 444 (consisting of an angular and a
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possibly representing a distinct genus in need of a new name. The results of their analyses are displayed in the cladogram below:
1557: 663:. This would mean that the halisaurines are more closely related to the mosasaurines than the russellosaurines (genera such as 1365:, the broadly rounded and posteriorly rounded external nares and the lack of anterior ridges on its frontal as well as its 985:
of Egypt, on the basis of disarticulated cranial elements, teeth, and vertebrae from the neck and back. After performing a
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Simões, Tiago R.; Vernygora, Oksana; Paparella, Ilaria; Jimenez-Huidobro, Paulina; Caldwell, Michael W. (2017-05-03).
1322:), which is Maastrichtian in age. Important subsequent specimens include NJSM 12146 (an incomplete cranium from the 146: 1774: 1867: 1070: 512:
are poorly differentiated which means that the genus lacked the hydrophalangy of more advanced mosasaurs. That
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The most recent major phylogenetic analysis of mosasaurs, conducted by Tiago R. Simões and colleagues in May
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at large. It was a medium-sized mosasaur about 3–4 m (9.8–13.1 ft) long. The length of the dubious
418:. According to modern rules, a difference of a letter is enough and the substitute name is unneeded, making " 2481: 1698:(Mosasauridae: Halisaurinae) from the lower Maastrichtian (Upper Cretaceous) of the Western Desert, Egypt". 1343: 928: 448:
was comparatively small by mosasaur standards. Though bigger than earlier and more basal mosasaurs, such as
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fragment. Though the specimen does not reveal much about the mosasaur, it was noted by D.A. Russell in his
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within the larger mosasaur family tree has long been considered controversial. The type specimen of
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Marsh, O.C. (1869). "Notice on some new mosasauroid reptiles from the Greensand of New Jersey".
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With a length of 3–4 m (9.8–13.1 ft), and weight of 3.33 tonnes, (3.02 metric tonnes)
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than the type species of their respective genera. They tentatively assigned these species to
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were problematic, as said genera have always been classified in separate subfamilies, the
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This species is differentiated from other species by the narrow and oblique shape of its
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being momentarily reassigned to the genus by Lingham-Soliar (1996). In 2005, the species
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respectively. Russell referred the genus to the Plioplatecarpinae on the basis of a
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appeared relatively early in the evolutionary history of the mosasaurs, during the
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Palaeontology (advance online publication) DOI:10.1080/14772019.2015.1113447
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as the basalmost genus of the subfamily. The analysis excluded the dubious
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http://www.tandfonline.com/doi/full/10.1080/14772019.2015.1113447#abstract
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in 1870, since he believed the name to already be preoccupied by the fish
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in age, though specimens of this species have been found across northern
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is difficult to tell due to the lack of remains, but was likely similar.
497:. As such, the genus retains many primitive characteristics, as does the 375: 367: 227: 198: 178: 93: 88: 73: 68: 58: 608:
Several discoveries throughout the 1980s and 1990s helped shed light on
2582: 1433: 430: 188: 133: 108: 83: 532:, was highly specialized to compensate for its lack of hydrophalangy. 1881: 1802: 1543:"Systematics and morphology of American mosasaurs (Reptilia, Sauria)" 1421: 963:(which significantly expands the known temporal range of the genus). 168: 2511: 1378: 944: 677: 477: 363: 2569: 960: 2515: 1763: 1350:, implying that the species ranged across the eastern coast of 939:
is debatable in most cases, unnamed species are known from the
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would have been dwarfed by many of its contemporaries, such as
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by Nathalie Bardet and colleagues. With the description of
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means "Arambourg's ocean lizard" and is named in honor of
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and the rest of the Halisaurinae as a sister group to the
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following an analysis by Takuya Konishi and colleagues (
601:-like suprastapedial process in specimens referred to 644:, the two genera were grouped into the new subfamily 279: 2524: 2469: 2346: 2308: 2273: 2246: 2201: 2160: 2126: 2106: 2068: 2048: 1987: 1960: 1912: 1829: 1443:preserve several features that distinguish it from 612:, with more complete specimens of the type species 1550:Bulletin of the Peabody Museum of Natural History 486:, the most completely known species of the genus. 1744:Sea Dragons: Predators Of The Prehistoric Oceans 1330:), USNM 442450 (an incomplete skeleton from the 632:, along with the description of the new species 558:Systematics and Morphology of American Mosasaurs 993:and closely related taxa, they suggested that 1775: 8: 2487:List of mosasaur-bearing stratigraphic units 548:is not comprehensive and only preserves an 2512: 2157: 2123: 2065: 1909: 1782: 1768: 1760: 508:As in other halisaurines, the flippers of 119: 31: 1634: 1616: 1397:due to his work on fossil vertebrates in 406:; "lizard"). It was renamed by Marsh to 1475: 1005:were more closely related to the genus 1338:). The species is also known from the 564:) that the angular resembles that of 7: 1523: 1521: 1510: 1508: 1506: 1504: 1502: 927:deposits that were once part of the 2642:Taxa named by Othniel Charles Marsh 1746:by Richard Ellis page (p. 214) 1206: 1182: 1158: 1122: 1086: 1062: 1038: 1028: 823: 799: 758: 734: 724: 1661:Journal of Vertebrate Paleontology 25: 697:) done during the description of 626:was reassigned to its own genus, 2498: 2497: 2039: 1461: 145: 1673:10.1080/02724634.1995.10011246 640:as a closely related genus to 1: 2637:Fossil taxa described in 1869 2477:Timeline of mosasaur research 1708:10.1016/j.cretres.2023.105719 996:Phosphorosaurus ponpetelegans 849:Phosphorosaurus ponpetelegans 700:Phosphorosaurus ponpetelegans 577:. The similarities with both 519:Phosphorosaurus ponpetelegans 1618:10.1371/journal.pone.0176773 1485:American Journal of Science 1395:Professor Camille Arambourg 2663: 2632:Mosasaurs of North America 1354:during the Middle to Late 899:At least three species of 536:Classification and species 2495: 2347:Related groups and genera 2037: 1797: 1722:"Fossilworks: Mosasaurus" 1493:10.2475/ajs.s2-48.144.392 1300:Halisaurus platyspondylus 1294:Halisaurus platyspondylus 1215:Halisaurus platyspondylus 1211: 1204: 1187: 1180: 1163: 1156: 1127: 1120: 1091: 1084: 1067: 1060: 1043: 1036: 845: 828: 821: 804: 797: 780: 767:Halisaurus platyspondylus 763: 756: 739: 732: 386:fragment discovered near 305: 300: 284:Halisaurus platyspondylus 278: 271: 142:Scientific classification 140: 127: 118: 34: 1752:by Jack M. Callaway and 1487:. s2-48 (144): 392–397. 1344:Merchantville Formations 1071:Phosphorosaurus ortliebi 832:Phosphorosaurus ortliebi 619:Phosphorosaurus ortliebi 528:, another member of the 27:Extinct genus of lizards 2482:List of mosasaur genera 1750:Ancient Marine Reptiles 1541:Russell, D. A. (1967). 1424:and potentially in the 1369:preserving nine teeth. 1306:, having been named by 1302:is the type species of 929:Western Interior Seaway 1387: 903:are considered valid; 686: 624:Halisaurus sternbergii 540:The exact position of 487: 464:and larger species of 2604:Paleobiology Database 1754:Elizabeth L. Nicholls 1391:Halisaurus arambourgi 1382: 1374:Halisaurus arambourgi 1308:Othniel Charles Marsh 1191:Halisaurus arambourgi 1047:Eonatator sternbergii 987:phylogenetic analyses 808:Eonatator sternbergii 743:Halisaurus arambourgi 681: 634:Halisaurus arambourgi 616:being discovered and 481: 402:; "sea") and σαῦρος ( 372:Othniel Charles Marsh 18:Halisaurus arambourgi 1002:Eonatator coellensis 784:Eonatator coellensis 128:Mounted skeleton in 1700:Cretaceous Research 1609:2017PLoSO..1276773S 1469:Paleontology portal 1316:New Egypt Formation 1314:fragment) from the 974:, a new species of 378:, consisting of an 1756:page (p. 283) 1388: 1324:Navesink Formation 687: 488: 2619: 2618: 2518:Taxon identifiers 2509: 2508: 2342: 2341: 2242: 2241: 2185:Plesioplatecarpus 2128:Plioplatecarpinae 2102: 2101: 2035: 2034: 1449:H. platyspondylus 1445:H. platyspondylus 1410:H. platyspondylus 1289: 1288: 1280: 1279: 1271: 1270: 1262: 1261: 1253: 1252: 1244: 1243: 1235: 1234: 1226: 1225: 905:H. platyspondylus 896: 895: 887: 886: 878: 877: 869: 868: 860: 859: 682:Front-flipper of 614:H. platyspondylus 591:Plioplatecarpinae 546:H. platyspondylus 355: 354: 341:H. platyspondylus 267: 16:(Redirected from 2654: 2612: 2611: 2599: 2598: 2586: 2585: 2573: 2572: 2560: 2559: 2558: 2545: 2544: 2543: 2513: 2501: 2500: 2470:Related articles 2460:Vallecillosaurus 2162:Plioplatecarpini 2158: 2124: 2066: 2043: 1962:Prognathodontini 1910: 1896:Plesiotylosaurus 1784: 1777: 1770: 1761: 1737: 1736: 1734: 1732: 1718: 1712: 1711: 1691: 1685: 1684: 1655: 1649: 1648: 1638: 1620: 1588: 1582: 1578: 1572: 1571: 1569: 1568: 1562: 1556:. Archived from 1547: 1538: 1532: 1525: 1516: 1512: 1497: 1496: 1480: 1471: 1466: 1465: 1464: 1332:Severn Formation 1207: 1183: 1167:Halisaurus hebae 1159: 1140: 1123: 1104: 1087: 1063: 1039: 1029: 983:Dakhla Formation 966:In 2023, Shaker 933:Second World War 824: 800: 759: 735: 725: 672:Plesiopatecarpus 349: 337: 329: 317: 281: 259: 252: 239: 226: 213: 150: 149: 123: 113: 50: 40:Temporal range: 32: 21: 2662: 2661: 2657: 2656: 2655: 2653: 2652: 2651: 2622: 2621: 2620: 2615: 2607: 2602: 2594: 2589: 2581: 2576: 2568: 2563: 2554: 2553: 2548: 2539: 2538: 2533: 2520: 2510: 2505: 2491: 2465: 2453:Portunatasaurus 2389:Dolichosauridae 2354:Aigialosauridae 2338: 2310:Yaguarasaurinae 2304: 2269: 2238: 2197: 2192:Plioplatecarpus 2171:Latoplatecarpus 2156: 2122: 2108:Russellosaurina 2098: 2093:Phosphorosaurus 2064: 2044: 2031: 1983: 1956: 1908: 1825: 1793: 1788: 1740: 1730: 1728: 1726:fossilworks.org 1720: 1719: 1715: 1693: 1692: 1688: 1657: 1656: 1652: 1603:(5): e0176773. 1590: 1589: 1585: 1579: 1575: 1566: 1564: 1560: 1545: 1540: 1539: 1535: 1526: 1519: 1513: 1500: 1482: 1481: 1477: 1467: 1462: 1460: 1457: 1439:The fossils of 1377: 1297: 1290: 1281: 1272: 1263: 1254: 1245: 1236: 1227: 1134: 1101:Phosphorosaurus 1098: 1019:Phosphorosaurus 980:Late Cretaceous 917:H. onchognathus 897: 888: 879: 870: 861: 713:H. onchognathus 603:H. onchognathus 538: 525:Phosphorosaurus 503:H. onchognathus 482:Restoration of 476: 343: 335: 333:H. onchognathus 323: 311: 296: 287: 258: 250: 237: 224: 211: 144: 114: 112: 111: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 45: 44: 42:Late Cretaceous 38: 28: 23: 22: 15: 12: 11: 5: 2660: 2658: 2650: 2649: 2644: 2639: 2634: 2624: 2623: 2617: 2616: 2614: 2613: 2600: 2587: 2574: 2561: 2546: 2530: 2528: 2522: 2521: 2516: 2507: 2506: 2496: 2493: 2492: 2490: 2489: 2484: 2479: 2473: 2471: 2467: 2466: 2464: 2463: 2456: 2449: 2442: 2435: 2434: 2433: 2426: 2419: 2412: 2405: 2398: 2386: 2385: 2384: 2377: 2370: 2363: 2350: 2348: 2344: 2343: 2340: 2339: 2337: 2336: 2329: 2326:Russellosaurus 2322: 2314: 2312: 2306: 2305: 2303: 2302: 2295: 2288: 2279: 2277: 2271: 2270: 2268: 2267: 2260: 2257:Pannoniasaurus 2252: 2250: 2244: 2243: 2240: 2239: 2237: 2236: 2229: 2222: 2215: 2207: 2205: 2199: 2198: 2196: 2195: 2188: 2181: 2174: 2166: 2164: 2155: 2154: 2147: 2140: 2132: 2130: 2121: 2120: 2112: 2110: 2104: 2103: 2100: 2099: 2097: 2096: 2089: 2082: 2074: 2072: 2063: 2062: 2054: 2052: 2046: 2045: 2038: 2036: 2033: 2032: 2030: 2029: 2022: 2015: 2008: 2001: 1998:Bentiabasaurus 1993: 1991: 1985: 1984: 1982: 1981: 1974: 1966: 1964: 1958: 1957: 1955: 1954: 1947: 1944:Igdamanosaurus 1940: 1933: 1926: 1918: 1916: 1907: 1906: 1899: 1892: 1885: 1878: 1871: 1864: 1857: 1850: 1843: 1835: 1833: 1827: 1826: 1824: 1823: 1817: 1811: 1805: 1798: 1795: 1794: 1789: 1787: 1786: 1779: 1772: 1764: 1758: 1757: 1747: 1739: 1738: 1713: 1686: 1667:(3): 532–544. 1650: 1583: 1573: 1533: 1517: 1498: 1474: 1473: 1472: 1456: 1453: 1376: 1371: 1296: 1291: 1287: 1286: 1283: 1282: 1278: 1277: 1274: 1273: 1269: 1268: 1265: 1264: 1260: 1259: 1256: 1255: 1251: 1250: 1247: 1246: 1242: 1241: 1238: 1237: 1233: 1232: 1229: 1228: 1224: 1223: 1220: 1219: 1210: 1205: 1203: 1200: 1199: 1196: 1195: 1186: 1181: 1179: 1176: 1175: 1172: 1171: 1162: 1157: 1155: 1152: 1151: 1148: 1147: 1126: 1121: 1119: 1116: 1115: 1112: 1111: 1090: 1085: 1083: 1080: 1079: 1076: 1075: 1066: 1061: 1059: 1056: 1055: 1052: 1051: 1042: 1037: 1035: 1027: 915:. The species 894: 893: 890: 889: 885: 884: 881: 880: 876: 875: 872: 871: 867: 866: 863: 862: 858: 857: 854: 853: 844: 841: 840: 837: 836: 827: 822: 820: 817: 816: 813: 812: 803: 798: 796: 793: 792: 789: 788: 779: 776: 775: 772: 771: 762: 757: 755: 752: 751: 748: 747: 738: 733: 731: 723: 715:and the genus 537: 534: 522:revealed that 475: 472: 424:junior synonym 362:is an extinct 353: 352: 351: 350: 338: 330: 318: 303: 302: 298: 297: 288: 276: 275: 269: 268: 248: 244: 243: 235: 231: 230: 222: 218: 217: 209: 202: 201: 196: 192: 191: 186: 182: 181: 176: 172: 171: 166: 162: 161: 156: 152: 151: 138: 137: 125: 124: 116: 115: 107: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 52: 51: 39: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 2659: 2648: 2645: 2643: 2640: 2638: 2635: 2633: 2630: 2629: 2627: 2610: 2605: 2601: 2597: 2592: 2588: 2584: 2579: 2575: 2571: 2566: 2562: 2557: 2551: 2547: 2542: 2536: 2532: 2531: 2529: 2527: 2523: 2519: 2514: 2504: 2494: 2488: 2485: 2483: 2480: 2478: 2475: 2474: 2472: 2468: 2462: 2461: 2457: 2455: 2454: 2450: 2448: 2447: 2443: 2441: 2440: 2436: 2432: 2431: 2430:Tetrapodophis 2427: 2425: 2424: 2420: 2418: 2417: 2413: 2411: 2410: 2406: 2404: 2403: 2402:Dolichosaurus 2399: 2397: 2396: 2392: 2391: 2390: 2387: 2383: 2382: 2378: 2376: 2375: 2371: 2369: 2368: 2364: 2362: 2361: 2360:Aigialosaurus 2357: 2356: 2355: 2352: 2351: 2349: 2345: 2335: 2334: 2333:Yaguarasaurus 2330: 2328: 2327: 2323: 2321: 2320: 2316: 2315: 2313: 2311: 2307: 2301: 2300: 2296: 2294: 2293: 2292:Taniwhasaurus 2289: 2287: 2286: 2281: 2280: 2278: 2276: 2272: 2266: 2265: 2261: 2259: 2258: 2254: 2253: 2251: 2249: 2248:Tethysaurinae 2245: 2235: 2234: 2230: 2228: 2227: 2223: 2221: 2220: 2219:Goronyosaurus 2216: 2214: 2213: 2209: 2208: 2206: 2204: 2200: 2194: 2193: 2189: 2187: 2186: 2182: 2180: 2179: 2175: 2173: 2172: 2168: 2167: 2165: 2163: 2159: 2153: 2152: 2148: 2146: 2145: 2141: 2139: 2138: 2134: 2133: 2131: 2129: 2125: 2119: 2118: 2114: 2113: 2111: 2109: 2105: 2095: 2094: 2090: 2088: 2087: 2083: 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1563:on 2016-06-24 1559: 1555: 1551: 1544: 1537: 1534: 1531: 1524: 1522: 1518: 1511: 1509: 1507: 1505: 1503: 1499: 1494: 1490: 1486: 1479: 1476: 1470: 1459: 1454: 1452: 1450: 1446: 1442: 1441:H. arambourgi 1437: 1435: 1431: 1427: 1423: 1419: 1418:Maastrichtian 1415: 1414:H. arambourgi 1411: 1406: 1404: 1400: 1396: 1392: 1386: 1385:H. arambourgi 1381: 1375: 1372: 1370: 1368: 1364: 1359: 1357: 1356:Maastrichtian 1353: 1352:North America 1349: 1345: 1341: 1337: 1333: 1329: 1325: 1321: 1317: 1313: 1309: 1305: 1301: 1295: 1292: 1285: 1284: 1276: 1275: 1267: 1266: 1258: 1257: 1249: 1248: 1240: 1239: 1231: 1230: 1222: 1221: 1218: 1217: 1216: 1209: 1208: 1202: 1201: 1198: 1197: 1194: 1193: 1192: 1185: 1184: 1178: 1177: 1174: 1173: 1170: 1169: 1168: 1161: 1160: 1154: 1153: 1150: 1149: 1146: 1145: 1144: 1138: 1133: 1132: 1125: 1124: 1118: 1117: 1114: 1113: 1110: 1109: 1108: 1107:ponpetelegans 1102: 1097: 1096: 1089: 1088: 1082: 1081: 1078: 1077: 1074: 1073: 1072: 1065: 1064: 1058: 1057: 1054: 1053: 1050: 1049: 1048: 1041: 1040: 1034: 1031: 1030: 1026: 1024: 1023:ponpetelegans 1020: 1016: 1012: 1008: 1004: 1003: 998: 997: 992: 988: 984: 981: 977: 973: 969: 964: 962: 958: 954: 950: 949:Maastrichtian 946: 942: 938: 934: 930: 926: 922: 919:, known from 918: 914: 910: 909:H. arambourgi 906: 902: 892: 891: 883: 882: 874: 873: 865: 864: 856: 855: 852: 851: 850: 843: 842: 839: 838: 835: 834: 833: 826: 825: 819: 818: 815: 814: 811: 810: 809: 802: 801: 795: 794: 791: 790: 787: 786: 785: 778: 777: 774: 773: 770: 769: 768: 761: 760: 754: 753: 750: 749: 746: 745: 744: 737: 736: 730: 727: 726: 722: 720: 719: 714: 710: 706: 702: 701: 696: 692: 685: 684:H. arambourgi 680: 676: 674: 673: 668: 667: 662: 658: 654: 649: 647: 643: 639: 635: 631: 630: 625: 621: 620: 615: 611: 606: 604: 600: 596: 592: 588: 587: 582: 581: 576: 575: 569: 568: 563: 559: 555: 551: 547: 543: 535: 533: 531: 527: 526: 521: 520: 515: 511: 506: 504: 500: 496: 492: 485: 484:H. arambourgi 480: 473: 471: 469: 468: 463: 462: 457: 453: 452: 447: 442: 440: 436: 435:United States 432: 427: 425: 421: 417: 416: 411: 410: 405: 401: 397: 396:Ancient Greek 393: 389: 385: 381: 377: 374:in 1869. The 373: 369: 365: 361: 360: 347: 344:Marsh, 1869 ( 342: 339: 336:Merriam, 1894 334: 331: 327: 322: 319: 315: 310: 309:H. arambourgi 307: 306: 304: 299: 295: 291: 286: 285: 277: 274: 270: 266: 262: 257: 256: 249: 246: 245: 242: 236: 233: 232: 229: 223: 220: 219: 216: 210: 207: 204: 203: 200: 197: 194: 193: 190: 187: 184: 183: 180: 177: 174: 173: 170: 167: 164: 163: 160: 157: 154: 153: 148: 143: 139: 135: 131: 126: 122: 117: 110: 105: 100: 95: 90: 85: 80: 75: 70: 65: 60: 55: 49: 46:86.3–66  43: 37: 33: 30: 19: 2525: 2458: 2451: 2444: 2437: 2428: 2421: 2414: 2407: 2400: 2393: 2381:Opetiosaurus 2379: 2372: 2365: 2358: 2331: 2324: 2317: 2297: 2290: 2283: 2275:Tylosaurinae 2262: 2255: 2231: 2224: 2217: 2212:Gavialimimus 2210: 2203:Selmasaurini 2190: 2183: 2176: 2169: 2151:Sarabosaurus 2149: 2144:Ectenosaurus 2142: 2137:Angolasaurus 2135: 2117:Haasiasaurus 2115: 2091: 2085: 2084: 2077: 2057: 2050:Halisaurinae 2024: 2017: 2010: 2005:Eremiasaurus 2003: 1996: 1976: 1971:Prognathodon 1969: 1949: 1942: 1935: 1928: 1921: 1914:Globidensini 1901: 1894: 1887: 1880: 1873: 1866: 1861:Gnathomortis 1859: 1852: 1845: 1840:Amphekepubis 1838: 1831:Mosasaurinae 1749: 1743: 1729:. 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Retrieved 1558:the original 1553: 1549: 1536: 1484: 1478: 1448: 1444: 1440: 1438: 1413: 1409: 1407: 1399:North Africa 1390: 1389: 1384: 1383:Skeleton of 1373: 1360: 1340:Mount Laurel 1303: 1299: 1298: 1293: 1214: 1213: 1212: 1190: 1189: 1188: 1166: 1165: 1164: 1142: 1141: 1136: 1130: 1129: 1128: 1106: 1105: 1100: 1094: 1093: 1092: 1069: 1068: 1045: 1044: 1022: 1018: 1014: 1010: 1006: 1000: 994: 990: 975: 971: 967: 965: 936: 916: 912: 908: 904: 900: 898: 847: 846: 830: 829: 806: 805: 782: 781: 766: 765: 764: 742: 741: 740: 729:Halisaurinae 716: 712: 708: 705:Halisaurinae 698: 688: 683: 670: 664: 661:Mosasaurinae 656: 655:, recovered 650: 646:Halisaurinae 641: 637: 633: 627: 623: 617: 613: 609: 607: 602: 598: 595:Mosasaurinae 584: 578: 572: 565: 557: 545: 541: 539: 530:Halisaurinae 523: 517: 513: 509: 507: 502: 499:Halisaurinae 490: 489: 483: 465: 459: 455: 454:, the sleek 449: 445: 443: 439:Halisaurinae 428: 419: 413: 408: 407: 403: 399: 358: 357: 356: 340: 332: 325: 320: 313: 308: 283: 282: 273:Type species 254: 253: 228:Mosasauridae 205: 35: 29: 2647:Mosasaurids 2550:Wikispecies 2446:Judeasaurus 2439:Adriosaurus 2423:Pontosaurus 2395:Coniasaurus 2374:Komensaurus 2367:Carsosaurus 2319:Romeosaurus 2264:Tethysaurus 2233:Selmasaurus 2178:Platecarpus 2070:Halisaurini 2026:Plotosaurus 2012:Moanasaurus 1989:Mosasaurini 1868:Jormungandr 1854:Dallasaurus 1731:17 December 1432:in central 1426:Middle East 1403:Middle East 1312:basicranium 1033:Halisaurini 689:Below is a 599:Platecarpus 580:Platecarpus 567:Platecarpus 554:basicranium 474:Description 451:Dallasaurus 420:Baptosaurus 409:Baptosaurus 388:Hornerstown 384:basicranium 241:Halisaurini 130:Thermopolis 2626:Categories 2556:Halisaurus 2526:Halisaurus 2416:Primitivus 2299:Tylosaurus 2285:Kaikaifilu 2086:Halisaurus 2019:Mosasaurus 1923:Carinodens 1903:Stelladens 1875:Kourisodon 1702:. 105719. 1696:Halisaurus 1567:2017-08-19 1455:References 1328:New Jersey 1320:New Jersey 1304:Halisaurus 1143:coellensis 1131:Halisaurus 1095:Halisaurus 1011:Halisaurus 1007:Halisaurus 991:Halisaurus 976:Halisaurus 953:California 937:Halisaurus 901:Halisaurus 709:Halisaurus 707:, showing 666:Tylosaurus 657:Halisaurus 642:Halisaurus 610:Halisaurus 542:Halisaurus 514:Halisaurus 510:Halisaurus 491:Halisaurus 461:Tylosaurus 456:Halisaurus 446:Halisaurus 415:Halosaurus 392:New Jersey 359:Halisaurus 255:Halisaurus 215:Mosasauria 36:Halisaurus 2409:Kaganaias 2226:Khinjaria 2079:Eonatator 2059:Pluridens 1930:Globidens 1847:Clidastes 1801:Kingdom: 1791:Mosasaurs 1681:0272-4634 1627:1932-6203 1430:Khouribga 1367:pterygoid 1137:Eonatator 978:from the 957:Santonian 941:Campanian 925:Santonian 921:Campanian 718:Pluridens 691:cladogram 638:Eonatator 629:Eonatator 586:Clidastes 574:Clidastes 495:Santonian 467:Clidastes 370:named by 165:Kingdom: 159:Eukaryota 2535:Wikidata 2503:Category 1951:Xenodens 1821:Squamata 1815:Reptilia 1809:Chordata 1807:Phylum: 1803:Animalia 1645:28467456 1597:PLOS ONE 1581:129–136. 1515:447–472. 1416:is Late 1401:and the 1363:quadrate 1348:Delaware 1336:Maryland 972:H. hebae 970:. named 955:and the 913:H. hebae 433:and the 376:holotype 368:mosasaur 321:H. hebae 301:Species 221:Family: 199:Squamata 189:Reptilia 179:Chordata 175:Phylum: 169:Animalia 155:Domain: 2596:1444477 2583:4819779 2570:4518658 2541:Q134622 1819:Order: 1813:Class: 1636:5415187 1605:Bibcode 1434:Morocco 1013:, with 675:) are. 550:angular 431:Morocco 380:angular 328:., 2023 324:Shaker 312:Bardet 247:Genus: 234:Tribe: 195:Order: 185:Class: 134:Wyoming 1882:Liodon 1679:  1643:  1633:  1625:  1422:Africa 947:, the 911:, and 552:and a 404:saûros 382:and a 316:, 2005 314:et al. 2609:36399 2591:IRMNG 1561:(PDF) 1546:(PDF) 1408:Like 968:et al 945:Texas 398:ἅλς ( 364:genus 326:et al 290:Marsh 261:Marsh 206:Clade 2578:GBIF 1733:2021 1677:ISSN 1641:PMID 1623:ISSN 1342:and 999:and 961:Peru 695:2015 669:and 653:2017 593:and 583:and 562:1967 422:" a 400:háls 346:type 294:1869 265:1869 54:PreꞒ 2565:EoL 1704:doi 1669:doi 1631:PMC 1613:doi 1489:doi 1451:). 1346:of 1334:of 1021:") 989:of 959:of 951:of 943:of 923:or 366:of 2628:: 2606:: 2593:: 2580:: 2567:: 2552:: 2537:: 1724:. 1675:. 1665:15 1663:. 1639:. 1629:. 1621:. 1611:. 1601:12 1599:. 1595:. 1554:23 1552:. 1548:. 1520:^ 1501:^ 1436:. 1412:, 1405:. 1358:. 1326:, 1139:") 1135:(" 1103:") 1099:(" 1017:(" 1015:H. 907:, 605:. 470:. 441:. 426:. 390:, 292:, 263:, 208:: 136:. 132:, 104:Pg 48:Ma 2282:? 1783:e 1776:t 1769:v 1735:. 1710:. 1706:: 1683:. 1671:: 1647:. 1615:: 1607:: 1570:. 1495:. 1491:: 1318:( 721:. 560:( 348:) 280:† 251:† 238:† 225:† 212:† 109:N 99:K 94:J 89:T 84:P 79:C 74:D 69:S 64:O 59:Ꞓ 20:)

Index

Halisaurus arambourgi
Late Cretaceous
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Thermopolis
Wyoming
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Reptilia
Squamata
Mosasauria
Mosasauridae
Halisaurini
Halisaurus
Marsh

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