2852:
2550:
142:
118:
2562:
2885:
2556:
496:
2029:
2034:
2950:
1110:
360:
2937:
3009:
1241:
3014:
1257:
977:
Benson, R.J. (2012). "Interrelationships of basal synapsids: cranial and postcranial morphological partitions suggest different topologies".
1294:
2851:
1182:"A Carboniferous synapsid with caniniform teeth and a reappraisal of mandibular size-shape heterodonty in the origin of mammals"
1077:
317:
and hence, mammals. It was at least 1.5 metres (5 ft) in length. It lived in present-day France during the Early
2561:
2549:
618:
belong to different taxa; one of these, ROM 43608, was named by
Spindler in 2020 as a new genus and species,
141:
1373:
919:
Laurin, M. (1993). "Anatomy and relationships of
Haptodus garnettensis, a Pennsylvanian synapsid from Kansas".
1124:
Spindler, F. (2019). "Re-evaluation of an early sphenacodontian synapsid from the Lower
Permian of England".
1400:
1287:
556:
536:
3004:
2905:
2753:
2555:
2028:
2976:
2660:
2634:
620:
559:
501:
394:
379:
307:
278:
544:
2718:
2686:
1249:
1193:
220:
2033:
2776:
1265:
2888:
2435:
2294:
2247:
1280:
888:"A new basal sphenacodontid synapsid from the Late Carboniferous of the Saar-Nahe Basin, Germany"
443:
136:
592:
In his 2015 dissertation revising the basal
Sphenacodontia, Frederik Spindler demonstrated that
2981:
2963:
2955:
2375:
2348:
1219:
1100:
810:
563:
548:
383:
2968:
535:
stages were referred to this species. All specimens came from the
Garnett Quarry, located in
2405:
2144:
2004:
1647:
1209:
1201:
1160:
1129:
1058:
1025:
986:
959:
928:
899:
2471:
2314:
2254:
2114:
2070:
1358:
678:
610:
398:
117:
1197:
2799:
2730:
2428:
2357:
2277:
2240:
2198:
2189:
1941:
1845:
1790:
1779:
1736:
1702:
1612:
1487:
1385:
1214:
1181:
770:
715:
703:
690:
650:
516:
419:
310:
199:
495:
2998:
2873:
2742:
2501:
2493:
2414:
2364:
2334:
2324:
2261:
2121:
2079:
1876:
1861:
1834:
1813:
1675:
1668:
1584:
1106:
794:
425:
403:
68:
30:
2878:
2868:
2832:
2692:
2649:
2623:
2485:
2421:
2226:
2212:
2205:
2128:
2100:
1827:
1743:
1682:
1626:
1619:
1577:
1570:
1563:
1512:
1062:
963:
932:
476:
456:
452:
409:
352:
266:
228:
1014:, two historical and previously mistaken basal caseasaurian synapsids from Europe"
990:
2707:
2675:
2478:
2457:
2233:
2093:
2086:
2051:
1994:
1984:
1912:
1820:
1715:
1543:
1521:
1502:
663:
528:
330:
43:
487:
might be a nomen dubium. His revision has been accepted by subsequent authors.
2822:
2812:
2765:
2592:
2579:
2513:
2443:
2391:
2175:
2107:
1954:
1869:
1854:
1764:
1757:
1729:
1654:
1640:
1133:
1126:
Earth and
Environmental Science Transactions of the Royal Society of Edinburgh
582:
578:
552:
531:) of an immature individual. More than 16 well-preserved specimens of various
390:
376:
256:
88:
53:
2928:
2609:
2464:
2165:
1890:
1750:
1661:
1605:
1598:
1536:
1466:
1453:
1412:
1303:
886:
Fröbisch, J.; Schoch, R.R.; Müller, J.; Schindler, T.; Schweiss, D. (2011).
746:
636:
314:
186:
153:
93:
37:
1223:
1102:
The basal
Sphenacodontia: systematic revision and evolutionary implications
1030:
1005:
483:
was not examined. Laurin also suggested that the type species of the genus
904:
887:
441:
in 1977, Currie (1979) synonymized all
European haptodontines, as well as
2922:
2450:
2219:
1974:
1964:
1897:
1883:
1806:
1799:
1554:
1529:
1333:
1327:
1180:
Huttenlocker, A. K.; Singh, S. A.; Henrici, A. C.; Sumida, S. S. (2021).
532:
520:
513:
372:
322:
173:
129:
83:
78:
63:
58:
48:
1205:
451:. Later, Laurin (1993 and 1994) considered Haptodontinae to represent a
2942:
1633:
1342:
586:
574:
524:
430:
326:
318:
300:
98:
73:
359:, is known only from a single, badly preserved specimen hosted in the
2782:
1772:
1321:
540:
414:
368:
163:
2899:
1591:
1165:
1148:
494:
364:
303:
437:, but retained these taxa as distinct species. After describing
2903:
2577:
2049:
1451:
1314:
1276:
1149:"A faunivorous early sphenacodontian synapsid with a diastema"
1272:
604:
and instead represent distinct genera. Spindler referred
614:
in 2019. He also showed that some specimens referred to
1078:"Lower Permian Pelycosaurs from the English Midlands"
2828:
2818:
2808:
2739:
2704:
2672:
2646:
2620:
2605:
2402:
2387:
2345:
2330:
2320:
2310:
2300:
2290:
2186:
2171:
2161:
2067:
2000:
1990:
1980:
1970:
1960:
1950:
1712:
1499:
1483:
273:
235:
2912:
2797:
2764:
2738:
2729:
2703:
2671:
2645:
2619:
2590:
2512:
2401:
2374:
2344:
2275:
2185:
2142:
2066:
1939:
1911:
1844:
1789:
1711:
1700:
1553:
1511:
1498:
1464:
1411:
1384:
1357:
1004:Spindler, F.; Falconnet, J.; Fröbisch, J. (2015).
881:
879:
1045:Laurin, M. (1993). "Anatomy and relationships of
401:(1940). They synonymized many genera, including
363:of Paris. It was collected at Les Télots, near
1288:
8:
1111:Freiberg University of Mining and Technology
950:, an Early Permian synapsid from Colorado".
371:, from a terrestrial horizon dating to the
2900:
2735:
2598:
2587:
2574:
2380:
2283:
2154:
2150:
2063:
2059:
2046:
1936:
1721:
1708:
1508:
1476:
1472:
1461:
1448:
1390:
1363:
1354:
1311:
1295:
1281:
1273:
116:
20:
1213:
1164:
1049:, a Pennsylvanian synapsid from Kansas".
1029:
903:
577:(Gz 1071) found in Pennsylvanian to
875:
505:, which may belong in a different genus
313:, a member of the clade that includes
1242:"Synapsida: Sphenacodontia: Haptodus"
946:Laurin, M. (1994). "Re-evaluation of
519: 14156, a partially articulated
7:
551:(Lansing Group), dating to the late
407:(from Margenne near Autun, France),
361:Muséum National d'Histoire Naturelle
979:Journal of Systematic Palaeontology
786:
762:
721:
696:
656:
646:
600:do not belong in the same genus as
1051:Journal of Vertebrate Paleontology
952:Journal of Vertebrate Paleontology
921:Journal of Vertebrate Paleontology
14:
2884:
2883:
2850:
2560:
2554:
2548:
2032:
2027:
140:
543:. They were collected from the
3010:Cisuralian synapsids of Europe
1063:10.1080/02724634.1993.10011501
964:10.1080/02724634.1994.10011544
933:10.1080/02724634.1993.10011501
447:, with the type of the genus,
1:
3015:Fossil taxa described in 1886
1018:Acta Palaeontologica Polonica
892:Acta Palaeontologica Polonica
991:10.1080/14772019.2011.631042
589:, by Roberta Paton in 1974.
3031:
1186:Royal Society Open Science
1153:Palaeontologia Electronica
393:was revised in details by
16:Extinct genus of synapsids
2863:
2848:
2601:
2586:
2573:
2546:
2383:
2286:
2157:
2153:
2062:
2058:
2045:
2025:
1935:
1724:
1479:
1475:
1460:
1447:
1393:
1366:
1353:
1310:
1134:10.1017/S175569101900015X
808:
791:
784:
767:
760:
743:
726:
719:
701:
694:
676:
661:
654:
253:
248:
234:
227:
137:Scientific classification
135:
124:
115:
23:
321:. It was a medium-sized
562:, about 305-303.9
512:was named based on the
502:"Haptodus" garnettensis
475:was considered to be a
382:, about 299-296.4
1031:10.4202/app.00221.2015
506:
337:Discovery and taxonomy
2977:Paleobiology Database
1147:Spindler, F. (2020).
1099:Spindler, F. (2015).
1047:Haptodus garnettensis
905:10.4202/app.2010.0039
621:Kenomagnathus scottae
510:Haptodus garnettensis
498:
395:Alfred Sherwood Romer
389:The taxonomy of many
1076:Paton, R.L. (1974).
125:Illustration of the
1206:10.1098/rsos.211237
1198:2021RSOS....811237H
639:after Huttenlocker
573:, was named from a
413:(from Nowa Ruda of
948:Cutleria wilmarthi
555:stage of the late
507:
459:, and revalidated
444:Cutleria wilmarthi
2992:
2991:
2964:Open Tree of Life
2906:Taxon identifiers
2897:
2896:
2859:
2858:
2846:
2845:
2842:
2841:
2793:
2792:
2569:
2568:
2544:
2543:
2540:
2539:
2536:
2535:
2532:
2531:
2376:Sphenacodontoidea
2349:Palaeohatteriidae
2271:
2270:
2138:
2137:
2041:
2040:
2023:
2022:
2019:
2018:
2015:
2014:
1931:
1930:
1907:
1906:
1696:
1695:
1692:
1691:
1443:
1442:
1439:
1438:
1435:
1434:
1431:
1430:
866:
865:
857:
856:
848:
847:
839:
838:
830:
829:
821:
820:
811:Sphenacodontoidea
616:"H." garnettensis
569:Another species,
564:million years old
549:Stanton Formation
384:million years old
292:
291:
286:
279:"H". garnettensis
270:
223:
3022:
2985:
2984:
2972:
2971:
2959:
2958:
2946:
2945:
2933:
2932:
2931:
2901:
2887:
2886:
2854:
2830:
2820:
2810:
2741:
2736:
2706:
2674:
2648:
2622:
2607:
2599:
2588:
2575:
2564:
2558:
2552:
2406:Sphenacodontidae
2404:
2389:
2381:
2347:
2332:
2322:
2312:
2302:
2292:
2284:
2188:
2173:
2163:
2155:
2151:
2145:Haptodontiformes
2069:
2064:
2060:
2047:
2036:
2031:
2005:Protoclepsydrops
2002:
1992:
1982:
1972:
1962:
1952:
1937:
1722:
1714:
1709:
1648:Lalieudorhynchus
1509:
1501:
1485:
1477:
1473:
1462:
1449:
1391:
1364:
1355:
1348:
1347:
1312:
1297:
1290:
1283:
1274:
1269:
1264:. Archived from
1253:
1248:. Archived from
1228:
1227:
1217:
1177:
1171:
1170:
1168:
1144:
1138:
1137:
1121:
1115:
1114:
1096:
1090:
1089:
1073:
1067:
1066:
1042:
1036:
1035:
1033:
1001:
995:
994:
974:
968:
967:
943:
937:
936:
916:
910:
909:
907:
883:
787:
763:
722:
697:
657:
647:
571:Haptodus grandis
545:Rock Lake Member
284:
275:
264:
237:
218:
211:
145:
144:
120:
110:
40:
29:Temporal range:
21:
3030:
3029:
3025:
3024:
3023:
3021:
3020:
3019:
2995:
2994:
2993:
2988:
2980:
2975:
2967:
2962:
2954:
2949:
2941:
2936:
2927:
2926:
2921:
2908:
2898:
2893:
2855:
2838:
2789:
2760:
2725:
2699:
2667:
2641:
2615:
2595:
2582:
2565:
2528:
2508:
2472:Secodontosaurus
2397:
2370:
2340:
2315:Hypselohaptodus
2280:
2267:
2255:Remigiomontanus
2181:
2147:
2134:
2115:Stereophallodon
2071:Ophiacodontidae
2054:
2037:
2011:
1927:
1903:
1840:
1785:
1705:
1688:
1549:
1494:
1469:
1456:
1427:
1407:
1380:
1374:Tetrapodomorpha
1359:Tetrapodomorpha
1349:
1317:
1316:
1306:
1301:
1256:
1240:
1237:
1232:
1231:
1179:
1178:
1174:
1146:
1145:
1141:
1123:
1122:
1118:
1098:
1097:
1093:
1075:
1074:
1070:
1044:
1043:
1039:
1003:
1002:
998:
976:
975:
971:
945:
944:
940:
918:
917:
913:
885:
884:
877:
872:
867:
858:
849:
840:
831:
822:
679:Ophiacodontidae
630:
611:Hypselohaptodus
608:to a new genus
594:H. garnettensis
537:Anderson County
493:
455:grade of basal
439:H. garnettensis
399:Llewellyn Price
349:Haptodus baylei
346:
343:Haptodus baylei
339:
244:
241:
239:Haptodus baylei
217:
209:
139:
111:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
36:299–296.4
35:
34:
27:
17:
12:
11:
5:
3028:
3026:
3018:
3017:
3012:
3007:
2997:
2996:
2990:
2989:
2987:
2986:
2973:
2960:
2947:
2934:
2918:
2916:
2910:
2909:
2904:
2895:
2894:
2892:
2891:
2881:
2876:
2871:
2864:
2861:
2860:
2857:
2856:
2849:
2847:
2844:
2843:
2840:
2839:
2837:
2836:
2826:
2816:
2805:
2803:
2800:Incertae sedis
2795:
2794:
2791:
2790:
2788:
2787:
2786:
2785:
2779:
2770:
2768:
2762:
2761:
2759:
2758:
2757:
2756:
2747:
2745:
2733:
2731:Eutheriodontia
2727:
2726:
2724:
2723:
2722:
2721:
2712:
2710:
2701:
2700:
2698:
2697:
2696:
2695:
2689:
2680:
2678:
2669:
2668:
2666:
2665:
2664:
2663:
2654:
2652:
2643:
2642:
2640:
2639:
2638:
2637:
2628:
2626:
2617:
2616:
2614:
2613:
2602:
2596:
2591:
2584:
2583:
2578:
2571:
2570:
2567:
2566:
2547:
2545:
2542:
2541:
2538:
2537:
2534:
2533:
2530:
2529:
2527:
2526:
2525:
2524:
2518:
2516:
2510:
2509:
2507:
2506:
2498:
2490:
2482:
2475:
2468:
2461:
2454:
2447:
2440:
2432:
2429:Ctenospondylus
2425:
2418:
2410:
2408:
2399:
2398:
2396:
2395:
2384:
2378:
2372:
2371:
2369:
2368:
2361:
2358:Palaeohatteria
2353:
2351:
2342:
2341:
2339:
2338:
2328:
2318:
2308:
2298:
2287:
2281:
2278:Sphenacodontia
2276:
2273:
2272:
2269:
2268:
2266:
2265:
2258:
2251:
2244:
2241:Melanedaphodon
2237:
2230:
2223:
2216:
2209:
2202:
2199:Bohemiclavulus
2194:
2192:
2190:Edaphosauridae
2183:
2182:
2180:
2179:
2169:
2158:
2148:
2143:
2140:
2139:
2136:
2135:
2133:
2132:
2125:
2118:
2111:
2104:
2097:
2090:
2083:
2075:
2073:
2056:
2055:
2050:
2043:
2042:
2039:
2038:
2026:
2024:
2021:
2020:
2017:
2016:
2013:
2012:
2010:
2009:
1998:
1988:
1978:
1968:
1958:
1947:
1945:
1942:Incertae sedis
1933:
1932:
1929:
1928:
1926:
1925:
1924:
1923:
1917:
1915:
1909:
1908:
1905:
1904:
1902:
1901:
1894:
1887:
1880:
1873:
1866:
1858:
1850:
1848:
1846:Varanodontinae
1842:
1841:
1839:
1838:
1831:
1824:
1817:
1810:
1803:
1795:
1793:
1791:Mesenosaurinae
1787:
1786:
1784:
1783:
1780:Thrausmosaurus
1776:
1769:
1761:
1754:
1747:
1740:
1737:Archaeovenator
1733:
1725:
1719:
1706:
1703:Eupelycosauria
1701:
1698:
1697:
1694:
1693:
1690:
1689:
1687:
1686:
1679:
1672:
1665:
1658:
1651:
1644:
1637:
1630:
1623:
1616:
1613:Cotylorhynchus
1609:
1602:
1595:
1588:
1581:
1574:
1567:
1559:
1557:
1551:
1550:
1548:
1547:
1540:
1533:
1526:
1517:
1515:
1506:
1496:
1495:
1493:
1492:
1488:Diadectomorpha
1480:
1470:
1465:
1458:
1457:
1452:
1445:
1444:
1441:
1440:
1437:
1436:
1433:
1432:
1429:
1428:
1426:
1425:
1424:
1423:
1417:
1415:
1409:
1408:
1406:
1405:
1404:
1403:
1401:Reptiliomorpha
1394:
1388:
1386:Reptiliomorpha
1382:
1381:
1379:
1378:
1377:
1376:
1367:
1361:
1351:
1350:
1346:
1345:
1336:
1330:
1324:
1315:
1308:
1307:
1302:
1300:
1299:
1292:
1285:
1277:
1271:
1270:
1268:on 2023-04-18.
1254:
1252:on 2006-01-02.
1236:
1235:External links
1233:
1230:
1229:
1192:(12): 211237.
1172:
1139:
1116:
1091:
1068:
1057:(2): 200–229.
1037:
1024:(3): 597–616.
996:
985:(4): 601–624.
969:
958:(1): 134–138.
938:
927:(2): 200–229.
911:
898:(1): 113–120.
874:
873:
871:
868:
864:
863:
860:
859:
855:
854:
851:
850:
846:
845:
842:
841:
837:
836:
833:
832:
828:
827:
824:
823:
819:
818:
815:
814:
807:
804:
803:
800:
799:
790:
785:
783:
780:
779:
776:
775:
771:Palaeohatteria
766:
761:
759:
756:
755:
752:
751:
742:
739:
738:
735:
734:
725:
720:
718:
716:Sphenacodontia
712:
711:
708:
707:
704:Edaphosauridae
700:
695:
693:
691:Sphenacomorpha
687:
686:
683:
682:
675:
672:
671:
668:
667:
660:
655:
653:
651:Eupelycosauria
645:
629:
626:
492:
489:
465:Palaeohatteria
420:Palaeohatteria
345:
340:
338:
335:
290:
289:
288:
287:
271:
265:Gaudry, 1886 (
251:
250:
246:
245:
242:
232:
231:
225:
224:
207:
203:
202:
200:Sphenacodontia
197:
190:
189:
184:
177:
176:
171:
167:
166:
161:
157:
156:
151:
147:
146:
133:
132:
122:
121:
113:
112:
104:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
41:
28:
15:
13:
10:
9:
6:
4:
3:
2:
3027:
3016:
3013:
3011:
3008:
3006:
3005:Sphenacodonts
3003:
3002:
3000:
2983:
2978:
2974:
2970:
2965:
2961:
2957:
2952:
2948:
2944:
2939:
2935:
2930:
2924:
2920:
2919:
2917:
2915:
2911:
2907:
2902:
2890:
2882:
2880:
2877:
2875:
2874:Phthinosuchia
2872:
2870:
2866:
2865:
2862:
2853:
2835:
2834:
2827:
2825:
2824:
2817:
2815:
2814:
2807:
2806:
2804:
2802:
2801:
2796:
2784:
2780:
2778:
2774:
2773:
2772:
2771:
2769:
2767:
2763:
2755:
2754:Therocephalia
2751:
2750:
2749:
2748:
2746:
2744:
2743:Therocephalia
2737:
2734:
2732:
2728:
2720:
2716:
2715:
2714:
2713:
2711:
2709:
2702:
2694:
2690:
2688:
2684:
2683:
2682:
2681:
2679:
2677:
2670:
2662:
2658:
2657:
2656:
2655:
2653:
2651:
2644:
2636:
2632:
2631:
2630:
2629:
2627:
2625:
2618:
2612:
2611:
2604:
2603:
2600:
2597:
2594:
2589:
2585:
2581:
2576:
2572:
2563:
2559:
2557:
2551:
2522:
2521:
2520:
2519:
2517:
2515:
2511:
2504:
2503:
2502:Tetraceratops
2499:
2496:
2495:
2494:Tappenosaurus
2491:
2488:
2487:
2483:
2481:
2480:
2476:
2474:
2473:
2469:
2467:
2466:
2462:
2460:
2459:
2455:
2453:
2452:
2448:
2446:
2445:
2441:
2438:
2437:
2433:
2431:
2430:
2426:
2424:
2423:
2419:
2417:
2416:
2415:Cryptovenator
2412:
2411:
2409:
2407:
2400:
2394:
2393:
2386:
2385:
2382:
2379:
2377:
2373:
2367:
2366:
2365:Pantelosaurus
2362:
2360:
2359:
2355:
2354:
2352:
2350:
2343:
2337:
2336:
2335:Tetraceratops
2329:
2327:
2326:
2325:Kenomagnathus
2319:
2317:
2316:
2309:
2307:
2306:
2299:
2297:
2296:
2289:
2288:
2285:
2282:
2279:
2274:
2264:
2263:
2262:Xyrospondylus
2259:
2257:
2256:
2252:
2250:
2249:
2245:
2243:
2242:
2238:
2236:
2235:
2231:
2229:
2228:
2224:
2222:
2221:
2217:
2215:
2214:
2210:
2208:
2207:
2203:
2201:
2200:
2196:
2195:
2193:
2191:
2184:
2178:
2177:
2170:
2168:
2167:
2160:
2159:
2156:
2152:
2149:
2146:
2141:
2131:
2130:
2126:
2124:
2123:
2122:Stereorhachis
2119:
2117:
2116:
2112:
2110:
2109:
2105:
2103:
2102:
2098:
2096:
2095:
2091:
2089:
2088:
2084:
2082:
2081:
2080:Archaeothyris
2077:
2076:
2074:
2072:
2065:
2061:
2057:
2053:
2048:
2044:
2035:
2030:
2007:
2006:
1999:
1997:
1996:
1989:
1987:
1986:
1979:
1977:
1976:
1969:
1967:
1966:
1959:
1957:
1956:
1949:
1948:
1946:
1944:
1943:
1938:
1934:
1921:
1920:
1919:
1918:
1916:
1914:
1910:
1900:
1899:
1895:
1893:
1892:
1888:
1886:
1885:
1881:
1879:
1878:
1877:Tambacarnifex
1874:
1872:
1871:
1867:
1864:
1863:
1862:Mycterosaurus
1859:
1857:
1856:
1852:
1851:
1849:
1847:
1843:
1837:
1836:
1835:Microvaranops
1832:
1830:
1829:
1825:
1823:
1822:
1818:
1816:
1815:
1814:Elliotsmithia
1811:
1809:
1808:
1804:
1802:
1801:
1797:
1796:
1794:
1792:
1788:
1782:
1781:
1777:
1775:
1774:
1770:
1767:
1766:
1762:
1760:
1759:
1755:
1753:
1752:
1748:
1746:
1745:
1741:
1739:
1738:
1734:
1732:
1731:
1727:
1726:
1723:
1720:
1717:
1710:
1707:
1704:
1699:
1685:
1684:
1680:
1678:
1677:
1676:Ruthenosaurus
1673:
1671:
1670:
1669:Phreatophasma
1666:
1664:
1663:
1659:
1657:
1656:
1652:
1650:
1649:
1645:
1643:
1642:
1638:
1636:
1635:
1631:
1629:
1628:
1624:
1622:
1621:
1617:
1615:
1614:
1610:
1608:
1607:
1603:
1601:
1600:
1596:
1594:
1593:
1589:
1587:
1586:
1585:Callibrachion
1582:
1580:
1579:
1575:
1573:
1572:
1568:
1566:
1565:
1561:
1560:
1558:
1556:
1552:
1546:
1545:
1541:
1539:
1538:
1534:
1532:
1531:
1527:
1524:
1523:
1519:
1518:
1516:
1514:
1510:
1507:
1504:
1497:
1490:
1489:
1482:
1481:
1478:
1474:
1471:
1468:
1463:
1459:
1455:
1450:
1446:
1421:
1420:
1419:
1418:
1416:
1414:
1410:
1402:
1398:
1397:
1396:
1395:
1392:
1389:
1387:
1383:
1375:
1371:
1370:
1369:
1368:
1365:
1362:
1360:
1356:
1352:
1344:
1340:
1337:
1335:
1331:
1329:
1325:
1323:
1319:
1318:
1313:
1309:
1305:
1298:
1293:
1291:
1286:
1284:
1279:
1278:
1275:
1267:
1263:
1259:
1258:"Pelycosaurs"
1255:
1251:
1247:
1243:
1239:
1238:
1234:
1225:
1221:
1216:
1211:
1207:
1203:
1199:
1195:
1191:
1187:
1183:
1176:
1173:
1167:
1166:10.26879/1023
1162:
1158:
1154:
1150:
1143:
1140:
1135:
1131:
1127:
1120:
1117:
1112:
1108:
1107:Dr. rer. nat.
1104:
1103:
1095:
1092:
1088:(3): 541–552.
1087:
1083:
1082:Palaeontology
1079:
1072:
1069:
1064:
1060:
1056:
1052:
1048:
1041:
1038:
1032:
1027:
1023:
1019:
1015:
1013:
1009:
1008:Callibrachion
1000:
997:
992:
988:
984:
980:
973:
970:
965:
961:
957:
953:
949:
942:
939:
934:
930:
926:
922:
915:
912:
906:
901:
897:
893:
889:
882:
880:
876:
869:
862:
861:
853:
852:
844:
843:
835:
834:
826:
825:
817:
816:
813:
812:
806:
805:
802:
801:
798:
797:
796:
795:Pantelosaurus
789:
788:
782:
781:
778:
777:
774:
773:
772:
765:
764:
758:
757:
754:
753:
750:
749:
748:
741:
740:
737:
736:
733:
732:
731:
724:
723:
717:
714:
713:
710:
709:
706:
705:
699:
698:
692:
689:
688:
685:
684:
681:
680:
674:
673:
670:
669:
666:
665:
659:
658:
652:
649:
648:
644:
642:
638:
634:
627:
625:
623:
622:
617:
613:
612:
607:
603:
599:
595:
590:
588:
584:
580:
576:
572:
567:
565:
561:
558:
557:Pennsylvanian
554:
550:
546:
542:
538:
534:
530:
526:
522:
518:
515:
511:
504:
503:
497:
491:Other species
490:
488:
486:
482:
478:
474:
473:Callibrachion
470:
469:Pantelosaurus
466:
462:
458:
457:sphenacodonts
454:
450:
446:
445:
440:
436:
432:
428:
427:
426:Pantelosaurus
422:
421:
416:
412:
411:
406:
405:
404:Callibrachion
400:
396:
392:
387:
385:
381:
378:
375:stage of the
374:
370:
366:
362:
358:
354:
350:
344:
341:
336:
334:
332:
328:
325:, feeding on
324:
320:
316:
312:
311:sphenacodonts
309:
305:
302:
298:
297:
282:
281:
280:
272:
268:
263:
262:
258:
255:
254:
252:
247:
240:
233:
230:
226:
222:
216:
215:
208:
205:
204:
201:
198:
195:
192:
191:
188:
185:
182:
179:
178:
175:
172:
169:
168:
165:
162:
159:
158:
155:
152:
149:
148:
143:
138:
134:
131:
128:
123:
119:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
31:Early Permian
26:
22:
19:
2913:
2879:Theriodontia
2869:Pelycosauria
2833:Niaftasuchus
2831:
2821:
2811:
2798:
2661:Dinocephalia
2650:Dinocephalia
2635:Biarmosuchia
2624:Biarmosuchia
2608:
2553:
2500:
2492:
2486:Steppesaurus
2484:
2477:
2470:
2463:
2456:
2449:
2442:
2434:
2427:
2422:Ctenorhachis
2420:
2413:
2390:
2363:
2356:
2333:
2323:
2313:
2304:
2303:
2293:
2260:
2253:
2246:
2239:
2232:
2227:Ianthasaurus
2225:
2218:
2213:Glaucosaurus
2211:
2206:Edaphosaurus
2204:
2197:
2174:
2164:
2129:Varanosaurus
2127:
2120:
2113:
2106:
2101:Echinerpeton
2099:
2092:
2085:
2078:
2003:
1993:
1983:
1973:
1963:
1953:
1940:
1896:
1889:
1882:
1875:
1868:
1860:
1853:
1833:
1828:Mesenosaurus
1826:
1819:
1812:
1805:
1798:
1778:
1771:
1763:
1756:
1749:
1744:Ascendonanus
1742:
1735:
1728:
1683:Trichasaurus
1681:
1674:
1667:
1660:
1653:
1646:
1639:
1632:
1627:Ennatosaurus
1625:
1620:Datheosaurus
1618:
1611:
1604:
1597:
1590:
1583:
1578:Arisierpeton
1576:
1571:Angelosaurus
1569:
1564:Alierasaurus
1562:
1542:
1535:
1528:
1520:
1513:Eothyrididae
1486:
1338:
1332:Superclass:
1266:the original
1261:
1250:the original
1245:
1189:
1185:
1175:
1156:
1152:
1142:
1125:
1119:
1101:
1094:
1085:
1081:
1071:
1054:
1050:
1046:
1040:
1021:
1017:
1012:Datheosaurus
1011:
1007:
999:
982:
978:
972:
955:
951:
947:
941:
924:
920:
914:
895:
891:
809:
793:
792:
769:
768:
745:
744:
729:
728:
727:
702:
677:
662:
640:
632:
631:
619:
615:
609:
606:"H." grandis
605:
601:
597:
593:
591:
581:deposits in
570:
568:
509:
508:
500:
499:The species
484:
481:Datheosaurus
480:
477:nomen dubium
472:
468:
464:
460:
453:polyphyletic
448:
442:
438:
434:
424:
418:
410:Datheosaurus
408:
402:
388:
356:
353:type species
348:
347:
342:
295:
294:
293:
285:Currie, 1977
277:
276:
260:
259:
243:Gaudry, 1886
238:
229:Type species
213:
212:
193:
180:
126:
24:
18:
2719:Gorgonopsia
2708:Gorgonopsia
2693:dicynodonts
2687:Anomodontia
2676:Anomodontia
2479:Sphenacodon
2458:Macromerion
2248:Ramodendron
2234:Lupeosaurus
2094:Clepsydrops
2087:Baldwinonus
2052:Metopophora
1995:Mastersonia
1985:Knoxosaurus
1913:Metopophora
1821:Heleosaurus
1716:Varanopidae
1544:Vaughnictis
1522:Asaphestera
1503:Caseasauria
664:Varanopidae
533:ontogenetic
529:postcranial
429:(both from
391:pelycosaurs
331:vertebrates
2999:Categories
2867:See also:
2823:Kamagorgon
2813:Dinosaurus
2777:Cynodontia
2766:Cynodontia
2593:Therapsida
2580:Therapsida
2523:see below↓
2514:Therapsida
2444:Dimetrodon
2392:Shashajaia
2176:Milosaurus
2108:Ophiacodon
1955:Dimacrodon
1922:see below↓
1870:Ruthiromia
1855:Aerosaurus
1765:Nitosaurus
1758:Dendromaia
1730:Apsisaurus
1655:Martensius
1641:Euromycter
1422:see below↓
870:References
643:. (2021):
598:H. grandis
583:Kenilworth
579:Cisuralian
553:Kasimovian
377:Cisuralian
329:and small
315:therapsids
2781:includes
2691:includes
2610:Raranimus
2465:Neosaurus
2166:Ianthodon
1891:Varanodon
1751:Eoscansor
1662:Oromycter
1606:Caseopsis
1599:Caseoides
1537:Oedaleops
1467:Synapsida
1454:Synapsida
1413:Synapsida
1334:Tetrapoda
1320:Kingdom:
1304:Synapsida
1262:Paleofile
747:Ianthodon
637:cladogram
628:Phylogeny
602:H. baylei
485:H. baylei
449:H. baylei
261:H. baylei
187:Synapsida
160:Kingdom:
154:Eukaryota
127:H. baylei
2923:Wikidata
2914:Haptodus
2889:Category
2451:Eosyodon
2436:Cutleria
2305:Haptodus
2295:Cutleria
2220:Gordodon
1975:Gorgodon
1965:Driveria
1898:Watongia
1884:Varanops
1807:Cabarzia
1800:Anningia
1555:Caseidae
1530:Eothyris
1328:Chordata
1326:Phylum:
1322:Animalia
1224:34925870
1128:: 1–11.
730:Haptodus
633:Haptodus
521:skeleton
514:holotype
471:, while
461:Cutleria
435:Haptodus
433:), with
373:Asselian
357:Haptodus
323:predator
296:Haptodus
249:Species
219:Gaudry,
214:Haptodus
174:Chordata
170:Phylum:
164:Animalia
150:Domain:
130:holotype
25:Haptodus
2969:4130012
2956:1365455
2943:4819414
2929:Q631703
2783:mammals
1634:Eocasea
1343:Amniota
1246:Palaeos
1215:8672069
1194:Bibcode
587:England
575:maxilla
547:of the
525:cranial
431:Germany
327:insects
319:Permian
301:extinct
206:Genus:
105:↓
1773:Pyozia
1222:
1212:
560:series
541:Kansas
415:Poland
397:&
380:series
369:France
351:, the
299:is an
2982:38905
2951:IRMNG
1592:Casea
1339:Clade
1159:(1).
641:et al
635:in a
365:Autun
308:basal
304:genus
194:Clade
181:Clade
2938:GBIF
2775:see
2752:see
2717:see
2685:see
2659:see
2633:see
1399:see
1372:see
1220:PMID
1010:and
596:and
527:and
479:and
467:and
423:and
267:type
221:1886
44:PreꞒ
1210:PMC
1202:doi
1161:doi
1130:doi
1109:).
1059:doi
1026:doi
987:doi
960:doi
929:doi
900:doi
417:),
367:of
355:of
306:of
3001::
2979::
2966::
2953::
2940::
2925::
1341::
1260:.
1244:.
1218:.
1208:.
1200:.
1188:.
1184:.
1157:23
1155:.
1151:.
1086:17
1084:.
1080:.
1055:13
1053:.
1022:61
1020:.
1016:.
983:10
981:.
956:14
954:.
925:13
923:.
896:56
894:.
890:.
878:^
624:.
585:,
566:.
539:,
517:RM
463:,
386:.
333:.
283:?
196::
183::
94:Pg
38:Ma
33:,
2829:†
2819:†
2809:†
2740:†
2705:†
2673:†
2647:†
2621:†
2606:†
2505:?
2497:?
2489:?
2439:?
2403:†
2388:†
2346:†
2331:†
2321:†
2311:†
2301:†
2291:†
2187:†
2172:†
2162:†
2068:†
2008:?
2001:†
1991:†
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99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:Ꞓ
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