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Palaeontinidae

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805: 475:, exhibit the most primitive wing forms in the family. The forewing was elliptical with the "nodal line" (the area where the wing bends during flight, also known as the "transverse flexion line") more or less dissecting through the center of the wing. The hindwing was short and broad. The bases of the forewings overlapped that of the hindwings like in modern butterflies. Taken together with their large bodies, these characteristics indicate that they were fast but moderately versatile fliers. 1816: 124: 565: 547: 608: 624: 529: 2647: 592: 102: 1830: 773:(2009), however, notes that palaeontinoids more closely resemble prosbolids in agreement with earlier studies by Wootton (1971), Shcherbakov (1984), and Shcherbakov and Popov (2002). They conclude that palaeontinoids descended directly from the family Prosbolidae rather than from tettigarctids. Modern cicadas therefore, did not descend directly from Palaeontinidae. 2659: 1867: 705:
supported Lameere's conclusion, noting that the wings of palaeontinid fossils lacked the characteristic scales of lepidopterans but instead had tubercules, pits, and cross-ridges like those found in modern cicadas. He also cited characteristics of wing venation that distinctly differs from that of
490:
had triangular forewings with the flexion line closer to the base. They had smaller and narrower hindwings that did not overlap with the forewing. These indicate that they were highly versatile fliers, able to fly with a wide range of speeds and agility like modern
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based on the geographic distribution of both groups, however other authors have argued that this association is likely to be spurious, given that that paleontinids also occur in areas with no ginkgophytes. Some authors have suggested that the decline of
468:
Palaeontinids, like modern cicadas, possess four membranous wings supported by veins. The length and width ratio of the wings can vary within the same species, sometimes as a result of fossil preservation. Early Jurassic palaeontinids, like
796:) is ancestral to palaeontinids. Both are distinct from the only other member of the superfamily, the more primitive and specialized family Mesogereonidae (Upper Triassic of Australia and South Africa). 735:
The name Cicadomorphidae was once proposed as a replacement for the name Palaeontinidae in 1956 by the Australian entomologist J.W. Evans. This was because of Handlirsch's earlier insistence that the
746:. However, Evans later conceded that retaining the name Palaeontinidae was preferable as the drawings Handlirsch based his conclusions on were from badly preserved specimens. 1545:- Badaowan Formation, Jiulongshan Formation, Daohugou, Karabastau Formation, Sagul Formation, Cheremkhovskaya Formation, Early to Late Jurassic, Central and Eastern Asia 2784: 2758: 2575: 2886: 2083:"A revision of Palaeontinidae (Insecta: Hemiptera: Cicadomorpha) from the Jurassic of China with descriptions of new taxa and new combinations" 2871: 2876: 101: 754:
Riek (1976) originally considered Palaeontinoidea to be the descendants of the family Cicadoprosbolidae (currently known as the family
2332: 1276: 1489:, Haifanggou Formation, Cheremkhovskaya Formation, Early to Middle Jurassic, Central and North Asia; Middle Jurassic, Central Asia 462: 1983:"Two new genera of fossil palaeontinids from the Middle Jurassic in Daohugou, Inner Mongolia, China (Hemiptera, Palaeontinidae)" 2375: 1946:"A new genus and species of Palaeontinidae (Insecta: Hemiptera: Cicadomorpha) from the Lower Cretaceous of southern England" 123: 2881: 2866: 1455: 461:. The patterns on their wings protected them as they perched on branches and fed on sap. They may also have served as 347: 2576:"†Palaeontinidae: After Grimaldi & Engel, 2005, Wang, Zhang & Fang, 2006, and Wang, Zhang & Szwedo, 2009" 2223:"Jurassic Palaeontinidae from China and the Higher Systematics of Palaeontinoidea (Insecta: Hemiptera: Cicadomorpha)" 2851: 804: 2861: 2856: 2663: 2272:(suppl. – Fossil Insects). Institute of Systematics and Evolution of Animals, Polish Academy of Sciences: 89–99. 110: 1337: 2598: 1770: 438:(flowering plants) during the Cretaceous could have been a factor in their extinction. Numerous newly evolved 1899: 571: 553: 1843: 1784: 961: 614: 2039:"New species of Palaeontinidae (Insecta: Cicadomorpha) from the Lower Cretaceous Crato Formation of Brazil" 630: 402:
had forewings that reached the length of 57 to 71 mm (2.2 to 2.8 in). They possessed an inflated
2675: 1087: 535: 369: 338: 2542:"Insectos del Cretácico inferior de Santa Maria de Meià (Lleida): Colección Lluís Maria Vidal i Carreras" 1848: 1698: 1333: 1204: 1099: 2810: 2599:"Contributions to Fossil Entomology: XI Fossil Insects from the Lower Purbecks, Durdlestone Bay, Dorset" 1797: 1680: 1510: 1482: 1329: 1208: 655:(butterflies and moths). Palaeontinids were then only known mostly from poorly preserved specimens like 1496: 598: 2828: 1673: 1600: 1103: 2745: 2462: 2176:"Permian Faunas of Homoptera (Hemiptera) in Relation to Phytogeography and the Permo-Triassic Crisis" 2038: 1821: 1750: 1524: 1486: 1375: 1308: 1095: 1059: 980: 702: 396:). They had small heads and broad wings. They superficially resemble moths. Large palaeontinids like 2258: 1637: 1609: 1465: 1361: 1182: 1141: 718: 507: 499:. They also possessed changes to the leading edge of their forewings, suggesting an overall gain in 471: 2713: 2651: 1586: 1259: 1113: 1041: 398: 1661: 1646: 1433: 1402: 1250: 1218: 873: 714: 690:
challenged this conclusion, claiming palaeontinids were more closely related to the extant family
486: 2618: 2541: 1708: 1572: 1538: 1447: 1419: 1393: 1294: 1196: 1127: 1033: 759: 657: 273: 246: 118: 2815: 1722: 1690: 1623: 1561: 1347: 1268: 1236: 1169: 1069: 1051: 1011: 1552: 1474: 2797: 2732: 2488: 2421: 2273: 2149: 2049: 2005: 1956: 1927: 1155: 836: 695: 663: 480: 407: 362: 2802: 2789: 2507: 2610: 2508:"A New Palaeontinid Species from the Lower Cretaceous of Brazil (Homoptera: Palaeontinidae)" 2478: 2470: 2413: 2347: 2234: 2141: 2097: 1997: 1284: 1226: 976: 898: 860: 816: 645: 317: 305: 226: 679:(ghost moths) instead. He said "There is little doubt that it belongs to the Hepialidae." 1077: 965: 824: 710: 687: 641: 208: 185: 2466: 2417: 1945: 1656:) Sulyukta Formation, Early Jurassic, Central Asia; Daohugou Middle Jurassic, North Asia 564: 546: 2483: 2450: 1835: 968: 957: 808: 781: 755: 672: 607: 359: 2614: 675:
supported the lepidopteran conclusion, though he believed they belonged to the family
623: 2845: 2622: 2299: 2239: 2222: 1921: 1023: 984: 913: 882: 852: 777: 439: 313: 59: 528: 2737: 2126: 1982: 1759: 1440:- Sagul Formation, Cheremkhovskaya Formation Early Jurassic, Central and North Asia 1321: 941: 789: 736: 725: 500: 366: 309: 195: 2145: 2082: 2771: 2707: 933: 758:), insects believed to be transitional between the ancestral cicada-like family 668: 652: 649: 591: 496: 426: 381: 351: 34: 2698: 2646: 2451:"An extraordinary palaeontinid from the Triassic of Korea and its significance" 2001: 1811: 1280: 949: 945: 937: 722: 676: 435: 431: 422: 235: 79: 44: 2425: 2333:"The Evolution of the Scorpion-flies and their Derivatives (Order Mecoptera)" 2277: 2175: 2153: 2053: 2009: 1960: 1923:
Lepidoptera Exotica; or, Descriptions and Illustrations of Exotic Lepidoptera
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in 1908. Like Butler, Handlirsch insisted that palaeontinids were members of
380:
ever recovered, having mistakenly identified it as a butterfly of the family
374:
Lepidoptera Exotica; or, Descriptions and Illustrations of Exotic Lepidoptera
1736: 785: 763: 743: 729: 699: 691: 415: 377: 175: 155: 135: 84: 2724: 2492: 2449:
Kye Soo Nam; Ying Wang; Dong Ren; Jong Heon Kim & Jacek Szwedo (2017).
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The trend of forewing elongation is most evident in members of the family
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Stuttgarter Beiträge zur Naturkunde, Serie B (Geologie und Paläontologie)
902: 443: 343: 74: 69: 54: 49: 39: 1829: 2763: 2127:"New data on Cretaceous Palaeontinidae (Insecta: Hemiptera) from China" 2081:
Bo Wang; Haichun Zhang; Yan Fang; Dejin Wang & Yutao Zhang (2007).
929: 909: 683: 458: 355: 325: 302: 89: 64: 2776: 2474: 709:
Palaeontinidae are currently classified under the extinct superfamily
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showing the approximate locations of some palaeontinid fossil sites.
447: 165: 145: 2669: 2101: 1340:, Early to Middle Jurassic, North Asia; Late Jurassic, Central Asia 365:
Edward Charlesworth. It was first described in 1873 by the English
2823: 2125:
Bo Wang; Haichun Zhang; Yan Fang; Dejin Wang; Shengzhu Ji (2008).
1426:- La Pedrera de Rúbies Formation Early Cretaceous, Western Europe 1000: 988: 953: 921: 901:(~115-113 Mya). They achieved their greatest diversity during the 864: 840: 812: 803: 793: 411: 403: 393: 321: 2750: 1531:- Cheremkhovskaya Formation, Early to Middle Jurassic, North Asia 1368:- La Pedrera de Rúbies Formation Early Cretaceous, Western Europe 492: 465:. The color patterns can vary slightly within the same species. 451: 2673: 2259:"Reconstructing insect flight performance from fossil evidence" 454:) may have also contributed significantly to their extinction. 410:(piercing and sucking mouthpart), indicating that they fed on 2307:
Proceedings: XIII International Congress of Entomology, Moscow
2521:. Kitakyushu Museum and Institute of Natural History: 99–104 1944:
Bo Wang; Haichun Zhang & Edmund A. Jarzembowski (2008).
1630:- Sulyukta Formation, Early to Middle Jurassic, Central Asia 1616:) Daohugou, Haifanggou Formation, Middle Jurassic, East Asia 1503:- Crato Formation, Early Cretaceous, Eastern South America 1134:- Crato Formation, Early Cretaceous, Eastern South America 1120:- Crato Formation, Early Cretaceous, Eastern South America 510:, an early offshoot and close relatives of palaeontinids. 2603:
The Quarterly Journal of the Geological Society of London
1743:- Early Cretaceous, Western Europe (too poorly preserved) 1593:- Sagul Formation, Early to Middle Jurassic, Central Asia 1517:- Sagul Formation, Early to Middle Jurassic, Central Asia 478:
In contrast, later palaeontinids like the Upper Jurassic
893:. 208.5 Mya) and the youngest members are from the late 392:
Palaeontinids had large bodies covered with bristles (
1907:
Proceedings of the Linnean Society of New South Wales
2515:
Bulletin of the Kitakyushu Museum of Natural History
1793: 1780: 1766: 1746: 1732: 1718: 1704: 1686: 1633: 1619: 1596: 1582: 1579:- Solnhofen Formation, Late Jurassic, Central Europe 1568: 1565:- Solnhofen Formation, Late Jurassic, Central Europe 1548: 1534: 1520: 1506: 1492: 1461: 1443: 1429: 1415: 1389: 1371: 1357: 1343: 1304: 1301:- Solnhofen Formation, Late Jurassic, Central Europe 1290: 1246: 1232: 1214: 1192: 1165: 1162:- Solnhofen Formation, Late Jurassic, Central Europe 1151: 1137: 1123: 1109: 1083: 1065: 1047: 1029: 1007: 667:. He claimed they were related to the extant family 281: 242: 2682: 2233:(Part 1). The Palaeontological Association: 53–64. 2221:Bo Wang; Haichun Zhang & Jacek Szwedo (2009). 776:Within Palaeontinoidea, the family Dunstaniidae ( 2048:(357). Staatliches Museum für Naturkunde: 1–10. 1287:Early Cretaceous, Western Europe & East Asia 2326: 2324: 1729:- Yixian Formation, Early Cretaceous, East Asia 1354:- Yixian Formation, Early Cretaceous, East Asia 27:Temporal range: Late Triassic-Early Cretaceous 2609:(1–2). Geological Society of London: 395–396. 2340:Annals of the Entomological Society of America 1939: 1937: 1715:- Weald Clay, Early Cretaceous, Western Europe 2346:(1). Entomological Society of America: 1–45. 8: 2549:Treballs del Museu de Geología de Barcelona 2216: 2214: 2212: 2210: 2208: 2206: 2204: 2670: 2444: 2442: 2370: 2368: 2252: 2250: 2120: 2118: 2076: 2074: 2072: 2070: 2037:Federica Menon & Sam W. Heads (2005). 1893: 1891: 1889: 1887: 1885: 100: 20: 2482: 2300:"The Evolution of Cicadoidea (Homoptera)" 2238: 2555:. Museo de Geología de Barcelona: 91–116 1479:Becker-Migdisova, 1950; Shcherbakov 1985 871:The oldest known member of the group is 376:. Butler claimed that it was the oldest 2032: 2030: 2028: 2026: 1859: 1080:Early Cretaceous, Eastern South America 671:(slug moths). The English entomologist 1386:- Daohugou, Middle Jurassic, East Asia 1211:, Daohugou Middle Jurassic, East Asia 1148:- Daohugou, Middle Jurassic, East Asia 457:Most species of palaeontinids exhibit 336:The first palaeontinid discovered was 320:. The family contains around 30 to 40 1412:) Daohugou Middle Jurassic, East Asia 1243:- Daohugou Middle Jurassic, East Asia 1189:) Daohugou Middle Jurassic, East Asia 981:Haifanggou (or Jiulongshan) Formation 964:for palaeontinid fossils include the 908:Palaeontinid fossils are abundant in 7: 2829:941187CC-FFE7-7A1B-1A40-FE3FB9A3FAB0 1920:Arthur Gardiner Butler (1869–1874). 1695:Becker-Migdisova & Wootton, 1965 800:Distribution and geologic time range 640:The family was first erected by the 421:Some authors have proposed that the 2418:10.1146/annurev.en.08.010163.000453 1713:Wang, Zhang & Jarzembowski 2008 1668:Daohugou Middle Jurassic, East Asia 14: 2615:10.1144/gsl.jgs.1854.010.01-02.43 1981:Wang Ying & Ren Dong (2007). 1870:. The EDNA Fossil Insect Database 1003:classified under Palaeontinidae: 2657: 2645: 2540:Xavier Martínez-Delclòs (1990). 2399:"The Phylogeny of the Homoptera" 2240:10.1111/j.1475-4983.2008.00826.x 1828: 1814: 1757:- Early Jurassic, Central Asia ( 1264:Whalley & Jarzembowski, 1985 916:. Fossils have been recorded in 622: 606: 590: 563: 545: 527: 463:secondary sexual characteristics 122: 1996:(1390). Magnolia Press: 41–49. 1955:(1751). Magnolia Press: 65–68. 1458:Middle Jurassic, Western Europe 1098:, Late Jurassic, Central Asia; 877:from South Korea dating to the 2887:Taxa named by Anton Handlirsch 1398:Wang, Zhang & Szwedo, 2009 1277:La Pedrera de Rúbies Formation 1062:, Early Jurassic, Central Asia 414:fluids like some other modern 1: 2574:Mikko Haaramo (May 5, 2009). 2146:10.1016/j.cretres.2008.01.007 1223:Nam, Wang, & Szwedo, 2017 1044:Late Jurassic, Central Europe 999:The following is the list of 2096:. Wiley InterScience: 1–18. 1026:, Middle Jurassic, East Asia 721:. They are classified under 521:Early Jurassic palaeontinids 388:Description and paleobiology 2872:Prehistoric insect families 2578:. Mikko's Phylogeny Archive 2406:Annual Review of Entomology 2266:Acta Zoologica Cracoviensia 1456:Taynton Limestone Formation 1229:Upper Triassic, South Korea 584:Late Jurassic palaeontinids 348:Taynton Limestone Formation 342:. It consisted of a single 265: 111:Synapocossus sciacchitanoae 16:Extinct family of true bugs 2903: 2877:Rhaetian first appearances 2174:D. E. Shcherbakov (2000). 1777:Middle Jurassic, East Asia 1683:, Australia, Late Jurassic 1106:, Late Jurassic North Asia 2412:. Annual Reviews: 77–94. 2257:Robin J. Wootton (2002). 2002:10.11646/zootaxa.1390.1.5 1926:. E. W. Jansen. pp.  1900:"The Panorpoid Complex 3" 1775:- Hong, 1986; Zhang, 1997 1681:Talbragar Fossil Fish Bed 1338:Cheremkhovskaya Formation 279: 272: 262: 257: 241: 234: 119:Scientific classification 117: 108: 99: 23: 2140:(4). Elsevier: 551–560. 1771:Palaeontinopsis sinensis 1701:, Jurassic, Central Asia 713:along with the families 312:. They existed from the 2506:Kyoichiro Ueda (1996). 2378:. Paleobiology Database 2183:Paleontological Journal 1868:"Family Palaeontinidae" 1844:Prehistoric Lepidoptera 1380:Wang & Zhang, 2007 1016:Wang & Zhang, 2007 694:(cicadas). The English- 572:Prolystra lithographica 554:Palaeocossus jurassicus 425:of palaeontinids to be 350:(Stonesfield Slate) of 2597:J.O. Westwood (1854). 2331:R.J. Tillyard (1935). 1898:R.J. Tillyard (1919). 1785:Fletcheriana jurassica 1642:Becker-Migdisova, 1949 1628:Becker-Migdisova, 1949 1591:Becker-Migdisova, 1949 1543:Becker-Migdisova, 1949 1515:Becker-Migdisova, 1949 1313:Wang & Zhang 2008 1056:Becker-Migdisova, 1962 868: 811:representation of the 762:and the modern family 615:Limacodites mesozoicus 370:Arthur Gardiner Butler 2811:Paleobiology Database 2189:(Suppl. 3): S251–S267 1614:Wang & Ren, 2007a 1577:Handlirsch, 1906–1908 1366:Gomez-Pallerola, 1984 1299:Handlirsch, 1906–1908 1255:Gomez-Pallerola, 1984 1241:Wang & Ren 2007ab 1209:Jiulongshan Formation 1038:Handlirsch, 1906–1908 807: 631:Beloptesis oppenheimi 484:and Early Cretaceous 324:and around a hundred 2666:at Wikimedia Commons 1822:Palaeontology portal 1096:Karabastau Formation 740:Palaeontina oolitica 703:Robert John Tillyard 536:Palaeontina oolitica 339:Palaeontina oolitica 297:, commonly known as 2467:2017NatSR...740691N 2397:J.W. Evans (1963). 2352:10.1093/aesa/28.1.1 2298:R. Wootton (1971). 2134:Cretaceous Research 1849:Prehistoric insects 1699:Kushmurun Formation 1334:Kushmurun Formation 1205:Dashankou Formation 1100:Glushkovo Formation 1042:Solnhofen Formation 442:animals (feathered 346:collected from the 2882:Aptian extinctions 2867:Cretaceous insects 2455:Scientific Reports 2090:Geological Journal 1798:Fletcheriana magna 1511:Phragmatoecicossus 1483:Sulyukta Formation 1330:Badaowan Formation 1174:Wang et al. 2006c 1132:Martins-Neto, 1998 869: 742:may not have been 459:cryptic coloration 2852:Extinct Hemiptera 2839: 2838: 2798:Open Tree of Life 2676:Taxon identifiers 2662:Media related to 2475:10.1038/srep40691 1803: 1790: 1776: 1756: 1742: 1728: 1714: 1696: 1679: 1667: 1652: 1643: 1629: 1615: 1606: 1592: 1578: 1558: 1544: 1530: 1516: 1502: 1497:Parawonnacottella 1480: 1471: 1453: 1439: 1425: 1408: 1399: 1385: 1384:Wang et al. 2007a 1367: 1353: 1327: 1318: 1300: 1274: 1265: 1256: 1242: 1224: 1202: 1201:Wang et al. 2006b 1188: 1187:Wang et al. 2007c 1179: 1178:Wang et al. 2006c 1161: 1147: 1146:Wang et al. 2006b 1133: 1119: 1118:Menon et al. 2005 1093: 1075: 1074:Menon et al. 2005 1057: 1039: 1021: 1020:Wang et al. 2007a 837:Serra del Montsec 698:entomologist and 599:Eocicada lameerei 363:natural historian 292: 291: 287: 230: 2894: 2862:Jurassic insects 2857:Triassic insects 2832: 2831: 2819: 2818: 2806: 2805: 2793: 2792: 2780: 2779: 2767: 2766: 2754: 2753: 2741: 2740: 2728: 2727: 2718: 2717: 2716: 2703: 2702: 2701: 2671: 2661: 2650:Data related to 2649: 2634: 2633: 2631: 2629: 2594: 2588: 2587: 2585: 2583: 2571: 2565: 2564: 2562: 2560: 2546: 2537: 2531: 2530: 2528: 2526: 2512: 2503: 2497: 2496: 2486: 2446: 2437: 2436: 2434: 2432: 2403: 2394: 2388: 2387: 2385: 2383: 2376:"Palaeontinidae" 2372: 2363: 2362: 2360: 2358: 2337: 2328: 2319: 2318: 2316: 2314: 2304: 2295: 2289: 2288: 2286: 2284: 2263: 2254: 2245: 2244: 2242: 2218: 2199: 2198: 2196: 2194: 2180: 2171: 2165: 2164: 2162: 2160: 2131: 2122: 2113: 2112: 2110: 2108: 2087: 2078: 2065: 2064: 2062: 2060: 2043: 2034: 2021: 2020: 2018: 2016: 1987: 1978: 1972: 1971: 1969: 1967: 1950: 1941: 1932: 1931: 1917: 1911: 1910: 1904: 1895: 1880: 1879: 1877: 1875: 1864: 1838: 1833: 1832: 1824: 1819: 1818: 1817: 1801: 1795: 1788: 1782: 1774: 1768: 1754: 1748: 1740: 1734: 1726: 1720: 1712: 1706: 1694: 1688: 1678:Chen et al. 2019 1677: 1674:Talbragarocossus 1666:Wang et al. 2013 1665: 1650: 1641: 1635: 1627: 1621: 1613: 1604: 1601:Sinopalaeocossus 1598: 1590: 1584: 1576: 1570: 1556: 1550: 1542: 1536: 1528: 1522: 1514: 1508: 1500: 1494: 1478: 1469: 1463: 1451: 1445: 1437: 1431: 1423: 1417: 1406: 1397: 1391: 1379: 1373: 1365: 1359: 1351: 1345: 1325: 1317:Wang et al. 2008 1312: 1306: 1298: 1292: 1285:Yixian Formation 1273:Ren et al., 1998 1272: 1263: 1254: 1248: 1240: 1234: 1227:Amisan Formation 1222: 1216: 1200: 1194: 1186: 1173: 1167: 1159: 1153: 1145: 1139: 1131: 1125: 1117: 1111: 1104:Ukurei Formation 1091: 1085: 1073: 1067: 1055: 1049: 1037: 1031: 1015: 1009: 977:Yixian Formation 899:Lower Cretaceous 861:Yixian Formation 817:Early Cretaceous 646:Anton Handlirsch 626: 610: 594: 567: 549: 531: 434:and the rise of 318:Early Cretaceous 285: 283: 244: 225: 220: 207: 127: 126: 104: 94: 31: 21: 2902: 2901: 2897: 2896: 2895: 2893: 2892: 2891: 2842: 2841: 2840: 2835: 2827: 2822: 2814: 2809: 2801: 2796: 2788: 2783: 2775: 2770: 2762: 2757: 2749: 2744: 2736: 2731: 2723: 2721: 2712: 2711: 2706: 2697: 2696: 2691: 2678: 2642: 2637: 2627: 2625: 2596: 2595: 2591: 2581: 2579: 2573: 2572: 2568: 2558: 2556: 2544: 2539: 2538: 2534: 2524: 2522: 2510: 2505: 2504: 2500: 2448: 2447: 2440: 2430: 2428: 2401: 2396: 2395: 2391: 2381: 2379: 2374: 2373: 2366: 2356: 2354: 2335: 2330: 2329: 2322: 2312: 2310: 2302: 2297: 2296: 2292: 2282: 2280: 2261: 2256: 2255: 2248: 2220: 2219: 2202: 2192: 2190: 2178: 2173: 2172: 2168: 2158: 2156: 2129: 2124: 2123: 2116: 2106: 2104: 2102:10.1002/gj.1092 2085: 2080: 2079: 2068: 2058: 2056: 2041: 2036: 2035: 2024: 2014: 2012: 1985: 1980: 1979: 1975: 1965: 1963: 1948: 1943: 1942: 1935: 1919: 1918: 1914: 1902: 1897: 1896: 1883: 1873: 1871: 1866: 1865: 1861: 1857: 1834: 1827: 1820: 1815: 1813: 1810: 1751:Palaeontinopsis 1557:Oppenheim, 1888 1529:Oppenheim, 1885 1525:Phragmatoecites 1487:Sagul Formation 1438:Oppenheim, 1885 1424:Handlirsch 1906 1376:Neimenggucossus 1352:Ren et al. 1998 1309:Martynovocossus 1160:Oppenheim, 1888 1078:Crato Formation 1060:Dzhil Formation 997: 966:Crato Formation 825:Crato Formation 809:Paleogeographic 802: 752: 711:Palaeontinoidea 706:lepidopterans. 688:Auguste Lameere 638: 637: 636: 635: 634: 627: 619: 618: 611: 603: 602: 595: 586: 585: 579: 578: 577: 576: 575: 568: 560: 559: 557: 550: 542: 541: 539: 532: 523: 522: 516: 390: 334: 284:Cicadomorphidae 253: 250: 224: 218: 209:Palaeontinoidea 205: 186:Auchenorrhyncha 121: 95: 93: 92: 87: 82: 77: 72: 67: 62: 57: 52: 47: 42: 37: 29: 28: 25: 17: 12: 11: 5: 2900: 2898: 2890: 2889: 2884: 2879: 2874: 2869: 2864: 2859: 2854: 2844: 2843: 2837: 2836: 2834: 2833: 2820: 2807: 2794: 2781: 2768: 2755: 2742: 2729: 2719: 2714:Palaeontinidae 2704: 2688: 2686: 2684:Palaeontinidae 2680: 2679: 2674: 2668: 2667: 2664:Palaeontinidae 2655: 2654:at Wikispecies 2652:Palaeontinidae 2641: 2640:External links 2638: 2636: 2635: 2589: 2566: 2551:(in Spanish). 2532: 2498: 2438: 2389: 2364: 2320: 2290: 2246: 2200: 2166: 2114: 2066: 2022: 1973: 1933: 1912: 1909:(44): 533–718. 1881: 1858: 1856: 1853: 1852: 1851: 1846: 1840: 1839: 1836:Insects portal 1825: 1809: 1806: 1805: 1804: 1791: 1778: 1764: 1755:Martynov, 1937 1744: 1741:Westwood, 1854 1730: 1716: 1702: 1684: 1669: 1657: 1638:Suljuktocossus 1631: 1617: 1610:Quadraticossus 1594: 1580: 1566: 1546: 1532: 1518: 1504: 1490: 1470:Martynov, 1937 1466:Palaeontinodes 1459: 1441: 1427: 1413: 1387: 1369: 1362:Montsecocossus 1355: 1341: 1326:Martynov, 1931 1302: 1288: 1244: 1230: 1212: 1190: 1183:Papilioncossus 1163: 1149: 1142:Daohugoucossus 1135: 1121: 1107: 1092:Martynov, 1926 1081: 1063: 1045: 1027: 996: 993: 958:United Kingdom 801: 798: 782:Lower Jurassic 756:Tettigarctidae 751: 748: 719:Mesogereonidae 673:Edward Meyrick 644:entomologist 628: 621: 620: 612: 605: 604: 596: 589: 588: 587: 583: 582: 581: 580: 569: 562: 561: 551: 544: 543: 533: 526: 525: 524: 520: 519: 518: 517: 515: 514:Classification 512: 508:Mesogereonidae 472:Suljuktocossus 389: 386: 333: 330: 295:Palaeontinidae 290: 289: 277: 276: 270: 269: 260: 259: 255: 254: 251: 239: 238: 232: 231: 222:Palaeontinidae 216: 212: 211: 203: 199: 198: 193: 189: 188: 183: 179: 178: 173: 169: 168: 163: 159: 158: 153: 149: 148: 143: 139: 138: 133: 129: 128: 115: 114: 106: 105: 97: 96: 88: 83: 78: 73: 68: 63: 58: 53: 48: 43: 38: 33: 32: 26: 24:Palaeontinidae 15: 13: 10: 9: 6: 4: 3: 2: 2899: 2888: 2885: 2883: 2880: 2878: 2875: 2873: 2870: 2868: 2865: 2863: 2860: 2858: 2855: 2853: 2850: 2849: 2847: 2830: 2825: 2821: 2817: 2812: 2808: 2804: 2799: 2795: 2791: 2786: 2782: 2778: 2773: 2769: 2765: 2760: 2756: 2752: 2747: 2743: 2739: 2734: 2730: 2726: 2720: 2715: 2709: 2705: 2700: 2694: 2690: 2689: 2687: 2685: 2681: 2677: 2672: 2665: 2660: 2656: 2653: 2648: 2644: 2643: 2639: 2624: 2620: 2616: 2612: 2608: 2604: 2600: 2593: 2590: 2577: 2570: 2567: 2554: 2550: 2543: 2536: 2533: 2520: 2516: 2509: 2502: 2499: 2494: 2490: 2485: 2480: 2476: 2472: 2468: 2464: 2460: 2456: 2452: 2445: 2443: 2439: 2427: 2423: 2419: 2415: 2411: 2407: 2400: 2393: 2390: 2377: 2371: 2369: 2365: 2353: 2349: 2345: 2341: 2334: 2327: 2325: 2321: 2308: 2301: 2294: 2291: 2279: 2275: 2271: 2267: 2260: 2253: 2251: 2247: 2241: 2236: 2232: 2228: 2227:Palaeontology 2224: 2217: 2215: 2213: 2211: 2209: 2207: 2205: 2201: 2188: 2184: 2177: 2170: 2167: 2155: 2151: 2147: 2143: 2139: 2135: 2128: 2121: 2119: 2115: 2103: 2099: 2095: 2091: 2084: 2077: 2075: 2073: 2071: 2067: 2055: 2051: 2047: 2040: 2033: 2031: 2029: 2027: 2023: 2011: 2007: 2003: 1999: 1995: 1991: 1984: 1977: 1974: 1962: 1958: 1954: 1947: 1940: 1938: 1934: 1929: 1925: 1924: 1916: 1913: 1908: 1901: 1894: 1892: 1890: 1888: 1886: 1882: 1869: 1863: 1860: 1854: 1850: 1847: 1845: 1842: 1841: 1837: 1831: 1826: 1823: 1812: 1807: 1800: 1799: 1792: 1789:- Zhang, 1997 1787: 1786: 1779: 1773: 1772: 1765: 1762: 1761: 1753: 1752: 1745: 1739: 1738: 1731: 1725: 1724: 1717: 1711: 1710: 1703: 1700: 1693: 1692: 1685: 1682: 1676: 1675: 1670: 1664: 1663: 1658: 1655: 1649: 1648: 1640: 1639: 1632: 1626: 1625: 1618: 1612: 1611: 1603: 1602: 1595: 1589: 1588: 1587:Shurabocossus 1581: 1575: 1574: 1567: 1564: 1563: 1555: 1554: 1547: 1541: 1540: 1533: 1527: 1526: 1519: 1513: 1512: 1505: 1499: 1498: 1491: 1488: 1484: 1477: 1476: 1468: 1467: 1460: 1457: 1450: 1449: 1442: 1436: 1435: 1428: 1422: 1421: 1414: 1411: 1405: 1404: 1396: 1395: 1388: 1383: 1378: 1377: 1370: 1364: 1363: 1356: 1350: 1349: 1342: 1339: 1335: 1331: 1324: 1323: 1316: 1311: 1310: 1303: 1297: 1296: 1289: 1286: 1282: 1278: 1271: 1270: 1262: 1261: 1260:Wonnacottella 1253: 1252: 1245: 1239: 1238: 1231: 1228: 1221: 1220: 1213: 1210: 1206: 1199: 1198: 1191: 1185: 1184: 1177: 1172: 1171: 1164: 1158: 1157: 1150: 1144: 1143: 1136: 1130: 1129: 1122: 1116: 1115: 1114:Colossocossus 1108: 1105: 1101: 1097: 1090: 1089: 1082: 1079: 1072: 1071: 1064: 1061: 1054: 1053: 1046: 1043: 1036: 1035: 1028: 1025: 1024:Daohugou Beds 1019: 1014: 1013: 1006: 1005: 1004: 1002: 994: 992: 990: 986: 985:Daohugou Beds 982: 978: 974: 970: 967: 963: 959: 955: 951: 947: 943: 939: 935: 931: 927: 923: 919: 915: 914:South America 911: 906: 904: 900: 896: 892: 888: 884: 883:Late Triassic 881:stage of the 880: 876: 875: 866: 862: 858: 854: 853:Transbaikalia 850: 846: 842: 838: 834: 830: 826: 822: 818: 814: 810: 806: 799: 797: 795: 791: 787: 783: 779: 778:Upper Permian 774: 772: 767: 765: 761: 757: 749: 747: 745: 741: 738: 733: 732:(true bugs). 731: 727: 724: 720: 716: 712: 707: 704: 701: 697: 693: 689: 686:entomologist 685: 680: 678: 674: 670: 666: 665: 660: 659: 654: 651: 647: 643: 633: 632: 625: 617: 616: 609: 601: 600: 593: 574: 573: 566: 556: 555: 548: 538: 537: 530: 513: 511: 509: 504: 502: 498: 494: 489: 488: 483: 482: 476: 474: 473: 466: 464: 460: 455: 453: 449: 445: 441: 440:insectivorous 437: 433: 428: 424: 419: 417: 413: 409: 405: 401: 400: 399:Colossocossus 395: 387: 385: 383: 379: 375: 371: 368: 364: 361: 357: 353: 349: 345: 341: 340: 331: 329: 327: 323: 319: 315: 314:Late Triassic 311: 307: 304: 300: 299:giant cicadas 296: 288: 278: 275: 271: 268: 267: 261: 256: 249: 248: 240: 237: 233: 228: 223: 217: 214: 213: 210: 204: 202:Superfamily: 201: 200: 197: 194: 191: 190: 187: 184: 181: 180: 177: 174: 171: 170: 167: 164: 161: 160: 157: 154: 151: 150: 147: 144: 141: 140: 137: 134: 131: 130: 125: 120: 116: 113: 112: 107: 103: 98: 91: 86: 81: 76: 71: 66: 61: 56: 51: 46: 41: 36: 30:Norian–Aptian 22: 19: 2683: 2626:. Retrieved 2606: 2602: 2592: 2580:. Retrieved 2569: 2557:. Retrieved 2552: 2548: 2535: 2523:. Retrieved 2518: 2514: 2501: 2458: 2454: 2429:. Retrieved 2409: 2405: 2392: 2380:. Retrieved 2355:. Retrieved 2343: 2339: 2311:. Retrieved 2309:(1): 318–319 2306: 2293: 2281:. Retrieved 2269: 2265: 2230: 2226: 2191:. Retrieved 2186: 2182: 2169: 2157:. Retrieved 2137: 2133: 2105:. Retrieved 2093: 2089: 2057:. Retrieved 2045: 2013:. Retrieved 1993: 1989: 1976: 1964:. Retrieved 1952: 1922: 1915: 1906: 1872:. Retrieved 1862: 1802:- Riek, 1976 1796: 1783: 1769: 1760:nomen dubium 1758: 1749: 1735: 1721: 1707: 1689: 1672: 1662:Synapocossus 1660: 1653: 1647:Fletcheriana 1645: 1636: 1622: 1608: 1599: 1585: 1571: 1560: 1551: 1537: 1523: 1509: 1495: 1473: 1464: 1452:Butler, 1873 1446: 1434:Palaeocossus 1432: 1418: 1409: 1403:Fletcheriana 1401: 1392: 1381: 1374: 1360: 1346: 1332:, Daohugou, 1322:Pseudocossus 1320: 1314: 1307: 1293: 1267: 1258: 1257:- (includes 1251:Ilerdocossus 1249: 1235: 1219:Hallakkungis 1217: 1195: 1181: 1180:- (includes 1175: 1168: 1154: 1140: 1126: 1112: 1088:Cicadomorpha 1086: 1068: 1050: 1032: 1017: 1010: 998: 960:. Important 942:Turkmenistan 907: 890: 886: 874:Hallakkungis 872: 870: 856: 844: 832: 820: 790:South Africa 775: 770: 768: 753: 739: 737:type species 734: 726:Cicadomorpha 715:Dunstaniidae 708: 681: 662: 656: 650:lepidopteran 639: 629: 613: 597: 570: 552: 534: 505: 497:sphinx moths 487:Ilerdocossus 485: 479: 477: 470: 467: 456: 450:, and early 446:, primitive 427:ginkgophytes 420: 397: 391: 373: 372:in his book 367:entomologist 337: 335: 310:cicadomorphs 298: 294: 293: 280: 263: 252:Butler, 1873 245: 221: 196:Cicadomorpha 192:Infraorder: 109: 18: 2772:iNaturalist 2708:Wikispecies 1709:Valdicossus 1651:Evans, 1956 1573:Protopsyche 1539:Plachutella 1448:Palaeontina 1420:Pachypsyche 1407:Evans, 1956 1394:Ningchengia 1295:Limacodites 1197:Gansucossus 1128:Cratocossus 1034:Archipsyche 969:Lagerstätte 934:Transbaikal 897:age of the 815:during the 760:Prosbolidae 730:hemipterans 669:Limacodidae 658:Palaeontina 653:Heteroneura 436:angiosperms 432:gymnosperms 423:host plants 416:hemipterans 406:and a long 382:Nymphalidae 352:Oxfordshire 286:Evans, 1956 247:Palaeontina 2846:Categories 1855:References 1723:Yanocossus 1691:Turgaiella 1624:Suljuktaja 1605:Hong, 1983 1562:Beloptesis 1501:Ueda, 1997 1348:Miracossus 1281:Weald Clay 1269:Liaocossus 1237:Hamicossus 1170:Eoiocossus 1070:Baeocossus 1052:Asiocossus 1012:Abrocossus 983:, and the 962:localities 956:, and the 950:Kazakhstan 946:Kyrgyzstan 938:Tajikistan 744:Hemipteran 723:infraorder 696:Australian 677:Hepialidae 558:(forewing) 540:(forewing) 236:Type genus 227:Handlirsch 182:Suborder: 156:Arthropoda 2623:129712238 2461:: 40691. 2426:0066-4170 2278:1734-915X 2154:0195-6671 2054:0341-0153 2010:1175-5334 1961:1175-5326 1737:Cyllonium 1727:Ren, 1995 1654:in partim 1553:Prolystra 1475:Ijacossus 1410:in partim 786:Australia 764:Cicadidae 750:Evolution 700:geologist 692:Cicadidae 444:theropods 378:butterfly 332:Discovery 176:Hemiptera 142:Kingdom: 136:Eukaryota 2751:23370914 2722:BioLib: 2699:Q7126461 2693:Wikidata 2628:July 13, 2582:July 15, 2559:July 21, 2525:July 21, 2493:28098190 2431:July 21, 2382:July 15, 2357:July 15, 2313:July 15, 2283:July 15, 2193:July 15, 2159:July 15, 2107:July 15, 2059:July 15, 2015:July 15, 1966:July 13, 1874:July 16, 1808:See also 1156:Eocicada 975:and the 936:region, 905:period. 903:Jurassic 889:. 227 – 664:Eocicada 642:Austrian 481:Eocicada 344:forewing 301:, is an 274:Synonyms 215:Family: 152:Phylum: 146:Animalia 132:Domain: 2803:4223563 2764:4783069 2725:1210637 2484:5241632 2463:Bibcode 1990:Zootaxa 1953:Zootaxa 930:Germany 910:Eurasia 728:of the 684:Belgian 448:mammals 408:rostrum 360:English 358:by the 356:England 326:species 316:to the 303:extinct 258:Genera 172:Order: 166:Insecta 162:Class: 2816:175586 2790:111501 2777:696646 2621:  2491:  2481:  2424:  2276:  2152:  2052:  2008:  1959:  1001:genera 995:Genera 973:Brazil 932:, the 926:Russia 918:Brazil 895:Aptian 879:Norian 855:; and 849:Baissa 829:Brazil 792:, and 322:genera 306:family 229:, 1906 2824:Plazi 2785:IRMNG 2738:8HBR6 2619:S2CID 2545:(PDF) 2511:(PDF) 2402:(PDF) 2336:(PDF) 2303:(PDF) 2262:(PDF) 2179:(PDF) 2130:(PDF) 2086:(PDF) 2042:(PDF) 1986:(PDF) 1949:(PDF) 1930:–127. 1903:(PDF) 1644:- (= 1607:- (= 1472:- (= 1400:- (= 1319:- (= 989:China 954:Spain 922:China 865:China 841:Spain 813:Earth 794:China 771:et al 769:Wang 493:wasps 452:birds 412:xylem 404:frons 394:setae 2759:GBIF 2630:2011 2584:2011 2561:2011 2527:2011 2489:PMID 2433:2011 2422:ISSN 2384:2011 2359:2011 2315:2011 2285:2011 2274:ISSN 2195:2011 2161:2011 2150:ISSN 2109:2011 2061:2011 2050:ISSN 2017:2011 2006:ISSN 1994:1390 1968:2011 1957:ISSN 1876:2011 1559:syn 1266:and 912:and 717:and 682:The 661:and 501:lift 495:and 266:text 264:See 35:PreꞒ 2746:EoL 2733:CoL 2611:doi 2479:PMC 2471:doi 2414:doi 2348:doi 2235:doi 2142:doi 2098:doi 1998:doi 1928:126 987:of 971:of 784:of 780:to 308:of 2848:: 2826:: 2813:: 2800:: 2787:: 2774:: 2761:: 2748:: 2735:: 2710:: 2695:: 2617:. 2607:10 2605:. 2601:. 2547:. 2519:16 2517:. 2513:. 2487:. 2477:. 2469:. 2457:. 2453:. 2441:^ 2420:. 2408:. 2404:. 2367:^ 2344:28 2342:. 2338:. 2323:^ 2305:. 2270:46 2268:. 2264:. 2249:^ 2231:52 2229:. 2225:. 2203:^ 2187:34 2185:. 2181:. 2148:. 2138:29 2136:. 2132:. 2117:^ 2094:43 2092:. 2088:. 2069:^ 2044:. 2025:^ 2004:. 1992:. 1988:. 1951:. 1936:^ 1905:. 1884:^ 1697:- 1485:, 1481:) 1454:- 1382:in 1336:, 1328:) 1315:in 1283:, 1279:, 1275:) 1225:- 1207:, 1203:- 1176:in 1102:, 1094:- 1076:- 1058:- 1040:- 1022:- 1018:in 991:. 979:, 952:, 948:, 944:, 940:, 928:, 924:, 920:, 891:ca 887:ca 863:, 859:- 851:, 847:- 843:; 839:, 835:- 831:; 827:, 823:- 788:, 766:. 503:. 418:. 384:. 354:, 328:. 85:Pg 2632:. 2613:: 2586:. 2563:. 2553:1 2529:. 2495:. 2473:: 2465:: 2459:7 2435:. 2416:: 2410:8 2386:. 2361:. 2350:: 2317:. 2287:. 2243:. 2237:: 2197:. 2163:. 2144:: 2111:. 2100:: 2063:. 2019:. 2000:: 1970:. 1878:. 1794:† 1781:† 1767:† 1763:) 1747:† 1733:† 1719:† 1705:† 1687:† 1671:† 1659:† 1634:† 1620:† 1597:† 1583:† 1569:† 1549:† 1535:† 1521:† 1507:† 1493:† 1462:† 1444:† 1430:† 1416:† 1390:† 1372:† 1358:† 1344:† 1305:† 1291:† 1247:† 1233:† 1215:† 1193:† 1166:† 1152:† 1138:† 1124:† 1110:† 1084:† 1066:† 1048:† 1030:† 1008:† 885:( 867:. 857:4 845:3 833:2 821:1 282:† 243:† 219:† 206:† 90:N 80:K 75:J 70:T 65:P 60:C 55:D 50:S 45:O 40:Ꞓ

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Synapocossus sciacchitanoae
Scientific classification
Edit this classification
Eukaryota
Animalia
Arthropoda
Insecta
Hemiptera
Auchenorrhyncha
Cicadomorpha
Palaeontinoidea
Palaeontinidae
Handlirsch
Type genus
Palaeontina
text
Synonyms

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