396:
1690:
1732:
606:. This area probably represented a rather shallow environment in the northwestern part of the basin. The fossils are usually fragmentary and preserved as internal or external moulds associated with angular claystone fragments within quartzose sandstones to siltstones. Accumulations of these fossil remains are commonly found in thick-bedded quartzose sandstones of light-grey and yellow or grey-brown color, which might have been deposited in areas sheltered from wave activity by sand bars.
1704:
289:
403:
412:
1718:
588:
deposition in the early
Paleozoic. While in most of the layers fossils are absent, certain levels contain a rich assemblage of shallow marine fauna interpreted as storm-induced beds. The concentration of a wide variety of excellently conserved fossils of the Letná Formation has granted it the name of
719:
Association, characterizing rather proximal and shallow environments of the Letná Formation. Most of the non-trilobite arthropods discovered in this formation, including forms with non-mineralized exoskeletons, are also found associated with
621:
Community, which probably represents a slightly deeper environment, is typically found in the poorly sorted greywackes of the higher horizons of the Letná Formation. These two communities constitute the
2188:
1866:
1952:
Pereira, Sofia; Marques da Silva, Carlos; Sá, Artur Abreu; Pires, Miguel; Marques Guedes, Armando; Budil, Petr; Laibl, Lukáš; Rábano, Isabel (2017), "The illaenid trilobites
2138:
598:
A rather diverse fauna composed of brachiopods, trilobites, echinoderms, conularids, and non-trilobite arthropods has been found in outcrops located between
2193:
503:
dominate, the formation, which is lean in fossils in many areas but exceptionally rich in what has been interpreted as storm beds, has been designated a
395:
2079:
728:. This suggests that the development of conditions conducive to soft-tissue fossilization was not uncommon in the environment represented by the
658:. The more distal, probably deeper parts of the basin are characterized by a poorly diversified assemblage of trilobites associated with rare
1885:"A new pustulose bivalve from the Late Ordovician of the Prague Basin (Czech Republic) and remarks on the diversification of pteriomorphids"
2143:
541:. The Letná Formation is the thickest unit in the sedimentary sequence with a thickness of more than 600 metres (2,000 ft) between
2117:Šnajdr, M., 1956. Trilobiti drabovských a letenských vrstev českého ordoviku. . Sborník Ústředního ústavu geologického, 22, 477-533
1689:
358:
2178:
2114:
V. Havlíček and J. Vaněk. 1966. The biostratigraphy of the
Ordovician of Bohemia. Sborník Geologických Věd, Paleontologie 8:7-69
1703:
2111:
F. C. Shaw. 1995. Ordovician trinucleid trilobites of the Prague Basin, Czech
Republic. Paleontological Society Memoir 40:1-23
1731:
2173:
2168:
2163:
2158:
2148:
2104:
R. J. Horný. 1997. Circumbilical retractor muscle attachment area in the
Ordovician trilobed bellerophontoidean gastropod
2072:"Ordovician of the Barrandian area: Reconstruction of the sedimentary basin, its benthic communities and ichnoassemblages"
2198:
689:
Fossilized gut remains have been observed in four of the about twenty trilobite species present in the Letná Formation:
351:
195:
2133:
241:
202:
499:
Because of the excellent preservation, including gut remains, of a wide assemblage of early
Paleozoic taxa in which
2153:
755:
581:
493:
2051:
1717:
2031:
2183:
1998:
1962:
1652:
458:
1994:"Fossilised guts in trilobites from the Upper Ordovician Letná Formation (Prague Basin, Czech Republic)"
1755:
1632:
1099:
707:
654:
1400:
1561:
1554:
538:
128:
1664:
1596:
1589:
1253:
1169:
909:
534:
138:
118:
1533:
1079:
987:
1582:
1527:
1519:
1491:
1323:
916:
1575:
1547:
1477:
1470:
1456:
1449:
1414:
1337:
1288:
1218:
1204:
1022:
1001:
994:
860:
853:
561:
518:
517:
The Letná Formation was first formally defined in 1966 by Havlíček and Vaněk and crops out in the
473:
446:
304:
1884:
1750:
1421:
1351:
1260:
1211:
1162:
1127:
1071:
1064:
1050:
1036:
1008:
902:
881:
609:
Five communities of marine fauna were identified in the formation. The rich brachiopod dominated
1960:(Barrande, 1846) from the Upper Ordovician of the Czech Republic, Portugal, Spain and Morocco",
1671:
1624:
1365:
1274:
1225:
1197:
1148:
1141:
1120:
1043:
938:
839:
630:
1568:
1407:
1379:
1372:
1155:
1134:
1113:
1085:
888:
867:
832:
617:
529:
in the northeast. The Prague Basin contains an
Ordovician to Devonian succession in the larger
521:, a northeast–southwest trending minibasin stretching over 100 kilometres (62 mi) between
457:
in the Czech capital, more specifically at Letná Hill, after which the formation is named. The
442:
35:
2007:
1330:
1316:
1309:
1302:
1281:
1183:
670:
624:
611:
1610:
1029:
959:
783:
464:
The more than 600 metres (2,000 ft) thick formation comprises a rhythmic alternation of
1979:
1971:
1896:
1484:
931:
776:
1938:
New echinoderm
Lagerstätte from the Letná Formation (Sandbian, Upper Ordovician) of Bohemia
1935:
Nohejlová, Martina; Lefebvre, Bertrand; Nardin, Elise; Fatka, Oldřich; Budil, Petr (2019),
1443:
530:
450:
434:
634:
Fauna. In more proximal environments, this fauna is replaced by the trilobite-dominated
1268:
591:
505:
294:
234:
2127:
2046:
1993:
1936:
1435:
1359:
1233:
973:
952:
945:
75:
27:
1386:
1106:
874:
825:
818:
790:
2071:
1900:
674:
Association are typical for the deepest portions of the Prague Basin. Intertidal (
615:
Community is associated with shallow water, well-sorted quartzose sandstones. The
1870:
1604:
1464:
1429:
895:
811:
797:
769:
695:
636:
603:
50:
1760:
1638:
1618:
1513:
1505:
1247:
1177:
846:
762:
664:
659:
648:
95:
60:
2011:
1975:
373:
360:
256:
243:
217:
204:
1984:
1541:
1295:
1016:
981:
925:
804:
749:
701:
676:
642:
577:
569:
565:
557:
553:
500:
481:
477:
469:
465:
183:
179:
169:
161:
100:
44:
2120:
580:
lenses in the upper part of the succession, deposited in a shallow marine
2025:
1498:
1394:
1345:
1239:
1191:
682:
526:
522:
438:
90:
85:
70:
65:
55:
31:
2108:
sp. n. (Mollusca). Bulletin of the Czech
Geological Survey 72(4):333-338
1093:
1057:
967:
454:
105:
80:
2047:"Trace fossils from the Letná Formation (Ordovician, Czech Republic)"
585:
546:
542:
489:
336:
276:
1646:
599:
573:
485:
165:
595:, giving insight into the diversification of marine ecosystems.
1883:
Polechová, Marika; Zicha, Ondřej; Rak, Štěpán (18 July 2023).
686:
ichnofacies) environments are also evidenced by ichnofossils.
1992:
Fatka, O.; Lerosey-Aubril, R.; Budil, P.; Rak, Š. (2013),
744:
The following fossils were reported from the formation:
1941:, 10th European Conference on Echinoderms, p. 69
552:
The formation consists of a rhythmic alternation of
1835:
1833:
1831:
1829:
350:
342:
331:
323:
315:
310:
300:
282:
272:
233:
194:
189:
175:
157:
152:
144:
134:
124:
114:
21:
1807:
1805:
461:is located at Malá Strana, Holešovice district.
1853:
1851:
1849:
1847:
1845:
1819:
1817:
8:
2189:Fossiliferous stratigraphic units of Europe
1777:
1775:
18:
2139:Geologic formations of the Czech Republic
1983:
826:Eccoptochile (Eccoptochiloides) tumescens
533:. The formation conformably overlies the
2080:Journal of the Czech Geological Society
1771:
1685:
411:
402:
711:. These four trilobites belong to the
662:. Lastly, trilobites of the so-called
2024:Kříž, Jiří; Steinová, Marika (2009),
819:Eccoptochile (Eccoptochile) clavigera
453:in the Czech Republic. The formation
7:
16:Geologic formation in Czech Republic
2194:Paleontology in the Czech Republic
868:Primaspis (Primaspis) primordialis
441:, or in the regional stratigraphy
14:
668:Biofacies and brachiopods of the
1730:
1716:
1702:
1688:
896:Zeliszkella (Zeliszkella) hawlei
419:Letná Formation (Czech Republic)
410:
401:
394:
287:
1956:(Vaněk & Vokáč, 1997) and
148:>600 m (2,000 ft)
1:
1901:10.1016/j.geobios.2023.04.001
680:ichnofacies) and near-shore (
1839:Noheljová et al., 2019, p.69
847:Placoparia (Hawleia) grandis
646:Association, which contains
445:) geologic formation of the
235:Approximate paleocoordinates
2144:Ordovician System of Europe
1922:Pereira et al., 2017, p.466
1590:Pseudoconularia grandissima
1401:Barrandicellopsis extenuata
2215:
1485:Echinosphaerites infaustus
1781:Kříž & Steinová, 2009
1696:Bicuspina multicostellata
1665:Chasmatoporella havliceki
1471:Ascocystites drabowiensis
1415:Dactylogonia blyskavensis
1289:Drabovinella draboviensis
1254:Mespilocystites bohemicus
389:
335:Malá Strana, Holešovice,
26:
2070:Mikuláš, Radek (1998b),
2052:Sborník Geologických Věd
2045:Mikuláš, Radek (1998a),
1976:10.3140/bull.geosci.1642
1913:Fatka et al., 2013, p.99
1857:Fatka et al., 2013, p.98
1823:Fatka et al., 2013, p.97
1799:Fatka et al., 2013, p.96
1710:Caleidocrinus multiramus
1583:Metaconularia pyramidata
1555:Archaeoconularia fecunda
1534:Dendrocystites sedgwicki
1282:Jezercia chrustenicensis
974:Caleidocrinus multiramus
2179:Shallow marine deposits
2032:Czech Geological Survey
1999:Bulletin of Geosciences
1963:Bulletin of Geosciences
1436:Conchicolites confertus
537:and is overlain by the
374:50.094583°N 14.417500°E
1653:Dendrocystis barrandei
1576:Exoconularia exquisita
1492:Macrocystella bohemica
1450:Hemicystites bohemicus
1275:Svobodaina ellipsoides
1107:Concavodonta ponderata
1790:Mikuláš, 1998b, p.143
1756:Folkeslunda Limestone
1639:Plumulites folliculum
1387:Rafinesquina occulata
1114:Grammysia catilloides
861:Platycoryphe bohemica
756:Actinopeltis completa
592:Konservat-Lagerstätte
525:in the southwest and
506:Konservat-Lagerstätte
2174:Open marine deposits
2169:Quartzite formations
2164:Limestone formations
2159:Siltstone formations
2149:Sandstone formations
1672:Versispira contraria
1611:Triopus draboviensis
1548:Anaconularia anomala
1422:Sinuitopsis neglecta
1352:Petrocrania obsoleta
1128:Modiolopsis veterana
1065:Tritonophon bohemica
1030:Bucania draboviensis
1023:Asinomphalus antiqua
995:Orbiculoidea grandis
960:Zonozoe drabowiensis
798:Dalmanitina socialis
791:Colpocoryphe grandis
696:Dalmanitina socialis
568:, intercalated with
488:deposited in marine
480:, intercalated with
379:50.094583; 14.417500
327:Havlíček & Vaněk
119:Geological formation
2199:Geography of Prague
1811:Mikuláš, 1998a, p.5
1520:Mitrocystites mitra
1506:Rhombifera bohemica
1499:Cystidea obscondita
1408:Cytherina graegaria
1205:Crescentilla pugnax
1198:Cerninella bohemica
1142:Praenucula bohemica
1121:Leptodesma patricia
1037:Bucanopsina calypso
1002:Schizocrania incola
939:Quasicaris bohemica
910:Heterocyclopyge sp.
882:Stenopareia panderi
840:Pharostoma pulchrum
833:Kloucekia phillipsi
805:Deanaspis goldfussi
770:Calymenella parvula
702:Deanaspis goldfussi
370: /
253: /
214: /
28:Stratigraphic range
1751:Fezouata Formation
1569:Conularia rugulosa
1380:B. multicostellata
1240:Ribeiria apusoides
1212:Cytheropsis testis
1135:Myoplusia bilunata
1100:Astarte convergens
1051:Cyrtodiscus procer
889:Vysocania moraveci
875:Selenopeltis buchi
708:Selenopeltis buchi
655:Selenopeltis buchi
2106:Tritonophon peeli
1738:Holopea praestans
1366:Aegiromena aquila
1324:Chrustenopora sp.
1303:Drabovia asperula
1296:Hirnantia ulrichi
1226:Trubinella latens
1086:Antispira praecox
1058:Holopea antiquata
1044:Bucanopsis comata
917:Prionocheilus sp.
854:Plaesiacomia rara
812:Eudolatites dubia
784:Cekovia transfuga
763:Birmanites ingens
691:Birmanites ingens
649:Birmanites ingens
584:, representing a
427:
426:
2206:
2154:Shale formations
2094:
2093:
2092:
2076:
2066:
2065:
2064:
2041:
2040:
2039:
2020:
2019:
2018:
1988:
1987:
1985:20.500.12468/767
1948:
1947:
1946:
1923:
1920:
1914:
1911:
1905:
1904:
1880:
1874:
1864:
1858:
1855:
1840:
1837:
1824:
1821:
1812:
1809:
1800:
1797:
1791:
1788:
1782:
1779:
1734:
1720:
1706:
1692:
1478:A. cf. micraster
1338:Saukrodictya sp.
1219:Hastatellina sp.
1184:Nothozoe pollens
777:Caryon bohemicus
539:Vinice Formation
414:
413:
405:
404:
398:
385:
384:
382:
381:
380:
375:
371:
368:
367:
366:
363:
293:
291:
290:
268:
267:
265:
264:
263:
258:
254:
251:
250:
249:
246:
229:
228:
226:
225:
224:
219:
215:
212:
211:
210:
207:
129:Vinice Formation
110:
47:
19:
2214:
2213:
2209:
2208:
2207:
2205:
2204:
2203:
2134:Letná Formation
2124:
2123:
2101:
2099:Further reading
2090:
2088:
2074:
2069:
2062:
2060:
2044:
2037:
2035:
2027:Letenský profil
2023:
2016:
2014:
1991:
1951:
1944:
1942:
1934:
1931:
1926:
1921:
1917:
1912:
1908:
1882:
1881:
1877:
1867:Letná Formation
1865:
1861:
1856:
1843:
1838:
1827:
1822:
1815:
1810:
1803:
1798:
1794:
1789:
1785:
1780:
1773:
1769:
1747:
1740:
1735:
1726:
1724:Duslia insignis
1721:
1712:
1707:
1698:
1693:
1684:
1679:
1444:Edrioasteroidea
1310:D. fascicostata
1261:Anatifopsis sp.
1170:Alena pustulosa
1163:Palaeoneilo sp.
1009:Schizotreta sp.
932:Duslia insignis
903:Girvanopyge sp.
742:
535:Libeň Formation
531:Bohemian Massif
515:
451:Bohemian Massif
435:Late Ordovician
431:Letná Formation
423:
422:
421:
420:
417:
416:
415:
407:
406:
378:
376:
372:
369:
364:
361:
359:
357:
356:
288:
286:
261:
259:
255:
252:
247:
244:
242:
240:
239:
222:
220:
216:
213:
208:
205:
203:
201:
200:
139:Libeň Formation
109:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
42:
41:
39:
22:Letná Formation
17:
12:
11:
5:
2212:
2210:
2202:
2201:
2196:
2191:
2186:
2184:Tidal deposits
2181:
2176:
2171:
2166:
2161:
2156:
2151:
2146:
2141:
2136:
2126:
2125:
2122:
2121:
2118:
2115:
2112:
2109:
2100:
2097:
2096:
2095:
2067:
2042:
2021:
1989:
1970:(4): 465–490,
1949:
1930:
1927:
1925:
1924:
1915:
1906:
1875:
1859:
1841:
1825:
1813:
1801:
1792:
1783:
1770:
1768:
1765:
1764:
1763:
1758:
1753:
1746:
1743:
1742:
1741:
1736:
1729:
1727:
1722:
1715:
1713:
1708:
1701:
1699:
1694:
1687:
1683:
1680:
1678:
1677:
1676:
1675:
1668:
1658:
1657:
1656:
1644:
1643:
1642:
1630:
1629:
1628:
1625:Serpulites sp.
1616:
1615:
1614:
1602:
1601:
1600:
1593:
1586:
1579:
1572:
1565:
1558:
1551:
1539:
1538:
1537:
1525:
1524:
1523:
1511:
1510:
1509:
1502:
1495:
1488:
1481:
1474:
1462:
1461:
1460:
1453:
1441:
1440:
1439:
1427:
1426:
1425:
1418:
1411:
1404:
1392:
1391:
1390:
1383:
1376:
1373:Bicuspina cava
1369:
1357:
1356:
1355:
1343:
1342:
1341:
1334:
1327:
1320:
1313:
1306:
1299:
1292:
1285:
1278:
1269:Rhynchonellata
1266:
1265:
1264:
1257:
1245:
1244:
1243:
1231:
1230:
1229:
1222:
1215:
1208:
1201:
1189:
1188:
1187:
1175:
1174:
1173:
1166:
1159:
1152:
1145:
1138:
1131:
1124:
1117:
1110:
1103:
1091:
1090:
1089:
1080:Paragastropoda
1077:
1076:
1075:
1068:
1061:
1054:
1047:
1040:
1033:
1026:
1014:
1013:
1012:
1005:
998:
991:
988:Lingula deleta
979:
978:
977:
965:
964:
963:
956:
949:
946:Furca bohemica
942:
935:
924:Non-trilobite
922:
921:
920:
913:
906:
899:
892:
885:
878:
871:
864:
857:
850:
843:
836:
829:
822:
815:
808:
801:
794:
787:
780:
773:
766:
759:
746:
741:
740:Fossil content
738:
514:
511:
425:
424:
418:
409:
408:
400:
399:
393:
392:
391:
390:
387:
386:
354:
348:
347:
344:
340:
339:
333:
329:
328:
325:
321:
320:
317:
313:
312:
308:
307:
302:
298:
297:
295:Czech Republic
284:
280:
279:
274:
270:
269:
237:
231:
230:
198:
192:
191:
187:
186:
177:
173:
172:
159:
155:
154:
150:
149:
146:
142:
141:
136:
132:
131:
126:
122:
121:
116:
112:
111:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
48:
24:
23:
15:
13:
10:
9:
6:
4:
3:
2:
2211:
2200:
2197:
2195:
2192:
2190:
2187:
2185:
2182:
2180:
2177:
2175:
2172:
2170:
2167:
2165:
2162:
2160:
2157:
2155:
2152:
2150:
2147:
2145:
2142:
2140:
2137:
2135:
2132:
2131:
2129:
2119:
2116:
2113:
2110:
2107:
2103:
2102:
2098:
2086:
2082:
2081:
2073:
2068:
2058:
2054:
2053:
2048:
2043:
2033:
2029:
2028:
2022:
2013:
2009:
2005:
2001:
2000:
1995:
1990:
1986:
1981:
1977:
1973:
1969:
1965:
1964:
1959:
1955:
1950:
1940:
1939:
1933:
1932:
1928:
1919:
1916:
1910:
1907:
1902:
1898:
1894:
1890:
1886:
1879:
1876:
1872:
1868:
1863:
1860:
1854:
1852:
1850:
1848:
1846:
1842:
1836:
1834:
1832:
1830:
1826:
1820:
1818:
1814:
1808:
1806:
1802:
1796:
1793:
1787:
1784:
1778:
1776:
1772:
1766:
1762:
1759:
1757:
1754:
1752:
1749:
1748:
1744:
1739:
1733:
1728:
1725:
1719:
1714:
1711:
1705:
1700:
1697:
1691:
1686:
1681:
1674:
1673:
1669:
1667:
1666:
1662:
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1659:
1655:
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1627:
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1620:
1617:
1613:
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1608:
1607:
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1603:
1599:
1598:
1594:
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1591:
1587:
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1584:
1580:
1578:
1577:
1573:
1571:
1570:
1566:
1564:
1563:
1559:
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1550:
1549:
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1543:
1540:
1536:
1535:
1531:
1530:
1529:
1526:
1522:
1521:
1517:
1516:
1515:
1512:
1508:
1507:
1503:
1501:
1500:
1496:
1494:
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1433:
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1428:
1424:
1423:
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1417:
1416:
1412:
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1405:
1403:
1402:
1398:
1397:
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1393:
1389:
1388:
1384:
1382:
1381:
1377:
1375:
1374:
1370:
1368:
1367:
1363:
1362:
1361:
1360:Strophomenata
1358:
1354:
1353:
1349:
1348:
1347:
1344:
1340:
1339:
1335:
1333:
1332:
1328:
1326:
1325:
1321:
1319:
1318:
1314:
1312:
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1307:
1305:
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1300:
1298:
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1293:
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1290:
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1279:
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1276:
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1270:
1267:
1263:
1262:
1258:
1256:
1255:
1251:
1250:
1249:
1246:
1242:
1241:
1237:
1236:
1235:
1234:Rostroconchia
1232:
1228:
1227:
1223:
1221:
1220:
1216:
1214:
1213:
1209:
1207:
1206:
1202:
1200:
1199:
1195:
1194:
1193:
1190:
1186:
1185:
1181:
1180:
1179:
1176:
1172:
1171:
1167:
1165:
1164:
1160:
1158:
1157:
1156:Deceptrix sp.
1153:
1151:
1150:
1146:
1144:
1143:
1139:
1137:
1136:
1132:
1130:
1129:
1125:
1123:
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1118:
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1111:
1109:
1108:
1104:
1102:
1101:
1097:
1096:
1095:
1092:
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1083:
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1078:
1074:
1073:
1069:
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1066:
1062:
1060:
1059:
1055:
1053:
1052:
1048:
1046:
1045:
1041:
1039:
1038:
1034:
1032:
1031:
1027:
1025:
1024:
1020:
1019:
1018:
1015:
1011:
1010:
1006:
1004:
1003:
999:
997:
996:
992:
990:
989:
985:
984:
983:
980:
976:
975:
971:
970:
969:
966:
962:
961:
957:
955:
954:
950:
948:
947:
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940:
936:
934:
933:
929:
928:
927:
923:
919:
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914:
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905:
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869:
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858:
856:
855:
851:
849:
848:
844:
842:
841:
837:
835:
834:
830:
828:
827:
823:
821:
820:
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813:
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800:
799:
795:
793:
792:
788:
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771:
767:
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764:
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758:
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753:
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751:
748:
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745:
739:
737:
736:Association.
735:
731:
727:
723:
718:
714:
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704:
703:
698:
697:
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687:
685:
684:
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678:
673:
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633:
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619:
614:
613:
607:
605:
601:
596:
594:
593:
587:
583:
579:
575:
571:
567:
563:
562:subgreywackes
559:
555:
550:
548:
544:
540:
536:
532:
528:
524:
520:
512:
510:
508:
507:
502:
497:
495:
491:
487:
483:
479:
475:
474:subgreywackes
471:
467:
462:
460:
459:type locality
456:
452:
448:
444:
440:
436:
432:
397:
388:
383:
365:014°25′03.0″E
355:
353:
349:
345:
341:
338:
334:
330:
326:
322:
318:
314:
309:
306:
303:
299:
296:
285:
281:
278:
275:
271:
266:
257:58.4°S 81.7°E
238:
236:
232:
227:
218:50.0°N 14.0°E
199:
197:
193:
188:
185:
181:
178:
174:
171:
167:
163:
160:
156:
151:
147:
143:
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133:
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117:
113:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
46:
43:460–450
37:
33:
29:
25:
20:
2105:
2089:, retrieved
2084:
2078:
2061:, retrieved
2056:
2050:
2036:, retrieved
2026:
2015:, retrieved
2003:
1997:
1967:
1961:
1957:
1953:
1943:, retrieved
1937:
1929:Bibliography
1918:
1909:
1892:
1888:
1878:
1862:
1795:
1786:
1737:
1723:
1709:
1695:
1670:
1663:
1651:
1637:
1623:
1609:
1595:
1588:
1581:
1574:
1567:
1560:
1553:
1546:
1532:
1518:
1504:
1497:
1490:
1483:
1476:
1469:
1455:
1448:
1434:
1420:
1413:
1406:
1399:
1385:
1378:
1371:
1364:
1350:
1336:
1331:Onniella sp.
1329:
1322:
1315:
1308:
1301:
1294:
1287:
1280:
1273:
1259:
1252:
1238:
1224:
1217:
1210:
1203:
1196:
1182:
1168:
1161:
1154:
1147:
1140:
1133:
1126:
1119:
1112:
1105:
1098:
1084:
1070:
1063:
1056:
1049:
1042:
1035:
1028:
1021:
1007:
1000:
993:
986:
972:
958:
951:
944:
937:
930:
915:
908:
901:
894:
887:
880:
873:
866:
859:
852:
845:
838:
831:
824:
817:
810:
803:
796:
789:
782:
775:
768:
761:
754:
743:
733:
729:
725:
722:D. goldfussi
721:
716:
712:
706:
700:
694:
690:
688:
681:
675:
669:
663:
653:
647:
641:
635:
629:
623:
616:
610:
608:
597:
590:
551:
519:Prague Basin
516:
504:
498:
494:environments
463:
447:Prague Basin
430:
428:
362:50°05′40.5″N
343:Year defined
311:Type section
305:Prague Basin
2034:, p. 1
1958:Octillaenus
1895:: 135–143.
1871:Fossilworks
1633:Machaeridia
1605:Merostomata
1562:A. insignis
1465:Eocrinoidea
1430:Tentaculita
1178:Crustaceans
730:Dalmanitina
726:D. socialis
713:Dalmanitina
660:graptolites
637:Dalmanitina
604:Chrustenice
582:environment
513:Description
377: /
352:Coordinates
262:-58.4; 81.7
260: /
221: /
196:Coordinates
2128:Categories
2091:2020-05-29
2063:2020-05-29
2038:2020-05-29
2017:2020-05-29
2006:: 95–104,
1945:2020-05-29
1767:References
1761:Soom Shale
1619:Polychaeta
1597:P. nobilis
1514:Stylophora
1457:H. velatus
1248:Blastoidea
1017:Gastropods
926:arthropods
750:Trilobites
665:Cyclopygid
631:Aegiromena
570:siltstones
566:quartzites
558:greywackes
554:sandstones
501:trilobites
482:siltstones
478:quartzites
470:greywackes
466:sandstones
319:Letná Hill
223:50.0; 14.0
2087:: 143–159
2012:1214-1119
1954:Vysocania
1542:Scyphozoa
1192:Ostracods
1149:P. dispar
982:Lingulata
953:Furca sp.
734:Deanaspis
717:Deanaspis
677:Skolithos
643:Deanaspis
618:Bicuspina
578:limestone
455:crops out
443:Berounian
316:Named for
184:quartzite
180:Limestone
170:siltstone
162:Sandstone
153:Lithology
145:Thickness
125:Underlies
36:Berounian
1745:See also
1395:Tergomya
1346:Craniata
1317:D. redux
1094:Bivalves
1072:T. peeli
968:Crinoids
683:Cruziana
671:Paterula
625:Drabovia
612:Drabovia
523:Rokycany
439:Sandbian
332:Location
324:Named by
190:Location
135:Overlies
32:Sandbian
1889:Geobios
1682:Gallery
724:and/or
283:Country
248:81°42′E
245:58°24′S
209:14°00′E
206:50°00′N
158:Primary
2059:: 5–25
2010:
1660:Other
1528:Soluta
586:flysch
574:shales
547:Prague
543:Beroun
492:-like
490:flysch
486:shales
337:Prague
301:Extent
292:
277:Prague
273:Region
2075:(PDF)
1647:Algae
600:Zdice
576:with
527:Úvaly
433:is a
176:Other
166:shale
2008:ISSN
1873:.org
705:and
652:and
602:and
572:and
560:and
545:and
484:and
472:and
429:The
346:1966
115:Type
51:PreꞒ
1980:hdl
1972:doi
1897:doi
1869:at
564:),
476:),
2130::
2085:43
2083:,
2077:,
2057:34
2055:,
2049:,
2030:,
2004:88
2002:,
1996:,
1978:,
1968:92
1966:,
1893:81
1891:.
1887:.
1844:^
1828:^
1816:^
1804:^
1774:^
699:,
693:,
589:a
549:.
509:.
496:.
449:,
182:,
168:,
164:,
101:Pg
45:Ma
30::
1982::
1974::
1903:.
1899::
732:-
715:-
640:-
628:-
556:(
468:(
437:(
106:N
96:K
91:J
86:T
81:P
76:C
71:D
66:S
61:O
56:Ꞓ
40:~
38:)
34:(
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