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Haifanggou Formation

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for not including enough specifics and detail in their paper, and also took issue with their radiometric dating of the Daohugou tuff. The tuff, Gao and Ren argued, contains crystals with a variety of diverse radiometric ages, some up to a billion years old, so using dates from only a few of these
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found that the beds formed between 164 and 158 million years ago, in the Middle to Late Jurassic. A 2012 study by Gao and Shubin agreed with this assessment, and reported an Argon–argon date of 164 plus or minus 4 million years ago for the Daohugou horizon.
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The age of the Daohugou bed has been debated, and a number of studies, using different methodologies, have reached conflicting conclusions. Various papers have placed the fossils here as being anywhere from the
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Gu J.-J., Montealegre-Z, F., Robert, D., Engel, M.S., Qiao G.-X., and Ren D. (2012). "Wing stridulation in a Jurassic katydid (Insecta, Orthoptera) produced low-pitched musical calls to attract females."
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Qing-Wen Ma; David K. Ferguson; Hai-Ming Liu; Jing-Xian Xu (2021). "Compressions of Sequoia (Cupressaceae sensu lato) from the Middle Jurassic of Daohugou, Ningcheng, Inner Mongolia, China".
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At least 760 insect species have been described from the Haifanggou Formation and associated deposits, making it one of the most diverse insect-bearing strata from the Jurassic period.
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Giribet, G., et al., 2012. An exquisitely preserved harvestman (Arthropoda, Arachnida, Opiliones) from the Middle Jurassic of China. Organisms, Diversity & Evolution, 12, p.51-56.
2422: 3486:"The new records of Coniopteris (Dicksoniaceae, Filicales) with in-situ spores from the Middle Jurassic of northeastern China and its spatio-temporal distribution in China" 4644: 3839: 2581:"The early assemblage of Middle–Late Jurassic Yanliao biota: checklist, bibliography and statistical analysis of described taxa from the Daohugou beds and coeval deposits" 381: 3718: 615:, argued that the 159-164 million years old Tiaojishan Formation underlies, rather than overlies, the Daohugou Beds. Unlike the earlier study by Gao and Ren, Wang 3224:"A chronostratigraphic and biostratigraphic framework for the Yanliao Biota of northeastern China: Implications for Jurassic terrestrial ecosystems and evolution" 2159:
Liu, Yongqing; Liu, Yanxue; Ji, S.; Yang, Z. (2006). "U-Pb zircon age for the Daohugou Biota at Ningcheng of Inner Mongolia and comments on related issues".
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Gao, K.; Ren, D. (2006). "Radiometric dating of ignimbrite from Inner Mongolia provides no indication of a post-Middle Jurassic age for the Daohugou Beds".
645:, a typical fish of the Jehol Biota, the Daohugou vertebrate assemblage is closer to that of the Early Cretaceous Jehol Biota than to any other biota." 3830: 2309:"(^40)Ar/(^39)Ar dating of ignimbrite from Inner Mongolia, northeastern China, indicates a post-Middle Jurassic age for the overlying Daohugou Beds" 353: 4649: 657:, and that it "belongs to the same cycle of volcanism and sedimentation as the Yixian Formation of the Jehol Group." However, a later study by Ji 4526: 3675:
Xin WANG; and Shijun WANG (2010). "Xingxueanthus: An Enigmatic Jurassic Seed Plant and Its Implications for the Origin of Angiospermy".
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crystals could not determine the overall age of the deposits. Gao and Ren went on to defend a Middle Jurassic age for the beds based on
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Li, Yuling; Chang, Su-Chin; Zhang, Haichun; Wang, Jun; Pei, Rui; Zheng, Daran; Fang, Yan; Hemming, Sidney R. (November 2023).
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Ren, D.; et al. (2002). "On the biostratigraphy of the Jurassic fossil beds at Daohugou near Ningcheng, Inner Mongolia".
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on the volcanic rocks overlying and underlying salamander-bearing layers (salamanders are often used as index fossils). Liu
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Wang, Yongjie; Labandeira, Conrad C.; Shih, Chungkun; Ding, Qiaoling; Wang, Chen; Zhao, Yunyun; Ren, Dong (2012-12-11).
210: 2393:"Biodiversity and palaeoclimate of the Middle Jurassic floras from the Tiaojishan Formation in western Liaoning, China" 3846: 256: 217: 2134: 4456: 4041: 4031: 2235:"An Updated Review of the Middle-Late Jurassic Yanliao Biota: Chronology, Taphonomy, Paleontology and Paleoecology" 595:
However, subsequent studies cast doubt on this relatively recent age. In a 2006 study, Gao & Ren criticized He
4634: 4619: 4451: 4393: 4135: 3965: 3955: 3892: 2491:"LA-ICPMS Zircon U-Pb Dating in the Jurassic Daohugou Beds and Correlative Strata in Ningcheng of Inner Mongolia" 4629: 4604: 4599: 4388: 4276: 2113: 2078: 525: 4221: 3078:"A new Middle Jurassic cockroach (Blattaria: Blattulidae) from the Jiulongshan Formation of Daohugou in China" 3808: 2842:"The Earliest Case of Extreme Sexual Display with Exaggerated Male Organs by Two Middle Jurassic Mecopterans" 4594: 4398: 4281: 4140: 3816: 3779: 608:), and the bed's relationship to a layer that is known to mark the Middle Jurassic-Late Jurassic boundary. 585: 4261: 2083: 2060: 1235: 502: 176: 3620:
Sokoloff, Dmitry D.; Remizowa, Margarita V.; El, Elena S.; Rudall, Paula J.; Bateman, Richard M. (2020).
2490: 1153: 580:, found them to be Early Cretaceous, only a few million years older than the overlying Jehol beds of the 4266: 3868: 3712: 3043: 2416: 1615: 1207: 3136:"Two new genera of giant lacewings (Insecta, Neuroptera, Ithonidae) from the Middle Jurassic of China" 819: 4036: 3684: 3532: 3497: 3429: 3277: 3235: 3186: 3135: 3089: 3036:"A new hangingfly (Insecta: Mecoptera: Bittacidae) from the Middle Jurassic of Inner Mongolia, China" 2914: 2855: 2663: 2533: 2453: 2357: 2320: 2246: 2168: 1191: 1137: 1109: 1082: 1026: 998: 970: 914: 886: 858: 830: 752: 619:
found an overall similarity between the fossil animals found in the Daohugou Beds and those from the
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period (122 ma). One of the first studies on the age of the Daohugou beds, published in 2004 by He
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show much resemblance to those of the Yixian Formation. In other words, despite the absence of
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Liaoning Stratigraphic Group. 1978. The Stratigraphic Tables of Liaoning. Geology Press 1-296
2979:"A golden orb-weaver spider (Araneae: Nephilidae: Nephila) from the Middle Jurassic of China" 2656:"New archidermapteran earwigs (Dermaptera) from the Middle Jurassic of Inner Mongolia, China" 4554: 4417: 4329: 4271: 4198: 4056: 4046: 3970: 3960: 3945: 3745: 3733: 3692: 3633: 3589: 3579: 3540: 3505: 3453: 3437: 3388: 3349: 3324: 3285: 3243: 3194: 3147: 3097: 3006: 2990: 2940: 2922: 2873: 2863: 2811: 2776: 2743: 2725: 2681: 2671: 2631: 2592: 2551: 2541: 2502: 2461: 2404: 2365: 2328: 2254: 2176: 2073: 987: 620: 581: 573: 487: 3734:"How deep is the conflict between molecular and fossil evidence on the age of angiosperms?" 1842: 4471: 4352: 4216: 4104: 4071: 4008: 3950: 3917: 3902: 3897: 3887: 3318: 1935: 1594: 1569: 903: 601: 569: 536: 3393: 3376: 2797:"Two Middle Jurassic hanging-flies (Insecta: Mecoptera: Bittacidae) from Northeast China" 3688: 3622:"Supposed Jurassic angiosperms lack pentamery, an important angiosperm-specific feature" 3536: 3501: 3458: 3433: 3417: 3281: 3239: 3190: 3093: 2918: 2859: 2796: 2748: 2713: 2667: 2537: 2457: 2361: 2324: 2250: 2172: 653:
concluded that the Daohugou probably represents the earliest evolutionary stages of the
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He, H.; Wang, X.; Zhou, Z.; Zhu, R.; Jin, F.; Wang, F.; Ding, X.; Boven, A. (2004).
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published their own study of the age of the Daohugou beds, this time using Zircon
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Proceedings of the National Academy of Sciences of the United States of America
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Gao, Wei; Xu, Yifan; Shih, Chungkun; Ren, Dong; Wang, Yongjie (2023-06-18).
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Xiong, Shurong; Engel, Michael; Xiao, Lifang; Ren, Dong (26 October 2021).
2565: 2206:[Dahugou bed and fossil record of its basal conglomerate section]. 1473: 777: 2636: 2619: 1727: 4479: 4357: 4296: 4232: 4109: 4096: 2846: 2333: 2308: 2233:
Xu, Xing; Zhou, Zhonghe; Sullivan, Corwin; Wang, Yuan; Ren, Dong (2016).
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Yin, Yuqing; Shih, Chungkun; Engel, Michael S.; Ren, Dong (2023-07-07).
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Jin, F.; Zhang, F.C.; Li, Z.H.; Zhang, J.Y.; Li, C.; Zhou, Z.H. (2008).
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argued that the key indicator of the Jehol biota are the index fossils
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Xin Wing; Shuying Duan; Baoyin Geng; Jinzhong Cui; Yong Yang (2007).
3377:"The Mesozoic Acipenseriformes in northeast China and adjacent areas" 3336:
Mao, Fangyuan; Li, Zhiheng; Hooker, Jerry J; Meng, Jin (2023-11-01).
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period's Asian prehistoric life, geological formations, fossil record
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Liang, Junhui; Wang, Ying; Shih, Chungkun; Ren, Dong (2023-01-02).
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The most prominent locality of the Haifanggou Formation are the
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Li, Fang-Yu; Xiu, Yun-Yan; Tan, Xiao; Tian, Ning (2023-06-26).
2903:"Jurassic mimicry between a hangingfly and a ginkgo from China" 2442:
and the early vertebrate fossil assemblages of the Jehol Biota"
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Lian, Xin-Neng; Cai, Chen-Yang; Huang, Di-Ying (2021-04-29).
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Wang, Qi; Shih, Chungkun; Ren, Dong; Carrier, David (2013).
470:), is a fossil-bearing rock deposit located near Daohugou ( 3416:
Wu, Feixiang; Janvier, Philippe; Zhang, Chi (2023-10-31).
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Selden, Paul A.; Shih, ChungKun; Ren, Dong (2011-10-23).
2522:"Late Jurassic salamandroid from western Liaoning, China" 2202:
Huang, D.; Cai, C.; Jiang, J.; Su, Y.; Liao, H. (2015).
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IOP Conference Series: Earth and Environmental Science
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Yanxue, Liu; Yongqing, Liu; Hong, Zhang (2010-09-07).
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Coiro, Mario; Doyle, James A.; Hilton, Jason (2019).
4545: 4470: 4425: 4416: 4343: 4295: 4207: 4149: 4095: 4022: 3994: 3936: 3878: 3867: 3034:Petrulevicius, J. F.; Huang, D-Y.; Ren, D. (2007). 338: 330: 320: 315: 297: 287: 248: 209: 204: 182: 172: 167: 151: 141: 131: 32: 3228:Palaeogeography, Palaeoclimatology, Palaeoecology 2421:: CS1 maint: DOI inactive as of September 2024 ( 3568:"Schmeissneria: A missing link to angiosperms?" 2835: 2833: 2773:Proceedings of the National Academy of Sciences 2526:Proceedings of the National Academy of Sciences 592:within the Daohugou Beds to determine its age. 3824: 2618:Zhang, K.; Yang, D.; Ren, D.; Ge, F. (2008). 2154: 2152: 8: 3717:: CS1 maint: multiple names: authors list ( 3418:"The rise of predation in Jurassic lampreys" 388: 360: 4422: 3875: 3831: 3817: 3809: 29: 4645:Fossiliferous stratigraphic units of Asia 3593: 3583: 3525:Palaeobiodiversity and Palaeoenvironments 3457: 3392: 3342:Zoological Journal of the Linnean Society 3198: 3010: 2944: 2926: 2877: 2867: 2790: 2788: 2747: 2729: 2685: 2675: 2635: 2555: 2545: 2465: 2332: 611:Another study, published in 2006 by Wang 3263:"Earliest known crown-group salamanders" 2039:Uncertain plant originally described as 2009:Uncertain plant originally described as 1979:Uncertain plant originally described as 1858: 710: 3677:Acta Geologica Sinica - English Edition 2495:Acta Geologica Sinica - English Edition 2350:Acta Geologica Sinica - English Edition 2239:Acta Geologica Sinica - English Edition 2148: 3710: 2414: 572:period (169 million years ago) to the 3615: 3613: 3490:Review of Palaeobotany and Palynology 3479: 3477: 3261:Gao, Ke-Qin; Shubin, Neil H. (2003). 3129: 3127: 2795:Li, Y-L.; Ren, D.; Shih, C-K (2008). 2707: 2705: 2649: 2647: 2127:, fossiliferous formation of Tanzania 396:Haifanggou Formation (Inner Mongolia) 7: 2520:Gao, K. -Q.; Shubin, N. H. (2012). 2507:10.1111/j.1755-6724.2006.tb00296.x 2370:10.1111/j.1755-6724.2006.tb00793.x 2104:, fossiliferous formations of the 2086:, fossiliferous formation of Niger 25: 2133:, fossiliferous formation of the 501:The formation consists of coarse 3697:10.1111/j.1755-6724.2010.00169.x 3200:10.1111/j.1475-4983.2008.00773.x 1953:The earliest known redwood tree 1841: 1647: 1308: 584:. The 2004 study primarily used 387: 380: 359: 352: 302: 4532:Plain and Bordered White Banner 2775:, (advance online publication) 2597:10.11646/palaeoentomology.4.2.1 4650:Paleontology in Inner Mongolia 3986:Darhan'muminggan United Banner 3510:10.1016/j.revpalbo.2023.104937 3394:10.1088/1755-1315/864/1/012005 483: 475: 466: 457: 449: 436: 427: 419: 1: 3152:10.1080/08912963.2023.2224377 2624:Acta Palaeontologica Polonica 2391:Wang, Y; et al. (2006). 27:Geological formation in China 3248:10.1016/j.palaeo.2023.111818 3051:(1): 145–152. Archived from 2869:10.1371/journal.pone.0071378 2313:Geophysical Research Letters 679:fossils. Later in 2006, Liu 627:"vertebrate fossils such as 368:Haifanggou Formation (China) 334:Liaoning Stratigraphic Group 250:Approximate paleocoordinates 3173:Huang, D.; Nel, A. (2008). 2397:Progress in Natural Science 2208:Acta Palaeontologica Sinica 764: 531:The formation dates to the 4666: 4457:Horqin Right Middle Banner 3545:10.1007/s12549-020-00454-z 3442:10.1038/s41467-023-42251-0 3354:10.1093/zoolinnean/zlad050 3102:10.1007/s11756-022-01300-7 2677:10.3897/zookeys.1065.72720 2409:10.1080/10020070612330087A 2131:Cañadón Calcáreo Formation 399:Show map of Inner Mongolia 4452:Horqin Right Front Banner 4394:Qahar Right Middle Banner 4136:Horqin Left Middle Banner 3966:Bayan'obo Mining District 3853: 3375:Li, X.; Chen, Y. (2021). 2816:10.11646/zootaxa.1929.1.2 2467:10.1007/s11434-008-0209-5 2181:10.1007/s11434-006-2165-2 2056:Geology of Inner Mongolia 1888: 1549: 623:. The authors stated that 346: 37: 4389:Qahar Right Front Banner 3572:BMC Evolutionary Biology 2446:Chinese Science Bulletin 2203: 2161:Chinese Science Bulletin 1691:A docodont mammaliaform 1670:A docodont mammaliaform 4610:Conglomerate formations 4590:Jurassic System of Asia 4399:Qahar Right Rear Banner 4141:Horqin Left Rear Banner 2928:10.1073/pnas.1205517109 2781:10.1073/pnas.1118372109 2731:10.3390/insects14070614 2547:10.1073/pnas.1009828109 2259:10.1111/1755-6724.13033 2065:Jurassic System of Asia 4527:Bordered Yellow Banner 4287:Morin'dawa Daur Banner 4262:New Barag Right Banner 3840:County-level divisions 3585:10.1186/1471-2148-7-14 2995:10.1098/rsbl.2011.0228 2411:(inactive 2024-09-18). 2210:(in Chinese (China)). 1788: 1731: 1699: 1619: 1587: 1562: 1530: 1477: 1445: 1413: 1381: 784: 773: 647: 4267:New Barag Left Banner 4222:Zhalainuo'er District 3422:Nature Communications 3044:African Invertebrates 2637:10.4202/app.2008.0112 2135:Cañadón Asfalto Basin 1787: 1751:an arboreal/climbing 1730: 1698: 1618: 1586: 1561: 1529: 1476: 1444: 1412: 1380: 780: 769: 696:Animal fossil content 625: 442:Jiulongshan Formation 440:), also known as the 18:Jiulongshan Formation 4625:Sandstone formations 4037:Yuanbaoshan District 2334:10.1029/2004gl020792 2114:Oxfordian formations 2079:Callovian formations 1755:mammal/mammaliaform 1236:Stictopolystoechotes 517:layers deposited in 412:Haifanggou Formation 146:Tiaojishan Formation 136:Geological formation 33:Haifanggou Formation 4640:Lacustrine deposits 4615:Mudstone formations 4517:West Ujimqin Banner 4512:East Ujimqin Banner 4335:Hanggin Rear Banner 4067:Bairin Right Banner 3689:2010AcGlS..84...47W 3626:The New Phytologist 3537:2021PdPe..101...25M 3502:2023RPaPa.31604937L 3434:2023NatCo..14.6652W 3290:10.1038/nature01491 3282:2003Natur.422..424G 3240:2023PPP...63011818L 3191:2008Palgy..51..715H 3094:2023Biolg..78.1429L 2919:2012PNAS..10920514W 2913:(50): 20514–20519. 2860:2013PLoSO...871378W 2668:2021ZooK.1065..125X 2538:2012PNAS..109.5767G 2458:2008SciBu..53.2820J 2438:"On the horizon of 2362:2006AcGlS..80...42G 2325:2004GeoRL..3120609H 2251:2016AcGlS..90.2229X 2173:2006ChSBu..51.2634L 2125:Tendaguru Formation 2084:Tiourarén Formation 1154:Pycnopolystoechotes 685:Uranium-lead dating 673:Huajiying Formation 480:traditional Chinese 454:traditional Chinese 424:traditional Chinese 268: /  229: /  39:Stratigraphic range 4507:Sonid Right Banner 4325:Urad Middle Banner 4164:Kangbashi District 4159:Dongsheng District 4062:Bairin Left Banner 3981:Tumed Right Banner 3140:Historical Biology 1789: 1732: 1700: 1620: 1588: 1563: 1531: 1478: 1446: 1414: 1382: 1267:Based largely on. 785: 774: 586:Argon–argon dating 472:simplified Chinese 446:simplified Chinese 416:simplified Chinese 4577: 4576: 4573: 4572: 4560:Alxa Right Banner 4537:Plain Blue Banner 4502:Sonid Left Banner 4412: 4411: 4384:Liangcheng County 4320:Urad Front Banner 4179:Otog Front Banner 4042:Songshan District 4032:Hongshan District 4004:Haibowan District 3956:Qingshan District 3928:Tumed Left Banner 3923:Qingshuihe County 3893:Xincheng District 3869:Prefecture-level 3847:Autonomous Region 3780:"PBDB Collection" 3750:10.1111/nph.15708 3638:10.1111/nph.15974 3276:(6930): 424–428. 2662:(1065): 125–139. 2452:(18): 2820–2827. 2290:Geol. Bull. China 2204:道虎沟化石层及其底部砾岩段化石记录 2167:(21): 2634–2644. 2047: 2046: 1849: 1848: 1713:M. mammaliaformis 1552:scansoriopterygid 1427:J. ningchengensis 1395:J. ningchengensis 1260: 1259: 1208:Sinopalaeodermata 1187:S. ningchengensis 960:Juracimbrophlebia 771:Juracimbrophlebia 761:tangle-veined fly 748:A. neimengguensis 408: 407: 371:Show map of China 159:(Separated by an 156:Beipiao Formation 16:(Redirected from 4657: 4635:Deltaic deposits 4620:Shale formations 4555:Alxa Left Banner 4423: 4330:Urad Rear Banner 4272:Old Barag Banner 4199:Ejin'horo Banner 4057:Ar'horqin Banner 4047:Ningcheng County 3971:Jiuyuan District 3961:Shiguai District 3946:Hondlon District 3876: 3833: 3826: 3819: 3810: 3794: 3793: 3791: 3790: 3776: 3770: 3769: 3729: 3723: 3722: 3716: 3708: 3672: 3666: 3665: 3617: 3608: 3607: 3597: 3587: 3563: 3557: 3556: 3520: 3514: 3513: 3481: 3472: 3471: 3461: 3413: 3407: 3406: 3396: 3372: 3366: 3365: 3333: 3327: 3325:Science Magazine 3316: 3310: 3309: 3267: 3258: 3252: 3251: 3219: 3213: 3212: 3202: 3170: 3164: 3163: 3146:(8): 1672–1676. 3131: 3122: 3121: 3088:(6): 1429–1432. 3073: 3067: 3066: 3064: 3063: 3057: 3040: 3031: 3025: 3024: 3014: 2974: 2968: 2965: 2959: 2958: 2948: 2930: 2898: 2892: 2891: 2881: 2871: 2837: 2828: 2827: 2801: 2792: 2783: 2768: 2762: 2761: 2751: 2733: 2709: 2700: 2699: 2689: 2679: 2651: 2642: 2641: 2639: 2615: 2609: 2608: 2585:Palaeoentomology 2576: 2570: 2569: 2559: 2549: 2517: 2511: 2510: 2486: 2480: 2479: 2469: 2433: 2427: 2426: 2420: 2412: 2388: 2382: 2381: 2345: 2339: 2338: 2336: 2304: 2298: 2297: 2285: 2279: 2278: 2245:(6): 2229–2243. 2230: 2224: 2223: 2199: 2193: 2192: 2156: 2074:Yixian Formation 1856: 1845: 1651: 1512:P. daohugouensis 1346:N. daohugouensis 1312: 1270: 988:Jurahylobittacus 820:Applanatiforceps 708: 621:Yixian Formation 582:Yixian Formation 574:Early Cretaceous 488:Ningcheng County 485: 477: 468: 459: 451: 438: 429: 421: 400: 391: 390: 384: 372: 363: 362: 356: 308: 306: 305: 283: 282: 280: 279: 278: 273: 269: 266: 265: 264: 261: 244: 243: 241: 240: 239: 234: 230: 227: 226: 225: 222: 127: 57: 30: 21: 4665: 4664: 4660: 4659: 4658: 4656: 4655: 4654: 4630:Coal formations 4605:Oxfordian Stage 4600:Callovian Stage 4580: 4579: 4578: 4569: 4541: 4466: 4408: 4353:Jining District 4339: 4291: 4217:Hailar District 4203: 4145: 4105:Horqin District 4091: 4072:Hexigten Banner 4018: 4009:Hainan District 3990: 3951:Donghe District 3932: 3918:Horinger County 3903:Saihan District 3898:Yuquan District 3888:Huimin District 3870: 3863: 3849: 3837: 3802: 3800:Further reading 3797: 3788: 3786: 3778: 3777: 3773: 3738:New Phytologist 3731: 3730: 3726: 3709: 3674: 3673: 3669: 3619: 3618: 3611: 3565: 3564: 3560: 3522: 3521: 3517: 3483: 3482: 3475: 3415: 3414: 3410: 3374: 3373: 3369: 3335: 3334: 3330: 3317: 3313: 3265: 3260: 3259: 3255: 3221: 3220: 3216: 3172: 3171: 3167: 3133: 3132: 3125: 3075: 3074: 3070: 3061: 3059: 3055: 3038: 3033: 3032: 3028: 2983:Biology Letters 2976: 2975: 2971: 2966: 2962: 2900: 2899: 2895: 2839: 2838: 2831: 2799: 2794: 2793: 2786: 2769: 2765: 2711: 2710: 2703: 2653: 2652: 2645: 2617: 2616: 2612: 2578: 2577: 2573: 2532:(15): 5767–72. 2519: 2518: 2514: 2488: 2487: 2483: 2435: 2434: 2430: 2413: 2390: 2389: 2385: 2347: 2346: 2342: 2306: 2305: 2301: 2287: 2286: 2282: 2232: 2231: 2227: 2205: 2201: 2200: 2196: 2158: 2157: 2150: 2146: 2102:Qigu Formations 2052: 1854: 1828:Y. ingensdentes 1633:C. lutrasimilis 1595:Volaticotherium 1570:Epidendrosaurus 1265: 1105:M. daohugoensis 904:Formosibittacus 775: 703: 698: 602:biostratigraphy 570:Middle Jurassic 565: 560: 537:Middle Jurassic 494:, northeastern 404: 403: 402: 401: 398: 397: 394: 393: 392: 375: 374: 373: 370: 369: 366: 365: 364: 303: 301: 276: 274: 270: 267: 262: 259: 257: 255: 254: 237: 235: 231: 228: 223: 220: 218: 216: 215: 158: 126: 125: 124: 123: 118: 113: 108: 103: 98: 93: 88: 83: 78: 73: 68: 63: 52: 51: 49: 28: 23: 22: 15: 12: 11: 5: 4663: 4661: 4653: 4652: 4647: 4642: 4637: 4632: 4627: 4622: 4617: 4612: 4607: 4602: 4597: 4595:Jurassic China 4592: 4582: 4581: 4575: 4574: 4571: 4570: 4568: 4567: 4562: 4557: 4551: 4549: 4543: 4542: 4540: 4539: 4534: 4529: 4524: 4519: 4514: 4509: 4504: 4499: 4494: 4489: 4483: 4476: 4474: 4468: 4467: 4465: 4464: 4459: 4454: 4449: 4444: 4438: 4431: 4429: 4420: 4414: 4413: 4410: 4409: 4407: 4406: 4401: 4396: 4391: 4386: 4381: 4376: 4374:Shangdu County 4371: 4366: 4361: 4355: 4349: 4347: 4341: 4340: 4338: 4337: 4332: 4327: 4322: 4317: 4315:Dengkou County 4312: 4307: 4305:Linhe District 4301: 4299: 4293: 4292: 4290: 4289: 4284: 4279: 4274: 4269: 4264: 4259: 4254: 4248: 4242: 4236: 4230: 4224: 4219: 4213: 4211: 4205: 4204: 4202: 4201: 4196: 4191: 4189:Hanggin Banner 4186: 4181: 4176: 4171: 4166: 4161: 4155: 4153: 4147: 4146: 4144: 4143: 4138: 4133: 4128: 4123: 4118: 4113: 4107: 4101: 4099: 4093: 4092: 4090: 4089: 4084: 4079: 4077:Ongniud Banner 4074: 4069: 4064: 4059: 4054: 4049: 4044: 4039: 4034: 4028: 4026: 4020: 4019: 4017: 4016: 4011: 4006: 4000: 3998: 3992: 3991: 3989: 3988: 3983: 3978: 3973: 3968: 3963: 3958: 3953: 3948: 3942: 3940: 3934: 3933: 3931: 3930: 3925: 3920: 3915: 3913:Wuchuan County 3910: 3905: 3900: 3895: 3890: 3884: 3882: 3873: 3865: 3864: 3854: 3851: 3850: 3844:Inner Mongolia 3838: 3836: 3835: 3828: 3821: 3813: 3807: 3806: 3801: 3798: 3796: 3795: 3784:paleobiodb.org 3771: 3724: 3667: 3632:(2): 420–426. 3609: 3558: 3515: 3473: 3408: 3367: 3348:(3): 832–859. 3328: 3311: 3253: 3214: 3185:(3): 715–719. 3165: 3123: 3068: 3026: 2989:(5): 775–778. 2969: 2960: 2893: 2829: 2784: 2763: 2701: 2643: 2630:(1): 161–164. 2610: 2591:(2): 095–136. 2571: 2512: 2501:(5): 733–742. 2481: 2428: 2403:(S1): 222–30. 2383: 2340: 2319:(20): L20609. 2299: 2280: 2225: 2194: 2147: 2145: 2142: 2141: 2140: 2139: 2138: 2128: 2122: 2111: 2110: 2109: 2087: 2076: 2071: 2058: 2051: 2048: 2045: 2044: 2037: 2035: 2030: 2025: 2017: 2016: 2014: 2007: 2005: 2000: 1995: 1987: 1986: 1984: 1977: 1975: 1970: 1965: 1957: 1956: 1954: 1951: 1949: 1944: 1939: 1931: 1930: 1928: 1925: 1923: 1918: 1912: 1911: 1909: 1906: 1904: 1899: 1894: 1886: 1885: 1882: 1879: 1876: 1873: 1870: 1866: 1865: 1853: 1850: 1847: 1846: 1839: 1833: 1830: 1825: 1817: 1816: 1814: 1807: 1804: 1799: 1791: 1790: 1781: 1774: 1772: 1767: 1759: 1758: 1756: 1749: 1747: 1742: 1734: 1733: 1724: 1717: 1715: 1710: 1702: 1701: 1692: 1689: 1687: 1682: 1674: 1673: 1671: 1668: 1666: 1661: 1653: 1652: 1645: 1638: 1635: 1630: 1622: 1621: 1612: 1605: 1603: 1598: 1590: 1589: 1580: 1578: 1576:ningchengensis 1573: 1565: 1564: 1555: 1548: 1546: 1541: 1533: 1532: 1523: 1516: 1514: 1509: 1501: 1500: 1498: 1495: 1493: 1488: 1485:Daohugoupterus 1480: 1479: 1470: 1463: 1461: 1459:P. wellnhoferi 1456: 1448: 1447: 1438: 1431: 1429: 1424: 1416: 1415: 1406: 1399: 1397: 1392: 1384: 1383: 1374: 1371: 1369: 1364: 1356: 1355: 1353: 1350: 1348: 1343: 1335: 1334: 1332: 1329: 1327: 1322: 1314: 1313: 1306: 1303: 1300: 1298:C. tianyiensis 1295: 1287: 1286: 1283: 1280: 1277: 1274: 1264: 1261: 1258: 1257: 1251: 1249: 1244: 1239: 1231: 1230: 1224: 1221: 1216: 1211: 1203: 1202: 1196: 1194: 1189: 1184: 1176: 1175: 1169: 1167: 1162: 1157: 1149: 1148: 1142: 1140: 1135: 1130: 1122: 1121: 1115: 1112: 1107: 1102: 1099:Mongolbittacus 1094: 1093: 1087: 1085: 1080: 1075: 1067: 1066: 1060: 1057: 1052: 1047: 1039: 1038: 1032: 1029: 1024: 1019: 1011: 1010: 1004: 1001: 996: 991: 983: 982: 976: 973: 968: 966:J. ginkgofolia 963: 955: 954: 948: 945: 940: 935: 927: 926: 920: 917: 912: 907: 899: 898: 892: 889: 884: 882:E. gracilentum 879: 871: 870: 864: 861: 856: 851: 843: 842: 836: 833: 828: 823: 815: 814: 808: 805: 800: 795: 787: 786: 763: 757: 755: 750: 745: 737: 736: 733: 730: 727: 724: 721: 717: 716: 702: 699: 697: 694: 637:and feathered 564: 561: 559: 556: 492:Inner Mongolia 467:Jiǔlóngshān zǔ 406: 405: 395: 386: 385: 379: 378: 377: 376: 367: 358: 357: 351: 350: 349: 348: 347: 344: 343: 340: 336: 335: 332: 328: 327: 322: 318: 317: 313: 312: 299: 295: 294: 292:Inner Mongolia 289: 285: 284: 272:44.5°N 126.0°E 252: 246: 245: 233:41.8°N 120.8°E 213: 207: 206: 202: 201: 184: 180: 179: 174: 170: 169: 165: 164: 153: 149: 148: 143: 139: 138: 133: 129: 128: 121: 120: 119: 114: 109: 104: 99: 94: 89: 84: 79: 74: 69: 64: 59: 58: 35: 34: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 4662: 4651: 4648: 4646: 4643: 4641: 4638: 4636: 4633: 4631: 4628: 4626: 4623: 4621: 4618: 4616: 4613: 4611: 4608: 4606: 4603: 4601: 4598: 4596: 4593: 4591: 4588: 4587: 4585: 4566: 4563: 4561: 4558: 4556: 4553: 4552: 4550: 4548: 4544: 4538: 4535: 4533: 4530: 4528: 4525: 4523: 4522:Taibus Banner 4520: 4518: 4515: 4513: 4510: 4508: 4505: 4503: 4500: 4498: 4495: 4493: 4492:Duolun County 4490: 4487: 4484: 4481: 4478: 4477: 4475: 4473: 4469: 4463: 4462:Jalaid Banner 4460: 4458: 4455: 4453: 4450: 4448: 4447:Tuquan County 4445: 4442: 4439: 4436: 4433: 4432: 4430: 4428: 4424: 4421: 4419: 4415: 4405: 4404:Dorbod Banner 4402: 4400: 4397: 4395: 4392: 4390: 4387: 4385: 4382: 4380: 4379:Xinghe County 4377: 4375: 4372: 4370: 4367: 4365: 4364:Zhuozi County 4362: 4359: 4356: 4354: 4351: 4350: 4348: 4346: 4342: 4336: 4333: 4331: 4328: 4326: 4323: 4321: 4318: 4316: 4313: 4311: 4310:Wuyuan County 4308: 4306: 4303: 4302: 4300: 4298: 4294: 4288: 4285: 4283: 4280: 4278: 4277:Oroqin Banner 4275: 4273: 4270: 4268: 4265: 4263: 4260: 4258: 4255: 4252: 4249: 4246: 4243: 4240: 4237: 4234: 4231: 4228: 4225: 4223: 4220: 4218: 4215: 4214: 4212: 4210: 4206: 4200: 4197: 4195: 4192: 4190: 4187: 4185: 4182: 4180: 4177: 4175: 4174:Jungar Banner 4172: 4170: 4167: 4165: 4162: 4160: 4157: 4156: 4154: 4152: 4148: 4142: 4139: 4137: 4134: 4132: 4129: 4127: 4126:Naiman Banner 4124: 4122: 4119: 4117: 4114: 4111: 4108: 4106: 4103: 4102: 4100: 4098: 4094: 4088: 4085: 4083: 4082:Harqin Banner 4080: 4078: 4075: 4073: 4070: 4068: 4065: 4063: 4060: 4058: 4055: 4053: 4050: 4048: 4045: 4043: 4040: 4038: 4035: 4033: 4030: 4029: 4027: 4025: 4021: 4015: 4014:Wuda District 4012: 4010: 4007: 4005: 4002: 4001: 3999: 3997: 3993: 3987: 3984: 3982: 3979: 3977: 3976:Guyang County 3974: 3972: 3969: 3967: 3964: 3962: 3959: 3957: 3954: 3952: 3949: 3947: 3944: 3943: 3941: 3939: 3935: 3929: 3926: 3924: 3921: 3919: 3916: 3914: 3911: 3909: 3908:Togtoh County 3906: 3904: 3901: 3899: 3896: 3894: 3891: 3889: 3886: 3885: 3883: 3881: 3877: 3874: 3872: 3866: 3861: 3857: 3852: 3848: 3845: 3841: 3834: 3829: 3827: 3822: 3820: 3815: 3814: 3811: 3804: 3803: 3799: 3785: 3781: 3775: 3772: 3767: 3763: 3759: 3755: 3751: 3747: 3743: 3739: 3735: 3728: 3725: 3720: 3714: 3706: 3702: 3698: 3694: 3690: 3686: 3682: 3678: 3671: 3668: 3663: 3659: 3655: 3651: 3647: 3643: 3639: 3635: 3631: 3627: 3623: 3616: 3614: 3610: 3605: 3601: 3596: 3591: 3586: 3581: 3577: 3573: 3569: 3562: 3559: 3554: 3550: 3546: 3542: 3538: 3534: 3530: 3526: 3519: 3516: 3511: 3507: 3503: 3499: 3495: 3491: 3487: 3480: 3478: 3474: 3469: 3465: 3460: 3455: 3451: 3447: 3443: 3439: 3435: 3431: 3427: 3423: 3419: 3412: 3409: 3404: 3400: 3395: 3390: 3387:(1): 012005. 3386: 3382: 3378: 3371: 3368: 3363: 3359: 3355: 3351: 3347: 3343: 3339: 3332: 3329: 3326: 3322: 3321: 3315: 3312: 3307: 3303: 3299: 3295: 3291: 3287: 3283: 3279: 3275: 3271: 3264: 3257: 3254: 3249: 3245: 3241: 3237: 3233: 3229: 3225: 3218: 3215: 3210: 3206: 3201: 3196: 3192: 3188: 3184: 3180: 3179:Palaeontology 3176: 3169: 3166: 3161: 3157: 3153: 3149: 3145: 3141: 3137: 3130: 3128: 3124: 3119: 3115: 3111: 3107: 3103: 3099: 3095: 3091: 3087: 3083: 3079: 3072: 3069: 3058:on 2012-03-21 3054: 3050: 3046: 3045: 3037: 3030: 3027: 3022: 3018: 3013: 3008: 3004: 3000: 2996: 2992: 2988: 2984: 2980: 2973: 2970: 2964: 2961: 2956: 2952: 2947: 2942: 2938: 2934: 2929: 2924: 2920: 2916: 2912: 2908: 2904: 2897: 2894: 2889: 2885: 2880: 2875: 2870: 2865: 2861: 2857: 2854:(8): e71378. 2853: 2849: 2848: 2843: 2836: 2834: 2830: 2825: 2821: 2817: 2813: 2809: 2805: 2798: 2791: 2789: 2785: 2782: 2778: 2774: 2767: 2764: 2759: 2755: 2750: 2745: 2741: 2737: 2732: 2727: 2723: 2719: 2715: 2708: 2706: 2702: 2697: 2693: 2688: 2683: 2678: 2673: 2669: 2665: 2661: 2657: 2650: 2648: 2644: 2638: 2633: 2629: 2625: 2621: 2614: 2611: 2606: 2602: 2598: 2594: 2590: 2586: 2582: 2575: 2572: 2567: 2563: 2558: 2553: 2548: 2543: 2539: 2535: 2531: 2527: 2523: 2516: 2513: 2508: 2504: 2500: 2496: 2492: 2485: 2482: 2477: 2473: 2468: 2463: 2459: 2455: 2451: 2447: 2443: 2441: 2432: 2429: 2424: 2418: 2410: 2406: 2402: 2398: 2394: 2387: 2384: 2379: 2375: 2371: 2367: 2363: 2359: 2355: 2351: 2344: 2341: 2335: 2330: 2326: 2322: 2318: 2314: 2310: 2303: 2300: 2295: 2291: 2284: 2281: 2276: 2272: 2268: 2264: 2260: 2256: 2252: 2248: 2244: 2240: 2236: 2229: 2226: 2221: 2217: 2213: 2209: 2198: 2195: 2190: 2186: 2182: 2178: 2174: 2170: 2166: 2162: 2155: 2153: 2149: 2143: 2136: 2132: 2129: 2126: 2123: 2120: 2117: 2116: 2115: 2112: 2107: 2106:Junggar Basin 2103: 2099: 2095: 2091: 2088: 2085: 2082: 2081: 2080: 2077: 2075: 2072: 2070: 2066: 2062: 2061:Jurassic Asia 2059: 2057: 2054: 2053: 2049: 2042: 2038: 2036: 2034: 2031: 2029: 2026: 2024: 2023: 2019: 2018: 2015: 2012: 2008: 2006: 2004: 2001: 1999: 1996: 1994: 1993: 1992:Xingxueanthus 1989: 1988: 1985: 1982: 1978: 1976: 1974: 1971: 1969: 1966: 1964: 1963: 1962:Schmeissneria 1959: 1958: 1955: 1952: 1950: 1948: 1945: 1943: 1942:S. jeholensis 1940: 1938: 1937: 1933: 1932: 1929: 1926: 1924: 1922: 1919: 1917: 1916:C. burejensis 1914: 1913: 1910: 1907: 1905: 1903: 1900: 1898: 1895: 1893: 1892: 1887: 1883: 1880: 1877: 1874: 1871: 1868: 1867: 1864: 1863: 1857: 1851: 1844: 1840: 1838: 1834: 1831: 1829: 1826: 1824: 1823: 1819: 1818: 1815: 1812: 1808: 1805: 1803: 1800: 1798: 1797: 1793: 1792: 1786: 1782: 1780:mammaliaform 1779: 1775: 1773: 1771: 1768: 1766: 1765: 1761: 1760: 1757: 1754: 1753:euharamiyidan 1750: 1748: 1746: 1743: 1741: 1740: 1736: 1735: 1729: 1725: 1722: 1718: 1716: 1714: 1711: 1709: 1708: 1704: 1703: 1697: 1693: 1690: 1688: 1686: 1685:A, scansorius 1683: 1681: 1680: 1676: 1675: 1672: 1669: 1667: 1665: 1662: 1660: 1659: 1655: 1654: 1650: 1646: 1644:mammaliaform 1643: 1639: 1636: 1634: 1631: 1629: 1628: 1624: 1623: 1617: 1613: 1610: 1609:eutriconodont 1606: 1604: 1602: 1599: 1597: 1596: 1592: 1591: 1585: 1581: 1579: 1577: 1574: 1572: 1571: 1567: 1566: 1560: 1556: 1553: 1547: 1545: 1542: 1540: 1539: 1538:Epidexipteryx 1535: 1534: 1528: 1524: 1521: 1517: 1515: 1513: 1510: 1508: 1507: 1503: 1502: 1499: 1496: 1494: 1492: 1489: 1487: 1486: 1482: 1481: 1475: 1471: 1468: 1467:darwinopteran 1464: 1462: 1460: 1457: 1455: 1454: 1453:Pterorhynchus 1450: 1449: 1443: 1439: 1436: 1432: 1430: 1428: 1425: 1423: 1422: 1418: 1417: 1411: 1407: 1404: 1400: 1398: 1396: 1393: 1391: 1390: 1386: 1385: 1379: 1375: 1373:A salamander 1372: 1370: 1368: 1365: 1363: 1362: 1358: 1357: 1354: 1352:A salamander 1351: 1349: 1347: 1344: 1342: 1341: 1340:Neimengtriton 1337: 1336: 1333: 1331:A salamander 1330: 1328: 1326: 1323: 1321: 1320: 1316: 1315: 1311: 1307: 1305:A salamander 1304: 1301: 1299: 1296: 1294: 1293: 1289: 1288: 1284: 1281: 1278: 1275: 1272: 1271: 1268: 1262: 1256: 1252: 1250: 1248: 1245: 1243: 1240: 1238: 1237: 1233: 1232: 1229: 1225: 1222: 1220: 1217: 1215: 1212: 1210: 1209: 1205: 1204: 1201: 1197: 1195: 1193: 1190: 1188: 1185: 1183: 1182: 1178: 1177: 1174: 1170: 1168: 1166: 1163: 1161: 1158: 1156: 1155: 1151: 1150: 1147: 1143: 1141: 1139: 1136: 1134: 1131: 1129: 1128: 1127:Pseudomantina 1124: 1123: 1120: 1116: 1113: 1111: 1108: 1106: 1103: 1101: 1100: 1096: 1095: 1092: 1088: 1086: 1084: 1081: 1079: 1076: 1074: 1073: 1072:Mongolarachne 1069: 1068: 1065: 1061: 1058: 1056: 1053: 1051: 1048: 1046: 1045: 1041: 1040: 1037: 1033: 1030: 1028: 1025: 1023: 1020: 1018: 1017: 1013: 1012: 1009: 1005: 1002: 1000: 997: 995: 992: 990: 989: 985: 984: 981: 977: 974: 972: 969: 967: 964: 962: 961: 957: 956: 953: 949: 946: 944: 941: 939: 936: 934: 933: 932:Fortiholcorpa 929: 928: 925: 921: 918: 916: 913: 911: 908: 906: 905: 901: 900: 897: 893: 890: 888: 885: 883: 880: 878: 877: 873: 872: 869: 865: 862: 860: 857: 855: 852: 850: 849: 845: 844: 841: 837: 834: 832: 829: 827: 824: 822: 821: 817: 816: 813: 809: 806: 804: 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Banner 4169:Dalad Banner 4131:Jarud Banner 4116:Kailu County 4087:Aohan Banner 4052:Linxi County 3787:. 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Index

Jiulongshan Formation
Stratigraphic range
Callovian
Oxfordian
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N
Geological formation
Tiaojishan Formation
Beipiao Formation
unconformity
Conglomerate
Sandstone
mudstone
shale
coal
Coordinates
41°48′N 120°48′E / 41.8°N 120.8°E / 41.8; 120.8
Approximate paleocoordinates
44°30′N 126°00′E / 44.5°N 126.0°E / 44.5; 126.0

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