406:
141:
1158:
375:
420:
119:
389:
was essentially a four-legged whale: however, it had webbed feet with small hooves on the tips of its toes, making it more capable of moving on land than modern seals. It featured sharp teeth and a long snout which suggests it fed on fish and/or crustaceans. From its caudal vertebrae, it has been
445:
Lambert, Olivier; Bianucci, Giovanni; Salas-Gismondi, Rodolfo; Di Celma, Claudio; Steurbaut, Etienne; Urbina, Mario; de Muizon, Christian (4 April 2019).
1261:
1210:
1276:
597:
532:
366:
reached the
Pacific Ocean and attained a near circumequatorial distribution while retaining functional weight-bearing limbs.
499:
562:
340:. Parts recovered include the jaw, front and hind legs, bits of spine, and tail. Olivier Lambert, a scientist at the
140:
447:"An amphibious whale from the Middle Eocene of Peru reveals early South Pacific dispersal of quadrupedal cetaceans"
341:
590:
1271:
1266:
1256:
1178:
1251:
1223:
309:
317:
1161:
583:
305:
504:
411:
359:
135:
1228:
721:
480:
470:
460:
425:
758:
451:
301:
1074:
975:
778:
1245:
840:
829:
801:
730:
710:
687:
676:
355:
69:
374:
1108:
1083:
1007:
984:
883:
869:
860:
815:
808:
769:
737:
537:
363:
268:
228:
215:
1201:
1138:
1090:
1056:
1035:
1000:
993:
939:
925:
849:
751:
696:
653:
643:
606:
337:
313:
290:
192:
44:
575:
118:
1124:
1117:
1049:
1042:
1021:
953:
946:
932:
897:
890:
876:
822:
794:
744:
703:
631:
465:
446:
401:
89:
54:
1131:
1063:
960:
911:
904:
787:
669:
662:
152:
94:
38:
484:
348:"fills in a crucial gap" about the evolution of whales and their spread.
1195:
1028:
625:
172:
84:
79:
64:
59:
49:
1215:
475:
1097:
1014:
321:
202:
99:
74:
637:
619:
391:
390:
suggested that it might have possessed a flattened tail similar to a
329:
298:
182:
162:
30:
1172:
373:
333:
286:
500:"Fossilized Remains of Ancient 4-Legged Whale Discovered in Peru"
563:"Unknown Species of Ancient Four-Legged Whale Uncovered in Peru"
325:
294:
1176:
579:
533:"Fossil of ancient four-legged whale with hooves discovered"
1185:
1107:
1073:
983:
974:
859:
839:
777:
768:
720:
686:
652:
354:is the first recorded quadrupedal whale from the
591:
8:
342:Royal Belgian Institute of Natural Sciences
1173:
980:
774:
598:
584:
576:
526:
524:
522:
304:. Its fossil was uncovered in 2011 in the
117:
20:
557:
555:
474:
464:
344:and lead author of the study, noted that
440:
438:
436:
432:
320:by a team consisting of members from
7:
14:
1157:
1156:
418:
404:
139:
1262:Eocene mammals of South America
531:Devlin, Hannah (4 April 2019).
1:
1277:Fossil taxa described in 2019
362:. The discovery reveals that
498:Wright, Pam (5 April 2019).
1293:
293:that lived in what is now
1152:
614:
466:10.1016/j.cub.2019.02.050
258:
253:
136:Scientific classification
134:
125:
116:
23:
16:Extinct genus of whales
383:
267:Lambert et al., 2019 (
1224:Paleobiology Database
377:
130:compared to a human.
378:Life restoration of
248:Lambert et al., 2019
505:The Weather Channel
459:(8): 1352β1359.e3.
412:Paleontology portal
360:Southern Hemisphere
126:Size silhouette of
384:
382:resting at a cliff
1239:
1238:
1179:Taxon identifiers
1170:
1169:
1148:
1147:
970:
969:
722:Remingtonocetidae
310:Paracas Formation
278:
277:
272:
249:
1284:
1232:
1231:
1219:
1218:
1206:
1205:
1204:
1174:
1160:
1159:
981:
775:
609:genera by family
600:
593:
586:
577:
571:
570:
559:
550:
549:
547:
545:
528:
517:
516:
514:
512:
495:
489:
488:
478:
468:
442:
428:
426:Cetaceans portal
423:
422:
421:
414:
409:
408:
407:
318:Playa Media Luna
266:
247:
240:
227:
214:
144:
143:
121:
111:
41:
29:Temporal range:
21:
1292:
1291:
1287:
1286:
1285:
1283:
1282:
1281:
1272:Fossils of Peru
1242:
1241:
1240:
1235:
1227:
1222:
1214:
1209:
1200:
1199:
1194:
1181:
1171:
1166:
1144:
1103:
1069:
966:
855:
835:
764:
759:Remingtonocetus
716:
682:
648:
610:
604:
574:
569:. 4 April 2019.
561:
560:
553:
543:
541:
530:
529:
520:
510:
508:
497:
496:
492:
452:Current Biology
444:
443:
434:
424:
419:
417:
410:
405:
403:
400:
372:
246:
238:
225:
212:
138:
112:
110:
109:
108:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
36:
35:
33:
27:
17:
12:
11:
5:
1290:
1288:
1280:
1279:
1274:
1269:
1267:Paleogene Peru
1264:
1259:
1254:
1244:
1243:
1237:
1236:
1234:
1233:
1220:
1207:
1191:
1189:
1183:
1182:
1177:
1168:
1167:
1165:
1164:
1153:
1150:
1149:
1146:
1145:
1143:
1142:
1135:
1128:
1121:
1113:
1111:
1105:
1104:
1102:
1101:
1094:
1087:
1079:
1077:
1075:Basilosaurinae
1071:
1070:
1068:
1067:
1060:
1053:
1046:
1039:
1032:
1025:
1018:
1011:
1004:
997:
989:
987:
978:
976:Basilosauridae
972:
971:
968:
967:
965:
964:
957:
950:
943:
936:
929:
922:
915:
908:
901:
894:
887:
880:
873:
865:
863:
857:
856:
854:
853:
845:
843:
837:
836:
834:
833:
826:
819:
812:
805:
798:
791:
783:
781:
779:Georgiacetinae
772:
766:
765:
763:
762:
755:
748:
741:
734:
726:
724:
718:
717:
715:
714:
707:
700:
692:
690:
684:
683:
681:
680:
673:
666:
658:
656:
650:
649:
647:
646:
640:
634:
628:
622:
615:
612:
611:
605:
603:
602:
595:
588:
580:
573:
572:
551:
518:
490:
431:
430:
429:
415:
399:
396:
371:
368:
306:Yumaque Member
276:
275:
274:
273:
256:
255:
251:
250:
236:
232:
231:
223:
219:
218:
210:
206:
205:
200:
196:
195:
190:
186:
185:
180:
176:
175:
170:
166:
165:
160:
156:
155:
150:
146:
145:
132:
131:
123:
122:
114:
113:
105:
104:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
42:
28:
15:
13:
10:
9:
6:
4:
3:
2:
1289:
1278:
1275:
1273:
1270:
1268:
1265:
1263:
1260:
1258:
1257:Lutetian life
1255:
1253:
1250:
1249:
1247:
1230:
1225:
1221:
1217:
1212:
1208:
1203:
1197:
1193:
1192:
1190:
1188:
1184:
1180:
1175:
1163:
1155:
1154:
1151:
1141:
1140:
1136:
1134:
1133:
1129:
1127:
1126:
1122:
1120:
1119:
1115:
1114:
1112:
1110:
1106:
1100:
1099:
1095:
1093:
1092:
1088:
1086:
1085:
1081:
1080:
1078:
1076:
1072:
1066:
1065:
1061:
1059:
1058:
1054:
1052:
1051:
1047:
1045:
1044:
1040:
1038:
1037:
1033:
1031:
1030:
1026:
1024:
1023:
1019:
1017:
1016:
1012:
1010:
1009:
1005:
1003:
1002:
998:
996:
995:
991:
990:
988:
986:
982:
979:
977:
973:
963:
962:
958:
956:
955:
951:
949:
948:
944:
942:
941:
937:
935:
934:
930:
928:
927:
923:
921:
920:
916:
914:
913:
909:
907:
906:
902:
900:
899:
895:
893:
892:
888:
886:
885:
881:
879:
878:
874:
872:
871:
867:
866:
864:
862:
858:
852:
851:
847:
846:
844:
842:
841:Makaracetinae
838:
832:
831:
830:Pontobasileus
827:
825:
824:
820:
818:
817:
813:
811:
810:
806:
804:
803:
802:Carolinacetus
799:
797:
796:
792:
790:
789:
785:
784:
782:
780:
776:
773:
771:
767:
761:
760:
756:
754:
753:
749:
747:
746:
742:
740:
739:
735:
733:
732:
731:Andrewsiphius
728:
727:
725:
723:
719:
713:
712:
711:Himalayacetus
708:
706:
705:
701:
699:
698:
694:
693:
691:
689:
688:Ambulocetidae
685:
679:
678:
677:Ichthyolestes
674:
672:
671:
667:
665:
664:
660:
659:
657:
655:
651:
645:
641:
639:
635:
633:
629:
627:
623:
621:
617:
616:
613:
608:
601:
596:
594:
589:
587:
582:
581:
578:
568:
564:
558:
556:
552:
540:
539:
534:
527:
525:
523:
519:
507:
506:
501:
494:
491:
486:
482:
477:
472:
467:
462:
458:
454:
453:
448:
441:
439:
437:
433:
427:
416:
413:
402:
397:
395:
393:
388:
381:
376:
369:
367:
365:
361:
357:
356:Pacific Ocean
353:
349:
347:
343:
339:
335:
331:
327:
323:
319:
315:
311:
307:
303:
300:
299:Middle Eocene
296:
292:
288:
284:
283:
270:
265:
264:
260:
259:
257:
252:
245:
244:
237:
234:
233:
230:
224:
221:
220:
217:
211:
208:
207:
204:
201:
198:
197:
194:
191:
188:
187:
184:
181:
178:
177:
174:
171:
168:
167:
164:
161:
158:
157:
154:
151:
148:
147:
142:
137:
133:
129:
124:
120:
115:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
40:
32:
31:Middle Eocene
26:
22:
19:
1252:Protocetidae
1186:
1137:
1130:
1123:
1116:
1109:Pachycetinae
1096:
1089:
1084:Basilosaurus
1082:
1062:
1055:
1048:
1041:
1034:
1027:
1020:
1013:
1008:Cynthiacetus
1006:
999:
992:
985:Dorudontinae
959:
952:
945:
938:
931:
924:
918:
917:
910:
903:
896:
889:
884:Crenatocetus
882:
875:
870:Aegyptocetus
868:
861:Protocetinae
848:
828:
821:
816:Natchitochia
814:
809:Georgiacetus
807:
800:
793:
786:
770:Protocetidae
757:
750:
743:
738:Attockicetus
736:
729:
709:
702:
695:
675:
668:
661:
644:Artiodactyla
566:
542:. Retrieved
538:The Guardian
536:
509:. Retrieved
503:
493:
476:11581/425570
456:
450:
386:
385:
379:
351:
350:
345:
281:
280:
279:
263:P. pacificus
262:
261:
242:
241:
229:Protocetinae
216:Protocetidae
199:Infraorder:
193:Artiodactyla
127:
24:
18:
1187:Peregocetus
1139:Platyosphys
1091:Basiloterus
1057:Supayacetus
1036:Pontogeneus
1001:Chrysocetus
994:Ancalecetus
940:Qaisracetus
926:Phiomicetus
919:Peregocetus
850:Makaracetus
752:Kutchicetus
697:Ambulocetus
654:Pakicetidae
607:Archaeocete
387:Peregocetus
380:Peregocetus
370:Description
364:protocetids
352:Peregocetus
346:Peregocetus
338:Netherlands
314:Pisco Basin
297:during the
291:early whale
282:Peregocetus
243:Peregocetus
222:Subfamily:
128:Peregocetus
25:Peregocetus
1246:Categories
1125:Pachycetus
1118:Antaecetus
1050:Stromerius
1043:Saghacetus
1022:Masracetus
954:Takracetus
947:Rodhocetus
933:Protocetus
898:Gaviacetus
891:Dhedacetus
877:Artiocetus
823:Pappocetus
795:Babiacetus
745:Dalanistes
704:Gandakasia
632:Vertebrata
630:Subphylum
398:References
336:, and the
37:42.6
1202:Q62834728
1132:Perucetus
1064:Zygorhiza
961:Togocetus
912:Maiacetus
905:Indocetus
788:Aegicetus
670:Nalacetus
663:Pakicetus
312:) of the
159:Kingdom:
153:Eukaryota
1216:10515616
1196:Wikidata
1162:Category
1029:Ocucajea
638:Mammalia
626:Chordata
620:Animalia
618:Kingdom
485:30955933
358:and the
254:Species
209:Family:
183:Mammalia
173:Chordata
169:Phylum:
163:Animalia
149:Domain:
1098:Eocetus
1015:Dorudon
624:Phylum
567:Gizmodo
544:4 April
511:5 April
322:Belgium
235:Genus:
203:Cetacea
189:Order:
179:Class:
106:↓
1229:384282
642:Order
636:Class
483:
392:beaver
330:France
334:Italy
302:epoch
287:genus
285:is a
1211:GBIF
546:2019
513:2019
481:PMID
326:Peru
295:Peru
269:type
45:Preκ
471:hdl
461:doi
316:at
289:of
1248::
1226::
1213::
1198::
565:.
554:^
535:.
521:^
502:.
479:.
469:.
457:29
455:.
449:.
435:^
394:.
332:,
328:,
324:,
95:Pg
39:Ma
599:e
592:t
585:v
548:.
515:.
487:.
473::
463::
308:(
271:)
239:β
226:β
213:β
100:N
90:K
85:J
80:T
75:P
70:C
65:D
60:S
55:O
50:κ
34:~
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
