344:
80:
408:
31:
299:
55:
465:. 2014. Turtles of the World, 7th edition: annotated checklist of taxonomy, synonymy, distribution with maps, and conservation status. In: Rhodin, A.G.J., Pritchard, P.C.H., van Dijk, P.P., Saumure, R.A., Buhlmann, K.A., Iverson, J.B., and Mittermeier, R.A. (Eds.). Conservation Biology of Freshwater Turtles and Tortoises: A Compilation Project of the IUCN/SSC Tortoise and Freshwater Turtle Specialist Group.
793:
335:) which inhabits the Karoo region to the west. However greater padlopers can be distinguished not only by their larger size, but also by their nostrils which are below their eye level, by their having four toes on both their front and back feet, by the larger scales on their forelimbs, and by the darker rings around the shell scutes of many individuals.
430:
This species does not survive for long in captivity, unless considerable effort is made to supply specimens with their natural food, that is, endemic plants from the summer-rainfall grasslands region of South Africa. This summer-rainfall tortoise also has specific temperature requirements. They spend
323:
Its shell ranges in colour from olive to reddish-brown, and is slightly flattened in both sexes. The shields tend to be separated by very thin white lines. In juveniles and adult males, the shields of the shell usually have slightly darker edges. Males can also be distinguished from females by being
374:
In addition, there is a sparse relict population of greater padlopers that extends into the high escarpment on the edge of the Karoo, where the climate is relatively humid. Although it has also been known as the
310:("padloper") tortoises, but it is nonetheless tiny, averaging slightly over 10 cm in length – though males tend to be smaller. It has relatively large buttock tubercles. Like its close relative,
4179:
395:
is a summer-rainfall species. Within its range it tends to favour rocky outcrops, and its population is relatively sparse. It is little studied, although it appears in
4096:
423:
species is strictly illegal and any captive specimens are systematically registered in noncommercial studbooks in South Africa and
Namibia, any commercial sale of
4135:
4215:
4070:
4109:
1038:
1227:
1221:
694:
1798:
4230:
497:
4225:
449:
Boulenger, George A. 1888. Description of a new land-tortoise from South Africa, from a specimen living in the
Society's Gardens.
849:
1080:
3831:
2914:
1209:
1203:
1075:
834:
466:
4114:
3958:
Turtles of the World 2017 Update: Annotated
Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status
1767:
turtles of the world 2017 update: Annotated checklist and atlas of taxonomy, synonymy, distribution, and conservation status
1463:
1278:
1448:
1438:
1100:
462:
3808:
3956:
1766:
3967:
2490:
1874:
1588:
1453:
1105:
842:
635:
4140:
2223:
1443:
1049:
687:
1458:
1215:
866:
79:
2869:
1791:
1369:
1090:
1059:
1028:
916:
1125:
998:
3725:
1738:
1288:
1137:
1131:
210:
1685:
1679:
1637:
3984:
3697:
2497:
2455:
2095:
1697:
1643:
1337:
1319:
1095:
1033:
992:
968:
962:
934:
928:
922:
897:
827:
4022:
1673:
1343:
1325:
553:
2441:
1943:
1754:
1748:
1709:
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1542:
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1178:
1085:
1065:
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858:
680:
478:
364:
343:
194:
1733:
1715:
1415:
1349:
1313:
1013:
1008:
1667:
4057:
4031:
3683:
2349:
1973:
1703:
1250:
986:
974:
1661:
1649:
1631:
3668:
2027:
1784:
1743:
1536:
1420:
1183:
957:
44:
1116:
3075:
3040:
2907:
2705:
2646:
2616:
1560:
1524:
1504:
1410:
1304:
1023:
385:
371:
border. It is restricted to regions of high altitude, with rainfall over 250 mm per annum.
219:
74:
2624:
2378:
4101:
4161:
4044:
3618:
2601:
2553:
2544:
2427:
2004:
1565:
1387:
1155:
1044:
890:
885:
494:
359:
is primarily the summer-rainfall grasslands, savanna and bushveld of the highveld plateau of
4220:
4174:
4166:
4127:
3410:
2798:
2773:
2734:
2682:
2413:
1958:
1611:
1481:
1173:
1053:
809:
517:
4148:
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3235:
3203:
2840:
2749:
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2111:
1856:
1725:
1593:
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1268:
1069:
501:
407:
360:
312:
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2719:
2654:
2194:
2185:
2154:
2062:
1951:
1865:
1580:
1360:
1293:
952:
431:
winter under vegetation and rocks (June–September) and lay up to three eggs in summer.
380:
328:
30:
4209:
4122:
3933:
3816:
3716:
3705:
3676:
3590:
3474:
3458:
3418:
3352:
3262:
3227:
3171:
3155:
3139:
3113:
3101:
3085:
3063:
3055:
2862:
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2406:
2267:
2214:
2162:
1570:
1473:
819:
752:
151:
64:
59:
298:
4049:
3868:
3793:
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3286:
3163:
3147:
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3024:
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2893:
2809:
2783:
2697:
2608:
2587:
2511:
2476:
2286:
2252:
2119:
2035:
1981:
1603:
1515:
1194:
908:
758:
661:
579:
287:
4192:
4083:
4016:
3846:
3801:
3647:
3632:
3554:
3546:
3538:
3514:
3394:
3378:
3362:
3278:
3270:
3219:
3195:
3187:
3179:
3047:
2999:
2922:
2847:
2712:
2668:
2594:
2580:
2532:
2389:
2307:
2245:
2238:
2231:
2146:
2088:
2080:
1997:
1989:
1936:
1926:
1552:
1402:
944:
877:
639:
416:
4007:
3925:
3838:
3824:
3785:
3562:
3482:
3450:
3434:
3402:
3340:
3313:
3294:
3211:
3016:
2936:
2929:
2877:
2854:
2764:
2631:
2363:
2170:
2103:
2020:
2012:
1915:
1906:
1237:
1165:
801:
746:
740:
621:
541:
141:
514:
The
Completely Illustrated Atlas of Reptiles and Amphibians for the Terrarium
3917:
3777:
3761:
3690:
3654:
3604:
3597:
3522:
3498:
3442:
3386:
3093:
2885:
2833:
2791:
2504:
2462:
2370:
2356:
2342:
2335:
2328:
2321:
2293:
2138:
1966:
283:
252:
91:
4153:
4001:
3732:
3661:
3581:
3570:
3466:
3131:
3008:
2900:
2742:
2727:
2689:
2638:
2564:
2469:
2448:
2434:
2420:
2276:
2203:
2072:
2051:
2043:
1825:
1622:
1491:
1260:
1147:
778:
722:
703:
351:
tortoise species in southern Africa, showing the greater padloper (brown)
280:
161:
111:
4075:
3876:
3625:
3530:
3426:
3370:
2950:
2943:
2825:
2675:
2661:
2518:
2483:
2398:
2300:
1840:
1831:
1430:
1379:
734:
728:
368:
356:
317:
275:
171:
121:
4088:
3611:
3254:
1846:
1819:
1807:
716:
324:
smaller, with longer tails. Males do not exhibit plastral concavity.
320:), it has only four toes on its front feet as well as its hind feet.
306:
As its name suggests, the greater padloper is the largest of all the
131:
101:
3978:
4187:
2818:
1885:
672:
482:
406:
342:
297:
2314:
396:
4062:
3982:
1780:
676:
1776:
544:
Homopus
Research Foundation. (accessed February 19, 2017).
419:
and poaching for the pet trade. As the trade in collected
379:, this species is not widespread in the Karoo, unlike the
427:
tortoises is almost without exception strictly illegal.
667:, World Chelonian Trust (retrieved August 20, 2013).
3991:
3908:
3901:
3894:
3856:
3742:
3715:
3580:
3350:
3328:
3321:
3312:
3245:
3111:
3073:
2998:
2991:
2960:
2808:
2772:
2763:
2563:
2542:
2388:
2275:
2266:
2213:
2193:
2184:
2129:
2061:
1925:
1914:
1905:
1873:
1864:
1855:
1724:
1620:
1602:
1579:
1551:
1513:
1490:
1472:
1429:
1401:
1378:
1358:
1302:
1259:
1236:
1192:
1164:
1146:
1114:
1007:
943:
907:
876:
818:
800:
767:
451:Proceedings of the Zoological Society of London
532:, Reptile & Amphibian Magazine, 1996-1997.
467:Chelonian Research Monographs 5(7):000.329–479
3955:Phylogenetic arrangement of turtles based on
1792:
1765:Phylogenetic arrangement of turtles based on
688:
567:Cape Tortoises: Their identification and care
8:
327:This species is sometimes confused with the
512:J. Obst, K. Richter, and U. Jacob. (1988).
469:, doi:10.3854/ crm.5.000.checklist.v7.2014.
389:) which is restricted to the Karoo region.
3979:
3905:
3898:
3325:
3318:
2995:
2769:
2272:
2190:
1922:
1911:
1879:
1870:
1861:
1799:
1785:
1777:
771:
695:
681:
673:
445:
443:
363:. It is found as far north as the central
302:1888 sketch – Zoological Society of London
53:
29:
20:
611:. Southern Book Publishers: Johannesburg.
495:IUCN Appendix 1. Regional Species Lists
439:
415:The grassland species is threatened by
1228:Saddle-backed Rodrigues giant tortoise
1222:Saddle-backed Mauritius giant tortoise
7:
607:Boycott, R. and O. Bourquin (1988).
4216:IUCN Red List least concern species
273:, is a small tortoise of the genus
14:
791:
78:
609:The South African Tortoise Book
1210:Domed Rodrigues giant tortoise
1204:Domed Mauritius giant tortoise
1076:Santiago Island giant tortoise
1:
1029:Chatham Island giant tortoise
622:"Dwarf Tortoise Conservation"
463:Turtle Taxonomy Working Group
411:Young female greater padloper
347:Distribution map of the five
1096:Volcán Darwin giant tortoise
1091:Volcán Alcedo giant tortoise
1081:Pinzón Island giant tortoise
3968:List of Testudines families
1589:Flat-backed spider tortoise
1464:Speke's hinge-back tortoise
1454:Lobatse hinge-back tortoise
1279:Goode's thornscrub tortoise
1106:Western Santa Cruz tortoise
1086:Sierra Negra giant tortoise
1034:Eastern Santa Cruz tortoise
588:Homopus Research Foundation
569:. Cape Nature Conservation.
4247:
4231:Reptiles described in 1888
1449:Home's hinge-back tortoise
1444:Forest hinge-back tortoise
1439:Bell's hinge-back tortoise
1101:Volcán Wolf giant tortoise
1050:Hood Island giant tortoise
1039:Fernandina Island tortoise
789:
530:The Herpetology Sourcebook
3951:
1882:
1814:
1763:
1459:Natal hinge-back tortoise
1024:Cerro Azul giant tortoise
867:Aldabrachelys grandidieri
774:
711:
367:, and as far east as the
225:
218:
200:
193:
75:Scientific classification
73:
51:
42:
37:
28:
23:
4226:Reptiles of South Africa
1370:Hesperotestudo percrassa
1126:Cheirogaster bacharidisi
1060:Santa Fe Island tortoise
1045:Floreana Island tortoise
999:Chelonoidis sombrerensis
917:African spurred tortoise
665:(Padloper Tortoise) Care
269:, commonly known as the
1289:Sonoran Desert tortoise
1216:Réunion giant tortoise
1138:Cheirogaster schafferi
1132:Cheirogaster gymnesica
969:Chelonoidis alburyorum
963:Yellow-footed tortoise
935:Centrochelys vulcanica
929:Centrochelys burchardi
923:Centrochelys atlantica
898:Astrochelys rogerbouri
828:Aldabra giant tortoise
556:, CapeNature web site.
412:
352:
303:
248:Mähn & Wilms, 2001
1749:Spur-thighed tortoise
1686:Stylemys nebrascensis
1680:Stylemys karakolensis
1638:Stylemys calaverensis
1543:Megalochelys sondaari
1531:Megalochelys cautleyi
1500:Asian forest tortoise
1246:Burmese star tortoise
1066:Pinta Island tortoise
859:Aldabrachelys abrupta
565:E.H.W. Baard (1994).
410:
399:and other databases.
346:
301:
1739:Kleinmann's tortoise
1698:Stylemys oregonensis
1644:Stylemys canetotiana
1338:Hadrianus schucherti
1320:Hadrianus majusculus
1251:Indian star tortoise
993:Chelonoidis sellovii
987:Chelonoidis monensis
975:Chelonoidis cubensis
279:, indigenous to the
1744:Marginated tortoise
1674:Stylemys frizaciana
1537:Megalochelys margae
1421:Travancore tortoise
1344:Hadrianus utahensis
1326:Hadrianus octonaria
1014:Chelonoidis complex
958:Red-footed tortoise
580:"Greater Padloper,
479:Karoo Cape tortoise
377:Karoo cape tortoise
232:Boulenger, 1888:251
45:Conservation status
16:Species of tortoise
3076:Nanhsiungchelyidae
3041:Macrocephalochelys
1755:Testudo brevitesta
1734:Hermann's tortoise
1710:Stylemys uintensis
1692:Stylemys neglectus
1656:Stylemys conspecta
1561:Geometric tortoise
1525:Megalochelys atlas
1505:Impressed tortoise
1416:Forsten's tortoise
1411:Elongated tortoise
1332:Hadrianus robustus
1009:Galápagos tortoise
981:Chelonoidis lutzae
554:Save Our Tortoises
500:2010-07-16 at the
413:
386:Homopus boulengeri
353:
316:(and unlike other
304:
245:Homopus fermoralis
4203:
4202:
4162:Open Tree of Life
4037:Homopus_femoralis
4023:Homopus femoralis
3993:Homopus femoralis
3985:Taxon identifiers
3976:
3975:
3947:
3946:
3943:
3942:
3890:
3889:
3886:
3885:
3308:
3307:
3304:
3303:
2987:
2986:
2759:
2758:
2262:
2261:
2180:
2179:
1901:
1900:
1897:
1896:
1893:
1892:
1774:
1773:
1769:. Key: †=extinct.
1716:Stylemys undabuna
1566:Serrated tortoise
1350:Hadrianus vialovi
1314:Hadrianus corsoni
1184:Speckled padloper
1156:Angulate tortoise
891:Radiated tortoise
886:Angonoka tortoise
787:
786:
582:Homopus femoralis
393:Homopus femoralis
266:Homopus femoralis
262:
261:
249:
241:
237:Testudo femoralis
233:
229:Homopus femoralis
204:Homopus femoralis
186:H. femoralis
68:
24:Greater padloper
4238:
4196:
4195:
4183:
4182:
4170:
4169:
4157:
4156:
4144:
4143:
4131:
4130:
4118:
4117:
4105:
4104:
4092:
4091:
4079:
4078:
4066:
4065:
4053:
4052:
4040:
4039:
4027:
4026:
4025:
4012:
4011:
4010:
3980:
3906:
3899:
3411:Chupacabrachelys
3326:
3319:
2996:
2774:Carettochelyidae
2770:
2273:
2191:
1923:
1912:
1880:
1871:
1862:
1801:
1794:
1787:
1778:
1668:Stylemys emiliae
1612:Leopard tortoise
1482:Pancake tortoise
1393:Greater padloper
810:Russian tortoise
795:
794:
772:
697:
690:
683:
674:
668:
658:
652:
650:
648:
647:
638:. Archived from
632:
626:
625:
618:
612:
605:
599:
598:
596:
594:
576:
570:
563:
557:
551:
545:
539:
533:
527:
521:
510:
504:
492:
486:
485:species gallery.
476:
470:
460:
454:
447:
271:greater padloper
247:
239:
231:
206:
83:
82:
62:
57:
56:
33:
21:
4246:
4245:
4241:
4240:
4239:
4237:
4236:
4235:
4206:
4205:
4204:
4199:
4191:
4186:
4178:
4173:
4165:
4160:
4152:
4149:Observation.org
4147:
4139:
4134:
4126:
4121:
4113:
4108:
4100:
4095:
4087:
4082:
4074:
4069:
4061:
4056:
4048:
4043:
4035:
4030:
4021:
4020:
4015:
4006:
4005:
4000:
3987:
3977:
3972:
3939:
3882:
3859:Sahonachelyidae
3852:
3738:
3711:
3576:
3507:Phosphatochelys
3346:
3300:
3241:
3236:Terlinguachelys
3204:Pneumatoarthrus
3107:
3069:
2983:
2956:
2804:
2755:
2559:
2538:
2526:Siebenrockiella
2384:
2258:
2209:
2176:
2125:
2112:Mesodermochelys
2057:
1918:
1889:
1851:
1810:
1805:
1775:
1770:
1759:
1720:
1704:Stylemys pygmea
1616:
1598:
1594:Spider tortoise
1575:
1547:
1509:
1486:
1468:
1425:
1397:
1388:Common padloper
1374:
1354:
1298:
1284:Gopher tortoise
1274:Desert tortoise
1269:Bolson tortoise
1255:
1232:
1188:
1160:
1142:
1110:
1070:Lonesome George
1003:
939:
903:
872:
850:A. g. hololissa
814:
796:
792:
783:
776:Species of the
763:
707:
701:
671:
659:
655:
645:
643:
634:
633:
629:
620:
619:
615:
606:
602:
592:
590:
578:
577:
573:
564:
560:
552:
548:
540:
536:
528:
524:
511:
507:
502:Wayback Machine
493:
489:
477:
473:
461:
457:
448:
441:
437:
405:
361:southern Africa
341:
296:
214:
208:
202:
189:
77:
69:
58:
54:
47:
17:
12:
11:
5:
4244:
4242:
4234:
4233:
4228:
4223:
4218:
4208:
4207:
4201:
4200:
4198:
4197:
4184:
4171:
4158:
4145:
4132:
4119:
4106:
4093:
4080:
4067:
4054:
4041:
4028:
4013:
3997:
3995:
3989:
3988:
3983:
3974:
3973:
3971:
3970:
3963:
3962:
3961:. † = extinct.
3952:
3949:
3948:
3945:
3944:
3941:
3940:
3938:
3937:
3929:
3921:
3912:
3910:
3903:
3896:
3892:
3891:
3888:
3887:
3884:
3883:
3881:
3880:
3872:
3863:
3861:
3854:
3853:
3851:
3850:
3842:
3835:
3828:
3820:
3812:
3805:
3797:
3789:
3781:
3773:
3765:
3757:
3754:Albertwoodemys
3748:
3746:
3744:Podocnemididae
3740:
3739:
3737:
3736:
3729:
3721:
3719:
3713:
3712:
3710:
3709:
3701:
3694:
3687:
3680:
3672:
3665:
3658:
3651:
3644:
3641:Lomalatachelys
3636:
3629:
3622:
3615:
3608:
3601:
3594:
3586:
3584:
3578:
3577:
3575:
3574:
3566:
3558:
3550:
3542:
3534:
3526:
3518:
3510:
3502:
3494:
3491:Labrostochelys
3486:
3478:
3470:
3462:
3454:
3446:
3438:
3430:
3422:
3414:
3406:
3398:
3390:
3382:
3374:
3366:
3357:
3355:
3348:
3347:
3345:
3344:
3335:
3333:
3331:Araripemydidae
3323:
3316:
3310:
3309:
3306:
3305:
3302:
3301:
3299:
3298:
3290:
3282:
3274:
3266:
3258:
3249:
3247:
3243:
3242:
3240:
3239:
3231:
3223:
3215:
3207:
3199:
3191:
3183:
3175:
3167:
3159:
3151:
3143:
3135:
3127:
3118:
3116:
3109:
3108:
3106:
3105:
3097:
3089:
3080:
3078:
3071:
3070:
3068:
3067:
3059:
3051:
3044:
3036:
3033:Emarginachelys
3028:
3020:
3012:
3004:
3002:
2993:
2989:
2988:
2985:
2984:
2982:
2981:
2978:Sinaspideretes
2973:
2964:
2962:
2958:
2957:
2955:
2954:
2947:
2940:
2933:
2926:
2918:
2911:
2904:
2897:
2889:
2881:
2873:
2866:
2858:
2851:
2844:
2837:
2829:
2822:
2814:
2812:
2806:
2805:
2803:
2802:
2795:
2787:
2778:
2776:
2767:
2761:
2760:
2757:
2756:
2754:
2753:
2746:
2738:
2731:
2723:
2716:
2709:
2701:
2693:
2686:
2679:
2672:
2665:
2658:
2655:Hesperotestudo
2650:
2642:
2635:
2628:
2620:
2612:
2605:
2598:
2591:
2584:
2577:
2569:
2567:
2561:
2560:
2558:
2557:
2549:
2547:
2545:Platysternidae
2540:
2539:
2537:
2536:
2529:
2522:
2515:
2508:
2501:
2494:
2487:
2480:
2473:
2466:
2459:
2452:
2445:
2438:
2431:
2424:
2417:
2410:
2402:
2394:
2392:
2386:
2385:
2383:
2382:
2374:
2367:
2360:
2353:
2346:
2339:
2332:
2325:
2318:
2311:
2304:
2297:
2290:
2281:
2279:
2270:
2264:
2263:
2260:
2259:
2257:
2256:
2249:
2242:
2235:
2227:
2219:
2217:
2211:
2210:
2208:
2207:
2199:
2197:
2195:Dermatemydidae
2188:
2186:Kinosternoidea
2182:
2181:
2178:
2177:
2175:
2174:
2166:
2158:
2155:Procolpochelys
2150:
2142:
2133:
2131:
2127:
2126:
2124:
2123:
2115:
2107:
2099:
2092:
2084:
2076:
2067:
2065:
2063:Dermochelyidae
2059:
2058:
2056:
2055:
2047:
2039:
2031:
2024:
2016:
2008:
2001:
1993:
1985:
1977:
1970:
1962:
1955:
1952:Carolinochelys
1947:
1940:
1931:
1929:
1920:
1909:
1903:
1902:
1899:
1898:
1895:
1894:
1891:
1890:
1883:
1877:
1868:
1859:
1853:
1852:
1850:
1849:
1843:
1834:
1828:
1822:
1815:
1812:
1811:
1806:
1804:
1803:
1796:
1789:
1781:
1772:
1771:
1764:
1761:
1760:
1758:
1757:
1751:
1746:
1741:
1736:
1730:
1728:
1722:
1721:
1719:
1718:
1712:
1706:
1700:
1694:
1688:
1682:
1676:
1670:
1664:
1662:Stylemys copei
1658:
1652:
1650:Stylemys capax
1646:
1640:
1634:
1632:Stylemys botti
1627:
1625:
1618:
1617:
1615:
1614:
1608:
1606:
1600:
1599:
1597:
1596:
1591:
1585:
1583:
1577:
1576:
1574:
1573:
1568:
1563:
1557:
1555:
1549:
1548:
1546:
1545:
1539:
1533:
1527:
1520:
1518:
1511:
1510:
1508:
1507:
1502:
1496:
1494:
1488:
1487:
1485:
1484:
1478:
1476:
1470:
1469:
1467:
1466:
1461:
1456:
1451:
1446:
1441:
1435:
1433:
1427:
1426:
1424:
1423:
1418:
1413:
1407:
1405:
1399:
1398:
1396:
1395:
1390:
1384:
1382:
1376:
1375:
1373:
1372:
1365:
1363:
1361:Hesperotestudo
1356:
1355:
1353:
1352:
1346:
1340:
1334:
1328:
1322:
1316:
1309:
1307:
1300:
1299:
1297:
1296:
1294:Texas tortoise
1291:
1286:
1281:
1276:
1271:
1265:
1263:
1257:
1256:
1254:
1253:
1248:
1242:
1240:
1234:
1233:
1231:
1230:
1224:
1218:
1212:
1206:
1199:
1197:
1190:
1189:
1187:
1186:
1181:
1179:Karoo padloper
1176:
1170:
1168:
1162:
1161:
1159:
1158:
1152:
1150:
1144:
1143:
1141:
1140:
1134:
1128:
1121:
1119:
1112:
1111:
1109:
1108:
1103:
1098:
1093:
1088:
1083:
1078:
1073:
1062:
1057:
1047:
1041:
1036:
1031:
1026:
1020:
1018:
1017:
1016:
1005:
1004:
1002:
1001:
995:
989:
983:
977:
971:
965:
960:
955:
953:Chaco tortoise
949:
947:
941:
940:
938:
937:
931:
925:
919:
913:
911:
905:
904:
902:
901:
893:
888:
882:
880:
874:
873:
871:
870:
862:
854:
846:
843:A. g. daudinii
838:
831:
824:
822:
816:
815:
813:
812:
806:
804:
798:
797:
790:
788:
785:
784:
775:
769:
765:
764:
762:
761:
755:
749:
743:
737:
731:
725:
719:
712:
709:
708:
702:
700:
699:
692:
685:
677:
670:
669:
653:
627:
613:
600:
571:
558:
546:
534:
522:
505:
487:
471:
455:
438:
436:
433:
404:
401:
381:Karoo padloper
340:
337:
329:Karoo padloper
295:
292:
260:
259:
258:
257:
242:
234:
223:
222:
216:
215:
209:
198:
197:
191:
190:
183:
181:
177:
176:
169:
165:
164:
159:
155:
154:
149:
145:
144:
139:
135:
134:
129:
125:
124:
119:
115:
114:
109:
105:
104:
99:
95:
94:
89:
85:
84:
71:
70:
52:
49:
48:
43:
40:
39:
35:
34:
26:
25:
15:
13:
10:
9:
6:
4:
3:
2:
4243:
4232:
4229:
4227:
4224:
4222:
4219:
4217:
4214:
4213:
4211:
4194:
4189:
4185:
4181:
4176:
4172:
4168:
4163:
4159:
4155:
4150:
4146:
4142:
4137:
4133:
4129:
4124:
4120:
4116:
4111:
4107:
4103:
4098:
4094:
4090:
4085:
4081:
4077:
4072:
4068:
4064:
4059:
4055:
4051:
4046:
4042:
4038:
4033:
4029:
4024:
4018:
4014:
4009:
4003:
3999:
3998:
3996:
3994:
3990:
3986:
3981:
3969:
3965:
3964:
3960:
3959:
3954:
3953:
3950:
3936:
3935:
3934:Tacuarembemys
3930:
3928:
3927:
3922:
3920:
3919:
3914:
3913:
3911:
3907:
3904:
3900:
3897:
3893:
3879:
3878:
3873:
3871:
3870:
3869:Sahonachelys
3865:
3864:
3862:
3860:
3855:
3849:
3848:
3843:
3841:
3840:
3836:
3834:
3833:
3832:Peltocephalus
3829:
3827:
3826:
3821:
3819:
3818:
3817:Lapparentemys
3813:
3811:
3810:
3806:
3804:
3803:
3798:
3796:
3795:
3790:
3788:
3787:
3782:
3780:
3779:
3774:
3772:
3771:
3766:
3764:
3763:
3758:
3756:
3755:
3750:
3749:
3747:
3745:
3741:
3735:
3734:
3730:
3728:
3727:
3723:
3722:
3720:
3718:
3717:Pelomedusidae
3714:
3708:
3707:
3706:Yaminuechelys
3702:
3700:
3699:
3695:
3693:
3692:
3688:
3686:
3685:
3681:
3679:
3678:
3677:Prochelidella
3673:
3671:
3670:
3666:
3664:
3663:
3659:
3657:
3656:
3652:
3650:
3649:
3645:
3643:
3642:
3637:
3635:
3634:
3630:
3628:
3627:
3623:
3621:
3620:
3616:
3614:
3613:
3609:
3607:
3606:
3602:
3600:
3599:
3595:
3593:
3592:
3591:Acanthochelys
3588:
3587:
3585:
3583:
3579:
3573:
3572:
3567:
3565:
3564:
3559:
3557:
3556:
3551:
3549:
3548:
3543:
3541:
3540:
3535:
3533:
3532:
3527:
3525:
3524:
3519:
3517:
3516:
3511:
3509:
3508:
3503:
3501:
3500:
3495:
3493:
3492:
3487:
3485:
3484:
3479:
3477:
3476:
3475:Kinkonychelys
3471:
3469:
3468:
3463:
3461:
3460:
3459:Itapecuruemys
3455:
3453:
3452:
3447:
3445:
3444:
3439:
3437:
3436:
3431:
3429:
3428:
3423:
3421:
3420:
3419:Eotaphrosphys
3415:
3413:
3412:
3407:
3405:
3404:
3399:
3397:
3396:
3391:
3389:
3388:
3383:
3381:
3380:
3375:
3373:
3372:
3367:
3365:
3364:
3359:
3358:
3356:
3354:
3353:Bothremydidae
3349:
3343:
3342:
3337:
3336:
3334:
3332:
3327:
3324:
3320:
3317:
3315:
3311:
3297:
3296:
3291:
3289:
3288:
3283:
3281:
3280:
3275:
3273:
3272:
3267:
3265:
3264:
3263:Argillochelys
3259:
3257:
3256:
3251:
3250:
3248:
3244:
3238:
3237:
3232:
3230:
3229:
3228:Santanachelys
3224:
3222:
3221:
3216:
3214:
3213:
3208:
3206:
3205:
3200:
3198:
3197:
3192:
3190:
3189:
3184:
3182:
3181:
3176:
3174:
3173:
3172:Desmatochelys
3168:
3166:
3165:
3160:
3158:
3157:
3156:Calcarichelys
3152:
3150:
3149:
3144:
3142:
3141:
3140:Atlantochelys
3136:
3134:
3133:
3128:
3126:
3125:
3120:
3119:
3117:
3115:
3114:Protostegidae
3110:
3104:
3103:
3102:Jiangxichelys
3098:
3096:
3095:
3090:
3088:
3087:
3086:Anomalochelys
3082:
3081:
3079:
3077:
3072:
3066:
3065:
3064:Protochelydra
3060:
3058:
3057:
3056:Planiplastron
3052:
3050:
3049:
3045:
3043:
3042:
3037:
3035:
3034:
3029:
3027:
3026:
3021:
3019:
3018:
3013:
3011:
3010:
3006:
3005:
3003:
3001:
2997:
2994:
2990:
2980:
2979:
2974:
2972:
2971:
2966:
2965:
2963:
2959:
2953:
2952:
2948:
2946:
2945:
2941:
2939:
2938:
2934:
2932:
2931:
2927:
2925:
2924:
2919:
2917:
2916:
2912:
2910:
2909:
2905:
2903:
2902:
2898:
2896:
2895:
2890:
2888:
2887:
2882:
2880:
2879:
2874:
2872:
2871:
2867:
2865:
2864:
2863:Drazinderetes
2859:
2857:
2856:
2852:
2850:
2849:
2845:
2843:
2842:
2838:
2836:
2835:
2830:
2828:
2827:
2823:
2821:
2820:
2816:
2815:
2813:
2811:
2807:
2801:
2800:
2799:Carettochelys
2796:
2794:
2793:
2788:
2786:
2785:
2780:
2779:
2777:
2775:
2771:
2768:
2766:
2762:
2752:
2751:
2747:
2745:
2744:
2739:
2737:
2736:
2732:
2730:
2729:
2724:
2722:
2721:
2717:
2715:
2714:
2710:
2708:
2707:
2702:
2700:
2699:
2694:
2692:
2691:
2687:
2685:
2684:
2683:Malacochersus
2680:
2678:
2677:
2673:
2671:
2670:
2666:
2664:
2663:
2659:
2657:
2656:
2651:
2649:
2648:
2643:
2641:
2640:
2636:
2634:
2633:
2629:
2627:
2626:
2621:
2619:
2618:
2613:
2611:
2610:
2606:
2604:
2603:
2599:
2597:
2596:
2592:
2590:
2589:
2585:
2583:
2582:
2578:
2576:
2575:
2574:Aldabrachelys
2571:
2570:
2568:
2566:
2562:
2556:
2555:
2551:
2550:
2548:
2546:
2541:
2535:
2534:
2530:
2528:
2527:
2523:
2521:
2520:
2516:
2514:
2513:
2509:
2507:
2506:
2502:
2500:
2499:
2495:
2493:
2492:
2488:
2486:
2485:
2481:
2479:
2478:
2474:
2472:
2471:
2467:
2465:
2464:
2460:
2458:
2457:
2456:Leucocephalon
2453:
2451:
2450:
2446:
2444:
2443:
2439:
2437:
2436:
2432:
2430:
2429:
2425:
2423:
2422:
2418:
2416:
2415:
2411:
2409:
2408:
2407:Banhxeochelys
2403:
2401:
2400:
2396:
2395:
2393:
2391:
2387:
2381:
2380:
2375:
2373:
2372:
2368:
2366:
2365:
2361:
2359:
2358:
2354:
2352:
2351:
2347:
2345:
2344:
2340:
2338:
2337:
2333:
2331:
2330:
2326:
2324:
2323:
2319:
2317:
2316:
2312:
2310:
2309:
2305:
2303:
2302:
2298:
2296:
2295:
2291:
2289:
2288:
2283:
2282:
2280:
2278:
2274:
2271:
2269:
2268:Testudinoidea
2265:
2255:
2254:
2250:
2248:
2247:
2243:
2241:
2240:
2236:
2234:
2233:
2228:
2226:
2225:
2221:
2220:
2218:
2216:
2215:Kinosternidae
2212:
2206:
2205:
2201:
2200:
2198:
2196:
2192:
2189:
2187:
2183:
2173:
2172:
2167:
2165:
2164:
2163:Protosphargis
2159:
2157:
2156:
2151:
2149:
2148:
2143:
2141:
2140:
2135:
2134:
2132:
2128:
2122:
2121:
2116:
2114:
2113:
2108:
2106:
2105:
2100:
2098:
2097:
2093:
2091:
2090:
2085:
2083:
2082:
2077:
2075:
2074:
2069:
2068:
2066:
2064:
2060:
2054:
2053:
2048:
2046:
2045:
2040:
2038:
2037:
2032:
2030:
2029:
2025:
2023:
2022:
2017:
2015:
2014:
2009:
2007:
2006:
2002:
2000:
1999:
1994:
1992:
1991:
1986:
1984:
1983:
1978:
1976:
1975:
1971:
1969:
1968:
1963:
1961:
1960:
1956:
1954:
1953:
1948:
1946:
1945:
1941:
1939:
1938:
1933:
1932:
1930:
1928:
1924:
1921:
1919:(Sea turtles)
1917:
1913:
1910:
1908:
1904:
1888:
1887:
1881:
1878:
1876:
1872:
1869:
1867:
1863:
1860:
1858:
1854:
1848:
1844:
1842:
1838:
1835:
1833:
1829:
1827:
1823:
1821:
1817:
1816:
1813:
1809:
1802:
1797:
1795:
1790:
1788:
1783:
1782:
1779:
1768:
1762:
1756:
1752:
1750:
1747:
1745:
1742:
1740:
1737:
1735:
1732:
1731:
1729:
1727:
1723:
1717:
1713:
1711:
1707:
1705:
1701:
1699:
1695:
1693:
1689:
1687:
1683:
1681:
1677:
1675:
1671:
1669:
1665:
1663:
1659:
1657:
1653:
1651:
1647:
1645:
1641:
1639:
1635:
1633:
1629:
1628:
1626:
1624:
1619:
1613:
1610:
1609:
1607:
1605:
1601:
1595:
1592:
1590:
1587:
1586:
1584:
1582:
1578:
1572:
1571:Tent tortoise
1569:
1567:
1564:
1562:
1559:
1558:
1556:
1554:
1550:
1544:
1540:
1538:
1534:
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835:A. g. arnoldi
832:
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820:Aldabrachelys
817:
811:
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803:
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751:Superfamily:
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748:
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651:=342&sm=1
642:on 2014-03-07
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333:H. boulengeri
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291:
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240:Strauch, 1890
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195:Binomial name
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152:Testudinoidea
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97:
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90:
87:
86:
81:
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72:
66:
61:
60:Least Concern
50:
46:
41:
38:Adult female
36:
32:
27:
22:
19:
3992:
3957:
3932:
3924:
3916:
3875:
3867:
3845:
3837:
3830:
3823:
3815:
3809:Erymnochelys
3807:
3800:
3794:Cerrejonemys
3792:
3784:
3776:
3770:Brontochelys
3768:
3760:
3752:
3731:
3724:
3704:
3696:
3689:
3684:Pseudemydura
3682:
3675:
3667:
3660:
3653:
3646:
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3631:
3624:
3617:
3610:
3603:
3596:
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3513:
3505:
3497:
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3473:
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3425:
3417:
3409:
3401:
3393:
3385:
3377:
3369:
3361:
3339:
3293:
3287:Prionochelys
3285:
3277:
3269:
3261:
3253:
3234:
3226:
3218:
3210:
3202:
3194:
3186:
3178:
3170:
3164:Cratochelone
3162:
3154:
3148:Bouliachelys
3146:
3138:
3130:
3124:Alienochelys
3122:
3100:
3092:
3084:
3062:
3054:
3046:
3039:
3031:
3025:Chelydropsis
3023:
3015:
3007:
2976:
2970:Basilochelys
2968:
2949:
2942:
2935:
2928:
2921:
2913:
2906:
2899:
2894:Khunnuchelys
2892:
2884:
2876:
2868:
2861:
2853:
2846:
2839:
2832:
2824:
2817:
2810:Trionychidae
2797:
2790:
2784:Allaeochelys
2782:
2748:
2741:
2735:Stigmochelys
2733:
2726:
2718:
2711:
2704:
2698:Megalochelys
2696:
2688:
2681:
2674:
2667:
2660:
2653:
2645:
2637:
2630:
2623:
2615:
2609:Cylindraspis
2607:
2600:
2593:
2588:Centrochelys
2586:
2579:
2572:
2565:Testudinidae
2554:Platysternon
2552:
2533:Vijayachelys
2531:
2524:
2517:
2512:Rhinoclemmys
2510:
2503:
2496:
2489:
2482:
2477:Melanochelys
2475:
2468:
2461:
2454:
2447:
2440:
2433:
2426:
2419:
2412:
2405:
2397:
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2362:
2355:
2348:
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2334:
2327:
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2306:
2299:
2292:
2287:Acherontemys
2285:
2253:Sternotherus
2251:
2244:
2237:
2230:
2222:
2202:
2169:
2161:
2153:
2145:
2137:
2120:Psephophorus
2118:
2110:
2102:
2094:
2087:
2079:
2071:
2050:
2042:
2036:Pacifichelys
2034:
2026:
2019:
2011:
2005:Lepidochelys
2003:
1996:
1988:
1982:Gigantatypus
1980:
1974:Eretmochelys
1972:
1965:
1957:
1950:
1942:
1935:
1916:Chelonioidea
1884:
1836:
1604:Stigmochelys
1516:Megalochelys
1392:
1195:Cylindraspis
1117:Cheirogaster
909:Centrochelys
896:
865:
857:
848:
841:
833:
830:(subspecies:
777:
759:Testudinidae
662:
660:Corton, M.,
656:
644:. Retrieved
640:the original
636:"CapeNature"
630:
616:
608:
603:
593:February 20,
591:. Retrieved
587:
581:
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518:T.F.H. press
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458:
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403:Conservation
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354:
348:
339:Distribution
332:
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322:
313:H. areolatus
311:
307:
305:
288:South Africa
274:
270:
265:
264:
263:
251:
244:
236:
228:
203:
201:
185:
184:
172:
162:Testudinidae
18:
4084:iNaturalist
4017:Wikispecies
3847:Stupendemys
3802:Cordichelys
3648:Mesoclemmys
3633:Hydromedusa
3555:Taphrosphys
3547:Sankuchemys
3539:Rhothonemys
3515:Polysternon
3395:Cearachelys
3379:Azabbaremys
3363:Araiochelys
3279:Ctenochelys
3271:Bashuchelys
3220:Rhinochelys
3196:Ocepechelon
3188:Notochelone
3180:Iserosaurus
3048:Macrochelys
3000:Chelydridae
2923:Palaeoamyda
2848:Cyclanorbis
2713:Psammobates
2706:Oligopherus
2669:Indotestudo
2617:Cymatholcus
2595:Chelonoidis
2581:Astrochelys
2390:Geoemydidae
2308:Deirochelys
2246:Staurotypus
2239:Kinosternon
2232:Hoplochelys
2147:Peritresius
2096:Dermochelys
2089:Cosmochelys
2081:Corsochelys
1998:Itilochelys
1990:Glarichelys
1937:Allopleuron
1927:Cheloniidae
1866:Superfamily
1553:Psammobates
1403:Indotestudo
945:Chelonoidis
878:Astrochelys
417:overgrazing
294:Description
4210:Categories
3926:Caririemys
3839:Podocnemis
3825:Latentemys
3786:Carbonemys
3726:Pelomedusa
3563:Ummulisani
3483:Kurmademys
3451:Inaechelys
3435:Galianemys
3403:Chedighaii
3341:Araripemys
3314:Pleurodira
3295:Toxochelys
3212:Protostega
3017:Chelydrops
2937:Pelodiscus
2930:Pelochelys
2878:Gilmoremys
2855:Cycloderma
2765:Trionychia
2632:Geochelone
2625:Floridemys
2491:Notochelys
2379:Wilburemys
2350:Malaclemys
2204:Dermatemys
2171:Puppigerus
2104:Eosphargis
2021:Miocaretta
2013:Mexichelys
1907:Cryptodira
1847:Testudines
1808:Testudines
1238:Geochelone
1166:Chersobius
802:Agrionemys
747:Cryptodira
745:Suborder:
741:Testudines
733:Subclass:
646:2014-03-07
435:References
365:Free State
284:grasslands
142:Cryptodira
138:Suborder:
132:Testudines
4180:femoralis
3966:See also
3918:Caribemys
3778:Caninemys
3762:Bauruemys
3691:Rheodytes
3655:Myuchelys
3598:Chelodina
3523:Puentemys
3499:Nigeremys
3443:Ilatardia
3387:Bothremys
3094:Basilemys
2908:Nilssonia
2886:Hutchemys
2834:Axestemys
2792:Anosteira
2647:Hadrianus
2505:Pangshura
2463:Malayemys
2428:Geoclemys
2371:Trachemys
2364:Terrapene
2357:Pseudemys
2343:Graptemys
2336:Glyptemys
2329:Emydoidea
2322:Actinemys
2294:Chrysemys
2139:Euclastes
1967:Eochelone
1818:Kingdom:
1305:Hadrianus
715:Kingdom:
542:Tortoises
453:1888:251.
318:padlopers
253:ex errore
211:Boulenger
180:Species:
98:Kingdom:
92:Eukaryota
4188:Species+
4102:10951485
4008:Q3080873
4002:Wikidata
3733:Pelusios
3698:Rhinemys
3669:Platemys
3662:Phrynops
3582:Chelidae
3571:Zolhafah
3467:Jainemys
3132:Archelon
3009:Chelydra
2901:Lissemys
2743:Stylemys
2728:Solitudo
2690:Manouria
2639:Gopherus
2602:Chersina
2470:Mauremys
2449:Heosemys
2442:Hardella
2435:Geoemyda
2421:Cyclemys
2277:Emydidae
2224:Claudius
2073:Arabemys
2052:Tasbacka
2044:Syllomus
1959:Chelonia
1857:Suborder
1841:Diapsida
1832:Reptilia
1826:Chordata
1824:Phylum:
1820:Animalia
1623:Stylemys
1492:Manouria
1261:Gopherus
1148:Chersina
779:tortoise
757:Family:
735:Anapsida
729:Reptilia
723:Chordata
721:Phylum:
717:Animalia
704:Tortoise
498:Archived
281:highveld
220:Synonyms
158:Family:
122:Reptilia
112:Chordata
108:Phylum:
102:Animalia
88:Domain:
65:IUCN 3.1
4221:Homopus
4076:5220253
3877:Sokatra
3626:Emydura
3531:Rosasia
3427:Foxemys
3371:Arenila
2951:Trionyx
2944:Rafetus
2870:Dogania
2826:Apalone
2750:Testudo
2676:Kinixys
2662:Homopus
2519:Sacalia
2498:Orlitia
2484:Morenia
2399:Batagur
2301:Clemmys
2028:Natator
1944:Caretta
1845:Order:
1830:Class:
1726:Testudo
1431:Kinixys
1380:Homopus
739:Order:
727:Class:
663:Homopus
425:Homopus
421:Homopus
369:Lesotho
357:habitat
349:Homopus
308:Homopus
276:Homopus
173:Homopus
168:Genus:
128:Order:
118:Class:
63: (
4167:777385
4154:100020
4141:285998
4128:170522
4115:551786
4063:793251
3909:
3902:
3895:
3619:Elusor
3612:Elseya
3605:Chelus
3322:
3255:Adocus
3246:
2992:
2961:
2841:Chitra
2543:
2130:
1875:Family
782:family
768:Genera
706:family
213:, 1888
4193:10479
4097:IRMNG
4089:40074
4050:3MHMD
2915:Palea
2819:Amyda
2720:Pyxis
2414:Cuora
1886:Genus
1837:Clade
1581:Pyxis
1054:Diego
483:CITES
4136:NCBI
4123:IUCN
4110:ITIS
4071:GBIF
2315:Emys
595:2017
397:ITIS
355:Its
4058:EoL
4045:CoL
4032:ADW
286:of
4212::
4190::
4177::
4175:RD
4164::
4151::
4138::
4125::
4112::
4099::
4086::
4073::
4060::
4047::
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1839::
1068:†(
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Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
