1359:
1279:
743:
1339:
453:
80:
713:
753:
1299:
443:
1369:
1309:
463:
1329:
1289:
703:
370:
723:
90:
1239:
733:
58:(BAP) listed 22 butterfly species. The 2011 Red List of British butterflies lists 4 species as "regionally extinct" (RE), 2 as "critically endangered", 8 as "endangered (E), 9 as "vulnerable" (V), 11 as "near threatened" (NT) and 28 as "least concern" (LC) in a UK context. In the list below, the categories are as taken from the 2022 Red List (RE 4, E 8, V 16, NT 5, LC 29). Range expansions according to the
1319:
1349:
2077:– throughout, except north-west England and north-west half of Scotland; expanding range in English Midlands, western England, English–Scottish borders, and Scotland; "infilling" southern Scotland
2893:
3083:
3128:
68:
lists 29 of Great
Britain's 58 breeding butterfly species as "High UK threat priority", with 9 of those with conservation priority status "Action urgent across UK range".
3138:
3016:
3240:
3680:
3660:
3421:
3351:
3245:
3161:
3103:
3260:
3133:
3665:
3670:
3558:
3371:
3113:
3614:
3396:
3108:
3046:
55:
3629:
3581:
3478:
3293:
1358:
3184:
3118:
3443:
3009:
2780:
51:
1863:– throughout England and Wales, southern Scotland (localized); expanding north and "infilling", but declined rapidly inland in East Anglia
579:– throughout England (except north) and Wales (except south-west, central west and far north-west); expanding range north and "infilling"
3591:
3204:
1823:– throughout southern third of Great Britain, Scottish Highlands; expanding range north and east in England and Scotland; has colonized
1278:
3543:
742:
3513:
3473:
3453:
3433:
3336:
3224:
3069:
2414:
2806:
Fox, Richard; Warren, Martin S.; Brereton, Tom M.; Roy, David B.; Robinson, Anna (2011). "A new Red List of
British butterflies".
2392:
Species included in the Great
Britain Lepidoptera numbering system, but believed never to have occurred naturally in a wild state
3619:
3448:
3219:
3093:
3685:
3639:
3624:
3609:
3528:
3523:
3498:
3356:
3002:
1388:
1338:
452:
3675:
3553:
3503:
3463:
3416:
3411:
3401:
3386:
3381:
3376:
3361:
3346:
3341:
3331:
3326:
3321:
3316:
3209:
3143:
3123:
2731:
2726:
3483:
1950:– throughout south-east half of the country, including West Country, but not most of East Anglia; expanding range northwards
712:
752:
3634:
3586:
3576:
3548:
3538:
3518:
3508:
3493:
3488:
3468:
3438:
3406:
3391:
3366:
3302:
3214:
3088:
3078:
79:
3533:
3458:
3428:
1298:
442:
3255:
1368:
1308:
462:
35:, including extinct, naturalised species and those of dubious origin. The list comprises butterfly species listed in
3286:
3179:
2641:
2459:
2336:
1806:
1430:
1332:
1513:
1328:
1288:
1282:
688:– throughout, except far north and north-west; expanding range in Scotland and "infilling" in England and Wales
109:
369:
2207:(now vagrant only, although sightings in southern England since 2007 suggest recolonisation may be occurring)
2085:
1372:
854:
54:
in 2004 found there has been a species decline of 71% of butterfly species between 1983 and 2003. The 2007
3250:
3025:
2880:
2720:
2423:
2004:
1871:
1352:
702:
674:
466:
65:
950:– southern and south-central England, south Wales coast, and east coast of Scotland (patchy distribution)
3279:
2377:
2308:
1482:
1460:
389:
373:
1452:– patchily distributed through southern England, Wales, north-west and north-east England, and Scotland
722:
2614:
2477:
2235:
1958:
1739:
1056:
736:
89:
2540:
2531:
2522:
2200:
1936:
1647:
1342:
1302:
958:
2973:
2858:
2823:
2695:
2513:
2450:
2432:
2405:
2368:
2280:
2271:
2116:
639:
456:
425:
331:
1238:
812:
estuary, with concentrations in south-west Wales, north Devon and south-west
Somerset, and west
3041:
2549:
2486:
2345:
2317:
2289:
2262:
2210:
1695:
1543:
1322:
832:
565:
446:
128:
2918:
1709:– throughout England and Wales; expanding range northwards and spreading in southern Scotland
17:
3036:
2850:
2815:
2704:
2659:
2632:
2623:
2590:
2585:
2244:
1849:
1769:
1717:
1603:
1292:
1006:
876:
794:
772:
716:
706:
513:
482:
421:
222:
207:
93:
2894:"BC UK Conservation Strategy Appendix 3: Conservation status of threatened UK butterflies"
2677:
2668:
2650:
2567:
2504:
2441:
2326:
2298:
2253:
2171:
1839:
1758:
1527:– south-western half of England and Wales; (re)expanding range east, including East Anglia
1250:
1242:
509:
3051:
2596:
2576:
2468:
2018:– throughout southern half of country, except central Wales; expanding range northwards
1625:
1420:
1124:
821:
746:
732:
539:
517:
407:
236:
101:
2197:(Great Britain subspecies extinct; continental subspecies introduced now also extinct)
3654:
2862:
2819:
2495:
2396:
2180:
1784:
1754:
1684:
1535:
997:
928:
868:– throughout much of England (except far south-west and north-west) and eastern Wales
521:
474:
341:
316:
286:
177:
147:
83:
32:
2827:
846:– throughout most of England and Wales, more thinly distributed north to River Clyde
2558:
2226:
2190:
2162:
2026:
1905:
1573:
1380:
1362:
1318:
1264:– central-south England; "pockets" in north-east, north-west and south-east England
1183:
906:
617:
381:
362:
192:
2361:(single record from Cambridgeshire, April 2011, coincident with influx of vagrant
1348:
3271:
2605:
2354:
1824:
1595:
1271:
1146:
1102:
1023:
976:
726:
595:
554:
139:
72:
2758:
Vol. 7 Part 1 (Hesperiidae to
Nymphalidae). Harley Books, Colchester, UK. 370p.
2686:
1565:
1231:
1216:
1198:
1187:
1168:
1074:
1038:
936:
756:
695:
531:
1798:
1669:
1312:
990:
817:
764:
275:
271:
263:
28:
2994:
2794:
Priority butterfly species listed in the UK Biodiversity Action Plan 2007
1994:
993:
587:
501:
435:
411:
308:
124:
123:– formerly thinly distributed in south and east, now confined to western
2854:
2362:
2063:
1889:
898:
226:
166:
2841:
Fox, Richard; Dennis, Emily B.; Brown, Andrew F.; Curson, Jon (2022).
2139:– central-west and north Wales, northern England and southern Scotland
809:
505:
267:
259:
255:
196:
2842:
2953:
Butterfly
Conservation Cambridgeshire & Essex Branch Newsletter
330:– thinly distributed through much of England and Wales, and in the
1788:
1367:
1357:
1347:
1337:
1327:
1317:
1307:
1297:
1287:
1277:
1237:
986:
813:
751:
741:
731:
721:
711:
701:
497:
461:
451:
441:
368:
355:– southern England north to north-east Wales, and south-east Wales
162:
88:
78:
1975:– much of English coast, inland in parts of south and East Anglia
300:– throughout England and Wales, and north to south-west Scotland
251:
169:, except far north-east and north-west; spreading north and west
3275:
2998:
191:– throughout south-east England, with scattered populations in
1687:; expanding range throughout Scotland, including Western Isles
1413:– widespread in Wales, Scotland, southern and northern England
609:– throughout, but thinly spread in north-west half of Scotland
2133:– patchy distribution throughout northern and central England
2043:– throughout (except for areas occupied by other subspecies)
2754:(Ed.), 1990. The Butterflies of Great Britain and Ireland.
553:– immigrant, though overwintering in south-west; north to
1041:, south and north coasts of Wales; expanding range north
2756:
37:
656:– throughout (except for area occupied by subspecies
3600:
3567:
3309:
3233:
3197:
3170:
3152:
3067:
3060:
2723:– Britain's butterfly and moth conservation society
979:
coast, and north Wales and north-east Wales borders
890:– confined to heavy clay soils along Chiltern hills
996:of south and south-east coasts, now restricted to
1499:– patchy distribution throughout, except Scotland
2877:2010 Atlas of Butterflies in Britain and Ireland
60:2010 Atlas of Butterflies in Britain and Ireland
1474:– patchy distribution in west England and Wales
1160:– south England; expanding range north and west
127:; re-establishment project ongoing (2018–2020)
3287:
3010:
1224:– introduced to various sites in west England
920:– throughout, except far north and north-west
8:
1683:– throughout, except Scottish Highlands and
2102:– throughout, except far north and Hebrides
3294:
3280:
3272:
3064:
3017:
3003:
2995:
2919:"Back from the Brink: Roots of Rockingham"
2796:. Butterfly Conservation, Wareham, UK. 1p.
1617:– resident and common immigrant throughout
1073:(Castle Eden argus) – England from north
2967:
2965:
2963:
2961:
1037:– throughout southern England, north to
3241:Endangered species of the British Isles
2843:"A new Red List of British butterflies"
2743:
1015:– extinct (formerly north-west England)
250:– restricted to southern England: east
2586:Small brown shoemaker (orange mapwing)
2055:– western Scotland, including Hebrides
428:to southern England and southern Wales
221:– confined to the south coast between
3681:Lists of animals of the British Isles
3661:Lists of insects in the British Isles
2309:Scarce or yellow-legged tortoiseshell
1832:– western Scotland and Inner Hebrides
1731:– patchy distribution, mostly in west
1274:– fritillaries, nymphalids and browns
7:
3246:Endemic species of the British Isles
2559:Albin's Hampstead eye (meadow argus)
1557:– southern England and eastern Wales
3261:Birds of Leicestershire and Rutland
2951:Williamson, B., 2011. Plain Tiger.
2942:Worcestershire Life June 2013. p71.
2771:. WildGuides, Old Basing, UK. 192p.
2152:Vagrant, extinct and exotic species
1761:(formerly widespread in south-east)
406:(endemic subspecies) – confined to
2767:Tomlinson, D. and R. Still, 2002.
25:
3666:Lists of insects of Great Britain
2847:Insect Conservation and Diversity
2808:Insect Conservation and Diversity
1086:across northern England and Wales
1082:– evidence of hybridization with
1048:across northern England and Wales
1044:– evidence of hybridization with
679:Anthocharis cardamines britannica
199:estuary; spreading north and west
3671:Lists of butterflies by location
2820:10.1111/j.1752-4598.2010.00117.x
975:– south-west and south England,
698:– hairstreaks, coppers and blues
1981:– eastern Scotland (near coast)
1389:Small pearl-bordered fritillary
837:Neozephyrus (Quercusia) quercus
3303:List of butterflies of Europe
2899:. Butterfly Conservation. 2016
2792:Butterfly Conservation, 2007.
2732:List of moths of Great Britain
2727:List of butterflies of Ireland
2635:or Mediterranean fritillary –
2333:(formerly introduced and bred)
1722:Euphydryas (Eurodryas) aurinia
859:Satyrium (Strymonidia) w-album
631:– throughout, except far north
1:
3129:Orthoptera and allied insects
816:. In 2009 eggs were found at
18:List of butterflies of Jersey
2875:Fox, R. and J. Asher, 2010.
1173:Celastrina argiolus britanna
881:Satyrium (Strymonidia) pruni
786:– throughout much of country
424:and occasional breeder from
3256:Flora and fauna of Cornwall
3109:Damselflies and dragonflies
2415:Oberthür's grizzled skipper
2009:Pyronia tithonus britanniae
56:UK Biodiversity Action Plan
3702:
1941:Melanargia galathea serena
3084:Aquatic heteropteran bugs
3032:
2883:, East Lulworth, UK. 68p.
2642:Great spangled fritillary
2337:Queen of Spain fritillary
1783:– West Country and Kent;
1431:Pearl-bordered fritillary
1107:Polyommatus icarus icarus
2294:Cupido (Everes) argiades
2272:Lang's short-tailed blue
1514:Silver-washed fritillary
1283:Silver-washed fritillary
1092:A. artaxerxes artaxerxes
570:Gonepteryx rhamni rhamni
487:Leptidea sinapis sinapis
278:; expanding distribution
195:and as far north as the
114:Carterocephalus palaemon
47:by Tomlinson and Still.
3139:Soldierflies and allies
2469:Moorland clouded yellow
2305:(imported on geraniums)
2236:Berger's clouded yellow
2106:C. pamphilus rhoumensis
2047:M. jurtina cassiteridum
1419:– western Scotland and
1215:(endemic subspecies) –
1190:; expanding range north
855:White-letter hairstreak
666:– east-central Scotland
622:Pieris (Artogeia) rapae
3686:Fauna of Great Britain
3352:Bosnia and Herzegovina
3026:Fauna of Great Britain
2881:Butterfly Conservation
2750:Emmet, A.M., J. Heath
2721:Butterfly Conservation
2696:Cassia's owl-butterfly
2100:C. pamphilus pamphilus
1872:Small mountain ringlet
1639:– immigrant throughout
1375:
1365:
1355:
1345:
1335:
1325:
1315:
1305:
1295:
1285:
1245:
1071:A. artaxerxes salmacis
1046:A. artaxerxes salmacis
759:
749:
739:
729:
719:
709:
644:Pieris (Artogeia) napi
469:
459:
449:
404:P. machaon britannicus
376:
237:Silver-spotted skipper
96:
86:
66:Butterfly Conservation
3676:Butterflies of Europe
2769:Britain's Butterflies
2378:American painted lady
2313:Nymphalis xanthomelas
2285:Iphiclides podalirius
2254:Western dappled white
2205:Nymphalis polychloros
2090:Coenonympha pamphilus
2068:Aphantopus hyperantus
1926:E. aethiops caledonia
1483:Dark green fritillary
1461:High brown fritillary
1371:
1361:
1351:
1341:
1331:
1321:
1311:
1301:
1291:
1281:
1241:
911:Lycaena phlaeas eleus
755:
745:
735:
725:
715:
705:
465:
455:
445:
374:Old World swallowtail
372:
152:Thymelicus sylvestris
92:
82:
45:Britain's Butterflies
2974:"Butterfly taxonomy"
2615:Aphrodite fritillary
2541:Green-underside blue
2482:Gonepteryx cleopatra
2428:Saliana longirostris
2382:Vanessa virginiensis
2240:Colias alfacariensis
2053:M. jurtina splendida
2041:M. jurtina insularis
1920:E. aethiops aethiops
1753:– southern coast of
1740:Glanville fritillary
1057:Northern brown argus
818:Feckenham Wylde Moor
737:Northern brown argus
438:– whites and yellows
3568:States with limited
3162:Non-marine molluscs
2705:Illioneus giant owl
2691:Arethusana arethusa
2606:Weaver's fritillary
2545:Glaucopsyche alexis
2536:Polyommatus dorylas
2523:Purple-edged copper
2227:Pale clouded yellow
2211:Almond-eyed ringlet
2201:Large tortoiseshell
1991:H. semele atlantica
1895:E. epiphron scotica
1648:Small tortoiseshell
1417:B. selene insularum
1303:Small tortoiseshell
1003:P. argus caernensis
959:Silver-studded blue
540:Dark clouded yellow
418:P. machaon gorganus
321:Erynnis tages tages
3047:Scottish Highlands
2855:10.1111/icad.12582
2700:Opsiphanes cassiae
2651:Spotted fritillary
2619:Speyeria aphrodite
2597:Indian red admiral
2514:Purple-shot copper
2500:Lycaena virgaureae
2455:Zerynthia polyxena
2437:Parnassius phoebus
2419:Pyrgus armoricanus
2410:Carcharodus alceae
2303:Cacyreus marshalli
2281:Scarce swallowtail
2276:Leptotes pirithous
2187:(now vagrant only)
2185:Cyaniris semiargus
2172:Black-veined white
2145:– rest of Scotland
2137:C. tullia polydama
2121:Coenonympha tullia
1897:– central Scotland
1886:E. epiphron mnemon
1587:– southern England
1376:
1366:
1356:
1346:
1336:
1326:
1316:
1306:
1296:
1286:
1246:
1213:P. arion eutyphron
1151:Lysandra bellargus
1138:– southern England
983:P. argus cretaceus
760:
750:
740:
730:
720:
710:
654:P. napi sabellicae
640:Green-veined white
470:
460:
457:Green-veined white
450:
426:Continental Europe
410:(formerly also in
377:
332:Scottish Highlands
182:Thymelicus lineola
97:
87:
27:This is a list of
3648:
3647:
3269:
3268:
3193:
3192:
2938:Hitchcock, James
2669:Woodland grayling
2527:Lycaena hippothoe
2518:Lycaena alciphron
2491:Rapala schistacea
2460:Tiger swallowtail
2401:Hylephila phyleus
2373:Parnassius apollo
2359:Danaus chrysippus
2322:Nymphalis antiopa
2318:Camberwell beauty
2290:Short-tailed blue
2267:Lampides boeticus
2215:Erebia alberganus
2143:C. tullia scotica
2049:– Isles of Scilly
1963:Hipparchia semele
1854:Lasiommata megera
1836:P. aegeria insula
1830:P. aegeria oblita
1821:P. aegeria tircis
1700:Polygonia c-album
1548:Limenitis camilla
1503:S. aglaja scotica
1465:Fabriciana adippe
1061:Aricia artaxerxes
833:Purple hairstreak
291:Ochlodes sylvanus
274:and the southern
212:Thymelicus acteon
129:Rockingham Forest
110:Chequered skipper
16:(Redirected from
3693:
3601:Dependencies and
3310:Sovereign states
3296:
3289:
3282:
3273:
3065:
3019:
3012:
3005:
2996:
2989:
2988:
2986:
2984:
2969:
2956:
2949:
2943:
2936:
2930:
2929:
2927:
2925:
2915:
2909:
2908:
2906:
2904:
2898:
2890:
2884:
2873:
2867:
2866:
2838:
2832:
2831:
2803:
2797:
2790:
2784:
2778:
2772:
2765:
2759:
2748:
2709:Caligo illioneus
2664:Lasiommata maera
2637:Argynnis pandora
2628:Fabriciana niobe
2624:Niobe fritillary
2591:Hypanartia lethe
2451:Southern festoon
2446:Zerynthia rumina
2350:Danaus plexippus
2341:Issoria lathonia
2331:Araschnia levana
2263:Long-tailed blue
2249:Pontia daplidice
1985:H. semele thyone
1973:H. semele semele
1774:Melitaea athalia
1770:Heath fritillary
1718:Marsh fritillary
1608:Vanessa atalanta
1497:S. aglaja aglaja
1411:B. selene selene
1251:Duke of Burgundy
1243:Duke of Burgundy
1129:Lysandra coridon
1013:P. argus masseyi
1007:Great Ormes Head
877:Black hairstreak
795:Brown hairstreak
773:Green hairstreak
717:Brown hairstreak
707:Green hairstreak
664:P. napi thomsoni
600:Pieris brassicae
566:Common brimstone
514:Northamptonshire
447:Common brimstone
342:Grizzled skipper
208:Lulworth skipper
94:Grizzled skipper
21:
3701:
3700:
3696:
3695:
3694:
3692:
3691:
3690:
3651:
3650:
3649:
3644:
3602:
3596:
3582:Northern Cyprus
3569:
3563:
3484:North Macedonia
3305:
3300:
3270:
3265:
3229:
3189:
3166:
3148:
3056:
3028:
3023:
2993:
2992:
2982:
2980:
2971:
2970:
2959:
2950:
2946:
2940:For Peat's Sake
2937:
2933:
2923:
2921:
2917:
2916:
2912:
2902:
2900:
2896:
2892:
2891:
2887:
2874:
2870:
2840:
2839:
2835:
2805:
2804:
2800:
2791:
2787:
2779:
2775:
2766:
2762:
2749:
2745:
2740:
2717:
2682:Chazara briseis
2673:Hipparchia fagi
2655:Melitaea didyma
2646:Speyeria cybele
2563:Junonia villida
2509:Lycaena tityrus
2464:Papilio glaucus
2442:Spanish festoon
2299:Geranium bronze
2258:Euchloe crameri
2176:Aporia crataegi
2154:
2131:C. tullia davus
2031:Maniola jurtina
1979:H. semele scota
1910:Erebia aethiops
1876:Erebia epiphron
1840:Isles of Scilly
1811:Pararge aegeria
1759:Channel Islands
1744:Melitaea cinxia
1518:Argynnis paphia
1487:Speyeria aglaja
1276:
1255:Hamearis lucina
1236:
1203:Phengaris arion
808:– south of the
777:Callophrys rubi
700:
510:Buckinghamshire
467:Male orange tip
440:
394:Papilio machaon
367:
77:
23:
22:
15:
12:
11:
5:
3699:
3697:
3689:
3688:
3683:
3678:
3673:
3668:
3663:
3653:
3652:
3646:
3645:
3643:
3642:
3637:
3632:
3627:
3622:
3617:
3612:
3606:
3604:
3603:other entities
3598:
3597:
3595:
3594:
3589:
3584:
3579:
3573:
3571:
3565:
3564:
3562:
3561:
3559:United Kingdom
3556:
3551:
3546:
3541:
3536:
3531:
3526:
3521:
3516:
3511:
3506:
3501:
3496:
3491:
3486:
3481:
3476:
3471:
3466:
3461:
3456:
3451:
3446:
3441:
3436:
3431:
3426:
3424:
3419:
3414:
3409:
3404:
3399:
3394:
3389:
3384:
3379:
3374:
3372:Czech Republic
3369:
3364:
3359:
3354:
3349:
3344:
3339:
3334:
3329:
3324:
3319:
3313:
3311:
3307:
3306:
3301:
3299:
3298:
3291:
3284:
3276:
3267:
3266:
3264:
3263:
3258:
3253:
3251:Birds of Wales
3248:
3243:
3237:
3235:
3231:
3230:
3228:
3227:
3222:
3217:
3212:
3207:
3201:
3199:
3195:
3194:
3191:
3190:
3188:
3187:
3182:
3176:
3174:
3168:
3167:
3165:
3164:
3158:
3156:
3150:
3149:
3147:
3146:
3141:
3136:
3131:
3126:
3121:
3116:
3111:
3106:
3101:
3096:
3091:
3086:
3081:
3075:
3073:
3062:
3058:
3057:
3055:
3054:
3049:
3044:
3039:
3033:
3030:
3029:
3024:
3022:
3021:
3014:
3007:
2999:
2991:
2990:
2978:UK Butterflies
2972:Eeles, Peter.
2957:
2944:
2931:
2910:
2885:
2868:
2833:
2814:(3): 159–172.
2798:
2785:
2773:
2760:
2742:
2741:
2739:
2736:
2735:
2734:
2729:
2724:
2716:
2713:
2712:
2711:
2702:
2693:
2687:False grayling
2684:
2675:
2666:
2657:
2648:
2639:
2630:
2621:
2612:
2603:
2601:Vanessa indica
2594:
2583:
2581:Colobura dirce
2574:
2572:Junonia oenone
2565:
2556:
2547:
2538:
2532:Turquoise blue
2529:
2520:
2511:
2502:
2493:
2484:
2475:
2473:Colias palaeno
2466:
2457:
2448:
2439:
2430:
2421:
2412:
2406:Mallow skipper
2403:
2385:
2384:
2375:
2366:
2352:
2343:
2334:
2324:
2315:
2306:
2296:
2287:
2278:
2269:
2260:
2251:
2242:
2233:
2218:
2217:
2208:
2198:
2195:Lycaena dispar
2188:
2178:
2169:
2153:
2150:
2149:
2148:
2147:
2146:
2140:
2134:
2125:
2124:
2112:
2111:
2110:
2109:
2103:
2094:
2093:
2081:
2080:
2079:
2078:
2072:
2071:
2059:
2058:
2057:
2056:
2050:
2044:
2035:
2034:
2022:
2021:
2020:
2019:
2013:
2012:
2000:
1999:
1998:
1997:
1988:
1982:
1976:
1967:
1966:
1954:
1953:
1952:
1951:
1945:
1944:
1932:
1931:
1930:
1929:
1923:
1914:
1913:
1901:
1900:
1899:
1898:
1892:
1880:
1879:
1867:
1866:
1865:
1864:
1858:
1857:
1845:
1844:
1843:
1842:
1833:
1827:
1815:
1814:
1794:
1793:
1792:
1791:
1778:
1777:
1765:
1764:
1763:
1762:
1748:
1747:
1735:
1734:
1733:
1732:
1726:
1725:
1713:
1712:
1711:
1710:
1704:
1703:
1691:
1690:
1689:
1688:
1678:
1677:
1665:
1664:
1663:
1662:
1656:
1655:
1652:Aglais urticae
1643:
1642:
1641:
1640:
1634:
1633:
1630:Vanessa cardui
1621:
1620:
1619:
1618:
1612:
1611:
1591:
1590:
1589:
1588:
1582:
1581:
1574:Purple emperor
1561:
1560:
1559:
1558:
1552:
1551:
1531:
1530:
1529:
1528:
1522:
1521:
1509:
1508:
1507:
1506:
1500:
1491:
1490:
1478:
1477:
1476:
1475:
1469:
1468:
1456:
1455:
1454:
1453:
1447:
1446:
1426:
1425:
1424:
1423:
1421:Inner Hebrides
1414:
1405:
1404:
1275:
1269:
1268:
1267:
1266:
1265:
1259:
1258:
1235:
1229:
1228:
1227:
1226:
1225:
1222:P. arion arion
1219:
1207:
1206:
1194:
1193:
1192:
1191:
1177:
1176:
1164:
1163:
1162:
1161:
1155:
1154:
1142:
1141:
1140:
1139:
1133:
1132:
1125:Chalkhill blue
1120:
1119:
1118:
1117:
1111:
1110:
1098:
1097:
1096:
1095:
1088:
1087:
1079:
1078:
1065:
1064:
1052:
1051:
1050:
1049:
1042:
1032:
1031:
1028:Aricia agestis
1019:
1018:
1017:
1016:
1010:
1000:
985:– formerly on
980:
973:P. argus argus
967:
966:
963:Plebejus argus
954:
953:
952:
951:
945:
944:
941:Cupido minimus
924:
923:
922:
921:
915:
914:
894:
893:
892:
891:
885:
884:
872:
871:
870:
869:
863:
862:
850:
849:
848:
847:
841:
840:
828:
827:
826:
825:
822:Worcestershire
803:
802:
799:Thecla betulae
790:
789:
788:
787:
781:
780:
747:Chalkhill blue
699:
693:
692:
691:
690:
689:
683:
682:
670:
669:
668:
667:
661:
648:
647:
635:
634:
633:
632:
626:
625:
613:
612:
611:
610:
604:
603:
583:
582:
581:
580:
574:
573:
561:
560:
559:
558:
548:
547:
544:Colias croceus
527:
526:
525:
524:
518:Worcestershire
491:
490:
439:
433:
432:
431:
430:
429:
415:
408:Norfolk Broads
398:
397:
366:
365:– swallowtails
360:
359:
358:
357:
356:
350:
349:
337:
336:
335:
334:
325:
324:
304:
303:
302:
301:
295:
294:
282:
281:
280:
279:
245:
244:
241:Hesperia comma
232:
231:
230:
229:
216:
215:
203:
202:
201:
200:
186:
185:
173:
172:
171:
170:
156:
155:
135:
134:
133:
132:
118:
117:
102:Heteropterinae
76:
70:
24:
14:
13:
10:
9:
6:
4:
3:
2:
3698:
3687:
3684:
3682:
3679:
3677:
3674:
3672:
3669:
3667:
3664:
3662:
3659:
3658:
3656:
3641:
3638:
3636:
3633:
3631:
3628:
3626:
3623:
3621:
3618:
3616:
3615:Faroe Islands
3613:
3611:
3608:
3607:
3605:
3599:
3593:
3590:
3588:
3587:South Ossetia
3585:
3583:
3580:
3578:
3575:
3574:
3572:
3566:
3560:
3557:
3555:
3552:
3550:
3547:
3545:
3542:
3540:
3537:
3535:
3532:
3530:
3527:
3525:
3522:
3520:
3517:
3515:
3512:
3510:
3507:
3505:
3502:
3500:
3497:
3495:
3492:
3490:
3487:
3485:
3482:
3480:
3477:
3475:
3472:
3470:
3467:
3465:
3462:
3460:
3457:
3455:
3452:
3450:
3447:
3445:
3444:Liechtenstein
3442:
3440:
3437:
3435:
3432:
3430:
3427:
3425:
3423:
3420:
3418:
3415:
3413:
3410:
3408:
3405:
3403:
3400:
3398:
3395:
3393:
3390:
3388:
3385:
3383:
3380:
3378:
3375:
3373:
3370:
3368:
3365:
3363:
3360:
3358:
3355:
3353:
3350:
3348:
3345:
3343:
3340:
3338:
3335:
3333:
3330:
3328:
3325:
3323:
3320:
3318:
3315:
3314:
3312:
3308:
3304:
3297:
3292:
3290:
3285:
3283:
3278:
3277:
3274:
3262:
3259:
3257:
3254:
3252:
3249:
3247:
3244:
3242:
3239:
3238:
3236:
3232:
3226:
3223:
3221:
3218:
3216:
3213:
3211:
3208:
3206:
3203:
3202:
3200:
3196:
3186:
3183:
3181:
3178:
3177:
3175:
3173:
3169:
3163:
3160:
3159:
3157:
3155:
3151:
3145:
3142:
3140:
3137:
3135:
3132:
3130:
3127:
3125:
3122:
3120:
3117:
3115:
3112:
3110:
3107:
3105:
3104:Conopid flies
3102:
3100:
3097:
3095:
3092:
3090:
3087:
3085:
3082:
3080:
3077:
3076:
3074:
3072:
3071:
3066:
3063:
3061:Invertebrates
3059:
3053:
3050:
3048:
3045:
3043:
3040:
3038:
3035:
3034:
3031:
3027:
3020:
3015:
3013:
3008:
3006:
3001:
3000:
2997:
2979:
2975:
2968:
2966:
2964:
2962:
2958:
2954:
2948:
2945:
2941:
2935:
2932:
2920:
2914:
2911:
2895:
2889:
2886:
2882:
2878:
2872:
2869:
2864:
2860:
2856:
2852:
2848:
2844:
2837:
2834:
2829:
2825:
2821:
2817:
2813:
2809:
2802:
2799:
2795:
2789:
2786:
2782:
2777:
2774:
2770:
2764:
2761:
2757:
2753:
2747:
2744:
2737:
2733:
2730:
2728:
2725:
2722:
2719:
2718:
2714:
2710:
2706:
2703:
2701:
2697:
2694:
2692:
2688:
2685:
2683:
2679:
2676:
2674:
2670:
2667:
2665:
2661:
2658:
2656:
2652:
2649:
2647:
2643:
2640:
2638:
2634:
2631:
2629:
2625:
2622:
2620:
2616:
2613:
2611:
2607:
2604:
2602:
2598:
2595:
2593:
2592:
2587:
2584:
2582:
2578:
2575:
2573:
2569:
2566:
2564:
2560:
2557:
2555:
2551:
2548:
2546:
2542:
2539:
2537:
2533:
2530:
2528:
2524:
2521:
2519:
2515:
2512:
2510:
2506:
2503:
2501:
2497:
2496:Scarce copper
2494:
2492:
2488:
2485:
2483:
2479:
2476:
2474:
2470:
2467:
2465:
2461:
2458:
2456:
2452:
2449:
2447:
2443:
2440:
2438:
2434:
2431:
2429:
2425:
2422:
2420:
2416:
2413:
2411:
2407:
2404:
2402:
2398:
2397:Fiery skipper
2395:
2394:
2393:
2390:
2389:
2383:
2379:
2376:
2374:
2370:
2367:
2364:
2360:
2356:
2353:
2351:
2347:
2344:
2342:
2338:
2335:
2332:
2328:
2325:
2323:
2319:
2316:
2314:
2310:
2307:
2304:
2300:
2297:
2295:
2291:
2288:
2286:
2282:
2279:
2277:
2273:
2270:
2268:
2264:
2261:
2259:
2255:
2252:
2250:
2246:
2243:
2241:
2237:
2234:
2232:
2228:
2225:
2224:
2223:
2222:
2216:
2212:
2209:
2206:
2202:
2199:
2196:
2192:
2189:
2186:
2182:
2181:Mazarine blue
2179:
2177:
2173:
2170:
2168:
2164:
2161:
2160:
2159:
2158:
2151:
2144:
2141:
2138:
2135:
2132:
2129:
2128:
2127:
2126:
2123:
2122:
2118:
2114:
2113:
2107:
2104:
2101:
2098:
2097:
2096:
2095:
2092:
2091:
2087:
2083:
2082:
2076:
2075:
2074:
2073:
2070:
2069:
2065:
2061:
2060:
2054:
2051:
2048:
2045:
2042:
2039:
2038:
2037:
2036:
2033:
2032:
2028:
2024:
2023:
2017:
2016:
2015:
2014:
2011:
2010:
2006:
2002:
2001:
1996:
1992:
1989:
1986:
1983:
1980:
1977:
1974:
1971:
1970:
1969:
1968:
1965:
1964:
1960:
1956:
1955:
1949:
1948:
1947:
1946:
1943:
1942:
1938:
1937:Marbled white
1934:
1933:
1927:
1924:
1921:
1918:
1917:
1916:
1915:
1912:
1911:
1907:
1903:
1902:
1896:
1893:
1891:
1887:
1884:
1883:
1882:
1881:
1878:
1877:
1873:
1869:
1868:
1862:
1861:
1860:
1859:
1856:
1855:
1851:
1847:
1846:
1841:
1837:
1834:
1831:
1828:
1826:
1822:
1819:
1818:
1817:
1816:
1813:
1812:
1808:
1807:Speckled wood
1804:
1803:
1802:
1801:
1800:
1790:
1786:
1785:re-introduced
1782:
1781:
1780:
1779:
1776:
1775:
1771:
1767:
1766:
1760:
1756:
1755:Isle of Wight
1752:
1751:
1750:
1749:
1746:
1745:
1741:
1737:
1736:
1730:
1729:
1728:
1727:
1724:
1723:
1719:
1715:
1714:
1708:
1707:
1706:
1705:
1702:
1701:
1697:
1693:
1692:
1686:
1685:Western Isles
1682:
1681:
1680:
1679:
1676:
1675:
1671:
1667:
1666:
1660:
1659:
1658:
1657:
1654:
1653:
1649:
1645:
1644:
1638:
1637:
1636:
1635:
1632:
1631:
1627:
1623:
1622:
1616:
1615:
1614:
1613:
1610:
1609:
1605:
1601:
1600:
1599:
1598:
1597:
1586:
1585:
1584:
1583:
1580:
1579:
1575:
1571:
1570:
1569:
1568:
1567:
1556:
1555:
1554:
1553:
1550:
1549:
1545:
1544:White admiral
1541:
1540:
1539:
1538:
1537:
1536:Limenitidinae
1526:
1525:
1524:
1523:
1520:
1519:
1515:
1511:
1510:
1504:
1501:
1498:
1495:
1494:
1493:
1492:
1489:
1488:
1484:
1480:
1479:
1473:
1472:
1471:
1470:
1467:
1466:
1462:
1458:
1457:
1451:
1450:
1449:
1448:
1445:
1444:
1440:
1436:
1432:
1428:
1427:
1422:
1418:
1415:
1412:
1409:
1408:
1407:
1406:
1403:
1402:
1398:
1394:
1390:
1386:
1385:
1384:
1383:
1382:
1374:
1370:
1364:
1360:
1354:
1350:
1344:
1343:Marbled white
1340:
1334:
1333:Speckled wood
1330:
1324:
1320:
1314:
1310:
1304:
1300:
1294:
1290:
1284:
1280:
1273:
1270:
1263:
1262:
1261:
1260:
1257:
1256:
1252:
1248:
1247:
1244:
1240:
1233:
1230:
1223:
1220:
1218:
1214:
1211:
1210:
1209:
1208:
1205:
1204:
1200:
1196:
1195:
1189:
1185:
1181:
1180:
1179:
1178:
1175:
1174:
1170:
1166:
1165:
1159:
1158:
1157:
1156:
1153:
1152:
1148:
1144:
1143:
1137:
1136:
1135:
1134:
1131:
1130:
1126:
1122:
1121:
1115:
1114:
1113:
1112:
1109:
1108:
1104:
1100:
1099:
1093:
1090:
1089:
1085:
1081:
1080:
1076:
1072:
1069:
1068:
1067:
1066:
1063:
1062:
1058:
1054:
1053:
1047:
1043:
1040:
1036:
1035:
1034:
1033:
1030:
1029:
1025:
1021:
1020:
1014:
1011:
1009:(north Wales)
1008:
1004:
1001:
999:
998:Portland Bill
995:
992:
988:
984:
981:
978:
974:
971:
970:
969:
968:
965:
964:
960:
956:
955:
949:
948:
947:
946:
943:
942:
938:
934:
933:
932:
931:
930:
929:Polyommatinae
919:
918:
917:
916:
913:
912:
908:
904:
903:
902:
901:
900:
889:
888:
887:
886:
883:
882:
878:
874:
873:
867:
866:
865:
864:
861:
860:
856:
852:
851:
845:
844:
843:
842:
839:
838:
834:
830:
829:
823:
819:
815:
811:
807:
806:
805:
804:
801:
800:
796:
792:
791:
785:
784:
783:
782:
779:
778:
774:
770:
769:
768:
767:
766:
758:
754:
748:
744:
738:
734:
728:
724:
718:
714:
708:
704:
697:
694:
687:
686:
685:
684:
681:
680:
676:
672:
671:
665:
662:
659:
655:
652:
651:
650:
649:
646:
645:
641:
637:
636:
630:
629:
628:
627:
624:
623:
619:
615:
614:
608:
607:
606:
605:
602:
601:
597:
593:
592:
591:
590:
589:
578:
577:
576:
575:
572:
571:
567:
563:
562:
557:in some years
556:
552:
551:
550:
549:
546:
545:
541:
537:
536:
535:
534:
533:
523:
522:Herefordshire
519:
515:
511:
507:
503:
499:
495:
494:
493:
492:
489:
488:
484:
480:
479:
478:
477:
476:
475:Dismorphiinae
468:
464:
458:
454:
448:
444:
437:
434:
427:
423:
419:
416:
413:
409:
405:
402:
401:
400:
399:
396:
395:
391:
387:
386:
385:
384:
383:
375:
371:
364:
361:
354:
353:
352:
351:
348:
347:
346:Pyrgus malvae
343:
339:
338:
333:
329:
328:
327:
326:
323:
322:
318:
317:Dingy skipper
314:
313:
312:
311:
310:
299:
298:
297:
296:
293:
292:
288:
287:Large skipper
284:
283:
277:
273:
269:
265:
261:
257:
253:
249:
248:
247:
246:
243:
242:
238:
234:
233:
228:
224:
220:
219:
218:
217:
214:
213:
209:
205:
204:
198:
194:
190:
189:
188:
187:
184:
183:
179:
178:Essex skipper
175:
174:
168:
164:
161:– throughout
160:
159:
158:
157:
154:
153:
149:
148:Small skipper
145:
144:
143:
142:
141:
130:
126:
122:
121:
120:
119:
116:
115:
111:
107:
106:
105:
104:
103:
95:
91:
85:
84:Dingy skipper
81:
74:
71:
69:
67:
63:
61:
57:
53:
48:
46:
42:
38:
34:
33:Great Britain
30:
19:
3592:Transnistria
3171:
3153:
3098:
3068:
2981:. Retrieved
2977:
2952:
2947:
2939:
2934:
2922:. Retrieved
2913:
2901:. Retrieved
2888:
2876:
2871:
2846:
2836:
2811:
2807:
2801:
2793:
2788:
2776:
2768:
2763:
2755:
2751:
2746:
2708:
2699:
2690:
2681:
2672:
2663:
2654:
2645:
2636:
2627:
2618:
2609:
2600:
2589:
2580:
2571:
2562:
2553:
2544:
2535:
2526:
2517:
2508:
2505:Sooty copper
2499:
2490:
2481:
2472:
2463:
2454:
2445:
2436:
2433:Small Apollo
2427:
2418:
2409:
2400:
2391:
2387:
2386:
2381:
2372:
2358:
2349:
2340:
2330:
2321:
2312:
2302:
2293:
2284:
2275:
2266:
2257:
2248:
2239:
2231:Colias hyale
2230:
2220:
2219:
2214:
2204:
2194:
2191:Large copper
2184:
2175:
2167:Erebia ligea
2166:
2156:
2155:
2142:
2136:
2130:
2120:
2115:
2105:
2099:
2089:
2084:
2067:
2062:
2052:
2046:
2040:
2030:
2027:Meadow brown
2025:
2008:
2003:
1990:
1984:
1978:
1972:
1962:
1957:
1940:
1935:
1925:
1919:
1909:
1906:Scotch argus
1904:
1894:
1885:
1875:
1870:
1853:
1848:
1835:
1829:
1820:
1810:
1805:
1796:
1795:
1773:
1768:
1743:
1738:
1721:
1716:
1699:
1694:
1673:
1668:
1661:– throughout
1651:
1646:
1629:
1626:Painted lady
1624:
1607:
1602:
1593:
1592:
1578:Apatura iris
1577:
1572:
1563:
1562:
1547:
1542:
1533:
1532:
1517:
1512:
1502:
1496:
1486:
1481:
1464:
1459:
1442:
1438:
1434:
1429:
1416:
1410:
1400:
1396:
1392:
1387:
1381:Heliconiinae
1378:
1377:
1363:Meadow brown
1254:
1249:
1234:– metalmarks
1221:
1212:
1202:
1197:
1184:Solway Firth
1172:
1167:
1150:
1145:
1128:
1123:
1116:– throughout
1106:
1101:
1091:
1083:
1070:
1060:
1055:
1045:
1027:
1022:
1012:
1002:
982:
972:
962:
957:
940:
935:
926:
925:
910:
907:Small copper
905:
896:
895:
880:
875:
858:
853:
836:
831:
798:
793:
776:
771:
762:
761:
678:
673:
663:
657:
653:
643:
638:
621:
616:
599:
594:
585:
584:
569:
564:
543:
538:
529:
528:
486:
481:
472:
471:
417:
403:
393:
388:
382:Papilioninae
379:
378:
363:Papilionidae
345:
340:
320:
315:
306:
305:
290:
285:
240:
235:
211:
206:
193:West Country
181:
176:
151:
146:
137:
136:
113:
108:
99:
98:
64:
59:
49:
44:
40:
36:
26:
3630:Isle of Man
3570:recognition
3544:Switzerland
3479:Netherlands
3198:Vertebrates
3172:Crustaceans
3134:Shield bugs
3099:Butterflies
2610:Boloria dia
2554:Dryas julia
2487:Slate flash
2424:Shy saliana
2355:Plain tiger
2163:Arran brown
2117:Large heath
2086:Small heath
1825:Isle of Man
1604:Red admiral
1596:Nymphalinae
1373:Small heath
1293:Red admiral
1272:Nymphalidae
1182:– north to
1147:Adonis blue
1103:Common blue
1024:Brown argus
977:East Anglia
820:reserve in
727:Brown argus
618:Small white
596:Large white
555:River Clyde
390:Swallowtail
140:Hesperiinae
73:Hesperiidae
50:A study by
29:butterflies
3655:Categories
3514:San Marino
3474:Montenegro
3454:Luxembourg
3434:Kazakhstan
3337:Azerbaijan
3205:Amphibians
3114:Hoverflies
2955:65: 15–16.
2738:References
2660:Large wall
2568:Blue pansy
2245:Bath white
2108:– Hebrides
2005:Gatekeeper
1928:– Scotland
1797:Subfamily
1594:Subfamily
1566:Apaturinae
1564:Subfamily
1534:Subfamily
1505:– Scotland
1443:euphrosyne
1379:Subfamily
1353:Gatekeeper
1232:Riodinidae
1199:Large blue
1188:River Tyne
1169:Holly blue
1094:– Scotland
1084:A. agestis
1075:Lancashire
1039:River Tees
937:Small blue
927:Subfamily
897:Subfamily
763:Subfamily
757:Holly blue
696:Lycaenidae
675:Orange tip
586:Subfamily
532:Coliadinae
530:Subfamily
483:Wood white
473:Subfamily
380:Subfamily
307:Subfamily
138:Subfamily
100:Subfamily
75:– skippers
3620:Gibraltar
3449:Lithuania
2863:249088681
2478:Cleopatra
1922:– Cumbria
1799:Satyrinae
1674:Aglais io
1439:Clossiana
1397:Clossiana
991:limestone
765:Theclinae
500:and west
276:Chilterns
272:Wiltshire
264:Hampshire
131:, England
39:by Emmet
3640:Svalbard
3625:Guernsey
3529:Slovenia
3524:Slovakia
3499:Portugal
3357:Bulgaria
3234:See also
3225:Reptiles
3185:Woodlice
3154:Molluscs
3119:Mayflies
3042:Scotland
2983:13 March
2828:86056452
2715:See also
2633:Cardinal
2221:Vagrants
1995:Hebrides
1959:Grayling
994:downland
658:thomsoni
588:Pierinae
502:Somerset
436:Pieridae
412:The Fens
309:Pyrginae
270:, south
266:, north
223:Weymouth
125:Scotland
3554:Ukraine
3504:Romania
3464:Moldova
3422:Ireland
3417:Iceland
3412:Hungary
3402:Germany
3397:Georgia
3387:Finland
3382:Estonia
3377:Denmark
3362:Croatia
3347:Belgium
3342:Belarus
3332:Austria
3327:Armenia
3322:Andorra
3317:Albania
3220:Mammals
3094:Beetles
3070:Insects
3037:England
2924:15 June
2903:15 June
2783:, 2004.
2388:Exotics
2363:Odonata
2346:Monarch
2157:Extinct
2064:Ringlet
1987:– Wales
1890:Cumbria
1670:Peacock
1435:Boloria
1393:Boloria
1313:Peacock
1217:extinct
899:Lycaena
422:migrant
420:– rare
254:, east
227:Swanage
167:England
3635:Jersey
3577:Kosovo
3549:Turkey
3539:Sweden
3519:Serbia
3509:Russia
3494:Poland
3489:Norway
3469:Monaco
3439:Latvia
3407:Greece
3392:France
3367:Cyprus
2861:
2826:
2752:et al.
2678:Hermit
2369:Apollo
1401:selene
810:Humber
516:; and
506:Surrey
268:Dorset
260:Surrey
256:Sussex
197:Humber
41:et al.
3610:Åland
3534:Spain
3459:Malta
3429:Italy
3210:Birds
3180:Crabs
3144:Wasps
3124:Moths
3052:Wales
2897:(PDF)
2859:S2CID
2824:S2CID
2577:Zebra
2550:Julia
1789:Essex
1696:Comma
1323:Comma
1077:north
987:chalk
814:Weald
498:Devon
163:Wales
3215:Fish
3089:Bees
3079:Ants
2985:2014
2926:2019
2905:2019
2781:NERC
1850:Wall
1757:and
1186:and
989:and
520:and
512:and
252:Kent
225:and
165:and
52:NERC
43:and
2851:doi
2816:doi
2327:Map
1787:to
31:of
3657::
2976:.
2960:^
2879:.
2857:.
2849:.
2845:.
2822:.
2810:.
2707:–
2698:–
2689:–
2680:–
2671:–
2662:–
2653:–
2644:–
2626:–
2617:–
2608:–
2599:–
2588:–
2579:–
2570:–
2561:–
2552:–
2543:–
2534:–
2525:–
2516:–
2507:–
2498:–
2489:–
2480:–
2471:–
2462:–
2453:–
2444:–
2435:–
2426:–
2417:–
2408:–
2399:–
2380:–
2371:–
2357:–
2348:–
2339:–
2329:–
2320:–
2311:–
2301:–
2292:–
2283:–
2274:–
2265:–
2256:–
2247:–
2238:–
2229:–
2213:–
2203:–
2193:–
2183:–
2174:–
2165:–
2119:–
2088:–
2066:–
2029:–
2007:–
1993:–
1961:–
1939:–
1908:–
1888:–
1874:–
1852:–
1838:–
1809:–
1772:–
1742:–
1720:–
1698:–
1672:–
1650:–
1628:–
1606:–
1576:–
1546:–
1516:–
1485:–
1463:–
1441:)
1433:–
1399:)
1391:–
1253:–
1201:–
1171:–
1149:–
1127:–
1105:–
1059:–
1026:–
1005:–
961:–
939:–
909:–
879:–
857:–
835:–
797:–
775:–
677:–
642:–
620:–
598:–
568:–
542:–
508:;
504:;
496:–
485:–
392:–
344:–
319:–
289:–
262:,
258:,
239:–
210:–
180:–
150:–
112:–
62:.
3295:e
3288:t
3281:v
3018:e
3011:t
3004:v
2987:.
2928:.
2907:.
2865:.
2853::
2830:.
2818::
2812:4
2365:)
1437:(
1395:(
824:.
660:)
414:)
20:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
