3897:
3903:
565:
138:
4453:
553:
160:
453:
874:
628:
4483:
818:
4459:
735:
1493:
884:
579:
skeleton (MPC-D 100/535) from the Barun Goyot
Formation at the Hermiin Tsav locality. In 2010 and 2011 this specimen was examined to analyze several borings (tunnel-like holes) left by invertebrae scavengers on joint areas. As of 2019, MPC-D 100/535 remains largely undescribed. In 2019 a partial
792:
and therefore, junior synonyms. Czepiński reexamined many of the specimens originally described by Maryańska and Osmólska, as well as the respective holotypes of these taxa, providing evidence that all traits used to separate them are, in fact, indistinguishably present on
749:. Its holotype (PIN 3142/299) is represented by a very small and juvenile skull that was collected from the Hermiin Tsav locality near the end of the 1970s by the Joint Soviet–Mongolian Paleontological Expedition. Though Alifanov used this skull to erect the new
1403:
field paleoenvironments, under a semi-arid climate. The formation is known for its vertebrate fossils in life-like poses, most of which are preserved in unstructured sandstone, indicating a catastrophic rapid burial.
2726:
3026:
Pei, R.; Qin, Yuying; Wen, Aishu; Zhao, Q.; Wang, Z.; Liu, Z.; Guo, W.; Liu, P.; Ye, W.; Wang, L.; Yin, Z.; Dai, R.; Xu, X. (2022). "A new troodontid from the Upper
Cretaceous Gobi Basin of inner Mongolia, China".
607:, and evaluated the implications of these specimens for correlation of fossil sites of the latter formation. He considered one of these specimens in particular, MPC-D 100/551B, as a potential evidence of an
2752:
467:
During the large field work of the Polish-Mongolian
Palaeontological Expeditions in the 1970s, abundant protoceratopsid specimens were discovered on eroded surfaces of the Hermiin Tsav locality of the
2168:
Averianov, A. O.; Lopatin, A. V. (2022). "A re-appraisal of
Parvicursor remotus from the Late Cretaceous of Mongolia: implications for the phylogeny and taxonomy of alvarezsaurid theropod dinosaurs".
426:
was among the smallest ceratopsians, growing up to 1–1.5 m (3.3–4.9 ft) in length, with a weight about 22.7–45 kg (50–99 lb). Although emerging late in the reign of the dinosaurs,
2653:"Djadokhta Formation correlative strata in Chinese Inner Mongolia: an overview of the stratigraphy, sedimentary geology, and paleontology and comparisons with the type locality in the pre-Altai Gobi"
592:. This specimen was discovered in 2007 also from the Hermiin Tsav locality of the Barun Goyot Formation, and includes a partially preserved skull with partial skeleton of an adult individual.
937:, were long considered valid and distinct taxa, and sometimes placed within Protoceratopsidae, Czepiński found the diagnostic features used to distinguish these taxa to be largely present in
1558:
Watabe, M., Suzuki, S., 2000a. Report on the Japan-Mongolia Joint
Paleontological Expedition to the Gobi desert, 1993. Hayashibara Museum of Natural SciencesResearch Bulletin 1, 17 29.
2690:
Eberth, D. A. (1993). "Depositional environments and facies transitions of dinosaur-bearing Upper
Cretaceous redbeds at Bayan Mandahu (Inner Mongolia, People's Republic of China)".
3070:
Gao, K.; Hou, L. (1996). "Systematics and taxonomic diversity of squamates from the Upper
Cretaceous Djadochta Formation, Bayan Mandahu, Gobi Desert, People's Republic of China".
2880:
Godefroit, P.; Currie, P. J.; Li, H.; Shang, C. Y.; Dong, Z.-M. (2008). "A new species of
Velociraptor (Dinosauria: Dromaeosauridae) from the Upper Cretaceous of northern China".
2006:
Eberth, D. A. (2018). "Stratigraphy and paleoenvironmental evolution of the dinosaur-rich
Baruungoyot-Nemegt succession (Upper Cretaceous), Nemegt Basin, southern Mongolia".
1762:
Alifanov, V. R. (2003). "Two new dinosaurs of the infraorder
Neoceratopsia (Ornithischia) from the Upper Cretaceous of the Nemegt Depression, Mongolian People's Republic".
1647:"The postcranial skeleton of Bagaceratops (Ornithischia: Neoceratopsia) from the Baruungoyot Formation (Upper Cretaceous) in Hermiin Tsav of southwestern Gobi, Mongolia"
831:
was a small-sized protoceratopsid, reaching adult dimensions of about 1–1.5 m (3.3–4.9 ft) in length and 22.7–45 kg (50–99 lb) in body mass based on
731:
based on the reasoning that all exhibit anatomical traits already seen on other specimens of this protoceratopsid, and some of them are likely products of preservation.
476:
723:
region of a large individual and was collected during expeditions led by the Sino-Canadian Dinosaur Project. In 2006 Mackoviky regarded all of these ceratopsians as
1817:
1603:"Trace fossils on dinosaur bones from Upper Cretaceous eolian deposits in Mongolia: Taphonomic interpretation of paleoecosystems in ancient desert environments"
2571:
Gradziński, R.; Jerzykiewicz, T. (1974). "Dinosaur- and mammal-bearing aeolian and associated deposits of the Upper Cretaceous in the Gobi Desert (Mongolia)".
4604:
2090:
Arbour, V. M.; Currie, P. J.; Badamgarav, D. (2014). "The ankylosaurid dinosaurs of the Upper Cretaceous Baruungoyot and Nemegt formations of Mongolia".
4624:
4619:
2487:
Funston, G. F.; Mendonca, S. E.; Currie, P. J.; Barsbold, R.; Barsbold, R. (2018). "Oviraptorosaur anatomy, diversity and ecology in the Nemegt Basin".
3107:"New Genus and Species of Djadochtatheriid Multituberculate (Allotheria, Mammalia) from the Upper Cretaceous Bayan Mandahu Formation of Inner Mongolia"
4563:
2923:
Xing, X.; Choinere, J. N.; Pittman, M.; Tan, Q. W.; Xiao, D.; Li, Z. Q.; Tan, L.; Clark, J. M.; Norell, M. A.; Hone, D. W. E; Sullivan, C. (2010).
2119:"A new ankylosaurid skeleton from the Upper Cretaceous Baruungoyot Formation of Mongolia: its implications for ankylosaurid postcranial evolution"
4614:
3146:"New exceptionally well-preserved specimens of "Zangerlia" neimongolensis from Bayan Mandahu, Inner Mongolia, and their taxonomic significance"
541:
463:
fossils have been reported from the Bayan Mandahu (bottom right), Hermiin Tsav, Khulsan (both left), and possibly Udyn Sayr (center) localities
2608:"Taxonomic composition and systematics of late Cretaceous lizard assemblages from Ukhaa Tolgod and adjacent localities, Mongolian Gobi Desert"
3896:
1966:
3218:
2821:
Xing, X.; Sullivan, Corwin; Pittman, M.; Choiniere, J. N.; Hone, D. W. E.; Upchurch, P.; Tan, Q.; Xiao, Dong; Lin, Tan; Han, F. (2011).
2117:
Park, J.-Y.; Lee, Y. N.; Currie, P. J.; Ryan, M. J.; Bell, P.; Sissons, R.; Koppelhus, E. B.; Barsbold, R.; Lee, S.; Kim, S.-H. (2021).
511:, which consists of a relatively medium-sized skull, and a vast majority of the specimens collected by the expeditions were assigned to
2523:
1858:"The Tiny Horned Dinosaur Gobiceratops minutus gen. et sp. nov. (Bagaceratopidae, Neoceratopsia) from the Upper Cretaceous of Mongolia"
3902:
4476:
2299:
2243:
1781:
You, H.; Dong, Z. (2003). "A New Protoceratopsid (Dinosauria: Neoceratopsia) from the Late Cretaceous of Inner Mongolia, China".
1506:
4541:
769:
1646:
4629:
1013:
754:
1175:
in sands are also common across the unit. In addition, structureless, medium-grained, fine-grained and very fine-grained
4452:
2069:"Upper Cretaceous Djadokhta, Barun Goyot and Nemegt formations of Mongolia, including remarks on previous subdivisions"
159:
2925:"A new dromaeosaurid (Dinosauria: Theropoda) from the Upper Cretaceous Wulansuhai Formation of Inner Mongolia, China"
2727:"A new Species of Protoceratops (Dinosauria, Neoceratopsia) from the Late Cretaceous of Inner Mongolia (P. R. China)"
3000:
Xing, X.; Qing-Wei, T.; Shuo, W.; Sullivan, C.; Hone, D. W. E.; Feng-Lu, H.; Qing-Yu, M.; Lin, T.; Dong, T. (2013).
1818:"Yamaceratops dorngobiensis, a New Primitive Ceratopsian (Dinosauria: Ornithischia) from the Cretaceous of Mongolia"
849:
were very similar (mostly in the postcranial skeleton), the former had a much derived (advanced) skull morphology.
480:
2753:"A new species of the ankylosaurid dinosaur Pinacosaurus from the Late Cretaceous of Inner Mongolia (P.R. China)"
1741:
2619:
2356:"Redescription and affinities of Hulsanpes perlei (Dinosauria, Theropoda) from the Upper Cretaceous of Mongolia"
3309:
3211:
2205:"Description and ecologic analysis of Hollanda luceria, a Late Cretaceous bird from the Gobi Desert (Mongolia)"
1718:
961:
the derived-most one. Below are the proposed phylogenetic relationships within Protoceratopsidae by Czepiński:
835:. It had a smaller frill, about ten grinding teeth per jaw, and more triangular skull than its close relative,
687:
of the Hermiin Tsav locality. He also coined the family Bagaceratopidae in order to contain these new taxa and
644:
3001:
2893:
1832:
1395:-deposited) sediments. It is likely that sediments at Bayan Mandahu were deposited by short-lived rivers and
3427:
2203:
Bell, A. K.; Chiappe, L. M.; Erickson, G. M.; Suzuki, S.; Watabe, M.; Barsbold, R.; Tsogtbaatar, K. (2010).
1988:
1360:
953:. Based on cranial characters such as presence or absence of premaxillary teeth and an antorbital fenestra,
704:
409:
4503:
1168:
989:
584:
was described by Bitnara Kim and colleagues, who noted no significant differences between the skeleton of
575:
In 1993, the Japan-Mongolia Joint Paleontological Expedition collected an articulated and nearly complete
564:
137:
1179:
predominate in sediments of the Barun Goyot Formation. The sediments of this formation were deposited in
4609:
4576:
4486:
3336:
3188:
1569:
1132:
468:
405:
4458:
499:
dedicated to describe this material where they named the new genus and type species of protoceratopsid
2961:"A new oviraptorid (Dinosauria: Theropoda) from the Upper Cretaceous of Bayan Mandahu, Inner Mongolia"
1686:"New protoceratopsid specimens improve the age correlation of the Upper Cretaceous Gobi Desert strata"
4550:
3157:
3079:
3036:
3002:"A new oviraptorid from the Upper Cretaceous of Nei Mongol,China, and its stratigraphic implications"
2972:
2834:
2699:
2664:
2651:
Jerzykiewicz, T.; Currie, P. J.; Eberth, D. A.; Johnston, P. A.; Koster, E. H.; Zheng, J.-J. (1993).
2580:
2496:
2441:
2216:
2042:
2015:
1869:
1790:
1614:
1534:
1172:
679:(PIN 4487/26; a partial small-sized skull), recovered from the Khulsan locality, and the holotype of
3592:
3204:
862:
604:
492:
488:
417:
2546:
4312:
4262:
3694:
3144:
Brinkman, D. B.; Tong, H.-Y.; Li, H.; Sun, Y.; Zhang, J.-S.; Godefroit, P.; Zhang, Z.-M. (2015).
3126:
3052:
2905:
2897:
2803:
2633:
2336:
2280:
2185:
1937:
1885:
1666:
1304:
588:
and the former, with the exception of the anatomy of the skull and the shape and location of the
520:
330:
154:
2778:
552:
434:—and kept the small body size that characterized early ceratopsians. Unlike its close relative,
4581:
857:
teeth, had paired (fused) nasal bones, and an oval-shaped fenestra (hole) was developed in the
745:
In 2008 Alifanov described and named another ceratopsian taxon from the Barun Goyot Formation,
692:
4395:
4333:
4296:
3770:
3657:
2862:
2469:
2387:
2328:
2272:
2150:
1991:
1962:
1742:"The fossil record, systematics and evolution of pachycephalosaurs and ceratopsians from Asia"
1192:
968:
941:
and thus becoming synonyms of this genus. Under this reasoning, Protoceratopsidae consists of
281:
1209:
is the most common taxon across the Barun Goyot Formation, which was also home to many other
675:
from the Barun Goyot Formation. Material assigned by Alifanov corresponds to the holotype of
4425:
4223:
4048:
3750:
3445:
3165:
3118:
3087:
3044:
2980:
2939:
2889:
2852:
2842:
2793:
2707:
2672:
2623:
2615:
2588:
2504:
2459:
2449:
2377:
2367:
2318:
2310:
2262:
2254:
2224:
2177:
2140:
2130:
2099:
2068:
2023:
1929:
1877:
1836:
1828:
1798:
1697:
1658:
1622:
1581:
1498:
1475:
1295:
1200:
2298:
Napoli, J. G.; Ruebenstahl, A. A.; Bhullar, B.-A. S.; Turner, A. H.; Norell, M. A. (2021).
4319:
4271:
4230:
4166:
4069:
4020:
3722:
3646:
3601:
3475:
3461:
3420:
3294:
3278:
1271:
1140:
873:
397:
386:
48:
797:
and they fall within the large intraspecific variation of this taxon. He also considered
3161:
3083:
3040:
2976:
2924:
2838:
2777:
Currie, P. J.; Badamgarav, D.; Koppelhus, E. B.; Sissons, R.; Vickaryous, M. K. (2011).
2703:
2668:
2584:
2500:
2445:
2220:
2019:
1873:
1794:
1618:
452:
4326:
4285:
4216:
4203:
4174:
4159:
4129:
4090:
4076:
3993:
3954:
3833:
3743:
3708:
3701:
3581:
3531:
3454:
3434:
2857:
2822:
2464:
2425:
2382:
2355:
2145:
2118:
1802:
1464:
1443:
1339:
1196:
1180:
724:
708:
538:
236:
2760:
Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre
2734:
Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre
1379:
is reddish-brown, poorly cemented, fine grained sandstone with some conglomerate, and
627:
487:). In 1975, two of the expedition's leading scientists, namely Polish paleontologists
4598:
4404:
4383:
4353:
4344:
4244:
4138:
4083:
4062:
4034:
4027:
4013:
4006:
3812:
3801:
3794:
3736:
3729:
3715:
3673:
3608:
3574:
3545:
3498:
3130:
3106:
3056:
2985:
2960:
2909:
2823:"A monodactyl nonavian dinosaur and the complex evolution of the alvarezsauroid hand"
2807:
2652:
2637:
2607:
2592:
2406:
2340:
2284:
2204:
2189:
1984:
Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages
1956:
1941:
1889:
1857:
1670:
1413:
1280:
1275:
1235:
1037:
913:
In 2019 Czepiński analyzed a vast majority of referred specimens to the ceratopsians
903:
837:
527:
436:
88:
1474:. Other paleofauna from this unit comprises a variety of squamates and mammals, and
921:, and concluded that most were in fact specimens of the former. Although the genera
444:
lacked premaxillary teeth (cylindrical, blunt teeth near the tip of the upper jaw).
4418:
4369:
4362:
4278:
4251:
4237:
4145:
4118:
4111:
4104:
4055:
4041:
3933:
3920:
3851:
3840:
3825:
3779:
3761:
3666:
3632:
3615:
3567:
3560:
3516:
3321:
3269:
1470:
1458:
1437:
1419:
1317:
1214:
809:, as it features a combination of basal (primitive) and derived (advanced) traits.
696:
307:
268:
220:
31:
3048:
2547:"Multituberculate succession in the Late Cretaceous of the Gobi Desert (Mongolia)"
2228:
2181:
1933:
2508:
2454:
2027:
1626:
4535:
4411:
4305:
4188:
4152:
3975:
3687:
3552:
2300:"A New Dromaeosaurid (Dinosauria: Coelurosauria) from Khulsan, Central Mongolia"
1662:
1431:
1329:
1313:
1308:
1245:
1239:
817:
653:
472:
431:
63:
3170:
3145:
2827:
Proceedings of the National Academy of Sciences of the United States of America
2135:
4181:
4097:
3947:
3680:
3468:
3402:
3389:
3348:
3227:
2944:
1881:
1586:
1488:
1454:
1449:
1425:
1284:
1260:
1230:
1210:
1188:
895:
854:
720:
608:
252:
108:
73:
4526:
2332:
2276:
1159:(yellowish, grey-brown, and rarely reddish) that are locally cemented. Sandy
2847:
1916:
Czepiński, Ł. (2019). "Ontogeny and variation of a protoceratopsid dinosaur
1376:
1372:
1334:
1323:
1299:
1290:
1254:
1218:
1176:
1164:
1160:
1144:
734:
496:
171:
113:
57:
17:
2866:
2473:
2407:"A taxonomic review of the Pachycephalosauridae (Dinosauria: Ornithischia)"
2391:
2244:"A New Dromaeosaurid from the Late Cretaceous Khulsan Locality of Mongolia"
2154:
1702:
1685:
2798:
883:
4520:
3982:
3968:
3961:
3524:
3260:
3251:
3122:
2430:
from the Baruungoyot and Nemegt Formations (Late Cretaceous) of Mongolia"
1408:
1384:
1348:
1266:
1152:
1136:
877:
684:
589:
504:
389:
204:
191:
103:
98:
83:
78:
68:
2901:
2779:"Hands, feet and behaviour in Pinacosaurus (Dinosauria: Ankylosauridae)"
910:
during the Cretaceous Period, which ended roughly 66 million years ago.
4568:
4376:
3538:
3482:
2372:
1380:
1224:
858:
667:
In 2003 Russian paleontologist Vladimir R. Alifanov named the new taxa
118:
93:
2103:
764:
A comprehensive study on the intraspecific variation in morphology of
3245:
2628:
2323:
2267:
1841:
1478:
1388:
1344:
899:
484:
456:
181:
4497:
3091:
2711:
2676:
1721:[A new Late Cretaceous protoceratopsid genus from Mongolia]
1602:
603:
from the Udyn Sayr and Zamyn Khond localities, respectively, of the
145:
Skeletal reconstruction (including elements previously described as
2725:
Lambert, O.; Godefroit, P.; Li, H.; Shang, C.-Y.; Dong, Z. (2001).
2314:
2258:
2751:
Godefroit, P.; Pereda-Suberbiola, X.; Li, H.; Dong, Z. M. (1999).
2424:
Fanti, F.; Currie, P. J.; Badamgarav, D.; Lalueza-Fox, C. (2012).
1184:
882:
872:
816:
801:
to be a distinct and separate genus of protoceratopsid, from both
733:
712:
683:(PIN 3142/4; an almost complete medium-sized skull), found in the
626:
571:
specimen MPC-D 100/551B, which belongs to an indeterminate species
563:
551:
451:
382:
699:
described and named the new genus and species of protoceratopsid
475:. This newly collected and rich fossil material is stored in the
430:
had a fairly primitive anatomy—when compared to the much derived
4555:
1400:
1396:
1392:
1250:
1156:
907:
393:
4501:
3918:
3387:
3238:
3200:
1601:
Saneyoshi, M.; Watabe, M.; Suzuki, S.; Tsogtbaatar, K. (2011).
1151:
is known. This formation is mostly characterized by series of
719:, IVPP V12513, represents a nearly complete skull lacking the
3196:
515:, including juvenile and sub-adult skulls. The generic name,
2620:
10.1206/0003-0090(2000)249<0001:TCASOL>2.0.CO;2
1147:) and has virtually yielded the bulk of material for which
824:
specimens compared to a 1.8 m (5.9 ft) tall human
768:
was conducted by Polish paleontologist Łukasz Czepiński in
656:
in 2000 explained this by extrapolating that the juvenile
459:
localities of Mongolia and the location of Bayan Mandahu;
2894:
10.1671/0272-4634(2008)28[432:ANSOVD]2.0.CO;2
2411:
New Mexico Museum of Natural History and Science Bulletin
2067:
Gradziński, R.; Jaworowska, Z. K.; Maryańska, T. (1977).
1833:
10.1206/0003-0082(2006)3530[1:YDANPC]2.0.CO;2
1183:(flat land consisting of sediments deposited by highland
753:, it had already been displayed for several years at the
1371:), is considered to be Late Cretaceous in age, roughly
27:
Protoceratopsid dinosaur genus from the Late Cretaceous
2524:"A new sauropod from the Upper Cretaceous of Mongolia"
1719:"Новый род протоцератопсид из позднего мела Монголии"
2242:
Turner, A. H.; Montanari, S.; Norell, M. A. (2021).
788:
represent additional specimens and growth stages of
703:from red beds at the Bayan Mandahu locality of the
648:
by Kurzanov in 1990, were considered to be juvenile
519:, means "small-horned face" and is derived from the
314:
4510:
4394:
4352:
4343:
4295:
4261:
4202:
4128:
3992:
3931:
3850:
3811:
3769:
3760:
3656:
3630:
3591:
3496:
3444:
3400:
3347:
3320:
3293:
412:. One specimen may argue the possible presence of
2959:Longrich, N. R.; Currie, P. J.; Dong, Z. (2010).
2489:Palaeogeography, Palaeoclimatology, Palaeoecology
2008:Palaeogeography, Palaeoclimatology, Palaeoecology
1607:Palaeogeography, Palaeoclimatology, Palaeoecology
738:Skeletal diagram featuring the known elements of
1955:Dodson, P. (1993). "Dinosaurs Rule The World".
1751:. Cambridge University Press. pp. 488–489.
772:, where he concluded that the previously named
2001:
1999:
1920:from the Late Cretaceous of the Gobi Desert".
1574:The Journal of the Geological Society of Japan
595:In 2020, Czepiński described new specimens of
3212:
3105:Wible, J. R.; Shelley, S. L.; Bi, S. (2019).
638:Juvenile remains, at first tentatively named
8:
2043:"Sedimentation of the Barun Goyot Formation"
1961:. Publications International. p. 132.
1749:The Age of Dinosaurs in Russia and Mongolia
1399:on an alluvial plain with a combination of
4498:
4349:
4208:
3998:
3939:
3928:
3915:
3817:
3766:
3638:
3508:
3504:
3412:
3408:
3397:
3384:
3326:
3299:
3290:
3235:
3219:
3205:
3197:
1911:
1909:
1907:
1905:
1903:
1901:
1899:
1651:Journal of the Geological Society of Korea
1528:
1526:
1524:
1522:
1411:, such as closely related protoceratopsid
136:
38:
3169:
2984:
2943:
2856:
2846:
2797:
2627:
2522:Kurzanov, S. M.; Bannikov, A. F. (1983).
2463:
2453:
2381:
2371:
2322:
2266:
2144:
2134:
2092:Zoological Journal of the Linnean Society
2041:Gradziński, R.; Jerzykiewicz, T. (1974).
1840:
1701:
1640:
1638:
1636:
1585:
1816:Makovicky, P. J.; Norell, M. A. (2006).
1535:"Protoceratopsidae (Dinosauria) of Asia"
898:, a group of herbivorous dinosaurs with
533:= meaning horn face. The specific name,
1783:Acta Geologica Sinica (English Edition)
1518:
957:is the basal-most protoceratopsid and
542:Anatoly Konstantinovich Rozhdestvensky
1645:Kim, B.; Yun, H.; Lee, Y.-N. (2019).
1568:Matsumoto, Y.; Saneyoshi, M. (2010).
1407:This formation has produced numerous
544:for his notorious work on dinosaurs.
400:, around 72 to 71 million years ago.
7:
1533:Maryańska, T.; Osmólska, H. (1975).
1333:. Other taxa are represented by the
1195:paleoenvironments, under relatively
404:remains have been reported from the
2170:Journal of Systematic Palaeontology
1053:
1029:
1005:
981:
974:
964:
537:, is named in honor of the Russian
381:(meaning "small-horned face") is a
3072:Canadian Journal of Earth Sciences
2882:Journal of Vertebrate Paleontology
2692:Canadian Journal of Earth Sciences
2657:Canadian Journal of Earth Sciences
1803:10.1111/j.1755-6724.2003.tb00745.x
25:
4605:Late Cretaceous dinosaurs of Asia
2606:Keqin, G.; Norell, M. A. (2000).
1982:Holtz, T. R.; Rey, L. V. (2007).
887:Skull cast of specimen PIN 3142/1
4482:
4481:
4457:
4451:
3901:
3895:
2986:10.1111/j.1475-4983.2010.00968.x
1507:Timeline of ceratopsian research
1491:
158:
580:skeleton (specimen KID 196) of
4625:Taxa named by Teresa Maryańska
4620:Taxa named by Halszka Osmólska
2545:Kielan-Jaworowska, Z. (1974).
1989:Supplementary Information 2012
861:—otherwise known as accessory
324:Maryanska & Osmolska, 1975
301:Maryanska & Osmolska, 1975
1:
4615:Fossil taxa described in 1975
3049:10.1016/j.cretres.2021.105052
2786:Acta Palaeontologica Polonica
2229:10.1016/j.cretres.2009.09.001
2182:10.1080/14772019.2021.2013965
1934:10.1080/08912963.2019.1593404
1690:Acta Palaeontologica Polonica
1014:Protoceratops hellenikorhinus
902:-like beaks which thrived in
755:Moscow Paleontological Museum
4542:Bagaceratops rozhdestvenskyi
4512:Bagaceratops rozhdestvenskyi
2593:10.1016/0037-0738(74)90021-9
2509:10.1016/j.palaeo.2017.10.023
2455:10.1371/journal.pone.0031330
2354:Cau, A.; Madzia, D. (2018).
2028:10.1016/j.palaeo.2017.11.018
1918:Bagaceratops rozhdestvenskyi
1627:10.1016/j.palaeo.2011.07.024
759:Bagaceratops rozhdestvenskyi
617:Bagaceratops rozhdestvenskyi
501:Bagaceratops rozhdestvenskyi
319:Bagaceratops rozhdestvenskyi
1663:10.14770/jgsk.2019.55.2.179
4646:
3171:10.1016/j.crpv.2014.12.005
2136:10.1038/s41598-021-83568-4
1570:"Bored dinosaur skeletons"
1343:, and a wide diversity of
481:Polish Academy of Sciences
29:
4471:
4449:
4211:
4001:
3942:
3927:
3914:
3892:
3820:
3641:
3511:
3507:
3415:
3411:
3396:
3383:
3329:
3302:
3289:
3234:
3111:Annals of Carnegie Museum
2945:10.11646/zootaxa.2403.1.1
2612:American Museum Novitates
2307:American Museum Novitates
2251:American Museum Novitates
1882:10.1134/S0031030108060087
1825:American Museum Novitates
1587:10.5575/geosoc.116.1.I_II
1391:-deposited) and aeolian (
1171:, and large-scale trough
1058:
1051:
1034:
1027:
1010:
1003:
986:
979:
972:
778:Lamaceratops tereschenkoi
669:Lamaceratops tereschenkoi
660:would grow into a mature
477:Institute of Paleobiology
348:Lamaceratops tereschenkoi
336:
329:
313:
306:
155:Scientific classification
153:
144:
135:
41:
2405:Sullivan, R. M. (2006).
1856:Alifanov, V. R. (2008).
1729:(in Russian) (4): 91−97.
1717:Kurzanov, S. M. (1990).
1155:, mostly light-coloured
782:Platyceratops tatarinovi
673:Platyceratops tatarinovi
645:Breviceratops kozlowskii
640:Protoceratops kozlowskii
364:Platyceratops tatarinovi
30:Not to be confused with
3428:Micropachycephalosaurus
2848:10.1073/pnas.1011052108
2554:Palaeontologia Polonica
2528:Paleontological Journal
2073:Acta Geologica Polonica
2050:Palaeontologia Polonica
1862:Paleontological Journal
1764:Paleontological Journal
1727:Paleontological Journal
1542:Palaeontologia Polonica
1383:. Other facies include
1361:Bayan Mandahu Formation
1355:Bayan Mandahu Formation
705:Bayan Mandahu Formation
410:Bayan Mandahu Formation
3150:Comptes Rendus Palevol
3009:Vertebrata PalAsiatica
1740:Sereno, P. C. (2000).
1703:10.4202/app.00701.2019
1684:Czepiński, Ł. (2020).
1163:(often red-coloured),
990:Protoceratops andrewsi
888:
880:
825:
742:
635:
613:Protoceratops andrewsi
572:
561:
464:
4577:Paleobiology Database
2799:10.4202/app.2010.0055
1143:in age (Middle-Upper
1133:Barun Goyot Formation
1127:Barun Goyot Formation
886:
876:
820:
737:
630:
567:
555:
526:= meaning small; and
469:Barun Goyot Formation
455:
406:Barun Goyot Formation
4630:Ornithischian genera
3123:10.2992/007.085.0401
1958:The Age of Dinosaurs
1173:cross-stratification
786:Magnirostris dodsoni
774:Gobiceratops minutus
747:Gobiceratops minitus
701:Magnirostris dodsoni
691:. Also during 2003,
548:Additional specimens
495:, published a large
448:History of discovery
356:Magnirostris dodsoni
340:Gobiceratops minutus
3593:Archaeoceratopsidae
3162:2015CRPal..14..577B
3084:1996CaJES..33..578G
3041:2022CrRes.13005052P
3029:Cretaceous Research
2977:2010Palgy..53..945L
2839:2011PNAS..108.2338X
2704:1993CaJES..30.2196E
2669:1993CaJES..30.2180J
2585:1974SedG...12..249G
2573:Sedimentary Geology
2501:2018PPP...494..101F
2446:2012PLoSO...731330F
2221:2010CrRes..31...16B
2209:Cretaceous Research
2020:2018PPP...494...29E
1874:2008PalJ...42..621A
1795:2003AcGlS..77..299Y
1619:2011PPP...311...38S
863:antorbital fenestra
715:). The holotype of
642:, and then renamed
605:Djadochta Formation
418:Djadochta Formation
359:You & Dong 2003
4313:Crittendenceratops
4263:Albertaceratopsini
3035:(105052): 105052.
2426:"New specimens of
2373:10.7717/peerj.4868
2123:Scientific Reports
1992:Weight Information
1922:Historical Biology
1305:pachycephalosaurid
889:
881:
826:
790:B. rozhdestvenskyi
766:B. rozhdestvenskyi
743:
636:
631:Holotype skull of
573:
562:
535:B. rozhdestvenskyi
465:
4592:
4591:
4504:Taxon identifiers
4495:
4494:
4467:
4466:
4447:
4446:
4443:
4442:
4439:
4438:
4435:
4434:
4396:Pachyrhinosaurini
4334:Yehuecauhceratops
4297:Nasutoceratopsini
4198:
4197:
3910:
3909:
3890:
3889:
3886:
3885:
3882:
3881:
3878:
3877:
3874:
3873:
3870:
3869:
3771:Protoceratopsidae
3658:Leptoceratopsidae
3626:
3625:
3492:
3491:
3379:
3378:
3375:
3374:
3371:
3370:
3367:
3366:
2766:(supp. B): 17–36.
2698:(10): 2196−2213.
2663:(10): 2180−2195.
2176:(16): 1097–1128.
2104:10.1111/zoj.12185
1968:978-0-7853-0443-2
1928:(10): 1394–1421.
1476:nanhsiungchelyids
1429:; dromaeosaurids
1201:semiarid climates
1139:, is regarded as
1118:
1117:
1109:
1108:
1100:
1099:
1091:
1090:
1082:
1081:
1073:
1072:
969:Protoceratopsidae
853:lacked primitive
374:
373:
368:
360:
352:
344:
302:
282:Protoceratopsidae
16:(Redirected from
4637:
4585:
4584:
4572:
4571:
4559:
4558:
4546:
4545:
4544:
4531:
4530:
4529:
4499:
4485:
4484:
4462:
4461:
4455:
4426:Pachyrhinosaurus
4350:
4224:Machairoceratops
4209:
4049:Coahuilaceratops
3999:
3940:
3929:
3916:
3905:
3900:
3899:
3818:
3767:
3751:Zhuchengceratops
3639:
3509:
3505:
3446:Chaoyangsauridae
3413:
3409:
3398:
3385:
3327:
3300:
3291:
3284:
3283:
3236:
3221:
3214:
3207:
3198:
3176:
3175:
3173:
3156:(6–7): 577−587.
3141:
3135:
3134:
3102:
3096:
3095:
3067:
3061:
3060:
3023:
3017:
3016:
3006:
2997:
2991:
2990:
2988:
2956:
2950:
2949:
2947:
2929:
2920:
2914:
2913:
2877:
2871:
2870:
2860:
2850:
2833:(6): 2338–2342.
2818:
2812:
2811:
2801:
2783:
2774:
2768:
2767:
2757:
2748:
2742:
2741:
2731:
2722:
2716:
2715:
2687:
2681:
2680:
2648:
2642:
2641:
2631:
2603:
2597:
2596:
2568:
2562:
2561:
2551:
2542:
2536:
2535:
2519:
2513:
2512:
2484:
2478:
2477:
2467:
2457:
2421:
2415:
2414:
2402:
2396:
2395:
2385:
2375:
2351:
2345:
2344:
2326:
2304:
2295:
2289:
2288:
2270:
2248:
2239:
2233:
2232:
2200:
2194:
2193:
2165:
2159:
2158:
2148:
2138:
2114:
2108:
2107:
2087:
2081:
2080:
2064:
2058:
2057:
2047:
2038:
2032:
2031:
2003:
1994:
1987:
1979:
1973:
1972:
1952:
1946:
1945:
1913:
1894:
1893:
1853:
1847:
1846:
1844:
1822:
1813:
1807:
1806:
1778:
1772:
1771:
1759:
1753:
1752:
1746:
1737:
1731:
1730:
1724:
1714:
1708:
1707:
1705:
1681:
1675:
1674:
1642:
1631:
1630:
1598:
1592:
1591:
1589:
1565:
1559:
1556:
1550:
1549:
1539:
1530:
1501:
1499:Dinosaurs portal
1496:
1495:
1494:
1423:; alvarezsaurid
1367:(now synonym of
1363:, which yielded
1296:halszkaraptorine
1213:, including the
1122:Paleoenvironment
1054:
1030:
1006:
982:
975:
965:
878:Life restoration
841:. Although both
611:transition from
493:Halszka Osmólska
489:Teresa Maryańska
366:
358:
350:
342:
316:
300:
293:
280:
267:
251:
235:
219:
163:
162:
140:
130:
60:
47:Temporal range:
39:
21:
4645:
4644:
4640:
4639:
4638:
4636:
4635:
4634:
4595:
4594:
4593:
4588:
4580:
4575:
4567:
4562:
4554:
4549:
4540:
4539:
4534:
4525:
4524:
4519:
4506:
4496:
4491:
4463:
4456:
4431:
4390:
4339:
4320:Furcatoceratops
4291:
4272:Albertaceratops
4257:
4231:Menefeeceratops
4194:
4167:Tatankaceratops
4124:
4070:Mercuriceratops
4021:Arrhinoceratops
3988:
3936:
3923:
3906:
3894:
3866:
3846:
3807:
3756:
3723:Montanoceratops
3652:
3647:Graciliceratops
3635:
3622:
3602:Archaeoceratops
3587:
3501:
3488:
3476:Xuanhuaceratops
3462:Hualianceratops
3440:
3421:Albalophosaurus
3405:
3392:
3363:
3343:
3316:
3310:Avemetatarsalia
3295:Avemetatarsalia
3285:
3279:Marginocephalia
3241:
3240:
3230:
3225:
3185:
3180:
3179:
3143:
3142:
3138:
3104:
3103:
3099:
3092:10.1139/e96-043
3069:
3068:
3064:
3025:
3024:
3020:
3004:
2999:
2998:
2994:
2958:
2957:
2953:
2927:
2922:
2921:
2917:
2879:
2878:
2874:
2820:
2819:
2815:
2781:
2776:
2775:
2771:
2755:
2750:
2749:
2745:
2729:
2724:
2723:
2719:
2712:10.1139/e93-191
2689:
2688:
2684:
2677:10.1139/e93-190
2650:
2649:
2645:
2605:
2604:
2600:
2570:
2569:
2565:
2549:
2544:
2543:
2539:
2521:
2520:
2516:
2486:
2485:
2481:
2423:
2422:
2418:
2404:
2403:
2399:
2353:
2352:
2348:
2302:
2297:
2296:
2292:
2246:
2241:
2240:
2236:
2202:
2201:
2197:
2167:
2166:
2162:
2116:
2115:
2111:
2089:
2088:
2084:
2066:
2065:
2061:
2045:
2040:
2039:
2035:
2005:
2004:
1997:
1986:. Random House.
1981:
1980:
1976:
1969:
1954:
1953:
1949:
1915:
1914:
1897:
1855:
1854:
1850:
1820:
1815:
1814:
1810:
1780:
1779:
1775:
1761:
1760:
1756:
1744:
1739:
1738:
1734:
1722:
1716:
1715:
1711:
1683:
1682:
1678:
1644:
1643:
1634:
1600:
1599:
1595:
1567:
1566:
1562:
1557:
1553:
1537:
1532:
1531:
1520:
1515:
1497:
1492:
1490:
1487:
1441:; oviraptorids
1417:; ankylosaurid
1375:. The dominant
1357:
1272:protoceratopsid
1141:Late Cretaceous
1129:
1124:
1119:
1110:
1101:
1092:
1083:
1074:
871:
815:
757:under the name
725:junior synonyms
625:
550:
503:. The selected
450:
398:Late Cretaceous
387:protoceratopsid
325:
322:
299:
291:
278:
265:
249:
233:
217:
157:
131:
129:
128:
127:
126:
121:
116:
111:
106:
101:
96:
91:
86:
81:
76:
71:
66:
55:
54:
52:
49:Late Cretaceous
45:
35:
28:
23:
22:
15:
12:
11:
5:
4643:
4641:
4633:
4632:
4627:
4622:
4617:
4612:
4607:
4597:
4596:
4590:
4589:
4587:
4586:
4573:
4560:
4547:
4532:
4516:
4514:
4508:
4507:
4502:
4493:
4492:
4490:
4489:
4479:
4472:
4469:
4468:
4465:
4464:
4450:
4448:
4445:
4444:
4441:
4440:
4437:
4436:
4433:
4432:
4430:
4429:
4422:
4415:
4408:
4400:
4398:
4392:
4391:
4389:
4388:
4380:
4373:
4366:
4358:
4356:
4347:
4341:
4340:
4338:
4337:
4330:
4327:Nasutoceratops
4323:
4316:
4309:
4301:
4299:
4293:
4292:
4290:
4289:
4286:Medusaceratops
4282:
4275:
4267:
4265:
4259:
4258:
4256:
4255:
4248:
4241:
4234:
4227:
4220:
4217:Diabloceratops
4212:
4206:
4204:Centrosaurinae
4200:
4199:
4196:
4195:
4193:
4192:
4185:
4178:
4175:Titanoceratops
4171:
4163:
4160:Regaliceratops
4156:
4149:
4142:
4134:
4132:
4130:Triceratopsini
4126:
4125:
4123:
4122:
4115:
4108:
4101:
4094:
4091:Sierraceratops
4087:
4080:
4077:Navajoceratops
4073:
4066:
4059:
4052:
4045:
4038:
4031:
4024:
4017:
4010:
4002:
3996:
3994:Chasmosaurinae
3990:
3989:
3987:
3986:
3979:
3972:
3965:
3958:
3955:Brachyceratops
3951:
3943:
3937:
3932:
3925:
3924:
3919:
3912:
3911:
3908:
3907:
3893:
3891:
3888:
3887:
3884:
3883:
3880:
3879:
3876:
3875:
3872:
3871:
3868:
3867:
3865:
3864:
3863:
3862:
3856:
3854:
3848:
3847:
3845:
3844:
3837:
3834:Turanoceratops
3830:
3821:
3815:
3809:
3808:
3806:
3805:
3798:
3791:
3784:
3775:
3773:
3764:
3758:
3757:
3755:
3754:
3747:
3744:Unescoceratops
3740:
3733:
3726:
3719:
3712:
3709:Ischioceratops
3705:
3702:Gryphoceratops
3698:
3691:
3684:
3677:
3670:
3662:
3660:
3654:
3653:
3651:
3650:
3642:
3636:
3631:
3628:
3627:
3624:
3623:
3621:
3620:
3612:
3605:
3597:
3595:
3589:
3588:
3586:
3585:
3582:Sasayamagnomus
3578:
3571:
3564:
3557:
3549:
3542:
3535:
3532:Auroraceratops
3528:
3521:
3512:
3502:
3497:
3494:
3493:
3490:
3489:
3487:
3486:
3479:
3472:
3465:
3458:
3455:Chaoyangsaurus
3450:
3448:
3442:
3441:
3439:
3438:
3435:Psittacosaurus
3431:
3424:
3416:
3406:
3401:
3394:
3393:
3388:
3381:
3380:
3377:
3376:
3373:
3372:
3369:
3368:
3365:
3364:
3362:
3361:
3360:
3359:
3353:
3351:
3345:
3344:
3342:
3341:
3340:
3339:
3330:
3324:
3318:
3317:
3315:
3314:
3313:
3312:
3303:
3297:
3287:
3286:
3282:
3281:
3272:
3263:
3254:
3248:
3239:
3232:
3231:
3226:
3224:
3223:
3216:
3209:
3201:
3195:
3194:
3184:
3183:External links
3181:
3178:
3177:
3136:
3117:(4): 285–327.
3097:
3078:(4): 578−598.
3062:
3018:
2992:
2971:(5): 945−960.
2951:
2915:
2888:(2): 432–438.
2872:
2813:
2792:(3): 489–504.
2769:
2743:
2717:
2682:
2643:
2614:(249): 1−118.
2598:
2579:(4): 249–278.
2563:
2537:
2514:
2479:
2416:
2413:(35): 347–365.
2397:
2346:
2315:10.1206/3982.1
2309:(3982): 1–47.
2290:
2259:10.1206/3965.1
2253:(3965): 1–48.
2234:
2195:
2160:
2129:(4101): 4101.
2109:
2098:(3): 631−652.
2082:
2059:
2033:
1995:
1974:
1967:
1947:
1895:
1868:(6): 621–633.
1848:
1827:(3530): 1–42.
1808:
1789:(3): 299−303.
1773:
1754:
1732:
1709:
1696:(3): 481−497.
1676:
1657:(2): 179−190.
1632:
1613:(1–2): 38−47.
1593:
1560:
1551:
1517:
1516:
1514:
1511:
1510:
1509:
1503:
1502:
1486:
1483:
1465:Papiliovenator
1444:Machairasaurus
1356:
1353:
1340:Quaesitosaurus
1281:dromaeosaurids
1236:alvarezsaurids
1181:alluvial plain
1128:
1125:
1123:
1120:
1116:
1115:
1112:
1111:
1107:
1106:
1103:
1102:
1098:
1097:
1094:
1093:
1089:
1088:
1085:
1084:
1080:
1079:
1076:
1075:
1071:
1070:
1067:
1066:
1057:
1052:
1050:
1047:
1046:
1043:
1042:
1033:
1028:
1026:
1023:
1022:
1019:
1018:
1009:
1004:
1002:
999:
998:
995:
994:
985:
980:
978:
973:
971:
963:
870:
869:Classification
867:
814:
811:
709:Inner Mongolia
624:
621:
549:
546:
539:paleontologist
509:ZPAL MgD-I/126
449:
446:
392:that lived in
372:
371:
370:
369:
367:Alifanov, 2003
361:
353:
351:Alifanov, 2003
345:
343:Alifanov, 2008
334:
333:
327:
326:
323:
311:
310:
304:
303:
289:
285:
284:
276:
272:
271:
263:
256:
255:
247:
240:
239:
237:Neornithischia
231:
224:
223:
215:
208:
207:
202:
195:
194:
189:
185:
184:
179:
175:
174:
169:
165:
164:
151:
150:
142:
141:
133:
132:
124:
123:
122:
117:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
61:
46:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
4642:
4631:
4628:
4626:
4623:
4621:
4618:
4616:
4613:
4611:
4608:
4606:
4603:
4602:
4600:
4583:
4578:
4574:
4570:
4565:
4561:
4557:
4552:
4548:
4543:
4537:
4533:
4528:
4522:
4518:
4517:
4515:
4513:
4509:
4505:
4500:
4488:
4480:
4478:
4474:
4473:
4470:
4460:
4454:
4428:
4427:
4423:
4421:
4420:
4416:
4414:
4413:
4409:
4407:
4406:
4405:Achelousaurus
4402:
4401:
4399:
4397:
4393:
4386:
4385:
4384:Styracosaurus
4381:
4379:
4378:
4374:
4372:
4371:
4367:
4365:
4364:
4360:
4359:
4357:
4355:
4354:Centrosaurini
4351:
4348:
4346:
4345:Eucentrosaura
4342:
4336:
4335:
4331:
4329:
4328:
4324:
4322:
4321:
4317:
4315:
4314:
4310:
4308:
4307:
4303:
4302:
4300:
4298:
4294:
4288:
4287:
4283:
4281:
4280:
4276:
4274:
4273:
4269:
4268:
4266:
4264:
4260:
4254:
4253:
4249:
4247:
4246:
4245:Wendiceratops
4242:
4240:
4239:
4235:
4233:
4232:
4228:
4226:
4225:
4221:
4219:
4218:
4214:
4213:
4210:
4207:
4205:
4201:
4191:
4190:
4186:
4184:
4183:
4179:
4177:
4176:
4172:
4169:
4168:
4164:
4162:
4161:
4157:
4155:
4154:
4150:
4148:
4147:
4143:
4141:
4140:
4139:Eotriceratops
4136:
4135:
4133:
4131:
4127:
4121:
4120:
4116:
4114:
4113:
4109:
4107:
4106:
4102:
4100:
4099:
4095:
4093:
4092:
4088:
4086:
4085:
4084:Pentaceratops
4081:
4079:
4078:
4074:
4072:
4071:
4067:
4065:
4064:
4063:Kosmoceratops
4060:
4058:
4057:
4053:
4051:
4050:
4046:
4044:
4043:
4039:
4037:
4036:
4035:Bravoceratops
4032:
4030:
4029:
4028:Bisticeratops
4025:
4023:
4022:
4018:
4016:
4015:
4014:Anchiceratops
4011:
4009:
4008:
4007:Agujaceratops
4004:
4003:
4000:
3997:
3995:
3991:
3985:
3984:
3980:
3978:
3977:
3973:
3971:
3970:
3966:
3964:
3963:
3959:
3957:
3956:
3952:
3950:
3949:
3945:
3944:
3941:
3938:
3935:
3930:
3926:
3922:
3917:
3913:
3904:
3898:
3860:
3859:
3858:
3857:
3855:
3853:
3849:
3843:
3842:
3838:
3836:
3835:
3831:
3828:
3827:
3823:
3822:
3819:
3816:
3814:
3813:Ceratopsoidea
3810:
3804:
3803:
3802:Protoceratops
3799:
3797:
3796:
3795:Breviceratops
3792:
3790:
3789:
3785:
3782:
3781:
3777:
3776:
3774:
3772:
3768:
3765:
3763:
3759:
3753:
3752:
3748:
3746:
3745:
3741:
3739:
3738:
3737:Udanoceratops
3734:
3732:
3731:
3730:Prenoceratops
3727:
3725:
3724:
3720:
3718:
3717:
3716:Leptoceratops
3713:
3711:
3710:
3706:
3704:
3703:
3699:
3697:
3696:
3692:
3690:
3689:
3685:
3683:
3682:
3678:
3676:
3675:
3674:Bainoceratops
3671:
3669:
3668:
3664:
3663:
3661:
3659:
3655:
3649:
3648:
3644:
3643:
3640:
3637:
3634:
3629:
3618:
3617:
3613:
3611:
3610:
3609:Helioceratops
3606:
3604:
3603:
3599:
3598:
3596:
3594:
3590:
3584:
3583:
3579:
3577:
3576:
3575:Mosaiceratops
3572:
3570:
3569:
3565:
3563:
3562:
3558:
3555:
3554:
3550:
3548:
3547:
3546:Koreaceratops
3543:
3541:
3540:
3536:
3534:
3533:
3529:
3527:
3526:
3522:
3519:
3518:
3514:
3513:
3510:
3506:
3503:
3500:
3499:Neoceratopsia
3495:
3485:
3484:
3480:
3478:
3477:
3473:
3471:
3470:
3466:
3464:
3463:
3459:
3457:
3456:
3452:
3451:
3449:
3447:
3443:
3437:
3436:
3432:
3430:
3429:
3425:
3423:
3422:
3418:
3417:
3414:
3410:
3407:
3404:
3399:
3395:
3391:
3386:
3382:
3357:
3356:
3355:
3354:
3352:
3350:
3346:
3338:
3334:
3333:
3332:
3331:
3328:
3325:
3323:
3319:
3311:
3307:
3306:
3305:
3304:
3301:
3298:
3296:
3292:
3288:
3280:
3276:
3273:
3271:
3267:
3264:
3262:
3258:
3255:
3253:
3249:
3247:
3243:
3242:
3237:
3233:
3229:
3222:
3217:
3215:
3210:
3208:
3203:
3202:
3199:
3193:
3192:
3187:
3186:
3182:
3172:
3167:
3163:
3159:
3155:
3151:
3147:
3140:
3137:
3132:
3128:
3124:
3120:
3116:
3112:
3108:
3101:
3098:
3093:
3089:
3085:
3081:
3077:
3073:
3066:
3063:
3058:
3054:
3050:
3046:
3042:
3038:
3034:
3030:
3022:
3019:
3014:
3010:
3003:
2996:
2993:
2987:
2982:
2978:
2974:
2970:
2966:
2965:Palaeontology
2962:
2955:
2952:
2946:
2941:
2937:
2933:
2926:
2919:
2916:
2911:
2907:
2903:
2899:
2895:
2891:
2887:
2883:
2876:
2873:
2868:
2864:
2859:
2854:
2849:
2844:
2840:
2836:
2832:
2828:
2824:
2817:
2814:
2809:
2805:
2800:
2795:
2791:
2787:
2780:
2773:
2770:
2765:
2761:
2754:
2747:
2744:
2739:
2735:
2728:
2721:
2718:
2713:
2709:
2705:
2701:
2697:
2693:
2686:
2683:
2678:
2674:
2670:
2666:
2662:
2658:
2654:
2647:
2644:
2639:
2635:
2630:
2625:
2621:
2617:
2613:
2609:
2602:
2599:
2594:
2590:
2586:
2582:
2578:
2574:
2567:
2564:
2559:
2555:
2548:
2541:
2538:
2533:
2529:
2525:
2518:
2515:
2510:
2506:
2502:
2498:
2494:
2490:
2483:
2480:
2475:
2471:
2466:
2461:
2456:
2451:
2447:
2443:
2440:(2): e31330.
2439:
2435:
2431:
2429:
2420:
2417:
2412:
2408:
2401:
2398:
2393:
2389:
2384:
2379:
2374:
2369:
2365:
2361:
2357:
2350:
2347:
2342:
2338:
2334:
2330:
2325:
2320:
2316:
2312:
2308:
2301:
2294:
2291:
2286:
2282:
2278:
2274:
2269:
2264:
2260:
2256:
2252:
2245:
2238:
2235:
2230:
2226:
2222:
2218:
2214:
2210:
2206:
2199:
2196:
2191:
2187:
2183:
2179:
2175:
2171:
2164:
2161:
2156:
2152:
2147:
2142:
2137:
2132:
2128:
2124:
2120:
2113:
2110:
2105:
2101:
2097:
2093:
2086:
2083:
2079:(3): 281–326.
2078:
2074:
2070:
2063:
2060:
2055:
2051:
2044:
2037:
2034:
2029:
2025:
2021:
2017:
2013:
2009:
2002:
2000:
1996:
1993:
1990:
1985:
1978:
1975:
1970:
1964:
1960:
1959:
1951:
1948:
1943:
1939:
1935:
1931:
1927:
1923:
1919:
1912:
1910:
1908:
1906:
1904:
1902:
1900:
1896:
1891:
1887:
1883:
1879:
1875:
1871:
1867:
1863:
1859:
1852:
1849:
1843:
1838:
1834:
1830:
1826:
1819:
1812:
1809:
1804:
1800:
1796:
1792:
1788:
1784:
1777:
1774:
1770:(5): 524–534.
1769:
1765:
1758:
1755:
1750:
1743:
1736:
1733:
1728:
1720:
1713:
1710:
1704:
1699:
1695:
1691:
1687:
1680:
1677:
1672:
1668:
1664:
1660:
1656:
1652:
1648:
1641:
1639:
1637:
1633:
1628:
1624:
1620:
1616:
1612:
1608:
1604:
1597:
1594:
1588:
1583:
1579:
1575:
1571:
1564:
1561:
1555:
1552:
1547:
1543:
1536:
1529:
1527:
1525:
1523:
1519:
1512:
1508:
1505:
1504:
1500:
1489:
1484:
1482:
1480:
1477:
1473:
1472:
1467:
1466:
1461:
1460:
1456:
1452:
1451:
1446:
1445:
1440:
1439:
1434:
1433:
1428:
1427:
1422:
1421:
1416:
1415:
1414:Protoceratops
1410:
1405:
1402:
1398:
1394:
1390:
1386:
1382:
1378:
1374:
1370:
1366:
1362:
1354:
1352:
1350:
1346:
1342:
1341:
1336:
1332:
1331:
1326:
1325:
1320:
1319:
1315:
1311:
1310:
1306:
1302:
1301:
1297:
1293:
1292:
1287:
1286:
1282:
1278:
1277:
1276:Breviceratops
1273:
1269:
1268:
1263:
1262:
1257:
1256:
1252:
1248:
1247:
1242:
1241:
1237:
1233:
1232:
1227:
1226:
1221:
1220:
1216:
1215:ankylosaurids
1212:
1208:
1204:
1202:
1198:
1194:
1190:
1186:
1182:
1178:
1174:
1170:
1169:conglomerates
1166:
1162:
1158:
1154:
1150:
1146:
1142:
1138:
1134:
1126:
1121:
1114:
1113:
1105:
1104:
1096:
1095:
1087:
1086:
1078:
1077:
1069:
1068:
1065:
1064:
1063:
1056:
1055:
1049:
1048:
1045:
1044:
1041:
1040:
1039:
1038:Breviceratops
1032:
1031:
1025:
1024:
1021:
1020:
1017:
1016:
1015:
1008:
1007:
1001:
1000:
997:
996:
993:
992:
991:
984:
983:
977:
976:
970:
967:
966:
962:
960:
956:
952:
951:Protoceratops
948:
947:Breviceratops
944:
940:
936:
935:Platyceratops
932:
928:
924:
920:
919:Breviceratops
916:
911:
909:
905:
904:North America
901:
897:
893:
885:
879:
875:
868:
866:
864:
860:
856:
852:
848:
847:Protoceratops
844:
840:
839:
838:Protoceratops
834:
830:
823:
819:
812:
810:
808:
807:Protoceratops
804:
800:
799:Breviceratops
796:
791:
787:
783:
779:
775:
771:
767:
762:
760:
756:
752:
748:
741:
736:
732:
730:
726:
722:
718:
714:
710:
706:
702:
698:
694:
690:
686:
682:
681:Platyceratops
678:
674:
670:
665:
663:
659:
658:Breviceratops
655:
651:
647:
646:
641:
634:
629:
622:
620:
618:
614:
610:
606:
602:
601:Protoceratops
598:
593:
591:
587:
586:Protoceratops
583:
578:
570:
566:
559:
554:
547:
545:
543:
540:
536:
532:
529:
525:
522:
518:
514:
510:
506:
502:
498:
494:
490:
486:
482:
478:
474:
470:
462:
458:
454:
447:
445:
443:
439:
438:
437:Protoceratops
433:
429:
425:
421:
419:
415:
411:
407:
403:
399:
395:
391:
388:
384:
380:
379:
365:
362:
357:
354:
349:
346:
341:
338:
337:
335:
332:
328:
321:
320:
312:
309:
305:
298:
297:
290:
287:
286:
283:
277:
274:
273:
270:
264:
261:
258:
257:
254:
248:
245:
242:
241:
238:
232:
229:
226:
225:
222:
216:
213:
210:
209:
206:
203:
200:
197:
196:
193:
190:
187:
186:
183:
180:
177:
176:
173:
170:
167:
166:
161:
156:
152:
148:
143:
139:
134:
120:
115:
110:
105:
100:
95:
90:
85:
80:
75:
70:
65:
59:
50:
44:
40:
37:
33:
19:
4610:Ceratopsians
4511:
4424:
4419:Stellasaurus
4417:
4410:
4403:
4382:
4375:
4370:Coronosaurus
4368:
4363:Centrosaurus
4361:
4332:
4325:
4318:
4311:
4304:
4284:
4279:Lokiceratops
4277:
4270:
4252:Xenoceratops
4250:
4243:
4238:Sinoceratops
4236:
4229:
4222:
4215:
4187:
4180:
4173:
4165:
4158:
4151:
4146:Nedoceratops
4144:
4137:
4119:Vagaceratops
4117:
4112:Utahceratops
4110:
4105:Terminocavus
4103:
4096:
4089:
4082:
4075:
4068:
4061:
4056:Judiceratops
4054:
4047:
4042:Chasmosaurus
4040:
4033:
4026:
4019:
4012:
4005:
3981:
3974:
3967:
3960:
3953:
3946:
3934:Ceratopsidae
3921:Ceratopsidae
3852:Ceratopsidae
3841:Zuniceratops
3839:
3832:
3826:Ajkaceratops
3824:
3800:
3793:
3788:Bagaceratops
3787:
3786:
3780:Ajkaceratops
3778:
3762:Coronosauria
3749:
3742:
3735:
3728:
3721:
3714:
3707:
3700:
3693:
3686:
3679:
3672:
3667:Asiaceratops
3665:
3645:
3633:Euceratopsia
3616:Yamaceratops
3614:
3607:
3600:
3580:
3573:
3568:Microceratus
3566:
3561:Liaoceratops
3559:
3551:
3544:
3537:
3530:
3523:
3517:Ajkaceratops
3515:
3481:
3474:
3467:
3460:
3453:
3433:
3426:
3419:
3337:Ornithischia
3322:Ornithischia
3274:
3270:Ornithischia
3265:
3256:
3191:Bagaceratops
3190:
3153:
3149:
3139:
3114:
3110:
3100:
3075:
3071:
3065:
3032:
3028:
3021:
3015:(2): 85–101.
3012:
3008:
2995:
2968:
2964:
2954:
2935:
2931:
2918:
2885:
2881:
2875:
2830:
2826:
2816:
2789:
2785:
2772:
2763:
2759:
2746:
2737:
2733:
2720:
2695:
2691:
2685:
2660:
2656:
2646:
2611:
2601:
2576:
2572:
2566:
2557:
2553:
2540:
2531:
2527:
2517:
2492:
2488:
2482:
2437:
2433:
2427:
2419:
2410:
2400:
2363:
2359:
2349:
2306:
2293:
2250:
2237:
2215:(1): 16−26.
2212:
2208:
2198:
2173:
2169:
2163:
2126:
2122:
2112:
2095:
2091:
2085:
2076:
2072:
2062:
2053:
2049:
2036:
2011:
2007:
1983:
1977:
1957:
1950:
1925:
1921:
1917:
1865:
1861:
1851:
1824:
1811:
1786:
1782:
1776:
1767:
1763:
1757:
1748:
1735:
1726:
1712:
1693:
1689:
1679:
1654:
1650:
1610:
1606:
1596:
1577:
1573:
1563:
1554:
1545:
1541:
1471:Philovenator
1469:
1463:
1459:Linhevenator
1457:
1448:
1442:
1438:Velociraptor
1436:
1430:
1424:
1420:Pinacosaurus
1418:
1412:
1406:
1369:Bagaceratops
1368:
1365:Magnirostris
1364:
1358:
1338:
1328:
1322:
1318:Conchoraptor
1316:
1314:oviraptorids
1307:
1298:
1289:
1283:
1274:
1265:
1259:
1253:
1244:
1238:
1229:
1223:
1217:
1207:Bagaceratops
1206:
1205:
1149:Bagaceratops
1148:
1130:
1062:Bagaceratops
1061:
1060:
1059:
1036:
1035:
1012:
1011:
988:
987:
959:Bagaceratops
958:
954:
950:
946:
943:Bagaceratops
942:
939:Bagaceratops
938:
934:
931:Magnirostris
930:
927:Lamaceratops
926:
923:Gobiceratops
922:
918:
915:Bagaceratops
914:
912:
894:belonged to
892:Bagaceratops
891:
890:
855:premaxillary
851:Bagaceratops
850:
846:
843:Bagaceratops
842:
836:
832:
829:Bagaceratops
828:
827:
822:Bagaceratops
821:
806:
803:Bagaceratops
802:
798:
795:Bagaceratops
794:
789:
785:
781:
777:
773:
765:
763:
758:
751:Gobiceratops
750:
746:
744:
740:Gobiceratops
739:
729:Bagaceratops
728:
717:Magnirostris
716:
700:
697:Dong Zhiming
689:Bagaceratops
688:
680:
677:Lamaceratops
676:
672:
668:
666:
662:Bagaceratops
661:
657:
650:Bagaceratops
649:
643:
639:
637:
633:Magnirostris
632:
616:
612:
600:
597:Bagaceratops
596:
594:
585:
582:Bagaceratops
581:
577:Bagaceratops
576:
574:
569:Bagaceratops
568:
558:Bagaceratops
557:
534:
530:
523:
517:Bagaceratops
516:
513:Bagaceratops
512:
508:
500:
466:
461:Bagaceratops
460:
442:Bagaceratops
441:
435:
428:Bagaceratops
427:
424:Bagaceratops
423:
422:
414:Bagaceratops
413:
402:Bagaceratops
401:
378:Bagaceratops
377:
376:
375:
363:
355:
347:
339:
318:
317:
308:Type species
296:Bagaceratops
295:
294:
269:Coronosauria
259:
243:
227:
221:Ornithischia
211:
198:
147:Magnirostris
146:
43:Bagaceratops
42:
36:
32:Vagaceratops
18:Lamaceratops
4536:Wikispecies
4412:Einiosaurus
4306:Avaceratops
4189:Triceratops
4153:Ojoceratops
3976:Monoclonius
3688:Ferrisaurus
3553:Kulceratops
2495:: 101–120.
2428:Nemegtomaia
1580:(1): I–II.
1455:troodontids
1432:Linheraptor
1337:titanosaur
1330:Nemegtomaia
1309:Tylocephale
1246:Parvicursor
1240:Khulsanurus
1211:vertebrates
1135:, based on
955:P. andrewsi
833:Maginostris
813:Description
654:Paul Sereno
556:Diagram of
473:Gobi Desert
432:ceratopsids
396:during the
56:72–71
4599:Categories
4475:See also:
4182:Torosaurus
4098:Spiclypeus
3948:Agathaumas
3861:see below↓
3681:Cerasinops
3469:Stenopelix
3403:Ceratopsia
3390:Ceratopsia
3358:see below↓
3349:Ceratopsia
3261:Dinosauria
3228:Ceratopsia
2938:(1): 1–9.
2056:: 111−146.
1548:: 134−143.
1513:References
1450:Wulatelong
1426:Linhenykus
1261:Gobipteryx
1231:Zaraapelta
1189:lacustrine
1177:sandstones
1165:siltstones
1161:claystones
896:Ceratopsia
609:anagenetic
253:Ceratopsia
205:Dinosauria
3244:Kingdom:
3189:HVNHM on
3131:210840006
3057:244186762
2910:129414074
2808:129291148
2638:129367764
2629:2246/1596
2366:: e4868.
2341:243849373
2333:0003-0082
2324:2246/7286
2285:231597229
2277:0003-0082
2268:2246/7251
2190:247222017
2014:: 29–50.
1942:132780322
1890:129893753
1842:2246/5808
1671:150321203
1409:dinosaurs
1377:lithology
1373:Campanian
1349:squamates
1324:Heyuannia
1300:Hulsanpes
1270:; fellow
1255:Gobipipus
1219:Saichania
1145:Campanian
1137:sediments
693:You Hailu
590:clavicles
521:Mongolian
497:monograph
390:dinosaurs
385:of small
178:Kingdom:
172:Eukaryota
4521:Wikidata
4487:Category
4477:Timeline
3983:Polyonax
3969:Dysganus
3962:Ceratops
3525:Aquilops
3252:Chordata
3250:Phylum:
3246:Animalia
2902:20490961
2867:21262806
2560:: 23–44.
2534:: 90−96.
2474:22347465
2434:PLOS ONE
2392:29868277
2155:33737515
1485:See also
1385:alluvial
1267:Hollanda
1153:red beds
685:red beds
623:Synonyms
531:ceratops
505:holotype
331:Synonyms
275:Family:
192:Chordata
188:Phylum:
182:Animalia
168:Domain:
4569:4965778
4556:4531819
4527:Q131256
4377:Spinops
3695:Gremlin
3483:Yinlong
3158:Bibcode
3080:Bibcode
3037:Bibcode
2973:Bibcode
2932:Zootaxa
2858:3038769
2835:Bibcode
2740:: 5−28.
2700:Bibcode
2665:Bibcode
2581:Bibcode
2497:Bibcode
2465:3275628
2442:Bibcode
2383:5978397
2217:Bibcode
2146:7973727
2016:Bibcode
1870:Bibcode
1791:Bibcode
1615:Bibcode
1479:turtles
1381:caliche
1345:mammals
1225:Tarchia
1193:aeolian
859:maxilla
560:KID 196
479:of the
416:in the
288:Genus:
125:↓
3129:
3055:
2908:
2900:
2865:
2855:
2806:
2636:
2472:
2462:
2390:
2380:
2339:
2331:
2283:
2275:
2188:
2153:
2143:
1965:
1940:
1888:
1669:
1468:, and
1453:; and
1389:stream
1312:; and
1191:, and
1185:rivers
949:, and
933:, and
900:parrot
485:Poland
457:Fossil
4582:52826
3275:Clade
3266:Clade
3257:Clade
3127:S2CID
3053:S2CID
3005:(PDF)
2928:(PDF)
2906:S2CID
2898:JSTOR
2804:S2CID
2782:(PDF)
2756:(PDF)
2730:(PDF)
2634:S2CID
2550:(PDF)
2360:PeerJ
2337:S2CID
2303:(PDF)
2281:S2CID
2247:(PDF)
2186:S2CID
2046:(PDF)
1938:S2CID
1886:S2CID
1821:(PDF)
1745:(PDF)
1723:(PDF)
1667:S2CID
1538:(PDF)
1397:lakes
1335:large
1251:birds
1157:sands
721:frill
713:China
528:Greek
383:genus
260:Clade
244:Clade
228:Clade
212:Clade
199:Clade
4564:GBIF
3335:see
3308:see
2936:2403
2863:PMID
2470:PMID
2388:PMID
2329:ISSN
2273:ISSN
2151:PMID
1963:ISBN
1447:and
1435:and
1401:dune
1393:wind
1359:The
1347:and
1327:and
1291:Shri
1288:and
1285:Kuru
1264:and
1243:and
1228:and
1197:arid
1131:The
917:and
908:Asia
906:and
845:and
805:and
784:and
770:2019
695:and
671:and
599:and
524:baga
491:and
408:and
394:Asia
64:PreꞒ
4551:EoL
3539:Beg
3166:doi
3119:doi
3088:doi
3045:doi
3033:130
2981:doi
2940:doi
2890:doi
2853:PMC
2843:doi
2831:108
2794:doi
2708:doi
2673:doi
2624:hdl
2616:doi
2589:doi
2505:doi
2493:494
2460:PMC
2450:doi
2378:PMC
2368:doi
2319:hdl
2311:doi
2263:hdl
2255:doi
2225:doi
2178:doi
2141:PMC
2131:doi
2100:doi
2096:172
2024:doi
2012:494
1930:doi
1878:doi
1837:hdl
1829:doi
1799:doi
1698:doi
1659:doi
1623:doi
1611:311
1582:doi
1578:116
1199:to
1187:),
727:of
615:to
507:is
4601::
4579::
4566::
4553::
4538::
4523::
3277::
3268::
3259::
3164:.
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3113:.
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74:O
69:Ꞓ
53:~
51:,
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