3245:
5377:
3238:
543:
1742:
2567:
2553:
877:
4829:
4033:
4815:
2546:
5525:
4822:
719:) would be a functional counterpart of the cutting edge of a beak. Thus, increasing the number of teeth would be an adaptation for cutting and ripping, as long as the space between adjacent teeth was preserved...while it would have the effect of working as a beak if spaces were filled with more teeth. The adaption to a cut-and-rip function therefore becomes an exaptation with a slicing effect, eventually leading to the cutting edge seen in most ornithomimosaurs."
1748:
3252:
150:
2560:
129:
5390:
4051:
4039:
4058:
5384:
4809:
508:
600:
4046:
539:, showing two different basic forms. The teeth in the front of the upper jaw were broad and D-shaped in cross-section, while those further back were blade-like, and on the whole the teeth in the upper jaw were larger than those in the lower. All of its teeth were unserrated, and had a constricted "waist" between the crown and the root. Interdental plates were lacking.
5537:
699:, "The phylogenetic hypothesis...supports an unexpected approach, involving exaptation, which might explain the evolutionary process towards the toothless condition in Ornithomimosauria. Until now, a progressive reduction in the number of teeth has been considered as the most likely explanation: the primitive
627:
might have been much like a modern-day crane, wading out in lakes or ponds using its claws and teeth to capture fish and then storing them in its skin flap. Some parts of the impressions revealed wrinkled skin, interpreted as lacking scales or feathers. Filament-like structures were also preserved;
522:
was a small ornithomimosaur, at about 1.9–2.5 m (6.2–8.2 ft) and 17–30 kg (37–66 lb). Its skull was unusually long and narrow, with a maximum length of about 4.5 times its maximum height. It was highly unusual among ornithomimosaurs in its large number of teeth: it had about 220
707:
hypothesis suggests an alternative evolutionary process based on a functional analysis of increasing numbers of teeth. A high number of teeth with enough interdental space and properly placed
615:
Soft-tissue remains preserved by the exceptional preservational environment of the La Hoyas lagerstätte revealed the presence of a small skin or keratin crest on the back of the head, and a
1114:
D. E. G. Briggs, P. R. Wilby, B. Pérez Pérez-Moreno, J. L. Sanz, M. Fregenal-Martinez, (1997). "The mineralization of dinosaur soft tissue in the Lower
Cretaceous of Las Hoyas, Spain."
491:. The only known specimen consists of the articulated front half of a skeleton and includes the skull, lower jaws, all the neck vertebrae and most of the back vertebrae, ribs,
499:, a complete right forelimb and most of the left forelimb. Remains of the soft parts are visible at the back of the skull, around the neck and around the front limbs.
5567:
5469:
596:
bones in the lower arm tightly adhered to each other. The hand was hook-like and had fingers of equal length equipped with rather straight claws.
3244:
5587:
3237:
1151:
Makovicky, P., Kobayashi, Y. and Currie, P. J. (2004). "Ornithomimosauria." pp. 137-150 in D. B. Weishampel, P. Dodson, et H. Osmólska (eds.),
5376:
1074:
683:. A study by Kobayashi and Lü in 2003 indicated that these two species formed a basal arrangement of steps leading towards the more advanced
973:
484:
1204:
5582:
2566:
715:) would be an adaptation for cutting and ripping. On the other hand, an excessive number of teeth with no interdental space (as in
652:
plates and uncinate processes, which imply flight musculature. These adaptations have been noted years later by the paleontologist
4032:
2552:
4814:
703:
theropods have up to 80 teeth with tall blade-like crowns, and the primitive ornithomimosaurs have only a few small teeth. The
1741:
1167:
Kobayashi, Y., and Lü, J.-C. (2003). "A new ornithomimid dinosaur with gregarious habits from the Late
Cretaceous of China."
2545:
4828:
1747:
5255:
4050:
5562:
3251:
149:
1140:
4821:
2559:
542:
5515:
5389:
394:
4057:
5577:
5572:
5557:
564:, which had far fewer (eleven total, and only in the lower jaw). The presence of such a large number of teeth in
1292:
708:
4038:
1006:
Pérez-Moreno, B. P.; Sanz, J. L.; Buscalioni, A. D.; Moratalla, J. J.; Ortega, F.; Rasskin-Gutman, D. (1994).
5383:
1197:
429:
628:
first interpreted as an integument, some of these were later seen as representing preserved muscle fibers.
5409:
410:
386:
5495:
3763:
3675:
876:
432:
is a reference to the large number of teeth possessed by this theropod and is derived from Greek πολύς (
347:
300:
402:
4808:
5456:
5341:
3452:
3420:
1022:
653:
460:
277:
406:
5529:
3776:
3563:
887:
The Las Hoyas lagerstätte has produced numerous other exquisitely preserved species, including the
4045:
5447:
5248:
3752:
3612:
3468:
3408:
2843:
2753:
2636:
1500:
1190:
144:
5500:
5482:
5130:
5047:
5021:
4860:
4847:
4725:
4266:
4218:
3050:
2400:
1479:
1080:
1070:
733:
476:
398:
370:
252:
236:
5487:
584:
leading to the toothless cutting edge found in later ornithomimosaurs. The arms and hands of
5541:
5234:
4991:
4901:
4690:
4606:
4468:
4387:
4359:
4319:
4200:
4170:
3688:
3441:
3305:
3162:
3062:
2911:
2818:
2776:
2343:
1653:
1030:
667:
was by the describers assigned to the
Ornithomimosauria, in the basalmost position. A later
358:
128:
41:
5297:
5035:
4998:
4930:
4785:
4778:
4590:
4532:
4505:
4495:
4454:
4289:
4148:
4132:
3950:
3922:
3900:
3787:
3632:
3339:
3113:
2996:
2974:
2895:
2724:
2701:
2694:
2672:
2658:
2651:
2612:
2463:
2310:
2237:
2226:
2162:
2050:
1937:
1514:
1423:
1277:
1252:
961:
637:
496:
351:
1026:
5262:
5225:
5206:
5168:
5157:
4954:
4667:
4626:
4617:
4599:
4579:
4475:
4189:
3934:
3796:
3623:
3605:
3433:
3400:
3386:
3347:
3121:
2982:
2939:
2932:
2854:
2830:
2783:
2742:
2679:
2665:
2393:
2378:
2205:
2039:
1903:
1871:
1860:
1837:
1799:
1660:
1630:
1554:
1493:
1392:
888:
820:
480:
5551:
5304:
5111:
5090:
4969:
4962:
4889:
4683:
4656:
4514:
4430:
4408:
4401:
4394:
4337:
4254:
4117:
4089:
4076:
4000:
3877:
3717:
3551:
3482:
3368:
3355:
3100:
3078:
3071:
3035:
3008:
2960:
2884:
2620:
2411:
2385:
2301:
2282:
2275:
2212:
2191:
2170:
2155:
2141:
2100:
2066:
2059:
1978:
1830:
1717:
1710:
1637:
1607:
1596:
1486:
1457:
1450:
1376:
1131:
Dinosaurs of the Air: The
Evolution and Loss of Flight in Dinosaurs and Birds, pg 237
893:
648:
or be a recent descendant from a flying animal. This is due to the presence of large
593:
464:
79:
5276:
5214:
5184:
5104:
5097:
5075:
5067:
4947:
4939:
4919:
4634:
4545:
4483:
4373:
4344:
4296:
4125:
4110:
3885:
3817:
3745:
3724:
3703:
3519:
3475:
3283:
3270:
3200:
3153:
3028:
2918:
2870:
2863:
2800:
2769:
2762:
2643:
2470:
2322:
2219:
2121:
2114:
2009:
1993:
1929:
1880:
1812:
1589:
1472:
1465:
1436:
1399:
1368:
1008:"A unique multitoothed ornithomimosaur dinosaur from the Lower Cretaceous of Spain"
918:
704:
684:
607:. The small crest in back of the head (colored in light grey) was probably made by
551:
507:
5441:
5330:
5311:
5241:
5191:
5177:
5119:
5060:
5005:
4882:
4676:
4552:
4521:
4437:
4423:
4155:
4103:
3971:
3893:
3856:
3738:
3710:
3663:
3583:
3461:
3318:
3169:
3142:
3042:
2989:
2925:
2904:
2877:
2628:
2441:
2434:
2420:
2246:
2093:
2084:
1986:
1957:
1887:
1852:
1845:
1645:
1623:
1616:
1575:
1567:
1543:
1528:
1507:
1415:
1407:
946:
796:
616:
599:
54:
926:
bones. Coarse sediments of the La Hoyas lagerstätte have produced bones of the
5269:
5198:
5143:
4764:
4563:
4380:
4366:
4351:
4303:
4237:
4163:
3985:
3978:
3964:
3870:
3803:
3731:
3695:
3648:
3590:
3576:
3543:
3535:
3528:
3509:
3332:
3297:
3085:
2686:
2477:
2427:
2364:
2335:
2290:
2268:
2261:
2253:
2177:
2073:
2032:
2025:
2017:
2001:
1921:
1792:
1668:
1521:
1384:
1243:
956:
951:
941:
927:
905:
772:
712:
581:
573:
560:
536:
524:
207:
99:
64:
5432:
1084:
17:
5290:
5283:
4983:
4976:
4649:
4641:
4572:
4225:
3943:
3489:
3393:
3128:
2713:
2598:
2585:
2512:
2448:
2371:
2350:
2107:
1970:
1778:
1765:
1683:
1535:
1354:
1341:
1304:
1213:
932:
899:
722:
700:
692:
668:
576:
for cutting and ripping, a "functional counterpart of the cutting edge of a
568:, coupled with a lack of interdental space, was interpreted by Pérez-Moreno
488:
220:
161:
104:
48:
1050:
Récords y curiosidades de los dinosaurios Terópodos y otros dinosauromorfos
1007:
679:
5426:
4874:
4282:
4275:
4247:
4140:
3640:
3021:
2357:
2184:
1945:
1582:
1261:
1237:
923:
913:
452:
374:
354:
194:
181:
94:
89:
74:
69:
59:
619:
similar to the much larger pouches found in modern pelicans, from which
5474:
4415:
4177:
3907:
3810:
3135:
2456:
649:
608:
532:
528:
492:
340:
336:
109:
84:
5083:
4771:
2198:
1231:
1064:
1034:
937:
645:
171:
5403:
3914:
3655:
3597:
875:
633:
598:
541:
506:
468:
418:
366:
362:
343:
5461:
1101:
Barsbold and Perle 1984 (Dinosauria; Theropoda) of
Mongolia." in
691:
has played an important and surprising role in understanding the
5347:
3957:
910:
589:
577:
428:, "mimic", in reference to the long snout and throat pouch. The
5407:
4845:
4074:
3268:
2583:
1763:
1339:
1224:
1186:
632:
was also the first ornithomimosaur discovered with a preserved
442:
434:
1182:
459:, part of the Las Hoyas Collection presently housed at the
558:
Only one other ornithomimosaur is known to possess teeth,
1105:
ed. Carpenter, K. 2005. Indiana
University Press:97-126.
1163:
1161:
1141:
1001:
999:
997:
995:
993:
991:
989:
5513:
1097:
Kobayashi, Y., and
Barsbold, R. (2005). "Anatomy of
936:. The lagerstätte beds have also yielded remains of
5416:
5329:
5224:
5167:
5129:
5046:
5019:
4929:
4899:
4858:
4756:
4724:
4666:
4616:
4589:
4562:
4531:
4504:
4493:
4452:
4317:
4265:
4236:
4198:
4187:
4087:
3999:
3933:
3855:
3786:
3762:
3674:
3622:
3562:
3518:
3507:
3451:
3419:
3366:
3316:
3281:
3199:
3152:
3099:
3061:
3007:
2958:
2894:
2853:
2829:
2798:
2752:
2722:
2711:
2596:
2511:
2410:
2321:
2300:
2236:
2139:
2083:
2049:
1956:
1901:
1870:
1810:
1776:
1702:
1682:
1606:
1553:
1434:
1352:
1303:
1276:
1155:. 2nd ed. University of California Press, Berkeley.
636:apparatus (specialized tongue bones in the neck).
695:of the Ornithomimosauria. To quote Pérez-Moreno
588:were more typical of ornithomimosaurs, with the
393:site. In 1994 this was named and described by
1198:
8:
1116:Journal of the Geological Society of London
1069:. Princeton University Press. p. 128.
5404:
5135:
5052:
5027:
4911:
4907:
4866:
4855:
4842:
4753:
4721:
4537:
4501:
4460:
4329:
4325:
4210:
4206:
4195:
4095:
4084:
4071:
3862:
3852:
3768:
3680:
3568:
3515:
3425:
3378:
3374:
3324:
3289:
3278:
3265:
3196:
3105:
3013:
2966:
2835:
2810:
2806:
2734:
2730:
2719:
2604:
2593:
2580:
2508:
2327:
2318:
2147:
1962:
1913:
1909:
1822:
1818:
1784:
1773:
1760:
1559:
1442:
1360:
1349:
1336:
1282:
1273:
1221:
1205:
1191:
1183:
127:
31:
479:locality in Cuenca Province, Spain, from
944:, as well as of the unique early mammal
687:(see cladogram below). The discovery of
5520:
985:
883:(foreground) among contemporary animals
1066:The Princeton Field Guide to Dinosaurs
1048:Molina-Pérez & Larramendi (2016).
523:very small teeth in total, with seven
389:discovered a theropod skeleton at the
7:
5568:Early Cretaceous dinosaurs of Europe
974:Timeline of ornithomimosaur research
365:. It is notable for possessing more
812:
788:
764:
740:
728:
417:. The generic name is derived from
25:
485:Calizas de La Huérguina Formation
377:), most of which were toothless.
5535:
5523:
5388:
5382:
5375:
4827:
4820:
4813:
4807:
4056:
4049:
4044:
4037:
4031:
3250:
3243:
3236:
2565:
2558:
2551:
2544:
1746:
1740:
148:
1052:. Spain: Larousse. p. 268.
922:remains, and a few fragmentary
603:Illustration of the skull from
725:after Kobayashi and Lü, 2003:
1:
5588:Fossil taxa described in 1994
1169:Acta Palaeontologica Polonica
395:Bernardino Pérez Pérez-Moreno
369:than any other member of the
443:
435:
644:might have been capable of
5604:
531:, and seventy-five in the
27:Extinct genus of dinosaurs
5372:
5138:
5055:
5030:
4914:
4910:
4869:
4854:
4841:
4805:
4752:
4720:
4540:
4463:
4332:
4328:
4213:
4209:
4098:
4083:
4070:
4029:
3865:
3851:
3771:
3683:
3571:
3428:
3381:
3377:
3327:
3292:
3277:
3264:
3233:
3195:
3108:
3016:
2969:
2838:
2813:
2809:
2737:
2733:
2607:
2592:
2579:
2541:
2507:
2330:
2150:
1965:
1916:
1912:
1825:
1821:
1787:
1772:
1759:
1737:
1562:
1445:
1363:
1348:
1335:
1285:
1272:
1220:
1103:The Carnivorous Dinosaurs
1063:Paul, Gregory S. (2016).
817:
810:
793:
786:
769:
762:
745:
738:
306:
299:
145:Scientific classification
143:
135:
126:
34:
409:, Francisco Ortega and
5583:La Huérguina Formation
1099:Harpymimus okladnikovi
884:
675:(2005) confirmed that
671:analysis by Makovicky
612:
555:
516:
515:, compared to a human.
475:was recovered at the
415:Pelecanimimus polyodon
313:Pelecanimimus polyodon
5496:Paleobiology Database
3764:Carcharodontosaurinae
3676:Carcharodontosauridae
879:
602:
545:
510:
440:), "many" and ὀδούς (
4757:Dubious coelurosaurs
3453:Metriacanthosaurinae
3421:Metriacanthosauridae
1703:Dubious neotheropods
640:has speculated that
527:teeth, about thirty
487:dating to the Lower
411:Diego Rasskin-Gutman
387:Armando Díaz Romeral
381:Discovery and naming
5563:Feathered dinosaurs
3777:Carcharodontosaurus
3564:Carcharodontosauria
1027:1994Natur.370..363P
5249:Archaeornithomimus
3753:Veterupristisaurus
3613:Veterupristisaurus
3469:Metriacanthosaurus
3409:Piatnitzkysauridae
2844:Eustreptospondylus
2754:Piatnitzkysauridae
2637:Chuandongocoelurus
1501:Notatesseraeraptor
885:
613:
556:
517:
511:Estimated size of
413:as a new species:
352:ornithomimosaurian
5511:
5510:
5483:Open Tree of Life
5410:Taxon identifiers
5401:
5400:
5397:
5396:
5370:
5369:
5366:
5365:
5362:
5361:
5358:
5357:
5325:
5324:
5321:
5320:
5131:Macrocheiriformes
5048:Ornithomimosauria
5022:Maniraptoriformes
5015:
5014:
4861:Maniraptoromorpha
4848:Maniraptoromorpha
4837:
4836:
4803:
4802:
4799:
4798:
4795:
4794:
4748:
4747:
4744:
4743:
4726:Maniraptoromorpha
4716:
4715:
4712:
4711:
4708:
4707:
4704:
4703:
4700:
4699:
4448:
4447:
4313:
4312:
4267:Proceratosauridae
4219:Chingkankousaurus
4066:
4065:
4027:
4026:
4023:
4022:
4019:
4018:
3995:
3994:
3847:
3846:
3843:
3842:
3839:
3838:
3835:
3834:
3831:
3830:
3827:
3826:
3503:
3502:
3499:
3498:
3260:
3259:
3231:
3230:
3227:
3226:
3223:
3222:
3219:
3218:
3191:
3190:
3187:
3186:
3183:
3182:
3179:
3178:
3095:
3094:
3051:Vallibonavenatrix
2954:
2953:
2950:
2949:
2794:
2793:
2575:
2574:
2539:
2538:
2535:
2534:
2531:
2530:
2503:
2502:
2499:
2498:
2495:
2494:
2491:
2490:
2487:
2486:
2135:
2134:
2131:
2130:
1897:
1896:
1755:
1754:
1735:
1734:
1731:
1730:
1727:
1726:
1678:
1677:
1331:
1330:
1327:
1326:
1323:
1322:
1076:978-1-78684-190-2
1021:(6488): 363−367.
909:, along with non-
868:
867:
859:
858:
850:
849:
841:
840:
832:
831:
734:Ornithomimosauria
580:," as well as an
535:. The teeth were
424:, "pelican", and
403:Angela Buscalioni
371:Ornithomimosauria
328:
327:
280:
253:Macrocheiriformes
237:Ornithomimosauria
16:(Redirected from
5595:
5578:Fossils of Spain
5573:Cretaceous Spain
5558:Ornithomimosaurs
5540:
5539:
5538:
5528:
5527:
5526:
5519:
5504:
5503:
5491:
5490:
5478:
5477:
5465:
5464:
5452:
5451:
5450:
5437:
5436:
5435:
5405:
5392:
5386:
5379:
5235:Aepyornithomimus
5136:
5053:
5028:
4992:Sinocalliopteryx
4912:
4908:
4902:Neocoelurosauria
4867:
4856:
4843:
4831:
4824:
4817:
4811:
4754:
4722:
4691:Zhuchengtyrannus
4607:Thanatotheristes
4538:
4502:
4469:Appalachiosaurus
4461:
4330:
4326:
4320:Pantyrannosauria
4211:
4207:
4201:Tyrannosauroidea
4196:
4171:Xinjiangovenator
4096:
4085:
4072:
4060:
4053:
4048:
4041:
4035:
3863:
3853:
3769:
3689:Acrocanthosaurus
3681:
3569:
3516:
3442:Yangchuanosaurus
3426:
3379:
3375:
3325:
3306:Lourinhanosaurus
3290:
3279:
3266:
3254:
3247:
3240:
3197:
3163:Sigilmassasaurus
3106:
3063:Ceratosuchopsini
3014:
2967:
2912:Dubreuillosaurus
2836:
2819:Streptospondylus
2811:
2807:
2777:Piatnitzkysaurus
2735:
2731:
2720:
2605:
2594:
2581:
2569:
2562:
2555:
2548:
2509:
2344:Ekrixinatosaurus
2328:
2319:
2148:
1963:
1914:
1910:
1823:
1819:
1785:
1774:
1761:
1750:
1744:
1654:Procompsognathus
1560:
1443:
1361:
1350:
1337:
1283:
1274:
1267:
1266:
1222:
1207:
1200:
1193:
1184:
1172:
1165:
1156:
1149:
1143:
1138:
1132:
1129:
1123:
1112:
1106:
1095:
1089:
1088:
1060:
1054:
1053:
1045:
1039:
1038:
1035:10.1038/370363a0
1012:
1003:
964:are also known.
813:
789:
765:
741:
729:
549:being chased by
483:beds within the
446:
438:
391:Las Hoyas Unit 3
359:Early Cretaceous
315:
311:
292:P. polyodon
271:
264:
251:
235:
153:
152:
131:
121:
51:
42:Early Cretaceous
40:Temporal range:
32:
21:
5603:
5602:
5598:
5597:
5596:
5594:
5593:
5592:
5548:
5547:
5546:
5536:
5534:
5524:
5522:
5514:
5512:
5507:
5499:
5494:
5486:
5481:
5473:
5468:
5460:
5455:
5446:
5445:
5440:
5431:
5430:
5425:
5412:
5402:
5393:
5380:
5354:
5317:
5298:Sinornithomimus
5220:
5163:
5125:
5042:
5036:Compsognathidae
5024:
5011:
4999:Sinosauropteryx
4931:Compsognathidae
4925:
4904:
4895:
4863:
4850:
4833:
4832:
4818:
4791:
4786:Shanyangosaurus
4779:Phaedrolosaurus
4740:
4696:
4662:
4612:
4591:Daspletosaurini
4585:
4558:
4533:Tyrannosaurinae
4527:
4506:Albertosaurinae
4498:
4496:Tyrannosauridae
4489:
4457:
4455:Eutyrannosauria
4444:
4322:
4309:
4290:Proceratosaurus
4261:
4232:
4203:
4192:
4183:
4149:Richardoestesia
4133:Chilantaisaurus
4092:
4079:
4062:
4061:
4042:
4015:
3991:
3951:Australovenator
3929:
3923:Chilantaisaurus
3901:Phuwiangvenator
3823:
3788:Giganotosaurini
3782:
3758:
3670:
3633:Chilantaisaurus
3618:
3558:
3512:
3495:
3447:
3415:
3371:
3362:
3340:Monolophosaurus
3321:
3312:
3286:
3273:
3256:
3255:
3241:
3215:
3175:
3148:
3114:Camarillasaurus
3091:
3057:
3003:
2997:Ostafrikasaurus
2975:Camarillasaurus
2963:
2946:
2896:Afrovenatorinae
2890:
2849:
2825:
2803:
2790:
2748:
2727:
2725:Megalosauroidea
2716:
2707:
2702:Vectaerovenator
2695:Szechuanosaurus
2673:Monolophosaurus
2659:Kaijiangosaurus
2652:Cryolophosaurus
2613:Calamospondylus
2601:
2588:
2571:
2570:
2563:
2556:
2549:
2527:
2483:
2464:Pycnonemosaurus
2406:
2317:
2311:Majungasaurinae
2296:
2238:Majungasaurinae
2232:
2227:Xenotarsosaurus
2163:Dryptosauroides
2144:
2127:
2079:
2051:Elaphrosaurinae
2045:
1952:
1938:Ornithomimoides
1906:
1893:
1866:
1815:
1806:
1781:
1768:
1751:
1745:
1723:
1698:
1674:
1602:
1549:
1515:Shuangbaisaurus
1439:
1430:
1357:
1344:
1319:
1299:
1293:Avemetatarsalia
1278:Avemetatarsalia
1268:
1253:Avemetatarsalia
1227:
1226:
1216:
1211:
1181:
1176:
1175:
1166:
1159:
1150:
1146:
1139:
1135:
1130:
1126:
1113:
1109:
1096:
1092:
1077:
1062:
1061:
1057:
1047:
1046:
1042:
1010:
1005:
1004:
987:
982:
970:
962:crocodylomorphs
889:enantiornithine
874:
869:
860:
851:
842:
833:
662:
654:Mickey Mortimer
638:Gregory S. Paul
623:took its name.
505:
497:pectoral girdle
461:Museo de Cuenca
383:
324:
317:
309:
308:
295:
270:
262:
249:
233:
147:
136:Restoration of
122:
120:
119:
118:
117:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
46:
45:
38:
28:
23:
22:
15:
12:
11:
5:
5601:
5599:
5591:
5590:
5585:
5580:
5575:
5570:
5565:
5560:
5550:
5549:
5545:
5544:
5532:
5509:
5508:
5506:
5505:
5492:
5479:
5466:
5453:
5438:
5422:
5420:
5414:
5413:
5408:
5399:
5398:
5395:
5394:
5374:
5373:
5371:
5368:
5367:
5364:
5363:
5360:
5359:
5356:
5355:
5353:
5352:
5351:
5350:
5344:
5335:
5333:
5327:
5326:
5323:
5322:
5319:
5318:
5316:
5315:
5308:
5301:
5294:
5287:
5280:
5273:
5266:
5263:Dromiceiomimus
5259:
5252:
5245:
5238:
5230:
5228:
5226:Ornithomimidae
5222:
5221:
5219:
5218:
5211:
5207:Paraxenisaurus
5203:
5195:
5188:
5181:
5173:
5171:
5169:Deinocheiridae
5165:
5164:
5162:
5161:
5158:Shenzhousaurus
5154:
5147:
5139:
5133:
5127:
5126:
5124:
5123:
5116:
5108:
5101:
5094:
5087:
5080:
5072:
5064:
5056:
5050:
5044:
5043:
5041:
5040:
5031:
5025:
5020:
5017:
5016:
5013:
5012:
5010:
5009:
5002:
4995:
4988:
4980:
4973:
4966:
4959:
4955:Beipiaognathus
4951:
4944:
4935:
4933:
4927:
4926:
4924:
4923:
4915:
4905:
4900:
4897:
4896:
4894:
4893:
4886:
4879:
4870:
4864:
4859:
4852:
4851:
4846:
4839:
4838:
4835:
4834:
4826:
4812:
4806:
4804:
4801:
4800:
4797:
4796:
4793:
4792:
4790:
4789:
4782:
4775:
4768:
4760:
4758:
4750:
4749:
4746:
4745:
4742:
4741:
4739:
4738:
4737:
4736:
4730:
4728:
4718:
4717:
4714:
4713:
4710:
4709:
4706:
4705:
4702:
4701:
4698:
4697:
4695:
4694:
4687:
4680:
4672:
4670:
4668:Tyrannosaurini
4664:
4663:
4661:
4660:
4653:
4646:
4638:
4631:
4627:Bistahieversor
4622:
4620:
4618:Teratophoneini
4614:
4613:
4611:
4610:
4603:
4600:Daspletosaurus
4595:
4593:
4587:
4586:
4584:
4583:
4580:Qianzhousaurus
4576:
4568:
4566:
4560:
4559:
4557:
4556:
4549:
4541:
4535:
4529:
4528:
4526:
4525:
4518:
4510:
4508:
4499:
4494:
4491:
4490:
4488:
4487:
4480:
4476:Bistahieversor
4472:
4464:
4458:
4453:
4450:
4449:
4446:
4445:
4443:
4442:
4434:
4427:
4420:
4412:
4405:
4398:
4391:
4384:
4377:
4370:
4363:
4356:
4348:
4341:
4333:
4323:
4318:
4315:
4314:
4311:
4310:
4308:
4307:
4300:
4293:
4286:
4279:
4271:
4269:
4263:
4262:
4260:
4259:
4251:
4243:
4241:
4234:
4233:
4231:
4230:
4222:
4214:
4204:
4199:
4193:
4190:Tyrannoraptora
4188:
4185:
4184:
4182:
4181:
4174:
4167:
4160:
4152:
4145:
4137:
4129:
4122:
4114:
4107:
4099:
4093:
4088:
4081:
4080:
4075:
4068:
4067:
4064:
4063:
4055:
4036:
4030:
4028:
4025:
4024:
4021:
4020:
4017:
4016:
4014:
4013:
4012:
4011:
4005:
4003:
3997:
3996:
3993:
3992:
3990:
3989:
3982:
3975:
3968:
3961:
3954:
3947:
3939:
3937:
3935:Megaraptoridae
3931:
3930:
3928:
3927:
3919:
3911:
3904:
3897:
3890:
3882:
3874:
3866:
3860:
3849:
3848:
3845:
3844:
3841:
3840:
3837:
3836:
3833:
3832:
3829:
3828:
3825:
3824:
3822:
3821:
3814:
3807:
3800:
3797:Giganotosaurus
3792:
3790:
3784:
3783:
3781:
3780:
3772:
3766:
3760:
3759:
3757:
3756:
3749:
3742:
3735:
3728:
3721:
3714:
3707:
3700:
3692:
3684:
3678:
3672:
3671:
3669:
3668:
3660:
3652:
3645:
3637:
3628:
3626:
3624:Neovenatoridae
3620:
3619:
3617:
3616:
3609:
3606:Ulughbegsaurus
3602:
3594:
3587:
3580:
3572:
3566:
3560:
3559:
3557:
3556:
3548:
3540:
3532:
3524:
3522:
3513:
3508:
3505:
3504:
3501:
3500:
3497:
3496:
3494:
3493:
3486:
3479:
3472:
3465:
3457:
3455:
3449:
3448:
3446:
3445:
3438:
3434:Xuanhanosaurus
3429:
3423:
3417:
3416:
3414:
3413:
3405:
3401:Xuanhanosaurus
3397:
3390:
3387:Asfaltovenator
3382:
3372:
3367:
3364:
3363:
3361:
3360:
3352:
3348:Megalosauridae
3344:
3336:
3328:
3322:
3317:
3314:
3313:
3311:
3310:
3302:
3293:
3287:
3282:
3275:
3274:
3269:
3262:
3261:
3258:
3257:
3249:
3235:
3234:
3232:
3229:
3228:
3225:
3224:
3221:
3220:
3217:
3216:
3214:
3213:
3212:
3211:
3205:
3203:
3193:
3192:
3189:
3188:
3185:
3184:
3181:
3180:
3177:
3176:
3174:
3173:
3166:
3158:
3156:
3150:
3149:
3147:
3146:
3139:
3132:
3125:
3122:Ichthyovenator
3118:
3109:
3103:
3097:
3096:
3093:
3092:
3090:
3089:
3082:
3075:
3067:
3065:
3059:
3058:
3056:
3055:
3047:
3039:
3032:
3025:
3017:
3011:
3005:
3004:
3002:
3001:
2993:
2986:
2983:Cristatusaurus
2979:
2970:
2964:
2959:
2956:
2955:
2952:
2951:
2948:
2947:
2945:
2944:
2940:Poekilopleuron
2936:
2933:Piveteausaurus
2929:
2922:
2915:
2908:
2900:
2898:
2892:
2891:
2889:
2888:
2881:
2874:
2867:
2859:
2857:
2855:Megalosaurinae
2851:
2850:
2848:
2847:
2839:
2833:
2831:Megalosauridae
2827:
2826:
2824:
2823:
2814:
2804:
2799:
2796:
2795:
2792:
2791:
2789:
2788:
2784:Xuanhanosaurus
2780:
2773:
2766:
2758:
2756:
2750:
2749:
2747:
2746:
2743:Yunyangosaurus
2738:
2728:
2723:
2717:
2712:
2709:
2708:
2706:
2705:
2698:
2691:
2683:
2680:Pandoravenator
2676:
2669:
2666:Kayentavenator
2662:
2655:
2648:
2640:
2633:
2625:
2617:
2608:
2602:
2597:
2590:
2589:
2584:
2577:
2576:
2573:
2572:
2564:
2557:
2550:
2543:
2542:
2540:
2537:
2536:
2533:
2532:
2529:
2528:
2526:
2525:
2524:
2523:
2517:
2515:
2505:
2504:
2501:
2500:
2497:
2496:
2493:
2492:
2489:
2488:
2485:
2484:
2482:
2481:
2474:
2467:
2460:
2453:
2445:
2438:
2431:
2424:
2416:
2414:
2408:
2407:
2405:
2404:
2397:
2394:Skorpiovenator
2390:
2382:
2375:
2368:
2361:
2354:
2347:
2340:
2331:
2325:
2316:
2315:
2306:
2304:
2298:
2297:
2295:
2294:
2287:
2279:
2272:
2265:
2258:
2250:
2242:
2240:
2234:
2233:
2231:
2230:
2223:
2216:
2209:
2206:Spectrovenator
2202:
2195:
2188:
2181:
2174:
2167:
2159:
2151:
2145:
2140:
2137:
2136:
2133:
2132:
2129:
2128:
2126:
2125:
2118:
2111:
2104:
2097:
2089:
2087:
2081:
2080:
2078:
2077:
2070:
2063:
2055:
2053:
2047:
2046:
2044:
2043:
2040:Spinostropheus
2036:
2029:
2022:
2014:
2006:
1998:
1990:
1983:
1975:
1966:
1960:
1954:
1953:
1951:
1950:
1942:
1934:
1926:
1917:
1907:
1904:Abelisauroidea
1902:
1899:
1898:
1895:
1894:
1892:
1891:
1884:
1876:
1874:
1872:Ceratosauridae
1868:
1867:
1865:
1864:
1861:Saltriovenator
1857:
1849:
1842:
1838:Fosterovenator
1834:
1826:
1816:
1811:
1808:
1807:
1805:
1804:
1800:Lophostropheus
1796:
1788:
1782:
1777:
1770:
1769:
1764:
1757:
1756:
1753:
1752:
1739:
1738:
1736:
1733:
1732:
1729:
1728:
1725:
1724:
1722:
1721:
1714:
1706:
1704:
1700:
1699:
1697:
1696:
1695:
1694:
1688:
1686:
1680:
1679:
1676:
1675:
1673:
1672:
1665:
1661:Pterospondylus
1657:
1650:
1642:
1634:
1631:Lucianovenator
1627:
1620:
1612:
1610:
1604:
1603:
1601:
1600:
1593:
1586:
1579:
1572:
1563:
1557:
1555:Coelophysoidea
1551:
1550:
1548:
1547:
1540:
1532:
1525:
1518:
1511:
1504:
1497:
1494:Lophostropheus
1490:
1483:
1476:
1469:
1462:
1454:
1446:
1440:
1435:
1432:
1431:
1429:
1428:
1420:
1412:
1404:
1396:
1393:Erythrovenator
1389:
1381:
1373:
1364:
1358:
1353:
1346:
1345:
1340:
1333:
1332:
1329:
1328:
1325:
1324:
1321:
1320:
1318:
1317:
1316:
1315:
1309:
1307:
1301:
1300:
1298:
1297:
1296:
1295:
1286:
1280:
1270:
1269:
1265:
1264:
1255:
1246:
1240:
1234:
1225:
1218:
1217:
1212:
1210:
1209:
1202:
1195:
1187:
1180:
1179:External links
1177:
1174:
1173:
1157:
1153:The Dinosauria
1144:
1133:
1124:
1107:
1090:
1075:
1055:
1040:
984:
983:
981:
978:
977:
976:
969:
966:
873:
870:
866:
865:
862:
861:
857:
856:
853:
852:
848:
847:
844:
843:
839:
838:
835:
834:
830:
829:
826:
825:
821:Ornithomimidae
816:
811:
809:
806:
805:
802:
801:
792:
787:
785:
782:
781:
778:
777:
768:
763:
761:
758:
757:
754:
753:
744:
739:
737:
727:
661:
660:Classification
658:
504:
501:
407:José Moratalla
399:José Luis Sanz
382:
379:
373:(or any other
350:("primitive")
339:mimic") is an
326:
325:
318:
304:
303:
297:
296:
288:
286:
282:
281:
260:
256:
255:
247:
240:
239:
231:
224:
223:
218:
211:
210:
205:
198:
197:
192:
185:
184:
179:
175:
174:
169:
165:
164:
159:
155:
154:
141:
140:
133:
132:
124:
123:
115:
114:
113:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
52:
39:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
5600:
5589:
5586:
5584:
5581:
5579:
5576:
5574:
5571:
5569:
5566:
5564:
5561:
5559:
5556:
5555:
5553:
5543:
5533:
5531:
5521:
5517:
5502:
5497:
5493:
5489:
5484:
5480:
5476:
5471:
5467:
5463:
5458:
5454:
5449:
5448:Pelecanimimus
5443:
5439:
5434:
5428:
5424:
5423:
5421:
5419:
5418:Pelecanimimus
5415:
5411:
5406:
5391:
5387:
5385:
5378:
5349:
5345:
5343:
5339:
5338:
5337:
5336:
5334:
5332:
5328:
5314:
5313:
5309:
5307:
5306:
5305:Struthiomimus
5302:
5300:
5299:
5295:
5293:
5292:
5288:
5286:
5285:
5281:
5279:
5278:
5274:
5272:
5271:
5267:
5265:
5264:
5260:
5258:
5257:
5256:"Coelosaurus"
5253:
5251:
5250:
5246:
5244:
5243:
5239:
5237:
5236:
5232:
5231:
5229:
5227:
5223:
5217:
5216:
5212:
5209:
5208:
5204:
5201:
5200:
5196:
5194:
5193:
5189:
5187:
5186:
5182:
5180:
5179:
5175:
5174:
5172:
5170:
5166:
5160:
5159:
5155:
5153:
5152:
5151:Pelecanimimus
5148:
5146:
5145:
5141:
5140:
5137:
5134:
5132:
5128:
5122:
5121:
5117:
5114:
5113:
5112:Thecocoelurus
5109:
5107:
5106:
5102:
5100:
5099:
5095:
5093:
5092:
5091:Kinnareemimus
5088:
5086:
5085:
5081:
5078:
5077:
5073:
5070:
5069:
5065:
5063:
5062:
5058:
5057:
5054:
5051:
5049:
5045:
5038:
5037:
5033:
5032:
5029:
5026:
5023:
5018:
5008:
5007:
5003:
5001:
5000:
4996:
4994:
4993:
4989:
4986:
4985:
4981:
4979:
4978:
4974:
4972:
4971:
4970:Huaxiagnathus
4967:
4965:
4964:
4963:Compsognathus
4960:
4957:
4956:
4952:
4950:
4949:
4945:
4942:
4941:
4937:
4936:
4934:
4932:
4928:
4922:
4921:
4917:
4916:
4913:
4909:
4906:
4903:
4898:
4892:
4891:
4890:Ornitholestes
4887:
4885:
4884:
4880:
4877:
4876:
4872:
4871:
4868:
4865:
4862:
4857:
4853:
4849:
4844:
4840:
4830:
4825:
4823:
4816:
4810:
4788:
4787:
4783:
4781:
4780:
4776:
4774:
4773:
4769:
4767:
4766:
4762:
4761:
4759:
4755:
4751:
4734:
4733:
4732:
4731:
4729:
4727:
4723:
4719:
4693:
4692:
4688:
4686:
4685:
4684:Tyrannosaurus
4681:
4679:
4678:
4674:
4673:
4671:
4669:
4665:
4659:
4658:
4657:Teratophoneus
4654:
4652:
4651:
4647:
4644:
4643:
4639:
4637:
4636:
4632:
4629:
4628:
4624:
4623:
4621:
4619:
4615:
4609:
4608:
4604:
4602:
4601:
4597:
4596:
4594:
4592:
4588:
4582:
4581:
4577:
4575:
4574:
4570:
4569:
4567:
4565:
4561:
4555:
4554:
4550:
4548:
4547:
4543:
4542:
4539:
4536:
4534:
4530:
4524:
4523:
4519:
4517:
4516:
4515:Albertosaurus
4512:
4511:
4509:
4507:
4503:
4500:
4497:
4492:
4486:
4485:
4481:
4478:
4477:
4473:
4471:
4470:
4466:
4465:
4462:
4459:
4456:
4451:
4440:
4439:
4435:
4433:
4432:
4431:Xiongguanlong
4428:
4426:
4425:
4421:
4418:
4417:
4413:
4411:
4410:
4409:Suskityrannus
4406:
4404:
4403:
4402:Stokesosaurus
4399:
4397:
4396:
4395:Santanaraptor
4392:
4390:
4389:
4385:
4383:
4382:
4378:
4376:
4375:
4371:
4369:
4368:
4364:
4362:
4361:
4357:
4354:
4353:
4349:
4347:
4346:
4342:
4340:
4339:
4338:Alectrosaurus
4335:
4334:
4331:
4327:
4324:
4321:
4316:
4306:
4305:
4301:
4299:
4298:
4294:
4292:
4291:
4287:
4285:
4284:
4280:
4278:
4277:
4273:
4272:
4270:
4268:
4264:
4257:
4256:
4255:Tanycolagreus
4252:
4250:
4249:
4245:
4244:
4242:
4239:
4235:
4228:
4227:
4223:
4221:
4220:
4216:
4215:
4212:
4208:
4205:
4202:
4197:
4194:
4191:
4186:
4180:
4179:
4175:
4173:
4172:
4168:
4166:
4165:
4161:
4158:
4157:
4153:
4151:
4150:
4146:
4143:
4142:
4138:
4135:
4134:
4130:
4128:
4127:
4123:
4120:
4119:
4118:Bahariasaurus
4115:
4113:
4112:
4108:
4106:
4105:
4101:
4100:
4097:
4094:
4091:
4090:Coelurosauria
4086:
4082:
4078:
4077:Coelurosauria
4073:
4069:
4059:
4054:
4052:
4047:
4040:
4034:
4009:
4008:
4007:
4006:
4004:
4002:
4001:Coelurosauria
3998:
3988:
3987:
3983:
3981:
3980:
3976:
3974:
3973:
3969:
3967:
3966:
3962:
3960:
3959:
3955:
3953:
3952:
3948:
3946:
3945:
3941:
3940:
3938:
3936:
3932:
3926:
3924:
3920:
3918:
3916:
3912:
3910:
3909:
3905:
3903:
3902:
3898:
3896:
3895:
3891:
3888:
3887:
3883:
3880:
3879:
3878:Bahariasaurus
3875:
3873:
3872:
3868:
3867:
3864:
3861:
3858:
3854:
3850:
3820:
3819:
3815:
3813:
3812:
3808:
3806:
3805:
3801:
3799:
3798:
3794:
3793:
3791:
3789:
3785:
3779:
3778:
3774:
3773:
3770:
3767:
3765:
3761:
3755:
3754:
3750:
3748:
3747:
3743:
3741:
3740:
3736:
3734:
3733:
3729:
3727:
3726:
3722:
3720:
3719:
3718:Kelmayisaurus
3715:
3713:
3712:
3708:
3706:
3705:
3701:
3698:
3697:
3693:
3691:
3690:
3686:
3685:
3682:
3679:
3677:
3673:
3666:
3665:
3661:
3658:
3657:
3653:
3651:
3650:
3646:
3643:
3642:
3638:
3635:
3634:
3630:
3629:
3627:
3625:
3621:
3615:
3614:
3610:
3608:
3607:
3603:
3600:
3599:
3595:
3593:
3592:
3588:
3586:
3585:
3581:
3579:
3578:
3574:
3573:
3570:
3567:
3565:
3561:
3554:
3553:
3552:Saurophaganax
3549:
3546:
3545:
3541:
3538:
3537:
3533:
3531:
3530:
3526:
3525:
3523:
3521:
3517:
3514:
3511:
3506:
3492:
3491:
3487:
3485:
3484:
3483:Siamotyrannus
3480:
3478:
3477:
3473:
3471:
3470:
3466:
3464:
3463:
3459:
3458:
3456:
3454:
3450:
3444:
3443:
3439:
3436:
3435:
3431:
3430:
3427:
3424:
3422:
3418:
3411:
3410:
3406:
3403:
3402:
3398:
3396:
3395:
3391:
3389:
3388:
3384:
3383:
3380:
3376:
3373:
3370:
3369:Allosauroidea
3365:
3358:
3357:
3356:Spinosauridae
3353:
3350:
3349:
3345:
3342:
3341:
3337:
3335:
3334:
3330:
3329:
3326:
3323:
3320:
3315:
3308:
3307:
3303:
3300:
3299:
3295:
3294:
3291:
3288:
3285:
3280:
3276:
3272:
3267:
3263:
3253:
3248:
3246:
3239:
3209:
3208:
3207:
3206:
3204:
3202:
3198:
3194:
3172:
3171:
3167:
3165:
3164:
3160:
3159:
3157:
3155:
3151:
3145:
3144:
3140:
3138:
3137:
3133:
3131:
3130:
3126:
3124:
3123:
3119:
3116:
3115:
3111:
3110:
3107:
3104:
3102:
3101:Spinosaurinae
3098:
3088:
3087:
3083:
3081:
3080:
3079:Riparovenator
3076:
3074:
3073:
3072:Ceratosuchops
3069:
3068:
3066:
3064:
3060:
3053:
3052:
3048:
3045:
3044:
3040:
3038:
3037:
3036:Riojavenatrix
3033:
3031:
3030:
3026:
3024:
3023:
3019:
3018:
3015:
3012:
3010:
3009:Baryonychinae
3006:
2999:
2998:
2994:
2992:
2991:
2987:
2985:
2984:
2980:
2977:
2976:
2972:
2971:
2968:
2965:
2962:
2961:Spinosauridae
2957:
2942:
2941:
2937:
2935:
2934:
2930:
2928:
2927:
2923:
2921:
2920:
2916:
2914:
2913:
2909:
2907:
2906:
2902:
2901:
2899:
2897:
2893:
2887:
2886:
2885:Wiehenvenator
2882:
2880:
2879:
2875:
2873:
2872:
2868:
2866:
2865:
2861:
2860:
2858:
2856:
2852:
2846:
2845:
2841:
2840:
2837:
2834:
2832:
2828:
2821:
2820:
2816:
2815:
2812:
2808:
2805:
2802:
2797:
2786:
2785:
2781:
2779:
2778:
2774:
2772:
2771:
2767:
2765:
2764:
2760:
2759:
2757:
2755:
2751:
2745:
2744:
2740:
2739:
2736:
2732:
2729:
2726:
2721:
2718:
2715:
2710:
2704:
2703:
2699:
2697:
2696:
2692:
2689:
2688:
2684:
2682:
2681:
2677:
2675:
2674:
2670:
2668:
2667:
2663:
2661:
2660:
2656:
2654:
2653:
2649:
2646:
2645:
2641:
2639:
2638:
2634:
2631:
2630:
2626:
2623:
2622:
2621:Chienkosaurus
2618:
2615:
2614:
2610:
2609:
2606:
2603:
2600:
2595:
2591:
2587:
2582:
2578:
2568:
2561:
2554:
2547:
2521:
2520:
2519:
2518:
2516:
2514:
2510:
2506:
2480:
2479:
2475:
2473:
2472:
2468:
2466:
2465:
2461:
2459:
2458:
2454:
2451:
2450:
2446:
2444:
2443:
2439:
2437:
2436:
2432:
2430:
2429:
2425:
2423:
2422:
2418:
2417:
2415:
2413:
2412:Furileusauria
2409:
2403:
2402:
2398:
2396:
2395:
2391:
2388:
2387:
2386:Rahiolisaurus
2383:
2381:
2380:
2376:
2374:
2373:
2369:
2367:
2366:
2362:
2360:
2359:
2355:
2353:
2352:
2348:
2346:
2345:
2341:
2338:
2337:
2333:
2332:
2329:
2326:
2324:
2320:
2313:
2312:
2308:
2307:
2305:
2303:
2302:Carnotaurinae
2299:
2293:
2292:
2288:
2285:
2284:
2283:Rahiolisaurus
2280:
2278:
2277:
2276:Majungasaurus
2273:
2271:
2270:
2266:
2264:
2263:
2259:
2256:
2255:
2251:
2249:
2248:
2244:
2243:
2241:
2239:
2235:
2229:
2228:
2224:
2222:
2221:
2217:
2215:
2214:
2213:Tarascosaurus
2210:
2208:
2207:
2203:
2201:
2200:
2196:
2194:
2193:
2189:
2187:
2186:
2182:
2180:
2179:
2175:
2173:
2172:
2171:Eoabelisaurus
2168:
2165:
2164:
2160:
2158:
2157:
2156:Chenanisaurus
2153:
2152:
2149:
2146:
2143:
2142:Abelisauridae
2138:
2124:
2123:
2119:
2117:
2116:
2112:
2110:
2109:
2105:
2103:
2102:
2101:Masiakasaurus
2098:
2096:
2095:
2091:
2090:
2088:
2086:
2082:
2076:
2075:
2071:
2069:
2068:
2067:Huinculsaurus
2064:
2062:
2061:
2060:Elaphrosaurus
2057:
2056:
2054:
2052:
2048:
2042:
2041:
2037:
2035:
2034:
2030:
2028:
2027:
2023:
2020:
2019:
2015:
2012:
2011:
2007:
2004:
2003:
1999:
1996:
1995:
1991:
1989:
1988:
1984:
1981:
1980:
1979:Austrocheirus
1976:
1973:
1972:
1968:
1967:
1964:
1961:
1959:
1955:
1948:
1947:
1943:
1940:
1939:
1935:
1932:
1931:
1927:
1924:
1923:
1919:
1918:
1915:
1911:
1908:
1905:
1900:
1890:
1889:
1885:
1883:
1882:
1878:
1877:
1875:
1873:
1869:
1863:
1862:
1858:
1855:
1854:
1850:
1848:
1847:
1843:
1840:
1839:
1835:
1833:
1832:
1831:Berberosaurus
1828:
1827:
1824:
1820:
1817:
1814:
1809:
1802:
1801:
1797:
1795:
1794:
1790:
1789:
1786:
1783:
1780:
1775:
1771:
1767:
1762:
1758:
1749:
1743:
1720:
1719:
1718:Halticosaurus
1715:
1713:
1712:
1711:Teinurosaurus
1708:
1707:
1705:
1701:
1692:
1691:
1690:
1689:
1687:
1685:
1681:
1671:
1670:
1666:
1663:
1662:
1658:
1656:
1655:
1651:
1648:
1647:
1643:
1640:
1639:
1638:Megapnosaurus
1635:
1633:
1632:
1628:
1626:
1625:
1621:
1619:
1618:
1614:
1613:
1611:
1609:
1608:Coelophysidae
1605:
1599:
1598:
1597:Powellvenator
1594:
1592:
1591:
1587:
1585:
1584:
1580:
1578:
1577:
1573:
1570:
1569:
1565:
1564:
1561:
1558:
1556:
1552:
1546:
1545:
1541:
1538:
1537:
1533:
1531:
1530:
1526:
1524:
1523:
1519:
1517:
1516:
1512:
1510:
1509:
1505:
1503:
1502:
1498:
1496:
1495:
1491:
1489:
1488:
1487:Liliensternus
1484:
1482:
1481:
1477:
1475:
1474:
1470:
1468:
1467:
1463:
1460:
1459:
1458:Dolichosuchus
1455:
1453:
1452:
1451:Dilophosaurus
1448:
1447:
1444:
1441:
1438:
1433:
1426:
1425:
1424:Herrerasauria
1421:
1418:
1417:
1413:
1410:
1409:
1405:
1402:
1401:
1397:
1395:
1394:
1390:
1387:
1386:
1382:
1379:
1378:
1377:Daemonosaurus
1374:
1371:
1370:
1366:
1365:
1362:
1359:
1356:
1351:
1347:
1343:
1338:
1334:
1313:
1312:
1311:
1310:
1308:
1306:
1302:
1294:
1290:
1289:
1288:
1287:
1284:
1281:
1279:
1275:
1271:
1263:
1259:
1256:
1254:
1250:
1247:
1245:
1241:
1239:
1235:
1233:
1229:
1228:
1223:
1219:
1215:
1208:
1203:
1201:
1196:
1194:
1189:
1188:
1185:
1178:
1170:
1164:
1162:
1158:
1154:
1148:
1145:
1142:
1137:
1134:
1128:
1125:
1121:
1117:
1111:
1108:
1104:
1100:
1094:
1091:
1086:
1082:
1078:
1072:
1068:
1067:
1059:
1056:
1051:
1044:
1041:
1036:
1032:
1028:
1024:
1020:
1016:
1009:
1002:
1000:
998:
996:
994:
992:
990:
986:
979:
975:
972:
971:
967:
965:
963:
959:
958:
953:
949:
948:
943:
939:
935:
934:
929:
925:
921:
920:
915:
912:
908:
907:
902:
901:
896:
895:
894:Iberomesornis
890:
882:
881:Pelecanimimus
878:
871:
864:
863:
855:
854:
846:
845:
837:
836:
828:
827:
824:
823:
822:
815:
814:
808:
807:
804:
803:
800:
799:
798:
791:
790:
784:
783:
780:
779:
776:
775:
774:
767:
766:
760:
759:
756:
755:
752:
751:
750:
749:Pelecanimimus
743:
742:
736:
735:
731:
730:
726:
724:
720:
718:
717:Pelecanimimus
714:
710:
706:
702:
698:
694:
690:
689:Pelecanimimus
686:
685:ornithomimids
682:
678:
677:Pelecanimimus
674:
670:
666:
665:Pelecanimimus
659:
657:
655:
651:
647:
643:
642:Pelecanimimus
639:
635:
631:
630:Pelecanimimus
626:
625:Pelecanimimus
622:
621:Pelecanimimus
618:
610:
606:
605:Pelecanimimus
601:
597:
595:
591:
587:
586:Pelecanimimus
583:
579:
575:
571:
567:
566:Pelecanimimus
563:
562:
554:
553:
548:
547:Pelecanimimus
544:
540:
538:
534:
530:
526:
521:
520:Pelecanimimus
514:
513:Pelecanimimus
509:
502:
500:
498:
494:
490:
486:
482:
478:
474:
473:Pelecanimimus
470:
466:
462:
458:
454:
449:
447:
445:
439:
437:
431:
430:specific name
427:
423:
420:
416:
412:
408:
404:
400:
396:
392:
388:
385:In July 1993
380:
378:
376:
372:
368:
364:
360:
356:
353:
349:
345:
342:
338:
334:
333:
332:Pelecanimimus
322:
319:Perez-Moreno
316:
314:
305:
302:
301:Binomial name
298:
294:
293:
287:
284:
283:
279:
275:
272:Perez-Moreno
269:
268:
267:Pelecanimimus
261:
258:
257:
254:
248:
245:
242:
241:
238:
232:
229:
226:
225:
222:
219:
216:
213:
212:
209:
206:
203:
200:
199:
196:
193:
190:
187:
186:
183:
180:
177:
176:
173:
170:
167:
166:
163:
160:
157:
156:
151:
146:
142:
139:
134:
130:
125:
111:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
50:
43:
37:
36:Pelecanimimus
33:
30:
19:
18:Pelicanimimus
5530:Paleontology
5417:
5381:
5310:
5303:
5296:
5289:
5282:
5277:Ornithomimus
5275:
5268:
5261:
5254:
5247:
5240:
5233:
5215:Tyrannomimus
5213:
5205:
5197:
5190:
5185:Deinocheirus
5183:
5176:
5156:
5150:
5149:
5142:
5118:
5110:
5105:Nqwebasaurus
5103:
5098:Nedcolbertia
5096:
5089:
5082:
5076:Calamosaurus
5074:
5068:Aviatyrannis
5066:
5059:
5034:
5004:
4997:
4990:
4982:
4975:
4968:
4961:
4953:
4948:Aristosuchus
4946:
4940:Aniksosaurus
4938:
4920:Aniksosaurus
4918:
4888:
4881:
4873:
4819:
4784:
4777:
4770:
4763:
4689:
4682:
4675:
4655:
4648:
4640:
4635:Dynamoterror
4633:
4625:
4605:
4598:
4578:
4571:
4551:
4546:Asiatyrannus
4544:
4520:
4513:
4484:Dryptosaurus
4482:
4474:
4467:
4436:
4429:
4422:
4414:
4407:
4400:
4393:
4386:
4379:
4374:Jinbeisaurus
4372:
4365:
4358:
4350:
4345:Aviatyrannis
4343:
4336:
4302:
4297:Sinotyrannus
4295:
4288:
4281:
4274:
4253:
4246:
4224:
4217:
4176:
4169:
4162:
4154:
4147:
4139:
4131:
4126:Bicentenaria
4124:
4116:
4111:Asiamericana
4109:
4102:
4043:
3984:
3977:
3970:
3963:
3956:
3949:
3942:
3921:
3913:
3906:
3899:
3892:
3886:Deltadromeus
3884:
3876:
3869:
3818:Tyrannotitan
3816:
3809:
3802:
3795:
3775:
3751:
3746:Taurovenator
3744:
3737:
3730:
3725:Lajasvenator
3723:
3716:
3709:
3704:Concavenator
3702:
3694:
3687:
3662:
3654:
3647:
3639:
3631:
3611:
3604:
3596:
3589:
3582:
3575:
3550:
3542:
3534:
3527:
3520:Allosauridae
3488:
3481:
3476:Shidaisaurus
3474:
3467:
3460:
3440:
3432:
3407:
3399:
3392:
3385:
3354:
3346:
3338:
3331:
3304:
3296:
3284:Avetheropoda
3271:Avetheropoda
3242:
3201:Avetheropoda
3168:
3161:
3154:Spinosaurini
3141:
3134:
3127:
3120:
3112:
3084:
3077:
3070:
3049:
3041:
3034:
3029:Protathlitis
3027:
3020:
2995:
2988:
2981:
2973:
2938:
2931:
2924:
2919:Leshansaurus
2917:
2910:
2903:
2883:
2876:
2871:Megalosaurus
2869:
2864:Duriavenator
2862:
2842:
2817:
2801:Megalosauria
2782:
2775:
2770:Marshosaurus
2768:
2763:Condorraptor
2761:
2741:
2700:
2693:
2685:
2678:
2671:
2664:
2657:
2650:
2644:Cruxicheiros
2642:
2635:
2627:
2619:
2611:
2476:
2471:Quilmesaurus
2469:
2462:
2455:
2447:
2440:
2433:
2426:
2419:
2399:
2392:
2384:
2377:
2370:
2363:
2356:
2349:
2342:
2334:
2323:Brachyrostra
2309:
2289:
2281:
2274:
2267:
2260:
2252:
2245:
2225:
2220:Tralkasaurus
2218:
2211:
2204:
2197:
2190:
2183:
2176:
2169:
2161:
2154:
2122:Vespersaurus
2120:
2115:Velocisaurus
2113:
2106:
2099:
2092:
2072:
2065:
2058:
2038:
2031:
2024:
2016:
2010:Deltadromeus
2008:
2000:
1994:Compsosuchus
1992:
1985:
1977:
1969:
1944:
1936:
1930:Lametasaurus
1928:
1920:
1886:
1881:Ceratosaurus
1879:
1859:
1851:
1844:
1836:
1829:
1813:Ceratosauria
1798:
1791:
1716:
1709:
1667:
1659:
1652:
1644:
1636:
1629:
1622:
1615:
1595:
1590:Podokesaurus
1588:
1581:
1574:
1566:
1542:
1534:
1527:
1520:
1513:
1506:
1499:
1492:
1485:
1478:
1473:Gojirasaurus
1471:
1466:Dracovenator
1464:
1456:
1449:
1437:Neotheropoda
1422:
1414:
1406:
1400:Guaibasaurus
1398:
1391:
1383:
1375:
1369:Chindesaurus
1367:
1257:
1248:
1168:
1152:
1147:
1136:
1127:
1119:
1115:
1110:
1102:
1098:
1093:
1065:
1058:
1049:
1043:
1018:
1014:
955:
945:
931:
919:Concavenator
917:
904:
898:
892:
886:
880:
872:Paleoecology
819:
818:
795:
794:
771:
770:
748:
747:
746:
732:
721:
716:
705:phylogenetic
696:
688:
680:
676:
672:
664:
663:
641:
629:
624:
620:
614:
604:
585:
569:
565:
559:
557:
552:Concavenator
550:
546:
525:premaxillary
519:
518:
512:
472:
457:LH 7777
456:
450:
441:
433:
425:
421:
414:
390:
384:
331:
330:
329:
320:
312:
307:
291:
290:
273:
266:
265:
243:
227:
214:
201:
188:
137:
35:
29:
5442:Wikispecies
5342:Maniraptora
5331:Maniraptora
5312:Tototlmimus
5242:Anserimimus
5192:Garudimimus
5178:Beishanlong
5120:Valdoraptor
5061:Arkansaurus
5006:Xunmenglong
4883:Juravenator
4677:Tarbosaurus
4553:Nanuqsaurus
4522:Gorgosaurus
4438:Megaraptora
4424:Timurlengia
4156:Sciurumimus
4104:Aratasaurus
3972:Murusraptor
3894:Fukuiraptor
3857:Megaraptora
3739:Shaochilong
3711:Eocarcharia
3664:Megaraptora
3584:Lusovenator
3462:Alpkarakush
3319:Carnosauria
3170:Spinosaurus
3143:Siamosaurus
3043:Suchosaurus
2990:Iberospinus
2926:Magnosaurus
2905:Afrovenator
2878:Torvosaurus
2629:Chilesaurus
2442:Carnotaurus
2435:Caletodraco
2421:Abelisaurus
2247:Arcovenator
2094:Laevisuchus
2085:Noasaurinae
1987:Berthasaura
1958:Noasauridae
1888:Genyodectes
1853:Lukousaurus
1846:Jubbulpuria
1646:Panguraptor
1624:Coelophysis
1617:Camposaurus
1576:Panguraptor
1568:Dracoraptor
1544:Zupaysaurus
1529:Tachiraptor
1508:Sarcosaurus
1408:Nhandumirim
1171:48:235-259.
947:Spinolestes
942:salamanders
797:Garudimimus
713:troodontids
617:gular pouch
503:Description
481:lagerstätte
448:) "tooth".
138:P. polyodon
5552:Categories
5270:Gallimimus
5199:Harpymimus
5144:Harpymimus
4765:Iliosuchus
4735:see below↓
4564:Alioramini
4381:Juratyrant
4367:Eotyrannus
4352:Bagaraatan
4304:Yutyrannus
4238:Coeluridae
4164:Vayuraptor
4010:see below↓
3986:Tratayenia
3979:Orkoraptor
3965:Megaraptor
3871:Aoniraptor
3804:Mapusaurus
3732:Sauroniops
3696:Altispinax
3649:Neovenator
3591:Siamraptor
3577:Datanglong
3544:Epanterias
3536:Antrodemus
3529:Allosaurus
3510:Allosauria
3333:Altispinax
3298:Gasosaurus
3210:see below↓
3086:Suchomimus
2687:Sinosaurus
2522:see below↓
2478:Viavenator
2428:Aucasaurus
2365:Ilokelesia
2336:Dahalokely
2291:Rajasaurus
2269:Indosaurus
2262:Genusaurus
2254:Dahalokely
2178:Indosuchus
2074:Limusaurus
2033:Ligabueino
2026:Kiyacursor
2018:Genusaurus
2002:Dahalokely
1922:Betasuchus
1793:Dornraptor
1693:see below↓
1669:Segisaurus
1522:Sinosaurus
1385:Eodromaeus
1314:see below↓
1262:Dinosauria
1244:Sauropsida
1122:: 587-588.
980:References
957:Europejara
952:pterosaurs
950:. Several
928:ornithopod
906:Eoalulavis
773:Harpymimus
681:Harpymimus
582:exaptation
574:adaptation
561:Harpymimus
537:heterodont
455:specimen,
335:(meaning "
208:Saurischia
195:Dinosauria
5542:Dinosaurs
5346:includes
5291:Rativates
5284:Qiupalong
4984:Scipionyx
4977:Mirischia
4650:Lythronax
4642:Labocania
4573:Alioramus
4226:Labocania
3944:Aerosteon
3490:Sinraptor
3394:Erectopus
3129:Irritator
2714:Orionides
2599:Tetanurae
2586:Tetanurae
2513:Tetanurae
2449:Elemgasem
2372:Llukalkan
2351:Elemgasem
2108:Noasaurus
1971:Afromimus
1779:Averostra
1766:Averostra
1684:Averostra
1536:Velocipes
1355:Theropoda
1342:Theropoda
1305:Theropoda
1230:Kingdom:
1214:Theropoda
1085:985402380
933:Iguanodon
930:dinosaur
900:Concornis
723:Cladogram
709:denticles
701:tetanuran
693:evolution
669:cladistic
529:maxillary
489:Barremian
422:pelecanus
357:from the
285:Species:
221:Theropoda
168:Kingdom:
162:Eukaryota
47:130
5462:47458714
5427:Wikidata
4875:Coelurus
4283:Kileskus
4276:Guanlong
4248:Coelurus
4141:Gualicho
3641:Gualicho
3022:Baryonyx
2358:Guemesia
2185:Kryptops
1946:Ozraptor
1583:Pendraig
1238:Chordata
1236:Phylum:
1232:Animalia
968:See also
924:sauropod
914:theropod
477:La Hoyas
453:holotype
375:theropod
355:dinosaur
182:Chordata
178:Phylum:
172:Animalia
158:Domain:
5516:Portals
5488:4127289
5475:4822643
5433:Q134151
4416:Timimus
4178:Zuolong
3908:Rapator
3811:Meraxes
3136:Oxalaia
2457:Koleken
1480:Lepidus
1242:Class:
1023:Bibcode
938:lizards
916:teeth,
711:(as in
650:sternal
609:keratin
533:dentary
493:sternum
341:extinct
337:pelican
310:†
289:†
259:Genus:
116:↓
5084:Hexing
4772:Kakuru
4360:Dilong
2401:Thanos
2379:Niebla
2199:Rugops
2192:Kurupi
1083:
1073:
1015:Nature
903:, and
891:birds
697:et al.
673:et al.
646:flight
594:radius
572:as an
570:et al.
495:, the
465:Cuenca
323:, 1994
321:et al.
274:et al.
5501:64117
5348:birds
4388:Moros
3915:Siats
3656:Siats
3598:Siats
1258:Clade
1249:Clade
1011:(PDF)
954:like
911:avian
634:hyoid
471:, of
469:Spain
444:odous
436:polys
426:mimus
419:Latin
367:teeth
363:Spain
348:basal
344:genus
244:Clade
228:Clade
215:Clade
202:Clade
189:Clade
5470:GBIF
5340:see
3958:Maip
1416:Tawa
1291:see
1081:OCLC
1071:ISBN
960:and
940:and
592:and
590:ulna
578:beak
451:The
278:1994
55:PreꞒ
5457:EoL
1120:154
1031:doi
1019:370
361:of
346:of
5554::
5498::
5485::
5472::
5459::
5444::
5429::
1260::
1251::
1160:^
1118:,
1079:.
1029:.
1017:.
1013:.
988:^
897:,
656:.
467:,
463:,
405:,
401:,
397:,
276:,
246::
230::
217::
204::
191::
105:Pg
49:Ma
44:,
5518::
5210:?
5202:?
5115:?
5079:?
5071:?
5039:?
4987:?
4958:?
4943:?
4878:?
4645:?
4630:?
4479:?
4441:?
4419:?
4355:?
4258:?
4240:?
4229:?
4159:?
4144:?
4136:?
4121:?
3925:?
3917:?
3889:?
3881:?
3859:?
3699:?
3667:?
3659:?
3644:?
3636:?
3601:?
3555:?
3547:?
3539:?
3437:?
3412:?
3404:?
3359:?
3351:?
3343:?
3309:?
3301:?
3117:?
3054:?
3046:?
3000:?
2978:?
2943:?
2822:?
2787:?
2690:?
2647:?
2632:?
2624:?
2616:?
2452:?
2389:?
2339:?
2314:?
2286:?
2257:?
2166:?
2021:?
2013:?
2005:?
1997:?
1982:?
1974:?
1949:?
1941:?
1933:?
1925:?
1856:?
1841:?
1803:?
1664:?
1649:?
1641:?
1571:?
1539:?
1461:?
1427:?
1419:?
1411:?
1403:?
1388:?
1380:?
1372:?
1206:e
1199:t
1192:v
1087:.
1037:.
1033::
1025::
611:.
263:†
250:†
234:†
110:N
100:K
95:J
90:T
85:P
80:C
75:D
70:S
65:O
60:Ꞓ
20:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
