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Peștera cu Oase

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mature female, designated as "Oase 3". However, additional finds and work have shown that the temporal bone derives from the same cranium as the "Oase 2" facial and parietal bones. The lack of archaeological signs such as torches, charcoal or tools could suggest that the human remains may have washed in the cave through fissures. The "Oase 2" and "Oase 3" confirm a pattern already known from the probably contemporaneous "Oase 1" mandible, indicating a mixture of archaic, early modern human and Neanderthal morphological features. Thus, the specimens exhibit a suite of derived "modern human" features like projecting chin, no brow ridge, a high and rounded brain case. Yet, these features are associated with several archaic aspects of the cranium and dentition that place them outside the range of variation for modern humans, like a large face, a large crest of bone behind the ear and big teeth that get even larger toward the back. This mosaic of Neanderthal and modern human resembles similar traits found in a 25,000 years old fossil of a child in
567: 97: 69: 355: 62: 585: 90: 451: 45: 1307: 753:"Oase 2" belongs to the same basal subclade of mitochondrial DNA haplogroup N as "Oase 1". When compared against all DNA samples on record, "Oase 2" and "Oase 1" share the closest genetic affinity with each other. "Oase 1" and "Oase 2" appear to be from related, but not necessarily identical populations. 866:
In this context, the particular importance of the "Peștera cu Oase" findings resides both in the mixture of modern human and archaic (Neanderthal) features and in the fact that they are sufficiently complete to be taxonomically diagnosed and directly dated. Thus, the Oase fossils overlap in time for
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In June 2003 a further research team with Ștefan Milota, Ricardo Rodrigo, and Mircea Gherase discovered additional human remains on the cave's surface. Thus, an entire anterior cranial skeleton was found along with a largely complete left temporal bone and a number of frontal, parietal and occipital
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Trinkaus, E., Zilhão, J., Rougier, H., Rodrigo, R., Milota, S., Gherase, M., Sarcinã, L., Moldovan, O., Bãltean, I., Codrea, V., Bailey, S. E., Franciscus, R. G., Ponce de Léon, M., Zollikofer, C. P. E. (2006) The Peștera cu Oase and early modern humans in Southeastern Europe. In N.J. Conard (Ed.),
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When modern humans entered Europe, they encountered people with the same cognitive capabilities and featuring identical levels of cultural achievement. In such a situation, the entire gamut of cultural interaction situations, from conflict to mutual avoidance and full admixture, must have ensued at
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While "Oase 1" lower jaw is fully mature, the facial skeleton is that of a mid-second-decade adolescent, therefore corresponding to a second individual, designated as "Oase 2". Further analyses have revealed that the left temporal bone represents a third individual, assessed as adolescent versus
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Since genetics does not reject the hypothesis of a Neanderthal-modern admixture, and morphological and archaeological evidence suggest that Neanderthal lineages survived into later Upper Paleolithic populations, "Peștera cu Oase" findings provide a strong argument in favor of an admixture model
883:) at ~34,000 radiocarbon years BP. Besides, the notion that the Oase people are very close to the time of contact with Neanderthals is consistent with their archaic traits, and finds additional support in the patterns of spatio-temporal distribution of the latest Neanderthal remains. 1207:
Qiaomei Fu; Mateja Hajdinjak; Oana Teodora Moldovan; Silviu Constantin; Swapan Mallick; Pontus Skoglund; Nick Patterson; Nadin Rohland; Iosif Lazaridis; Birgit Nickel; Bence Viola; Kay Prüfer; Matthias Meyer; Janet Kelso; David Reich; Svante Pääbo (13 August 2015).
284:), presented unusual arrangements such as the placement of some remains on raised rocks, suggesting a certain human involvement in the accumulated deposits. In fact, speleologists Ștefan Milota, Adrian Bîlgăr and Laurențiu Sarcina discovered a complete 890:
Arguing with chronological overlapping and morphological blending, this model assumes significant Neanderthal/modern human admixture, suggesting that already on their arrival in Europe, modern humans met, intermixed and interbred with Neanderthals.
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differences have suggested a major physical anthropological discontinuity and hence, a complete population replacement at the Middle-to-Upper Palaeolithic transition, leading to what one might call "Out of Africa with Complete Replacement" model.
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the local and regional level. But the overall result in the long-term continental perspective was that of biological and cultural blending, the imbalance in the size of the gene reservoirs involved explaining the eventual loss of Neanderthal
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A skull found in Peștera cu Oase in 2004/5 bears features of both modern humans and Neanderthals. Radiocarbon dating indicates that the skull is 37,800 years old, making it amongst the oldest modern human fossils ever found in Europe.
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Around 6% of "Oase 2"'s genome is Neanderthal in origin, which is lower than for "Oase 1"; however, this is still much higher than expected based on its age and what is seen in other Upper Palaeolithic genomes.
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populations than with modern Europeans. However, Oase shared equal alleles with Mesolithic Europeans and East Eurasians suggesting non pre LGM-European admixture in modern Europeans, part of it being from the
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backgrounds, respectively, for whom Poznik et al. proposed the creation of a new subclade, named "K2a1"). (Earlier research by Fu et al. reported that Oase 1 belonged to a
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Fu, Qiaomei; Hajdinjak, Mateja; Moldovan, Oana Teodora; Constantin, Silviu; Mallick, Swapan; Skoglund, Pontus; Patterson, Nick; Rohland, Nadin; Lazaridis, Iosif (2015-08-13).
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Peștera cu Oase is subject to ongoing investigation. The on-site findings from the 2005 campaign are currently cross-examined at the Romanian "Emil Racoviță" Institute of
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and together with the linkage disequilibrium patterns indicates that Oase 1 had a relatively-recent Neanderthal ancestor – about four to six generations earlier.
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The marked contrast between the morphological modernity of "early modern" humans and even late "classical Neanderthal" trait-packages, as well as mitochondrial
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Trinkaus, E., Milota, Ș., Rodrigo, R., Gherase, M., Moldovan, O. (2003), Early Modern Human Cranial remains from the Peștera cu Oase, Romania,
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G. David Poznik et al., 2016, "Punctuated bursts in human male demography inferred from 1,244 worldwide Y-chromosome sequences"
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Trinkaus, E.; Moldovan, O.; Milota, Ș.; Bîlgăr, A.; Sarcina, L.; Athreya, S.; Bailey, S. E.; Rodrigo, R.; et al. (2003).
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The autosomal DNA of Oase 1 by Fu et al. (2015) indicates that he (as it was a male) may have shared more alleles with modern
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However, more recent direct dating of fossils has demonstrated that early modern human remains were instead of the mid/late
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ancestor, with an estimated 5-11% Neanderthal autosomal DNA. The specimen's 12th chromosome was 50% Neanderthal.
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Mellars, P (Feb 2006). "A new radiocarbon revolution and the dispersal of modern humans in Eurasia".
1221: 1158: 1053: 910: 832: 820: 809: 805: 541: 267: 5300: 776:. Neither "Oase 2" nor "Oase 1" are particularly close genetically to any modern human populations. 574:
Forensic facial reconstruction of Oase 2. Exhibited in the Neanderthal Museum in Mettmann, Germany.
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About 6-9% of the genome is Neanderthal in origin. This is the highest percentage of archaic
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Fu Q. et al., "An early modern human from Romania with a recent Neanderthal ancestor",
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Research by Poznik et al. (2016) suggests that Oase 1 Y-DNA belongs to haplogroup
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ancestry that was carried to Europe by Anatolian farmers and Yamnaya pastoralists.
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This article contains quotations from this source, which is available under the
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Cave of Altamira and Paleolithic Cave Art of Northern Spain World Heritage Site
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Researchers sequenced the genome of "Oase 2" (41,500–39,500 years old) to high
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DNA analysis of Oase 1 since 2015 has made a number of significant findings.
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team exploring the karstic system of Miniș Valley, in the southwestern
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Phylogenetic position of ancient Upper Paleolithic Eurasian specimens.
298:(The Cave with Bones) and the human mandible as "Oase 1" (also dubbed 228:(EEMH) remains, between 42,000 and 37,000 years old, have been found. 4255: 3529: 3524: 2452: 1450:"Early modern humans from the Peștera Muierii, Baia de Fier, Romania" 1337: 880: 656: 626: 606: 510:(also K2a*), 45,000-year-old remains from Siberia, and upstream from 408: 381: 247: 244: 335:"Oase 1" exhibits morphological traits from early modern humans and 1324:
Ponce, De, León, Ms; Zollikofer, Cp (Aug 2001). "Peștera cu Oase".
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Zilhão, J. (2006). "Neandertals and Moderns Mixed and It Matters".
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Human population history and its interplay with natural selection
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Rock art of the Iberian Mediterranean Basin World Heritage Site
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on 21 bone/tooth samples and 29 associated sediment samples),
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Caves and Ice Age Art in the Swabian Jura World Heritage Site
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Creative Commons Attribution 4.0 International (CC BY 4.0)
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and a rare lineage found in two living males (from ethnic
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According to Fu, Oase-1's maternal lineage is related to
954:"Fossil Teeth Put Humans in Europe Earlier Than Thought" 937:
524, 216–219 (13 August 2015), doi:10.1038/nature14558.
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between regional Neanderthals and early modern humans.
1147:"An early modern human from Peștera cu Oase, Romania" 867:
some 3000 years with late Neanderthals like those of
875:) dated to ~32,000 radiocarbon years BP or less for 345: 5607: 5591: 5575: 5557: 5536: 5518: 5502: 5474: 5278: 4997: 4981: 4940: 4884: 4868: 4852: 4834: 4793: 4752: 4731: 4710: 4664: 4638: 4622: 4606: 4561: 4540: 4524: 4498: 4327: 4301: 4285: 4264: 4248: 4202: 4181: 4165: 4144: 4123: 4107: 4091: 4070: 4054: 4008: 3987: 3961: 3949: 3923: 3892: 3856: 3838: 3822: 3806: 3790: 3774: 3728: 3707: 3691: 3640: 3559: 3538: 3482: 3451: 3405: 3389: 3363: 3302: 3286: 3240: 3224: 3208: 3082: 3066: 3045: 2788: 2757: 2731: 2700: 2654: 2623: 2572: 2551: 2535: 2517: 2501: 2485: 2469: 2451: 2388: 2367: 2306: 2278: 2202: 1878: 1862: 1821: 1805: 1764: 1728: 1707: 1652: 1616: 835:(AMS radiocarbon dating on 25 bone/tooth samples). 208:, meaning "The Cave with Bones") is a system of 12 184: 179: 142: 128: 116: 804:, (uranium-series analysis on 22 bone samples), 311:of the Oase fossils give an age of 37,800 years 893: 1507:Trinkaus, Erik (2005). "Early Modern Humans". 299: 293: 1553: 1448:Soficaru, A.; Dobo, A.; Trinkaus, E. (2006). 212:galleries and chambers located near the city 8: 1374:: CS1 maint: multiple names: authors list ( 977: 975: 973: 506:. That is, Oase 1 possesses SNPS similar to 243:In 2015 genetics research revealed that the 30: 5275: 5262: 4603: 4586: 4051: 4038: 3205: 3192: 1613: 1600: 1568:Prehistoric cave sites, rock shelters and 1560: 1546: 1538: 565: 556: 29: 1483: 1473: 1241: 1180: 1170: 1081: 947: 945: 943: 764:. "Oase 1" shows a genetic affinity for " 4335:Cradle of Humankind, World Heritage Site 1105: 1103: 1101: 808:, (uranium-series dating on 7 samples), 292:. The karstic chamber was designated as 1521:10.1146/annurev.anthro.34.030905.154913 1001:When Neanderthals and modern humans met 926: 900:lineages among later and extant humans. 96: 68: 27:Cave and archaeological site in Romania 5675:Upper Paleolithic Homo sapiens fossils 1367: 312: 1003:, pp. 145-164. Tübingen: Kerns Verlag 863:, hence much younger than supposed. 203: 7: 1202: 1200: 240:, by Cidália Duarte, et al. (1999). 236:or in the 31,000 years old site of 952:Wilford, John Noble (2 Nov 2011). 25: 5690:Prehistory of Southeastern Europe 2723:Mala Balanica and Velika Balanica 1886:Vézère Valley World Heritage Site 1681:Neolithic flint mines of Spiennes 819:dating on 8 bone/tooth samples), 1305: 583: 449: 353: 95: 88: 67: 60: 43: 5670:2002 archaeological discoveries 1013:Anderson, Andrea (8 May 2015). 315:. From a location close to the 790:Australian National University 437:Initial Upper Paleolithic wave 1: 1509:Annual Review of Anthropology 491:Oase 1 belongs to an extinct 5705:Early European modern humans 5680:Prehistoric sites in Romania 539:mitochondrial DNA haplogroup 497:mitochondrial DNA haplogroup 2911:Other caves with decoration 1993:Other caves with decoration 1915:Les Eyzies-de-Tayac-Sireuil 1591:Caves containing pictograms 831:on 37 bone/tooth samples), 226:European early modern human 224:, where some of the oldest 5721: 2895:Other World Heritage Sites 1982:Other World Heritage Sites 1581:Paleoanthropological sites 983:Journal of Human Evolution 205:[ˈpeʃterakuˈo̯ase] 5631: 5274: 5261: 4602: 4585: 4050: 4037: 3204: 3191: 1612: 1599: 1576: 1112:Evolutionary Anthropology 847:Implications for research 580:Scientific classification 578: 573: 564: 559: 474:anatomically modern human 104:Peștera cu Oase (Romania) 54: 42: 35: 5559:Northern Mariana Islands 1280:Siska, Veronika (2018). 722:H. neanderthalensis 1475:10.1073/pnas.0608443103 1293:University of Cambridge 1172:10.1073/pnas.2035108100 825:stable isotope analysis 794:electron spin resonance 201:Romanian pronunciation: 4083:Manyana Rock Paintings 2134:La Chapelle-aux-Saints 1270:, no. 48, pp. 593–599. 902: 300: 294: 4646:Actun Tunichil Muknal 802:University of Bristol 798:uranium-series dating 234:Abrigo do Lagar Velho 220:county, southwestern 135:county, southwestern 4220:Ifri N'Amr Ou Moussa 3160:Red Lady of Paviland 2052:Mayrières supérieure 833:University of Vienna 821:Max Planck Institute 810:University of Oxford 806:University of Bergen 526:of Y-DNA haplogroup 268:Carpathian Mountains 262:In February 2002, a 31:Peștera cu Oase Cave 5035:Colorado Millennial 3673:Shiraho Saonetabaru 2477:Bukuroshja e Fjetur 1466:2006PNAS..10317196S 1460:(46): 17196–17201. 1411:10.1038/nature04521 1403:2006Natur.439..931M 1234:10.1038/nature14558 1226:2015Natur.524..216F 1163:2003PNAS..10011231T 1157:(20): 11231–11236. 1066:10.1038/nature14558 1058:2015Natur.524..216F 916:Prehistoric Romania 742:(20-fold) from its 160: /  107:Show map of Romania 76:location in Romania 32: 5695:Peopling of Europe 4853:Dominican Republic 4614:Cueva de las Manos 3510:Huto and Kamarband 2920:Barranc del Migdia 2806:Caves in Cantabria 2230:Hohlenstein-Stadel 2037:Grotte de Gabillou 2022:La Chaire a Calvin 1301:10.17863/CAM.31536 1124:10.1002/evan.20110 304:"John of Anina"). 79:Show map of Europe 5657: 5656: 5627: 5626: 5623: 5622: 5257: 5256: 5253: 5252: 4770:Piedras del Tunjo 4702:Toca da Tira Peia 4581: 4580: 4577: 4576: 4033: 4032: 4029: 4028: 3187: 3186: 3183: 3182: 2823:Hornos de la Peña 1220:(7564): 216–219. 1052:(7564): 216–219. 1019:www.genomeweb.com 911:Peștera Muierilor 766:Peştera Muierii 2 736: 735: 462: 461: 321:Danubian corridor 309:radiocarbon dates 194: 193: 164:45.017°N 21.833°E 16:(Redirected from 5712: 5685:Caves of Romania 5576:Papua New Guinea 5276: 5263: 5215:Stanfield-Worley 5190:Rockhouse Cliffs 4604: 4587: 4455:Howieson's Poort 4152:Enkapune Ya Muto 4052: 4039: 3505:Eshkaft-e Siahoo 3206: 3193: 2960:Peñas de Cabrera 2260:Kleine Feldhofer 2220:Geissenklösterle 2072:Roc-aux-Sorciers 1614: 1601: 1562: 1555: 1548: 1539: 1533: 1532: 1504: 1498: 1497: 1487: 1477: 1445: 1439: 1438: 1386: 1380: 1379: 1373: 1365: 1338:10.1038/35087573 1321: 1315: 1309: 1304: 1286: 1277: 1271: 1262: 1256: 1255: 1245: 1204: 1195: 1194: 1184: 1174: 1142: 1136: 1135: 1107: 1096: 1095: 1085: 1037: 1031: 1030: 1028: 1026: 1010: 1004: 996: 990: 979: 968: 967: 965: 964: 949: 938: 931: 780:Current research 762:Native Americans 588: 587: 569: 557: 493:Y-DNA haplogroup 453: 452: 442: 440: 431: 429: 422: 420: 413: 411: 404: 402: 395: 393: 386: 384: 377: 375: 368: 366: 357: 346: 314: 303: 297: 276:Late Pleistocene 207: 202: 175: 174: 172: 171: 170: 165: 161: 158: 157: 156: 153: 108: 99: 98: 92: 80: 71: 70: 64: 47: 33: 21: 5720: 5719: 5715: 5714: 5713: 5711: 5710: 5709: 5660: 5659: 5658: 5653: 5619: 5603: 5587: 5571: 5553: 5532: 5514: 5498: 5470: 5386:Mackintosh 90/1 5306:Beginner's Luck 5270: 5249: 5065:Dutchess Quarry 5005:Arnold Research 4993: 4977: 4936: 4880: 4864: 4848: 4830: 4789: 4748: 4727: 4706: 4660: 4656:Midnight Terror 4634: 4618: 4598: 4573: 4557: 4536: 4520: 4494: 4323: 4297: 4281: 4260: 4244: 4235:Kelif el Boroud 4198: 4177: 4161: 4140: 4119: 4103: 4087: 4066: 4046: 4025: 4004: 3983: 3957: 3945: 3919: 3888: 3852: 3834: 3818: 3802: 3786: 3770: 3724: 3703: 3687: 3636: 3555: 3534: 3478: 3447: 3401: 3385: 3359: 3298: 3282: 3236: 3220: 3200: 3179: 3078: 3062: 3041: 2940:los Murciélagos 2888:Roca dels Moros 2784: 2753: 2727: 2696: 2667:Chertovy Vorota 2650: 2619: 2568: 2547: 2531: 2519:North Macedonia 2513: 2497: 2481: 2465: 2447: 2384: 2363: 2302: 2274: 2198: 1961:Abri du Poisson 1926:Les Combarelles 1874: 1858: 1817: 1801: 1760: 1724: 1703: 1699:Trou de l'Abîme 1648: 1608: 1595: 1572: 1566: 1536: 1506: 1505: 1501: 1447: 1446: 1442: 1397:(7079): 931–5. 1388: 1387: 1383: 1366: 1332:(6846): 534–8. 1323: 1322: 1318: 1284: 1279: 1278: 1274: 1267:Nature Genetics 1263: 1259: 1206: 1205: 1198: 1144: 1143: 1139: 1109: 1108: 1099: 1039: 1038: 1034: 1024: 1022: 1012: 1011: 1007: 997: 993: 980: 971: 962: 960: 951: 950: 941: 932: 928: 924: 907: 849: 782: 732: 582: 553: 495:and an extinct 458: 457: 456: 454: 446: 445: 444: 443: 439:"East-Eurasian" 438: 436: 434: 432: 427: 425: 423: 418: 416: 414: 409: 407: 405: 400: 398: 396: 391: 389: 387: 382: 380: 378: 373: 371: 369: 364: 362: 358: 332:bone segments. 329: 295:Peștera cu Oase 260: 200: 197:Peștera cu Oase 168: 166: 162: 159: 154: 151: 149: 147: 146: 112: 111: 110: 109: 106: 105: 102: 101: 100: 83: 82: 81: 78: 77: 74: 73: 72: 50: 38: 37:Peștera cu Oase 28: 23: 22: 18:Peştera cu Oase 15: 12: 11: 5: 5718: 5716: 5708: 5707: 5702: 5697: 5692: 5687: 5682: 5677: 5672: 5662: 5661: 5655: 5654: 5652: 5651: 5646: 5641: 5636: 5632: 5629: 5628: 5625: 5624: 5621: 5620: 5618: 5617: 5611: 5609: 5605: 5604: 5602: 5601: 5595: 5593: 5589: 5588: 5586: 5585: 5579: 5577: 5573: 5572: 5570: 5569: 5563: 5561: 5555: 5554: 5552: 5551: 5546: 5540: 5538: 5534: 5533: 5531: 5530: 5524: 5522: 5516: 5515: 5513: 5512: 5506: 5504: 5500: 5499: 5497: 5496: 5491: 5486: 5480: 5478: 5472: 5471: 5469: 5468: 5463: 5458: 5453: 5448: 5443: 5438: 5433: 5428: 5423: 5418: 5413: 5408: 5403: 5398: 5393: 5388: 5383: 5378: 5373: 5368: 5363: 5358: 5353: 5348: 5343: 5338: 5333: 5328: 5323: 5318: 5313: 5308: 5303: 5301:Beeton Shelter 5298: 5293: 5288: 5282: 5280: 5272: 5271: 5266: 5259: 5258: 5255: 5254: 5251: 5250: 5248: 5247: 5242: 5237: 5232: 5227: 5222: 5217: 5212: 5207: 5202: 5197: 5192: 5187: 5182: 5177: 5172: 5167: 5162: 5157: 5152: 5147: 5142: 5137: 5132: 5127: 5122: 5117: 5112: 5107: 5102: 5097: 5092: 5087: 5082: 5077: 5072: 5067: 5062: 5057: 5052: 5047: 5042: 5037: 5032: 5027: 5022: 5017: 5012: 5007: 5001: 4999: 4995: 4994: 4992: 4991: 4985: 4983: 4979: 4978: 4976: 4975: 4973:T'uqu T'uquyuq 4970: 4965: 4960: 4955: 4950: 4944: 4942: 4938: 4937: 4935: 4934: 4929: 4924: 4919: 4914: 4909: 4904: 4899: 4894: 4888: 4886: 4882: 4881: 4879: 4878: 4872: 4870: 4866: 4865: 4863: 4862: 4856: 4854: 4850: 4849: 4847: 4846: 4840: 4838: 4832: 4831: 4829: 4828: 4823: 4818: 4813: 4808: 4803: 4797: 4795: 4791: 4790: 4788: 4787: 4782: 4777: 4772: 4767: 4762: 4756: 4754: 4750: 4749: 4747: 4746: 4741: 4735: 4733: 4729: 4728: 4726: 4725: 4720: 4714: 4712: 4708: 4707: 4705: 4704: 4699: 4694: 4689: 4684: 4679: 4674: 4668: 4666: 4662: 4661: 4659: 4658: 4653: 4648: 4642: 4640: 4636: 4635: 4633: 4632: 4626: 4624: 4620: 4619: 4617: 4616: 4610: 4608: 4600: 4599: 4590: 4583: 4582: 4579: 4578: 4575: 4574: 4572: 4571: 4565: 4563: 4559: 4558: 4556: 4555: 4550: 4544: 4542: 4538: 4537: 4535: 4534: 4528: 4526: 4522: 4521: 4519: 4518: 4513: 4508: 4502: 4500: 4496: 4495: 4493: 4492: 4487: 4482: 4477: 4475:Pinnacle Point 4472: 4467: 4462: 4457: 4452: 4447: 4442: 4437: 4432: 4427: 4422: 4417: 4413: 4412: 4407: 4402: 4397: 4392: 4387: 4382: 4377: 4372: 4367: 4362: 4357: 4352: 4347: 4342: 4337: 4331: 4329: 4325: 4324: 4322: 4321: 4316: 4311: 4305: 4303: 4299: 4298: 4296: 4295: 4289: 4287: 4283: 4282: 4280: 4279: 4277:The White Lady 4274: 4268: 4266: 4262: 4261: 4259: 4258: 4252: 4250: 4246: 4245: 4243: 4242: 4237: 4232: 4227: 4222: 4217: 4212: 4206: 4204: 4200: 4199: 4197: 4196: 4191: 4185: 4183: 4179: 4178: 4176: 4175: 4169: 4167: 4163: 4162: 4160: 4159: 4154: 4148: 4146: 4142: 4141: 4139: 4138: 4133: 4127: 4125: 4121: 4120: 4118: 4117: 4111: 4109: 4105: 4104: 4102: 4101: 4095: 4093: 4089: 4088: 4086: 4085: 4080: 4074: 4072: 4068: 4067: 4065: 4064: 4058: 4056: 4048: 4047: 4042: 4035: 4034: 4031: 4030: 4027: 4026: 4024: 4023: 4018: 4012: 4010: 4006: 4005: 4003: 4002: 3997: 3991: 3989: 3985: 3984: 3982: 3981: 3976: 3971: 3965: 3963: 3959: 3958: 3953: 3951: 3947: 3946: 3944: 3943: 3938: 3933: 3927: 3925: 3921: 3920: 3918: 3917: 3912: 3907: 3902: 3896: 3894: 3890: 3889: 3887: 3886: 3881: 3876: 3871: 3866: 3860: 3858: 3854: 3853: 3851: 3850: 3844: 3842: 3836: 3835: 3833: 3832: 3826: 3824: 3820: 3819: 3817: 3816: 3810: 3808: 3804: 3803: 3801: 3800: 3798:Khoit Tsenkher 3794: 3792: 3788: 3787: 3785: 3784: 3778: 3776: 3772: 3771: 3769: 3768: 3763: 3758: 3753: 3748: 3743: 3738: 3732: 3730: 3726: 3725: 3723: 3722: 3717: 3711: 3709: 3705: 3704: 3702: 3701: 3695: 3693: 3689: 3688: 3686: 3685: 3680: 3675: 3670: 3665: 3660: 3655: 3650: 3644: 3642: 3638: 3637: 3635: 3634: 3629: 3624: 3619: 3614: 3609: 3604: 3599: 3594: 3589: 3584: 3579: 3574: 3569: 3563: 3561: 3557: 3556: 3554: 3553: 3548: 3542: 3540: 3536: 3535: 3533: 3532: 3527: 3522: 3517: 3512: 3507: 3502: 3497: 3492: 3486: 3484: 3480: 3479: 3477: 3476: 3471: 3466: 3461: 3455: 3453: 3449: 3448: 3446: 3445: 3440: 3435: 3430: 3425: 3420: 3415: 3409: 3407: 3403: 3402: 3400: 3399: 3393: 3391: 3387: 3386: 3384: 3383: 3378: 3373: 3367: 3365: 3361: 3360: 3358: 3357: 3352: 3347: 3342: 3337: 3332: 3327: 3322: 3317: 3312: 3306: 3304: 3300: 3299: 3297: 3296: 3290: 3288: 3284: 3283: 3281: 3280: 3275: 3270: 3265: 3260: 3255: 3250: 3244: 3242: 3238: 3237: 3235: 3234: 3228: 3226: 3222: 3221: 3219: 3218: 3212: 3210: 3202: 3201: 3196: 3189: 3188: 3185: 3184: 3181: 3180: 3178: 3177: 3172: 3167: 3162: 3157: 3152: 3147: 3142: 3137: 3132: 3127: 3122: 3117: 3112: 3110:Creswell Crags 3107: 3102: 3097: 3092: 3090:Aveline's Hole 3086: 3084: 3083:United Kingdom 3080: 3079: 3077: 3076: 3070: 3068: 3064: 3063: 3061: 3060: 3055: 3049: 3047: 3043: 3042: 3040: 3039: 3034: 3032:Santa Catalina 3029: 3024: 3019: 3014: 3009: 3004: 2999: 2994: 2989: 2984: 2978: 2977: 2972: 2967: 2962: 2957: 2952: 2947: 2942: 2937: 2932: 2927: 2922: 2917: 2912: 2908: 2907: 2902: 2897: 2891: 2890: 2885: 2880: 2873: 2872: 2867: 2862: 2857: 2852: 2851: 2850: 2847: 2846: 2845: 2840: 2837: 2834: 2827:Monte Castillo 2824: 2821: 2816: 2813: 2803: 2798: 2792: 2790: 2786: 2785: 2783: 2782: 2777: 2772: 2767: 2761: 2759: 2755: 2754: 2752: 2751: 2746: 2741: 2735: 2733: 2729: 2728: 2726: 2725: 2720: 2715: 2710: 2708:Hadži-Prodan's 2704: 2702: 2698: 2697: 2695: 2694: 2689: 2684: 2679: 2674: 2669: 2664: 2658: 2656: 2652: 2651: 2649: 2648: 2643: 2638: 2633: 2627: 2625: 2621: 2620: 2618: 2617: 2612: 2607: 2602: 2597: 2592: 2587: 2582: 2576: 2574: 2570: 2569: 2567: 2566: 2561: 2555: 2553: 2549: 2548: 2546: 2545: 2539: 2537: 2533: 2532: 2530: 2529: 2523: 2521: 2515: 2514: 2512: 2511: 2505: 2503: 2499: 2498: 2496: 2495: 2489: 2487: 2483: 2482: 2480: 2479: 2473: 2471: 2467: 2466: 2464: 2463: 2457: 2455: 2449: 2448: 2446: 2445: 2440: 2435: 2430: 2423: 2418: 2413: 2408: 2403: 2398: 2392: 2390: 2386: 2385: 2383: 2382: 2377: 2371: 2369: 2365: 2364: 2362: 2361: 2356: 2351: 2346: 2341: 2336: 2331: 2326: 2321: 2316: 2310: 2308: 2304: 2303: 2301: 2300: 2295: 2290: 2284: 2282: 2276: 2275: 2273: 2272: 2267: 2262: 2257: 2252: 2247: 2243: 2242: 2237: 2232: 2227: 2222: 2217: 2212: 2206: 2204: 2200: 2199: 2197: 2196: 2191: 2186: 2181: 2176: 2171: 2166: 2161: 2156: 2151: 2146: 2141: 2136: 2131: 2128: 2123: 2118: 2115: 2112: 2109: 2106: 2101: 2096: 2090: 2089: 2084: 2079: 2074: 2069: 2064: 2059: 2054: 2049: 2044: 2039: 2034: 2029: 2024: 2019: 2014: 2009: 2004: 1999: 1994: 1990: 1989: 1984: 1978: 1977: 1972: 1967: 1962: 1959: 1954: 1951: 1946: 1945:Laugerie-Haute 1943: 1941:Laugerie-Basse 1938: 1933: 1928: 1923: 1922:Abri Chadourne 1920: 1917: 1912: 1911: 1910: 1905: 1899: 1894: 1891: 1888: 1882: 1880: 1876: 1875: 1873: 1872: 1866: 1864: 1860: 1859: 1857: 1856: 1851: 1846: 1841: 1836: 1831: 1825: 1823: 1822:Czech Republic 1819: 1818: 1816: 1815: 1809: 1807: 1803: 1802: 1800: 1799: 1794: 1792:Romuald's Cave 1789: 1784: 1779: 1774: 1768: 1766: 1762: 1761: 1759: 1758: 1753: 1748: 1743: 1738: 1732: 1730: 1726: 1725: 1723: 1722: 1717: 1711: 1709: 1705: 1704: 1702: 1701: 1696: 1691: 1686: 1683: 1678: 1673: 1668: 1663: 1660: 1656: 1654: 1650: 1649: 1647: 1646: 1641: 1636: 1631: 1626: 1620: 1618: 1610: 1609: 1604: 1597: 1596: 1594: 1593: 1588: 1586:Cave paintings 1583: 1577: 1574: 1573: 1570:cave paintings 1567: 1565: 1564: 1557: 1550: 1542: 1535: 1534: 1499: 1440: 1381: 1316: 1272: 1257: 1196: 1137: 1118:(5): 183–195. 1097: 1032: 1005: 991: 989::245 –253 969: 958:New York Times 939: 925: 923: 920: 919: 918: 913: 906: 903: 848: 845: 781: 778: 774:Northern China 734: 733: 718: 716: 712: 711: 704: 700: 699: 694: 690: 689: 684: 680: 679: 674: 670: 669: 664: 660: 659: 654: 650: 649: 644: 640: 639: 634: 630: 629: 624: 620: 619: 614: 610: 609: 604: 600: 599: 594: 590: 589: 576: 575: 571: 570: 562: 561: 552: 549: 548: 547: 546: 545: 535: 508:Ust'-Ishim man 489: 486:Basal Eurasian 477: 460: 459: 448: 447: 433: 424: 415: 406: 397: 388: 379: 370: 361: 360: 359: 352: 351: 350: 349: 337:archaic humans 328: 325: 286:human mandible 281:Ursus spelaeus 259: 256: 192: 191: 186: 182: 181: 177: 176: 169:45.017; 21.833 144: 140: 139: 130: 126: 125: 118: 114: 113: 103: 94: 93: 87: 86: 85: 84: 75: 66: 65: 59: 58: 57: 56: 55: 52: 51: 48: 40: 39: 36: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 5717: 5706: 5703: 5701: 5698: 5696: 5693: 5691: 5688: 5686: 5683: 5681: 5678: 5676: 5673: 5671: 5668: 5667: 5665: 5650: 5647: 5645: 5644:Cave painting 5642: 5640: 5639:List of caves 5637: 5634: 5633: 5630: 5616: 5613: 5612: 5610: 5606: 5600: 5597: 5596: 5594: 5590: 5584: 5581: 5580: 5578: 5574: 5568: 5565: 5564: 5562: 5560: 5556: 5550: 5547: 5545: 5542: 5541: 5539: 5535: 5529: 5526: 5525: 5523: 5521: 5520:New Caledonia 5517: 5511: 5508: 5507: 5505: 5501: 5495: 5492: 5490: 5487: 5485: 5482: 5481: 5479: 5477: 5473: 5467: 5464: 5462: 5459: 5457: 5454: 5452: 5449: 5447: 5444: 5442: 5439: 5437: 5434: 5432: 5429: 5427: 5426:New Guinea II 5424: 5422: 5419: 5417: 5414: 5412: 5409: 5407: 5404: 5402: 5401:Mannalargenna 5399: 5397: 5394: 5392: 5389: 5387: 5384: 5382: 5379: 5377: 5374: 5372: 5369: 5367: 5364: 5362: 5359: 5357: 5354: 5352: 5349: 5347: 5344: 5342: 5339: 5337: 5334: 5332: 5329: 5327: 5324: 5322: 5319: 5317: 5314: 5312: 5309: 5307: 5304: 5302: 5299: 5297: 5294: 5292: 5289: 5287: 5284: 5283: 5281: 5277: 5273: 5269: 5264: 5260: 5246: 5243: 5241: 5238: 5236: 5233: 5231: 5228: 5226: 5223: 5221: 5218: 5216: 5213: 5211: 5208: 5206: 5203: 5201: 5198: 5196: 5193: 5191: 5188: 5186: 5183: 5181: 5178: 5176: 5173: 5171: 5170:On Your Knees 5168: 5166: 5163: 5161: 5158: 5156: 5153: 5151: 5148: 5146: 5143: 5141: 5138: 5136: 5133: 5131: 5128: 5126: 5123: 5121: 5118: 5116: 5113: 5111: 5108: 5106: 5103: 5101: 5098: 5096: 5095:Hidden Valley 5093: 5091: 5088: 5086: 5083: 5081: 5078: 5076: 5073: 5071: 5068: 5066: 5063: 5061: 5058: 5056: 5053: 5051: 5048: 5046: 5043: 5041: 5038: 5036: 5033: 5031: 5028: 5026: 5023: 5021: 5018: 5016: 5013: 5011: 5008: 5006: 5003: 5002: 5000: 4998:United States 4996: 4990: 4987: 4986: 4984: 4980: 4974: 4971: 4969: 4966: 4964: 4961: 4959: 4956: 4954: 4951: 4949: 4946: 4945: 4943: 4939: 4933: 4930: 4928: 4925: 4923: 4920: 4918: 4917:Guilá Naquitz 4915: 4913: 4910: 4908: 4905: 4903: 4902:Dzibilchaltún 4900: 4898: 4895: 4893: 4890: 4889: 4887: 4883: 4877: 4874: 4873: 4871: 4867: 4861: 4858: 4857: 4855: 4851: 4845: 4842: 4841: 4839: 4837: 4833: 4827: 4824: 4822: 4819: 4817: 4814: 4812: 4809: 4807: 4804: 4802: 4799: 4798: 4796: 4792: 4786: 4783: 4781: 4778: 4776: 4773: 4771: 4768: 4766: 4763: 4761: 4758: 4757: 4755: 4751: 4745: 4742: 4740: 4737: 4736: 4734: 4730: 4724: 4721: 4719: 4716: 4715: 4713: 4709: 4703: 4700: 4698: 4695: 4693: 4690: 4688: 4685: 4683: 4682:Pedra Pintada 4680: 4678: 4675: 4673: 4670: 4669: 4667: 4663: 4657: 4654: 4652: 4649: 4647: 4644: 4643: 4641: 4637: 4631: 4628: 4627: 4625: 4621: 4615: 4612: 4611: 4609: 4605: 4601: 4597: 4596:South America 4593: 4588: 4584: 4570: 4567: 4566: 4564: 4560: 4554: 4551: 4549: 4546: 4545: 4543: 4539: 4533: 4530: 4529: 4527: 4523: 4517: 4514: 4512: 4509: 4507: 4504: 4503: 4501: 4497: 4491: 4488: 4486: 4483: 4481: 4478: 4476: 4473: 4471: 4468: 4466: 4463: 4461: 4460:Klasies River 4458: 4456: 4453: 4451: 4448: 4446: 4443: 4441: 4438: 4436: 4435:Byneskranskop 4433: 4431: 4428: 4426: 4423: 4421: 4418: 4415: 4414: 4411: 4408: 4406: 4403: 4401: 4398: 4396: 4393: 4391: 4388: 4386: 4383: 4381: 4378: 4376: 4373: 4371: 4368: 4366: 4363: 4361: 4358: 4356: 4353: 4351: 4348: 4346: 4343: 4341: 4338: 4336: 4333: 4332: 4330: 4326: 4320: 4317: 4315: 4312: 4310: 4307: 4306: 4304: 4300: 4294: 4291: 4290: 4288: 4284: 4278: 4275: 4273: 4270: 4269: 4267: 4263: 4257: 4254: 4253: 4251: 4247: 4241: 4238: 4236: 4233: 4231: 4228: 4226: 4225:Ifri Oudadane 4223: 4221: 4218: 4216: 4213: 4211: 4208: 4207: 4205: 4201: 4195: 4194:Uan Muhuggiag 4192: 4190: 4187: 4186: 4184: 4180: 4174: 4171: 4170: 4168: 4164: 4158: 4155: 4153: 4150: 4149: 4147: 4143: 4137: 4134: 4132: 4129: 4128: 4126: 4122: 4116: 4113: 4112: 4110: 4106: 4100: 4097: 4096: 4094: 4090: 4084: 4081: 4079: 4076: 4075: 4073: 4069: 4063: 4060: 4059: 4057: 4053: 4049: 4045: 4040: 4036: 4022: 4019: 4017: 4014: 4013: 4011: 4007: 4001: 3998: 3996: 3993: 3992: 3990: 3986: 3980: 3977: 3975: 3972: 3970: 3967: 3966: 3964: 3960: 3956: 3952: 3948: 3942: 3939: 3937: 3934: 3932: 3929: 3928: 3926: 3922: 3916: 3913: 3911: 3908: 3906: 3903: 3901: 3900:Batadombalena 3898: 3897: 3895: 3891: 3885: 3882: 3880: 3877: 3875: 3872: 3870: 3867: 3865: 3862: 3861: 3859: 3855: 3849: 3846: 3845: 3843: 3841: 3837: 3831: 3828: 3827: 3825: 3821: 3815: 3812: 3811: 3809: 3805: 3799: 3796: 3795: 3793: 3789: 3783: 3780: 3779: 3777: 3773: 3767: 3764: 3762: 3759: 3757: 3756:Ras Baalbek I 3754: 3752: 3749: 3747: 3744: 3742: 3739: 3737: 3734: 3733: 3731: 3727: 3721: 3718: 3716: 3713: 3712: 3710: 3706: 3700: 3697: 3696: 3694: 3690: 3684: 3681: 3679: 3676: 3674: 3671: 3669: 3666: 3664: 3661: 3659: 3656: 3654: 3651: 3649: 3646: 3645: 3643: 3639: 3633: 3630: 3628: 3625: 3623: 3620: 3618: 3615: 3613: 3610: 3608: 3605: 3603: 3600: 3598: 3595: 3593: 3592:Nahal Me'arot 3590: 3588: 3585: 3583: 3580: 3578: 3575: 3573: 3570: 3568: 3565: 3564: 3562: 3558: 3552: 3549: 3547: 3544: 3543: 3541: 3537: 3531: 3528: 3526: 3523: 3521: 3518: 3516: 3513: 3511: 3508: 3506: 3503: 3501: 3498: 3496: 3493: 3491: 3488: 3487: 3485: 3481: 3475: 3472: 3470: 3467: 3465: 3462: 3460: 3457: 3456: 3454: 3450: 3444: 3441: 3439: 3436: 3434: 3431: 3429: 3426: 3424: 3421: 3419: 3416: 3414: 3411: 3410: 3408: 3404: 3398: 3395: 3394: 3392: 3388: 3382: 3379: 3377: 3374: 3372: 3369: 3368: 3366: 3362: 3356: 3353: 3351: 3348: 3346: 3343: 3341: 3338: 3336: 3333: 3331: 3328: 3326: 3323: 3321: 3318: 3316: 3313: 3311: 3308: 3307: 3305: 3301: 3295: 3292: 3291: 3289: 3285: 3279: 3276: 3274: 3271: 3269: 3266: 3264: 3261: 3259: 3256: 3254: 3251: 3249: 3246: 3245: 3243: 3239: 3233: 3230: 3229: 3227: 3223: 3217: 3214: 3213: 3211: 3207: 3203: 3199: 3194: 3190: 3176: 3175:Windmill Hill 3173: 3171: 3168: 3166: 3163: 3161: 3158: 3156: 3153: 3151: 3148: 3146: 3143: 3141: 3138: 3136: 3133: 3131: 3128: 3126: 3123: 3121: 3120:Heathery Burn 3118: 3116: 3113: 3111: 3108: 3106: 3103: 3101: 3098: 3096: 3093: 3091: 3088: 3087: 3085: 3081: 3075: 3072: 3071: 3069: 3065: 3059: 3056: 3054: 3051: 3050: 3048: 3044: 3038: 3035: 3033: 3030: 3028: 3025: 3023: 3020: 3018: 3015: 3013: 3010: 3008: 3005: 3003: 3000: 2998: 2995: 2993: 2990: 2988: 2985: 2983: 2980: 2979: 2976: 2973: 2971: 2968: 2966: 2963: 2961: 2958: 2956: 2953: 2951: 2948: 2946: 2943: 2941: 2938: 2936: 2933: 2931: 2928: 2926: 2923: 2921: 2918: 2916: 2913: 2910: 2909: 2906: 2903: 2901: 2898: 2896: 2893: 2892: 2889: 2886: 2884: 2881: 2878: 2875: 2874: 2871: 2868: 2866: 2863: 2861: 2858: 2856: 2855:Tito Bustillo 2853: 2848: 2844: 2841: 2838: 2836:Las Chimeneas 2835: 2833: 2830: 2829: 2828: 2825: 2822: 2820: 2817: 2814: 2812: 2809: 2808: 2807: 2804: 2802: 2799: 2797: 2794: 2793: 2791: 2787: 2781: 2778: 2776: 2773: 2771: 2768: 2766: 2763: 2762: 2760: 2756: 2750: 2747: 2745: 2742: 2740: 2737: 2736: 2734: 2730: 2724: 2721: 2719: 2716: 2714: 2711: 2709: 2706: 2705: 2703: 2699: 2693: 2690: 2688: 2685: 2683: 2680: 2678: 2675: 2673: 2670: 2668: 2665: 2663: 2662:Akhshtyrskaya 2660: 2659: 2657: 2653: 2647: 2644: 2642: 2639: 2637: 2634: 2632: 2629: 2628: 2626: 2622: 2616: 2613: 2611: 2610:Pego do Diabo 2608: 2606: 2603: 2601: 2598: 2596: 2593: 2591: 2588: 2586: 2583: 2581: 2578: 2577: 2575: 2571: 2565: 2562: 2560: 2557: 2556: 2554: 2550: 2544: 2541: 2540: 2538: 2534: 2528: 2525: 2524: 2522: 2520: 2516: 2510: 2507: 2506: 2504: 2500: 2494: 2491: 2490: 2488: 2484: 2478: 2475: 2474: 2472: 2468: 2462: 2459: 2458: 2456: 2454: 2450: 2444: 2441: 2439: 2436: 2434: 2431: 2429: 2428: 2424: 2422: 2419: 2417: 2414: 2412: 2409: 2407: 2404: 2402: 2401:Arene Candide 2399: 2397: 2394: 2393: 2391: 2387: 2381: 2378: 2376: 2373: 2372: 2370: 2366: 2360: 2357: 2355: 2352: 2350: 2347: 2345: 2342: 2340: 2337: 2335: 2332: 2330: 2327: 2325: 2322: 2320: 2317: 2315: 2312: 2311: 2309: 2305: 2299: 2296: 2294: 2291: 2289: 2288:Devil's Tower 2286: 2285: 2283: 2281: 2277: 2271: 2268: 2266: 2263: 2261: 2258: 2256: 2253: 2251: 2248: 2245: 2244: 2241: 2238: 2236: 2233: 2231: 2228: 2226: 2223: 2221: 2218: 2216: 2213: 2211: 2208: 2207: 2205: 2201: 2195: 2192: 2190: 2187: 2185: 2182: 2180: 2177: 2175: 2172: 2170: 2167: 2165: 2162: 2160: 2157: 2155: 2152: 2150: 2147: 2145: 2142: 2140: 2137: 2135: 2132: 2129: 2127: 2124: 2122: 2119: 2116: 2113: 2110: 2107: 2105: 2102: 2100: 2097: 2095: 2092: 2091: 2088: 2085: 2083: 2080: 2078: 2075: 2073: 2070: 2068: 2065: 2063: 2062:Pair-non-Pair 2060: 2058: 2055: 2053: 2050: 2048: 2047:Le Mas-d'Azil 2045: 2043: 2040: 2038: 2035: 2033: 2030: 2028: 2025: 2023: 2020: 2018: 2015: 2013: 2010: 2008: 2005: 2003: 2000: 1998: 1997:Arcy-sur-Cure 1995: 1992: 1991: 1988: 1985: 1983: 1980: 1979: 1976: 1973: 1971: 1968: 1966: 1963: 1960: 1958: 1955: 1952: 1950: 1947: 1944: 1942: 1939: 1937: 1936:Font-de-Gaume 1934: 1932: 1929: 1927: 1924: 1921: 1918: 1916: 1913: 1909: 1906: 1904:Abri Castanet 1903: 1902: 1901:Castel Merle 1900: 1898: 1895: 1892: 1889: 1887: 1884: 1883: 1881: 1877: 1871: 1868: 1867: 1865: 1861: 1855: 1852: 1850: 1847: 1845: 1842: 1840: 1837: 1835: 1832: 1830: 1827: 1826: 1824: 1820: 1814: 1811: 1810: 1808: 1804: 1798: 1795: 1793: 1790: 1788: 1785: 1783: 1780: 1778: 1775: 1773: 1770: 1769: 1767: 1763: 1757: 1754: 1752: 1749: 1747: 1744: 1742: 1739: 1737: 1734: 1733: 1731: 1727: 1721: 1718: 1716: 1713: 1712: 1710: 1706: 1700: 1697: 1695: 1692: 1690: 1687: 1684: 1682: 1679: 1677: 1674: 1672: 1669: 1667: 1664: 1661: 1658: 1657: 1655: 1651: 1645: 1642: 1640: 1637: 1635: 1632: 1630: 1627: 1625: 1622: 1621: 1619: 1615: 1611: 1607: 1602: 1598: 1592: 1589: 1587: 1584: 1582: 1579: 1578: 1575: 1571: 1563: 1558: 1556: 1551: 1549: 1544: 1543: 1540: 1530: 1526: 1522: 1518: 1514: 1510: 1503: 1500: 1495: 1491: 1486: 1481: 1476: 1471: 1467: 1463: 1459: 1455: 1451: 1444: 1441: 1436: 1432: 1428: 1424: 1420: 1416: 1412: 1408: 1404: 1400: 1396: 1392: 1385: 1382: 1377: 1371: 1363: 1359: 1355: 1351: 1347: 1343: 1339: 1335: 1331: 1327: 1320: 1317: 1313: 1308: 1302: 1298: 1294: 1290: 1283: 1276: 1273: 1269: 1268: 1261: 1258: 1253: 1249: 1244: 1239: 1235: 1231: 1227: 1223: 1219: 1215: 1211: 1203: 1201: 1197: 1192: 1188: 1183: 1178: 1173: 1168: 1164: 1160: 1156: 1152: 1148: 1141: 1138: 1133: 1129: 1125: 1121: 1117: 1113: 1106: 1104: 1102: 1098: 1093: 1089: 1084: 1079: 1075: 1071: 1067: 1063: 1059: 1055: 1051: 1047: 1043: 1036: 1033: 1020: 1016: 1009: 1006: 1002: 995: 992: 988: 984: 978: 976: 974: 970: 959: 955: 948: 946: 944: 940: 936: 930: 927: 921: 917: 914: 912: 909: 908: 904: 901: 899: 892: 888: 884: 882: 878: 877:Arcy-sur-Cure 874: 870: 864: 862: 857: 854: 846: 844: 842: 841:Erik Trinkaus 836: 834: 830: 826: 822: 818: 815: 811: 807: 803: 799: 795: 791: 787: 779: 777: 775: 771: 767: 763: 759: 754: 751: 747: 745: 741: 731: 730: 729: 724: 723: 717: 714: 713: 710: 709: 705: 702: 701: 698: 695: 692: 691: 688: 685: 682: 681: 678: 675: 672: 671: 668: 665: 662: 661: 658: 655: 652: 651: 648: 645: 642: 641: 638: 635: 632: 631: 628: 625: 622: 621: 618: 615: 612: 611: 608: 605: 602: 601: 598: 595: 592: 591: 586: 581: 577: 572: 568: 563: 558: 555: 550: 543: 540: 536: 533: 529: 525: 521: 517: 513: 512:Haplogroup NO 509: 505: 501: 500: 498: 494: 490: 487: 482: 478: 475: 471: 470:introgression 467: 466: 465: 441: 430: 421: 412: 403: 394: 385: 376: 367: 356: 348: 347: 344: 342: 338: 333: 326: 324: 322: 318: 310: 305: 302: 301:Ion din Anina 296: 291: 287: 283: 282: 277: 273: 269: 265: 264:speleological 257: 255: 253: 250:had a recent 249: 246: 241: 239: 235: 229: 227: 223: 219: 218:Caraș-Severin 215: 211: 206: 198: 190: 187: 183: 178: 173: 145: 141: 138: 134: 133:Caraș-Severin 131: 127: 123: 119: 115: 91: 63: 53: 46: 41: 34: 19: 5451:Wargata Mina 5411:Murrawijinie 5346:Eagles Reach 5341:Devil's Lair 5245:Wilson Butte 5030:Cherry Creek 5020:Bull Thistle 4765:Chiribiquete 4723:Charlie Lake 4651:Barton Creek 4465:Melkhoutboom 4405:Sterkfontein 4395:Plovers Lake 4328:South Africa 4230:Jebel Irhoud 4215:Ifri N'Ammar 3950:Turkmenistan 3915:Hunugalagala 3699:Iraq ed-Dubb 3464:Jeriji Saléh 3125:High Pasture 2682:Shulgan-Tash 2645: 2605:Pedra Furada 2426: 2265:Lichtenstein 2255:Brillenhöhle 2184:Le Regourdou 2144:La Ferrassie 2139:Combe Grenal 2082:Trois Frères 2017:Fontéchevade 1970:La Madeleine 1624:Drachenhöhle 1512: 1508: 1502: 1457: 1453: 1443: 1394: 1390: 1384: 1370:cite journal 1329: 1325: 1319: 1292: 1288: 1275: 1266: 1260: 1217: 1213: 1154: 1150: 1140: 1115: 1111: 1049: 1045: 1035: 1023:. Retrieved 1021:. Genome Web 1018: 1008: 1000: 994: 986: 982: 961:. Retrieved 957: 934: 929: 894: 889: 885: 869:Vindija Cave 865: 858: 850: 837: 827:and ancient 783: 770:Tianyuan man 755: 752: 748: 744:petrous bone 737: 726: 720: 719: 707: 653:Infraorder: 554: 472:found in an 463: 339:, including 334: 330: 306: 290:paleosurface 279: 261: 242: 230: 196: 195: 49:Oase 2 skull 5537:New Zealand 5361:Gwion Gwion 5230:Trail Creek 5155:Meadowcroft 5115:Last Supper 5045:Daugherty's 4907:Las Flechas 4692:Rei do Mato 4416:Other caves 4400:Rising Star 4375:Makapansgat 4340:Bolt's Farm 4157:Njoro River 4000:Teshik-Tash 3995:Obi-Rakhmat 3979:Yarımburgaz 3879:Lapuz Lapuz 3857:Philippines 3766:Nachcharini 3761:Ras El Kelb 3720:Tam Pa Ling 3715:Tham An Mah 3597:Nahal Hemar 3345:Zhoukoudian 3320:Guanyindong 3294:Laang Spean 3216:Darra-e Kur 3209:Afghanistan 3155:Portbraddon 3145:Marble Arch 3058:Wildkirchli 3046:Switzerland 2982:Other caves 2955:Ojo Guareña 2935:Maltravieso 2930:Los Casares 2905:Siega Verde 2870:Los Aviones 2865:Santimamiñe 2839:Las Monedas 2832:El Castillo 2739:Čertova pec 2687:Mezmaiskaya 2595:Lagar Velho 2527:Cocev Kamen 2406:Balzi Rossi 2246:Other caves 2235:Sirgenstein 2164:Le Moustier 2154:Fontbrégoua 2114:Bonne-Femme 2094:Other caves 1813:Aetokremnos 1756:Utroba Cave 1662:Claminforge 1659:Belle-Roche 1515:: 207–230. 817:radiocarbon 673:Subfamily: 657:Simiiformes 341:Neanderthal 307:The latest 278:cave bear ( 252:Neanderthal 189:Paleolithic 167: / 143:Coordinates 5664:Categories 5649:Speleology 5466:Wellington 5421:Naracoorte 5406:Mudgegonga 5391:Madjedbebe 5356:Gabarnmung 5296:Ballawinne 5286:Abrakurrie 5185:Pictograph 4963:Qillqatani 4958:Pikimachay 4953:Guitarrero 4948:Chivateros 4927:Oxtotitlán 4780:Tequendama 4490:Wonderwerk 4470:Nelson Bay 4450:Elands Bay 4410:Swartkrans 4355:Gladysvale 4302:Somaliland 4249:Mozambique 4189:Haua Fteah 3988:Uzbekistan 3653:Minatogawa 3648:Kosegasawa 3602:Nahal Oren 3546:Hazar Merd 3515:Qaleh Bozi 3500:Do-Ashkaft 3397:Satsurblia 3364:East Timor 3241:Azerbaijan 3130:Kendrick's 3095:Bontnewydd 3017:Don Gaspar 2970:Praileaitz 2925:Las Caldas 2843:La Pasiega 2770:Divje Babe 2692:Okladnikov 2677:Ignatievka 2600:Pala Pinta 2509:Għar Dalam 2486:Luxembourg 2461:St Brelade 2443:Villabruna 2354:Stravomyti 2344:Megalakkos 2329:Eileithyia 2314:Alepotrypa 2225:Hohle Fels 2111:Balauzière 2067:Pech Merle 1975:Rouffignac 1949:La Micoque 1931:Cro-Magnon 1890:Bara Bahau 1834:Býčí skála 1782:Vela Spila 1741:Devetashka 1736:Bacho Kiro 1295:(Thesis). 963:2012-04-19 922:References 786:Speleology 728:H. sapiens 693:Subtribe: 647:Haplorhini 643:Suborder: 481:East Asian 419:Bacho Kiro 401:Ust'-Ishim 343:features. 317:Iron Gates 5700:Cave bear 5615:Nanumanga 5599:Falemauga 5510:Makauwahi 5279:Australia 5235:Trinchera 5110:La Grange 5080:Gatecliff 5075:Franktown 5070:Fort Rock 5050:Dry Creek 4968:Toquepala 4932:Sac Actun 4922:Ox Bel Ha 4912:Frightful 4897:Coxcatlan 4876:Long Mile 4630:Quadiriki 4607:Argentina 4445:Diepkloof 4430:Boomplaas 4385:Minnaar's 4370:Kromdraai 4319:Laas Geel 4314:Dhaymoole 4309:Dhambalin 4272:Apollo 11 4173:Liphofung 4099:Shum Laka 4062:Gueldaman 4016:Con Moong 3893:Sri Lanka 3840:Palestine 3814:Padah-Lin 3751:Ksar Akil 3683:Yamashita 3668:Pinza-Abu 3474:Pettakere 3469:Liang Bua 3452:Indonesia 3438:Pahargarh 3423:Bhimbetka 3381:Lene Hara 3371:Jerimalai 3350:Zengpiyan 3340:Yuchanyan 3140:Long Hole 3037:del Valle 3012:dels Bous 2965:la Pileta 2900:Atapuerca 2815:Covalanas 2713:Pešturina 2641:Muierilor 2631:Coliboaia 2543:Svarthola 2359:Theopetra 2349:Petralona 2334:Franchthi 2324:Asphendou 2280:Gibraltar 2250:Baumann's 2240:Vogelherd 2215:Bockstein 2194:Vallonnet 2189:Rochereil 2179:Raymonden 2169:Noisetier 2121:Bruniquel 2042:Marsoulas 2032:Lombrives 2027:La Marche 1953:La Mouthe 1919:Abri Audi 1897:Cap Blanc 1839:Koněprusy 1829:Amatérská 1797:Veternica 1746:Kozarnika 1720:Ledenjača 1644:Tischofer 1634:Lurgrotte 1419:0028-0836 1346:0028-0836 1074:0028-0836 715:Species: 677:Homininae 667:Hominidae 603:Kingdom: 597:Eukaryota 392:Zlatý kůň 216:, in the 5456:Warratyi 5441:Tarragal 5431:Nunamira 5416:Murujuga 5381:Kutikina 5376:Koonalda 5371:Koongine 5331:Cliefden 5326:Cave Bay 5225:Tomaquag 5210:Sisyphus 5140:Mantle's 5130:Lovelock 5125:LoDaisKa 5105:Humboldt 4989:Werehpai 4982:Suriname 4892:Chan Hol 4811:Centella 4801:Ambrosio 4753:Colombia 4718:Bluefish 4562:Zimbabwe 4499:Tanzania 4485:Stadsaal 4390:Motsetsi 4360:Gondolin 4350:Drimolen 4345:Cooper's 4240:Taforalt 4210:Hercules 4136:Swimmers 4108:DR Congo 4092:Cameroon 4071:Botswana 3941:Tham Lod 3931:Pha Taem 3924:Thailand 3905:Belilena 3823:Pakistan 3791:Mongolia 3775:Malaysia 3736:Antelias 3678:Tobayama 3632:Zuttiyeh 3551:Shanidar 3413:Ambadevi 3330:Tianyuan 3310:Baishiya 3287:Cambodia 3027:El Mirón 3022:Guanches 2997:Armintxe 2915:Bacinete 2860:Altxerri 2849:El Pendo 2819:La Garma 2801:Altamira 2758:Slovenia 2749:Jasovská 2732:Slovakia 2718:Risovača 2672:Denisova 2636:Cuciulat 2590:Furninha 2585:Escoural 2573:Portugal 2564:Paradise 2559:Obłazowa 2493:Loschbur 2433:Paglicci 2416:Corbeddu 2298:Vanguard 2293:Gorham's 2174:La Quina 2117:Bouillon 2104:Aurignac 1908:Reverdit 1893:Bernifal 1772:Grapčeva 1729:Bulgaria 1689:Scladina 1676:Naulette 1639:Salzofen 1494:17085588 1427:16495989 1354:11484052 1314:license. 1252:26098372 1191:14504393 1132:18565967 1092:26098372 905:See also 861:Holocene 740:coverage 697:Hominina 687:Hominini 663:Family: 637:Primates 627:Mammalia 617:Chordata 613:Phylum: 607:Animalia 593:Domain: 524:subclade 428:Tianyuan 374:Kostenki 258:The cave 117:Location 5567:Chugai' 5549:Ruakuri 5484:Gadao's 5461:Warreen 5396:Mammoth 5366:Jenolan 5311:Blanche 5291:Acheron 5268:Oceania 5240:Ventana 5220:Tainter 5195:Russell 5180:Pendejo 5175:Paisley 5135:Mammoth 5055:Durango 5015:Bonfire 4869:Jamaica 4836:Curaçao 4775:Sáchica 4760:El Abra 4744:Milodón 4687:Peruaçu 4677:Maquiné 4569:Bambata 4548:Kalemba 4420:Blombos 4365:Haasgat 4286:Nigeria 4265:Namibia 4203:Morocco 4166:Lesotho 4078:Tsodilo 4055:Algeria 4009:Vietnam 3969:Belbaşı 3955:Dzhebel 3910:Fa Hien 3869:Kalanay 3830:Sanghao 3807:Myanmar 3746:Kaukaba 3729:Lebanon 3617:Raqefet 3587:Misliya 3572:HaYonim 3520:Warwasi 3495:Darband 3490:Bisitun 3459:Harimau 3433:Gudiyam 3428:Edakkal 3390:Georgia 3335:Xianren 3263:Damjili 3258:Buzeyir 3232:Areni-1 3225:Armenia 3150:Oldbury 3115:Gough's 3100:Cathole 3074:Verteba 3067:Ukraine 2624:Romania 2615:Salemas 2580:Aroeira 2411:Cavallo 2396:Addaura 2375:Baradla 2368:Hungary 2319:Apidima 2203:Germany 2159:Lazaret 2087:Villars 2007:Cosquer 1987:Chauvet 1965:Lascaux 1863:Finland 1787:Vindija 1777:Krapina 1765:Croatia 1685:Ramioul 1653:Belgium 1629:Gudenus 1617:Austria 1529:9039428 1485:1859909 1462:Bibcode 1435:4416359 1399:Bibcode 1362:4394515 1243:4537386 1222:Bibcode 1159:Bibcode 1083:4537386 1054:Bibcode 873:Croatia 703:Genus: 683:Tribe: 633:Order: 623:Class: 560:Oase 2 319:in the 288:on the 222:Romania 210:karstic 185:Periods 180:History 155:21°50′E 152:45°01′N 137:Romania 5608:Tuvalu 5544:Moncks 5528:Pindai 5503:Hawaii 5494:Talagi 5489:Mahlac 5351:Fossil 5336:Cloggs 5321:Burrup 5200:Sandia 5145:Marmes 5090:Hidden 5085:Graham 5040:Danger 5025:Burnet 4885:Mexico 4860:Pomier 4821:Patana 4806:Calero 4785:Tibitó 4711:Canada 4672:Gentio 4665:Brazil 4639:Belize 4553:Mumbwa 4541:Zambia 4525:Uganda 4506:Kondoa 4480:Sibudu 4425:Border 4380:Malapa 4256:Ngalue 4131:Beasts 4115:Matupi 4044:Africa 3974:Karain 3962:Turkey 3936:Spirit 3874:Minori 3864:Callao 3848:Shuqba 3692:Jordan 3663:Ōzakai 3658:Muroya 3607:Qafzeh 3577:Kebara 3560:Israel 3530:Yafteh 3525:Wezmeh 3355:Zhiren 3268:Tağlar 3170:Thor's 3105:Coygan 3053:Bichon 3007:Bedmar 2975:Sidrón 2811:Chufín 2744:Domica 2701:Serbia 2655:Russia 2552:Poland 2536:Norway 2470:Kosovo 2453:Jersey 2438:Romito 2380:Szelim 2339:Kleidi 2307:Greece 2012:Cussac 2002:Gargas 1957:Pataud 1879:France 1849:Mladeč 1806:Cyprus 1751:Magura 1715:Badanj 1708:Bosnia 1606:Europe 1527:  1492:  1482:  1433:  1425:  1417:  1391:Nature 1360:  1352:  1344:  1326:Nature 1250:  1240:  1214:Nature 1189:  1182:208740 1179:  1130:  1090:  1080:  1072:  1046:Nature 935:Nature 881:France 758:Asians 551:Oase 2 516:Telugu 383:Sungir 327:Oase 1 248:fossil 245:Oase 1 238:Mladeč 129:Region 5592:Samoa 5446:Ubirr 5436:ORS 7 5205:Shoup 5165:Mummy 5160:Modoc 5150:Martz 5100:Hogup 5010:Baker 4826:Pluma 4732:Chile 4697:Santo 4623:Aruba 4592:North 4532:Nyero 4516:Mumba 4440:Cango 4182:Libya 4145:Kenya 4124:Egypt 4021:Ngườm 3884:Tabon 3741:Jeita 3641:Japan 3627:Tabun 3622:Skhul 3612:Qesem 3582:Manot 3443:Tikla 3418:Belum 3406:India 3376:Laili 3325:Luobi 3315:Fuyan 3303:China 3278:Gazma 3253:Azykh 3248:Allar 3135:Kents 3002:Axlor 2992:Antón 2987:Ángel 2945:Nerja 2883:Araña 2789:Spain 2780:Potok 2775:Pekel 2765:Betal 2502:Malta 2427:Felci 2389:Italy 2270:Ofnet 2130:Cauna 2126:Calès 2099:Arago 2077:Renne 2057:Niaux 1854:Šipka 1844:Kůlna 1671:Goyet 1666:Engis 1525:S2CID 1431:S2CID 1358:S2CID 1285:(PDF) 1128:S2CID 1025:9 May 898:mtDNA 772:from 520:Malay 365:Goyet 272:Anina 270:near 214:Anina 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Index

Peştera cu Oase
Peștera cu Oase, Oase 2
Peștera cu Oase Cave Romania
Peștera cu Oase Cave Romania
Anina
Caraș-Severin
Romania
45°01′N 21°50′E / 45.017°N 21.833°E / 45.017; 21.833
Paleolithic
[ˈpeʃterakuˈo̯ase]
karstic
Anina
Caraș-Severin
Romania
European early modern human
Abrigo do Lagar Velho
Mladeč
Oase 1
fossil
Neanderthal
speleological
Carpathian Mountains
Anina
Late Pleistocene
Ursus spelaeus
human mandible
paleosurface
radiocarbon dates
Iron Gates
Danubian corridor

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