3524:). The names Hippomorpha and Ceratomorpha were introduced in 1937 by Horace Elmer Wood, in response to criticism of the name "Solidungula" that he proposed three years previously. It had been based on the grouping of horses and Tridactyla and on the rhinoceros/tapir complex. The extinct brontotheriidae were also classified under Hippomorpha and therefore possess a close relationship to horses. Some researchers accept this assignment because of similar dental features, but there is also the view that a very basal position within the odd-toed ungulates places them rather in the group of
3553:. The name "Tapiromorpha" goes back to Ernst Haeckel, who coined it in 1873, but it was long considered synonymous to Ceratomorpha because Wood had not considered it in 1937 when Ceratomorpha were named, since the term had been used quite differently in the past. Also in 1984, Robert M. Schoch used the conceptually similar term Moropomorpha, which today applies synonymously to Tapiromorpha. Included within the Tapiromorpha are the now extinct Isectolophidae, a sister group of the Ancylopoda-Ceratomorpha group and thus the most primitive members of this relationship complex.
3733:, which occupied similar ecological niches and had a much more efficient digestive system, is also associated with the decline in diversity of odd-toed ungulates. A significant cause for the decline of perissodactyls was climate change during the Miocene, leading to a cooler and drier climate accompanied by the spread of open landscapes. However, some lines flourished, such as the horses and rhinos; anatomical adaptations made it possible for them to consume tougher grass food. This led to open land forms that dominated newly created landscapes. With the emergence of the
2698:
3917:
3641:
1149:, meaning offspring are born already quite independent: for example, young horses can begin to follow the mother after a few hours. The young are nursed for a relatively long time, often into their second year, with rhinos reaching sexual maturity around eight or ten years old, but horses and tapirs maturing around two to four years old. Perissodactyls are long-lived, with several species, such as rhinos, reaching an age of almost 50 years in captivity.
269:
776:
1046:, a heat exchange that reduces the dependence of the temperature of the brain on that of the body. As a result, perissodactyls have limited thermoregulatory flexibility compared to artiodactyls which has restricted them to habitats of low seasonality and rich in food and water, such as tropical forests. In contrast, artiodactyls occupy a wide range of habits ranging from the Arctic Circle to deserts and tropical savannahs.
7628:
557:
2653:
3756:
2671:
846:
227:
38:
1055:
3721:, one of the largest mammals ever to walk the earth, evolved during this era. They weighed up to 20 metric tons (22 short tons) and lived throughout the Oligocene in Eurasia. About 20 million years ago, at the onset of the Miocene, the perissodactyls first reached Africa when it became connected to Eurasia because of the closing of the
755:, had a withers height of only 30 to 60 cm (12 to 24 in). Apart from dwarf varieties of the domestic horse and donkey, living perissodactyls reach a body length of 180–420 cm (71–165 in) and a weight of 150 to 4,500 kg (330 to 9,920 lb). While rhinos have only sparse hair and exhibit a thick
1134:
2728:. They were generally characterized in their late phase by a bony horn at the transition from the nose to the frontal bone and flat molars suitable for chewing soft plant food. The Brontotheroidea, which were almost exclusively confined to North America and Asia, died out at the beginning of the Upper
3932:
of both species began several millennia BCE. Due to the motorisation of agriculture and the spread of automobile traffic, such use has declined sharply in
Western industrial countries; riding is usually undertaken more as a hobby or sport. In less developed regions of the world, traditional uses for
3535:
were seen as members of
Hippomorpha, and presented as such in 1941. William Berryman Scott thought that, as claw-bearing perissodactyls, they belong in the new suborder Ancylopoda (where Ceratomorpha and Hippomorpha as odd-toed ungulates were combined in the group of Chelopoda). The term Ancylopoda,
3580:
of East Asia is often considered to be one of the oldest close relatives of the ungulates. Its 8 cm skull must have belonged to a very small and primitive animal with a π-shaped crown pattern on the enamel of its rear molars similar to that of perissodactyls and their relatives, especially the
1000:
shedding, prematurely reintroducing an infected horse may risk exposing other horses to the infection, even though the shedding horse appears well and may have previously returned negative samples. The function of the guttural pouch has been difficult to determine, but it is now believed to play a
783:
The main axes of both the front and rear feet pass through the third toe, which is always the largest. The remaining toes have been reduced in size to varying degrees. Tapirs, which are adapted to walking on soft ground, have four toes on their fore feet and three on their hind feet. Living rhinos
1421:
Odd-toed ungulates (Perissodactyla) comprise three living families with around 17 species—in horses, however, the exact count is still controversial. Rhinos and tapirs are more closely related to each other than to horses. The separation of horses from other perissodactyls took place according to
4841:
Frido Welker; Matthew J. Collins; Jessica A. Thomas; Marc
Wadsley; Selina Brace; Enrico Cappellini; Samuel T. Turvey; Marcelo Reguero; Javier N. Gelfo; Alejandro Kramarz; Joachim Burger; Thomas Jane Oates; David A. Ashford; Peter D. Ashton; Keri Rowsell; Duncan M. Porter; Benedikt Kessler; Roman
1108:
Perissodactyls inhabit a number of different habitats, leading to different lifestyles. Tapirs are solitary and inhabit mainly tropical rainforests. Rhinos tend to live alone in rather dry savannas, and in Asia, wet marsh or forest areas. Horses inhabit open areas such as grasslands, steppes, or
3544:
saw all three major groups of odd-toed ungulates as peers, based on the extremely long and independent phylogenetic development of the three lines. In the 1980s, Jeremy J. Hooker saw a general similarity between
Ancylopoda and Ceratomorpha based on dentition, especially in the earliest members,
3511:
in 1848, the present-day representatives were considered equal in rank. In the first half of the 20th century, a more systematic differentiation of odd-toed ungulates began, based on a consideration of fossil forms, and they were placed in two major suborders: Hippomorpha and
Ceratomorpha. The
3619:
The alignment of hyopsodontids and phenacodontids to
Perissodactyla in general suggests an older Laurasian origin and distribution for the clade, dispersed across the northern continents already in the early Paleocene. These forms already show a fairly well-developed molar morphology, with no
3711:, known from North America, reached a withers height of 2.5 m (8 ft 2 in) and could have weighed just over 3 metric tons (3.3 short tons). The decline of brontotheres at the end of the Eocene is associated with competition arising from the advent of more successful herbivores.
784:
have three toes on both the front and hind feet. Modern equines possess only a single toe; however, their feet are equipped with hooves, which almost completely cover the toe. Rhinos and tapirs, by contrast, have hooves covering only the leading edge of the toes, with the bottom being soft.
1377:, a very diverse group of mammals living from the Paleocene to the Pleistocene in South America, whose systematic unity is largely unexplained. Some of these were classified based on their paleogeographic distribution. However, a close relationship can be worked out to perissodactyls by
1066:
Most extant perissodactyl species occupy a small fraction of their original range. Members of this group are now found only in
Central and South America, eastern and southern Africa, and central, southern, and southeastern Asia. During the peak of odd-toed ungulate existence, from the
1099:
have reduced the surviving perissodactyl species to fragmented populations. In contrast, domesticated horses and donkeys have gained a worldwide distribution, and feral animals of both species are now also found in regions outside their original range, such as in
Australia.
1029:
is very long, reaching up to 26 m (85 ft) in horses. Extraction of nutrients from food is relatively inefficient, which probably explains why no odd-toed ungulates are small; nutritional requirements per unit of body weight are lower for large animals, as their
3819:. He also separated the rhinos from the rodents, but did not combine the three families now known as the odd-toed ungulates. In the transition to the 19th century, the individual perissodactyl genera were associated with various other groups, such as the
4907:
Ross MacPhee; Frido Welker; Jessica Thomas; Selina Brace; Enrico
Cappellini; Samuel Turvey; Ian Barnes; Marcelo Reguero; Javier Gelfo; Alejandro Kramarz (2014). "Ancient protein sequencing Resolves litoptern and notoungulate superordinal affinities".
4842:
Fischer; Carsten
Baessmann; Stephanie Kaspar; Jesper V. Olsen; Patrick Kiley; James A. Elliott; Christian D. Kelstrup; Victoria Mullin; Michael Hofreiter; Eske Willerslev; Jean-Jacques Hublin; Ludovic Orlando; Ian Barnes; Ross DE MacPhee (2015).
5776:
Kenneth D. Rose; Thierry Smith; Rajendra S. Rana; Ashok Sahni; H. Singh; A. Pieter Missiaen (2006). "Early Eocene (Ypresian) Continental vertebrate assemblage from India, with description of a new anthracobunid (Mammalia, tethytheria )".
5367:
Gong, Y.; Wang, Y.; et al. (December 2019). "Dietary adaptations and palaeoecology of Lophialetidae (Mammalia, Tapiroidea) from the Eocene of the Erlian Basin, China: combined evidence from mesowear and stable isotope analyses".
3714:
More successful lines of odd-toed ungulates emerged at the end of the Eocene when dense jungles gave way to steppe, such as the chalicotheriid rhinos, and their immediate relatives; their development also began with very small forms.
1405:. This kinship group is included among the Euungulata, which also contains the even-toed ungulates and whales (Artiodactyla). The separation of the Litopterna-Notoungulata group from the perissodactyls probably took place before the
2845:, are now understood to be part of the odd-toed ungulate assemblage. Phenacodontids seem to be stem-perissodactyls, while hyopsodontids are closely related to horses and brontotheres, despite their more primitive overall appearance.
2786:
Rhinocerotoidea (rhino relatives) included a large variety of forms from the Eocene up to the Oligocene, including dog-size leaf feeders, semiaquatic animals, and also huge long-necked animals. Only a few had horns on the nose. The
2821:
reached their greatest diversity in the Eocene, when several families lived in Eurasia and North America. They retained a primitive physique and were noted for developing a trunk. The extinct families within this group include the
3608:, indicating that it is actually a member of a sister group. Ancestors of Perissodactyla may have arrived via an island bridge from the Afro-Arab landmass onto the Indian subcontinent as it drifted north towards Asia. A study on
3603:
clade. The saddle-shaped configuration of the navicular joints and the mesaxonic construction of the front and hind feet also indicates a close relationship to Tethytheria. However, this construction deviates from that of
3741:, perissodactyls and other megafauna were given access to one of their last habitable continents: South America. However, many perissodactyls became extinct at the end of the ice ages, including American horses and the
5668:
Robert Milton Schoch (1984). "Two unusual specimens of the Yale Peabody Museum Helaletes in collections, and some comments on the ancestry of the Tapiridae (Perissodactyla, Mammalia)". Peabody Museum, Yale University:
995:
colonising their guttural pouch can continue to intermittently shed the bacteria for several months, and should be isolated from other horses during this time to prevent transmission. Due to the intermittent nature of
4684:
Dan Graur; Manolo Gouy; Laurent Duret (1997). "Evolutionary Affinities of the Order Perissodactyla and the Phylogenetic Status of the Superordinal taxa Ungulata and Altungulata Molecular Phylogenetics and Evolution".
1409:. "Condylarths" can probably be considered the starting point for the development of the two groups, as they represent a heterogeneous group of primitive ungulates that mainly inhabited the northern hemisphere in the
3907:
was held well into the 20th century. Only with the advent of molecular genetic research methods had it been recognized that the hyrax was not closely related to perissodactyls but rather to elephants and manatees.
1144:
Odd-toed ungulates are characterized by a long gestation period and a small litter size, usually delivering a single young. The gestation period is 330–500 days, being longest in rhinos. Newborn perissodactyls are
4671:
Ballenger, Liz; Myers, Phil. "Rhinocerotidae (rhinoceroses)". Animal Diversity Web. Retrieved 2021-03-27."Ballenger, Liz; Myers, Phil. "Rhinocerotidae (rhinoceroses)". Animal Diversity Web. Retrieved 2021-03-27.
2867:. However, later studies have shown that, while anthracobunids are definite perissodactyls, desmostylians have enough mixed characters to suggest that a position among the Afrotheria is not out of the question.
2753:. In contrast, the horse family (Equidae) flourished and spread. Over time this group saw a reduction in toe number, extension of the limbs, and the progressive adjustment of the teeth for eating hard grasses.
6168:
Pieter Missiaen; Gregg F. Gunnell; Philip D. Gingerich (2011). "New brontotheriidae (Mammalia, Perissodactyla) from the Early and Middle Eocene of Pakistan with implications for mammalian paleobiogeography".
1109:
semi-deserts, and live together in groups. Odd-toed ungulates are exclusively herbivores that feed, to varying degrees, on grass, leaves, and other plant parts. A distinction is often made between primarily
689:(even-toed ungulates) bear most of their weight equally on four or two (an even number) of the five toes: their third and fourth toes. Another difference between the two is that perissodactyls digest plant
681:). They typically have reduced the weight-bearing toes to three or one of the five original toes, though tapirs retain four toes on their front feet. The nonweight-bearing toes are either present, absent,
3687:
from the group of brontotheriidae. Initially, the members of the different lineages looked quite similar, with an arched back and generally four toes on the front and three on the hind feet.
3747:. Whether over-hunting by humans (overkill hypothesis), climatic change, or a combination of both factors was responsible for the extinction of ice age mega-fauna, remains controversial.
6397:Étienne Geoffroy Saint-Hilaire; Georges Cuvier (1795). "Memoire sur une nouvelle division of Mammifères, et sur les principes qui doivent servir de base dans cette sorte de travail".
794:
ungulates with only one digit or hoof have decreased mobility in their limbs, which allows for faster running speeds and agility. Differences in limb structure and physiology between
3540:
in 1889, had been established for chalicotheres. However, further morphological studies from the 1960s showed a middle position of Ancylopoda between Hippomorpha and Ceratomorpha.
2859:, but they may actually represent stem-perissodactyls. They are an early lineage of mammals that took to the water, spreading across semi-aquatic to fully marine niches in the
6497:
Rodolphe Tabuce; Laurent Marivaux; Mohammed Adaci; Mustapha Bensalah; Jean-Louis Hartenberger; Mohammed Mahboubi; Fateh Mebrouk; Paul Tafforeau; Jean-Jacques Jaeger (2007).
906:
can be very small or completely absent, as in the two African species of rhinoceros. In horses, usually only the males possess canines. The surface shape and height of the
3854:(1777–1850) classified ungulates by the structure of their feet, differentiating those with an even number of toes from those with an odd number. He moved the horses as
3653:
The Perissodactyla appeared relatively abruptly at the beginning of the Lower Paleocene about 63 million years ago, both in North America and Asia, in the form of
5890:
Rose, Kenneth D.; Holbrook, Luke T.; Kumar, Kishor; Rana, Rajendra S.; Ahrens, Heather E.; Dunn, Rachel H.; Folie, Annelise; Jones, Katrina E.; Smith, Thierry (2019).
4155:
5468:
Luke T. Holbrook; Joshua Lapergola (2011). "A new genus of perissodactyl (Mammalia) from the Bridgerian of Wyoming, with comments on basal perissodactyl phylogeny".
7835:
910:
is heavily dependent on whether soft leaves or hard grass make up the main component of their diets. Three or four cheek teeth are present on each jaw half, so the
3948:
3699:, the first representative of the rhino and tapir line. All were small compared to later forms and lived as fruit and foliage eaters in forests. The first of the
7887:
3936:
In contrast, the existence in the wild of almost all other odd-toed ungulates species has declined dramatically because of hunting and habitat destruction. The
6037:
Ross Secord; Jonathan I. Bloch; Stephen GB Chester; Doug M. Boyer; Aaron R. Wood; Scott L. Wing; Mary J. Kraus; Francesca A. McInerney; John Krigbaum (2012).
4998:
Mario A. Cozzuol; Camila L. Clozato; Elizete C. Holanda; Flávio HG Rodrigues; Samuel Nienow; Benoit de Thoisy; Rodrigo Redondo; AF Fabrício R. Santos (2013).
5831:
Kenneth D. Rose; Luke T. Holbrook; Rajendra S. Rana; Kishor Kumar; Katrina E. Jones; Heather E. Ahrens; Pieter Missiaen; Ashok Sahni; Thierry Smith (2014).
7961:
4925:"Phylogenetic Relationships of the Five Extant Rhinoceros species (Rhinocerotidae, Perissodactyla) Based on Mitochondrial Cytochrome b and 12S rRNA gene"
1180:
and other "ungulates". A close family relationship with hyraxes was suspected based on similarities in the construction of the ear and the course of the
3629:
1406:
5759:"New Early Eocene cambaythere (Perissodactyla, Mammalia) from the Vastan Lignite Mine (Gujarat, India) and on evaluation of cambaythere relationships"
3729:
also entered the ancient settlement areas of odd-toed ungulates, creating competition that led to the extinction of some of their lines. The rise of
7809:
7159:
4137:
7848:
5892:"Anatomy, Relationships, and Paleobiology of Cambaytherium (Mammalia, Perissodactylamorpha, Anthracobunia) from the lower Eocene of western India"
6338:
Ludovic Orlando; Jessica L. Metcalf; Maria T. Alberdi; Miguel Telles Antunes-Dominique Bonjean; Marcel Otte; Fabiana Martin; et al. (2009).
4591:
1239:). In a discredited alternative scenario, a close relationship exists between perissodactyls, carnivores, and bats, this assembly comprising the
3507:
Relationships within the large group of odd-toed ungulates are not fully understood. Initially, after the establishment of "Perissodactyla" by
4383:
6651:
6244:
4630:
3839:, tapirs and hyrax. The horses were still generally regarded as a group separate from other mammals and were often classified under the name
1025:
is present. The stomach of perissodactyls is simply built, while the cecum accommodates up to 90 L (24 US gal) in horses. The
4295:"What are the most accurate categories for mammal tarsus arrangement? A review with attention to South American Notoungulata and Litopterna"
6127:"New Early Eocene tapiromorph perissodactyls from the Ghazij formation of Pakistan, with implications for mammalian biochronology in Asia"
3891:, which he used for what are today called the odd-toed ungulates, including their extinct relatives, but explicitly excluding the hyrax.
991:
infection) is a highly transmissible respiratory infection of horses that can cause pus to accumulate in the guttural pouch; horses with
7976:
4445:
4366:
1373:
According to studies published in March 2015, odd-toed ungulates are in a close family relationship with at least some of the so-called
749:, is considered one of the largest land mammals of all time. At the other extreme, an early member of the order, the prehistoric horse
3851:
4410:
6689:
6635:
6618:
6603:
6588:
6573:
6412:
Georges Cuvier (1817). "Le Règne Animal distribue d'après son organisation pour servir de base à l'histoire naturelle des animaux".
6269:
6227:
Christine Janis (2008). "An evolutionary history of browsing and grazing ungulates". In Iain J. Gordon; Herbert H. T. Prins (eds.).
4213:
121:
2767:. The Chalicotheriidae developed claws instead of hooves and considerable extension of the forelegs. The best-known genera include
5931:"Convergence of Afrotherian and Laurasiatherian Ungulate-Like Mammals: First Morphological Evidence from the Paleocene of Morocco"
5125:"Convergence of Afrotherian and Laurasiatherian Ungulate-Like Mammals: First Morphological Evidence from the Paleocene of Morocco"
3933:
these animals are, however, still widespread. To a lesser extent, horses and donkeys are also kept for their meat and their milk.
3928:
The domestic horse and the donkey play an important role in human history, particularly as transport, work and pack animals. The
2582:
1075:, perissodactyls were distributed over much of the globe, the major exceptions being Australia and Antarctica. Horses and tapirs
6038:
1374:
55:
5406:
Cooper, L. N.; Seiffert, E. R.; Clementz, M.; Madar, S. I.; Bajpai, S.; Hussain, S. T.; Thewissen, J. G. M. (8 October 2014).
7866:
2516:
2507:
2456:
102:
59:
7853:
5990:"Eutherians experienced elevated evolutionary rates in the immediate aftermath of the Cretaceous–Palaeogene mass extinction"
4163:
3566:
The evolutionary development of Perissodactyla is well documented in the fossil record. Numerous finds are evidence of the
74:
7874:
268:
7152:
2692:
There are many perissodactyl fossils of multivariant form. The major lines of development include the following groups:
2626:
2341:
1381:
and comparison with fossil collagen from remnants of phylogenetically young members of the Meridiungulata (specifically
1031:
727:
Despite their very different appearances, they were recognized as related families in the 19th century by the zoologist
865:, giving them an elongated head. The various forms of snout between families are due to differences in the form of the
7861:
7731:
7708:
6626:
6208:(Perissodactyla, Mammalia) in den eozänen Geiseltalschichten" [Tooth morphology and stratigraphic distribution of
4203:
81:
5814:"Comments on 'Early Eocene Land Mammals from Vastan Lignite Mine, District Surat (Gujarat), western India' by Bajpai"
1401:). Both kinship groups, the odd-toed ungulates and the Litopterna-Notoungulata, are now in the higher-level taxon of
7981:
7788:
2697:
1076:
4077:
3899:, but it was sometimes also used for the true odd-toed ungulates as a subcategory (rhinos, horses, tapirs), while
48:
7892:
7801:
7137:
4045:
2545:
2536:
2444:
4240:"Evolutionary and Functional Anatomy of the Pelvic Limb in Fossil and Recent Equidae (Perissodactyla, Mammalia)"
88:
3812:
2588:
2498:
2486:
1929:
6564:
Martin S. Fischer: Mesaxonia (Perissodactyla) Perissodactyla. In: Wilfried Westheide, Reinhard Rieger (eds.):
5790:
4462:
959:) can cause serious damage to the tissues of the pouch, as well as surrounding structures including important
7966:
7632:
7145:
4037:
2811:
2477:
980:
874:
5220:
Ravel, Anthony; Orliac, Maeva (2014). "The inner ear morphology of the 'condylarthran' Hyopsodus lepidus".
5032:
4102:
Ravel, Anthony; Orliac, Maeva (2014). "The inner ear morphology of the 'condylarthran' Hyopsodus lepidus".
1219:
as a deeply nested subclade) are the sister taxon of the Perissodactyla; together, the two groups form the
70:
7971:
7648:
5641:
JJ Hooker (1984). "A primitive ceratomorph (Perissodactyla, Mammalia) from the Early Tertiary of Europe".
2468:
1158:
1026:
964:
628:
4141:
7923:
6682:
6455:
5674:
4056:
3956:
3882:
7291:
5626:, a new early Eocene chalicothere (Mammalia, Perissodactyla), and a revision of Eocene chalicotheres".
3941:
2086:
7736:
4599:
3158:
7902:
7757:
7695:
7594:
6351:
6050:
5942:
5891:
5844:
5419:
5259:(Mammalia, Perissodactyla) from Zambrana (Álava, Basque-Cantabrian Region). Phylogenetic analysis of
5136:
5077:
4855:
4792:
4014:
2815:). The rhinos (Rhinocerotidae) emerged in the Middle Eocene; five species survive to the present day.
2379:
1972:
1954:
1192:
1138:
1122:
1058:
Restriction of their habitat and poaching threaten the survival of most rhino species, including the
323:
251:
7686:
7484:
6039:"Evolution of the Earliest Horses Driven by Climate Change in the Paleocene-Eocene Thermal Maximum"
5589:
The Mammalian Fauna of the White River Oligocene Transactions of the American Philosophical Society
4552:"Perissodactyl | Definition, Characteristics, Species, Examples, Habitat, & Facts | Britannica"
3964:
3537:
3028:
2602:
2332:
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2267:
1877:
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802:. For example, often shorter, thicker, bones belong to the largest and heaviest ungulates like the
7387:
6447:
6286:
3999:
3640:
3021:
2911:
2352:
7442:
6186:
6074:
5911:
5794:
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5485:
5385:
5313:
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4889:
4761:
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4119:
4006:
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3567:
2617:
2167:
1805:
1378:
1169:
892:
263:
3916:
3620:
intermediary forms as evidence of the course of its development. The close relationship between
2294:
940:, is a characteristic feature of Perissodactyla. The guttural pouch is of particular concern in
7928:
6204:
Hlawatsch, Kerstin; Erfurt, Jörg (2007). "Zahnmorphologie und stratigraphische Verbreitung von
4843:
4781:"Pegasoferae, an unexpected mammalian clade revealed by tracking ancient retroposon insertions"
3815:(1723–1806) first separated the tapirs and hippos in 1762 with the introduction of the concept
3034:
779:
Seven figures showing the bones, blood vessels, ligaments and arteries of the hoof and pastern.
708:
The order was considerably more diverse in the past, with notable extinct groups including the
7910:
7744:
7451:
7050:
6657:
6647:
6631:
6614:
6599:
6584:
6569:
6528:
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5872:
5447:
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4881:
4820:
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4626:
4533:
4515:
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4267:
4259:
4209:
4010:
3734:
3104:
2956:
2834:
2756:
2556:
1787:
1402:
1291:
1080:
1059:
976:
522:
425:
243:
5737:"Early Eocene Land Mammals from Vastan Lignite Mine, District Surat (Gujarat), western India"
7938:
7915:
7840:
7576:
7532:
7508:
7460:
7322:
7276:
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6812:
6749:
6675:
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6232:
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6105:
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5950:
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5786:
5736:
5650:
5596:
5547:
5477:
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5427:
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5303:
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5229:
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5011:
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4251:
4228:
Hall, Brian K. (2006). Fins into Limbs. University of Chicago Press. ISBN 978-0-226-31337-5.
4111:
4033:
3987:
3972:
3968:
3960:
3541:
3290:
2714:
Brontotherioidea were among the earliest known large mammals, consisting of the families of
2565:
2527:
2435:
2415:
2397:
2312:
2276:
2176:
2107:
1895:
1762:
1114:
1088:
775:
717:
654:
574:
561:
433:
95:
5508:
George Gaylord (1945). "The Principles of Classification and a Classification of Mammals".
3671:. The distant ancestors of tapirs appeared not too long after that in the Ghazij lineup in
833:, which greatly restricts the mobility of the foot. The thigh is relatively short, and the
6964:
6865:
6833:
6468:
5833:"Early Eocene fossils suggest that the mammalian order Perissodactyla originated in India"
5687:
3717:
3654:
3521:
3379:
3190:
3173:
3119:
3054:
3047:
2986:
2917:
2900:
2881:
2852:
2842:
2797:
2740:
2725:
2715:
2706:
2702:
2203:
2194:
2125:
2116:
1737:
1712:
1196:
972:
933:
741:
713:
709:
493:
390:
365:
7082:(Rats, guinea pigs, squirrels, beavers, chinchillas, porcupines, capybaras and relatives)
5735:
Sunil Bajpai; Vivesh Kapur; Debasis P. Das; BN Tiwari; N. Saravanan; Ritu Sharma (2005).
4647:"Tapirs (extant/living species; Tapirus spp.) Fact Sheet: Reproduction & Development"
821:
that clearly distinguishes this group from other mammals is the articulation between the
724:, with the paraceratheres including the largest known land mammals to have ever existed.
6355:
6054:
5946:
5848:
5423:
5140:
5081:
4859:
4796:
4239:
885:
7796:
7783:
7700:
7585:
7378:
7209:
7010:
6956:
6857:
6841:
6523:
6498:
6374:
6339:
6305:
6014:
5989:
5965:
5930:
5654:
5442:
5407:
5352:
5335:
5159:
5124:
5100:
5065:
4844:"Ancient protein resolve the evolutionary history of Darwin's South American ungulates"
4815:
4780:
4528:
4495:
4327:
4255:
4018:
3983:
3828:
3658:
3621:
3435:
3418:
3273:
3237:
3015:
2999:
2949:
2933:
2894:
2838:
2792:
2769:
2764:
2428:
2388:
2323:
2285:
2185:
2144:
1855:
1536:
1326:
1204:
1181:
960:
956:
929:
535:
530:
514:
472:
452:
407:
357:
7713:
5572:
Robert M. Schoch (1989). "A brief historical review of perissodactyl classification".
4924:
3624:
mammals and perissoodactyls in particular is of interest since the latter appeared in
3057:(a clade of stem-perissodactyls; from the Early Palaeocene to the Middle Eocene epoch)
759:, tapirs and horses have dense, short coats. Most species are grey or brown, although
7955:
7603:
7499:
7252:
7001:
6873:
6482:
Donald R. Prothero; Robert M. Schoch (1989). "Classification of the Perissodactyla".
6190:
6110:
6093:
5915:
5813:
5758:
5408:"Anthracobunids from the Middle Eocene of India and Pakistan Are Stem Perissodactyls"
5389:
5066:"Anthracobunids from the Middle Eocene of India and Pakistan Are Stem Perissodactyls"
4983:
4966:
4738:
3991:
3929:
3871:
3768:
3743:
3625:
3613:
3583:
3513:
3394:
2993:
2864:
2788:
2736:
2658:
2406:
1426:
some 56 million years ago, while the rhinos and tapirs split off in the lower-middle
1118:
948:
907:
870:
721:
506:
177:
5798:
5489:
5317:
5241:
4765:
4279:
4123:
3591:
in the Cambay shale of western India indicate an origin in Asia dating to the Lower
873:
has projecting cusps in the eye sockets and a wide contact with the nasal bone. The
556:
7749:
7475:
7415:
7369:
7018:
6948:
6849:
6804:
6499:"Early Tertiary mammals from North Africa reinforce the molecular Afrotheria clade"
6078:
5292:"Reevaluation of some ungulate mammals from the Eocene Pondaung Formation, Myanmar"
4923:
Christelle Tougard; Thomas Delefosse; Catherine Hänni; Claudine Montgelard (2001).
4893:
4029:
3867:
3832:
3722:
3532:
3508:
3313:
3222:
2970:
2860:
2760:
2675:
2652:
2246:
1631:
1613:
1504:
1398:
1383:
1332:
1277:
1212:
1146:
1087:. Their North American counterparts died out around 10,000 years ago, leaving only
1043:
944:
903:
818:
728:
682:
338:
235:
5907:
5276:
4551:
3903:
stood for the entire order, including the hyrax. The assumption that hyraxes were
6613:, Part 3: Mammalia. 2nd edition. Gustav Fischer Verlag, Jena and Stuttgart 1989.
5955:
5481:
5432:
5336:"Comparative osteology of early Tertiary tapiromorphs (Mammalia, Perissodactyla)"
5233:
5149:
5090:
4115:
3835:), including in it not only the rhinos and elephants, but also the hippos, pigs,
1168:
Traditionally, the odd-toed ungulates were classified with other mammals such as
1001:
role in cooling blood in the internal carotid artery before it enters the brain.
7879:
7822:
7680:
7225:
7058:
6980:
6898:
6236:
4721:
3820:
3755:
3704:
3672:
3600:
3545:
leading to the unification in 1984 of the two submissions in the interim order,
3340:
3092:
2963:
2848:
2823:
2780:
2670:
2663:
1588:
1554:
1479:
1240:
1022:
968:
845:
830:
686:
480:
442:
226:
152:
37:
5551:
4177:
1054:
7424:
7282:
7087:
7061:(Pigs, camels, hippos, deer, buffalo, gazelles, giraffes, whales and dolphins)
6940:
6783:
6740:
6724:
5205:
5188:
4411:"Strangles in Horses - Respiratory Diseases of Horses - MSD Veterinary Manual"
3995:
3804:
3663:
3517:
3325:
3258:
3008:
2942:
2856:
2830:
2818:
2720:
2630:
2592:
2490:
2460:
2448:
2259:
2207:
2148:
2078:
2065:
1388:
1336:
1262:
1200:
937:
866:
822:
803:
694:
674:
464:
416:
197:
162:
17:
5757:
Sunil Bajpai; Vivesh Kapur; JGM Thewissen; Debasis P. Das; BN Tiwari (2006).
5189:"Evolutionary Transitions in the Fossil Record of Terrestrial Hoofed Mammals"
4519:
4318:
4263:
3866:, coming close to the concept of the odd-toed ungulate as a systematic unit.
3516:); the Ceratomorpha consist of tapirs and rhinos plus their extinct members (
1203:, so are not closely related to the perissodactyls. These in turn are in the
7770:
7095:
7042:
6919:
6911:
6791:
6757:
6364:
6062:
5016:
4999:
4805:
3887:
3700:
3695:
3661:. The oldest finds from an extant group originate among other sources, from
3577:
3572:
3357:
2750:
1697:
1423:
1410:
1236:
1232:
1188:
1177:
1072:
984:
911:
826:
756:
746:
690:
485:
280:
213:
202:
146:
7671:
6532:
6514:
6383:
6153:
6070:
6023:
6005:
5974:
5876:
5867:
5451:
5168:
5109:
5049:
4951:
4943:
4885:
4824:
4757:
4698:
4537:
4511:
4336:
3649:, an early relative of the horse, is one of the oldest-known perissodactyls
3632:, implying rapid ecological radiation and dispersal after mass extinction.
6143:
6126:
6094:"The systematics and taxonomy of the early Eocene equids (Perissodactyla)"
4706:
4271:
4244:
Anatomia, Histologia, Embryologia: Journal of Veterinary Medicine Series C
3791:), the only tapir then known in Europe. Linnaeus classified this tapir as
920:
917:
7723:
7665:
7203:
7197:
7191:
7034:
6890:
6718:
5255:
Badiola, A.; Pareda-Suberbiola, X.; Cuesta, M. (2003). "A new species of
3738:
3730:
3689:
3645:
2745:
2679:
1220:
1208:
1084:
1010:
878:
858:
834:
795:
787:
751:
646:
300:
217:
192:
187:
172:
167:
157:
6568:. Spektrum Akademischer Verlag, Heidelberg and Berlin 2004, pp 646–655,
5308:
5291:
4867:
3693:, which is considered a member of the horse family, outwardly resembled
3050:(a family of stem-perissodactyls; from the Early to Middle Eocene epoch)
7567:
7523:
7234:
7111:
6988:
6820:
6730:
6667:
5857:
5832:
5608:
5381:
4876:
4294:
3836:
3726:
3668:
3136:
2775:
2058:
1464:
1393:
1323:
1216:
1014:
941:
899:
888:
854:
799:
791:
698:
658:
650:
381:
207:
182:
6340:"Revising the recent evolutionary history of equids using ancient DNA"
4748:
4310:
3667:, an ancestor of the horses from the Willswood lineup in northwestern
3595:
roughly 54.5 million years ago. Their teeth also show similarities to
1133:
7827:
7340:
7331:
7316:
7185:
7103:
7079:
6927:
6774:
6712:
6699:
6182:
4025:
3937:
3921:
3808:
3800:
3592:
3570:
of this group, which was once much more varied and widely dispersed.
2795:
developed long limbs and long necks that were most pronounced in the
2759:
represented another characteristic group, consisting of the families
2729:
2255:
2158:
2135:
2074:
1427:
1092:
1068:
814:
666:
310:
290:
7642:
5600:
3870:(1804–1892) quoted Blainville in his study on fossil mammals of the
2718:(synonym Titanotheriidae), the most well-known representative being
7814:
7775:
6319:
Matthew Colbert (2007). "New Fossil Discoveries and the History of
1187:
Molecular genetic studies, however, have shown the ungulates to be
790:
have stances that require them to stand on the tips of their toes.
7550:
7363:
7349:
7302:
7261:
6972:
6630:, 3rd edition. The Johns Hopkins University Press, Baltimore 2005
5988:
Halliday Thomas John Dixon; Upchurch Paul; Goswami Anjali (1833).
5705:, a perissodactyl-like mammal from the Late Palaeocene of China".
4052:
3915:
3776:
3754:
3639:
2809:), the second largest known land mammal ever to have lived (after
2696:
2669:
2651:
2372:
2217:
2096:
1228:
1173:
1132:
1053:
1018:
862:
844:
774:
764:
760:
702:
678:
670:
662:
555:
4489:
4487:
739:
The largest odd-toed ungulates are rhinoceroses, and the extinct
6661:
6212:(Perissodactyla, Mammalia) in the Eocene Geiseltal layers].
5263:
and first mention in the Late Eocene of the Iberian Peninsula".
3512:
Hippomorpha comprises today's horses and their extinct members (
1207:, a superorder that had its origin in the former supercontinent
1191:, meaning that in some cases the similarities are the result of
849:
Tapirs are the only extant group of perissodactyls with a trunk.
810:
7646:
7141:
6671:
6583:. 6th edition. Johns Hopkins University Press, Baltimore 1999,
6231:. Ecological Studies. Vol. 195. Springer. pp. 21–45.
5701:
Malcolm C. McKenna; Chow Minna; Suyin Ting; Luo Zhexi (1989). "
5123:
Gheerbrant, Emmanuel; Filippo, Andrea; Schmitt, Arnaud (2016).
5064:
Cooper L. N.; Seiffert E. R.; Clementz M.; et al. (2014).
7026:
6547:
3549:. At the same time, he expanded the Ancylopoda to include the
1224:
1095:
were pushed to extinction in 19th century Europe. Hunting and
947:, due to its frequent involvement in some serious infections.
31:
7762:
898:
The number and form of the teeth vary according to diet. The
5929:
Gheerbrant Emmanuel; Filippo Andrea; Schmitt Arnaud (2016).
4779:
Hidenori Nishihara; Masami Hasegawa; Norihiro Okada (2006).
6646:(6th ed.), Baltimore: Johns Hopkins University Press,
4162:. National Geographic Society. 14 July 2015. Archived from
1091:
with a range extending to what is now southern Mexico. The
1013:, hindgut fermenters store digested food that has left the
619:
595:
580:
4579:. Chicago: University of Chicago Press. 2013. p. 273.
3827:. In 1795, Étienne Geoffroy Saint-Hilaire (1772–1844) and
2743:
are known mainly from Europe; their most famous member is
1009:
All perissodactyls are hindgut fermenters. In contrast to
6448:"Tableau Elementaire de l'histoire naturelle des animaux"
5791:
10.1671/0272-4634(2006)26[219:eeycva]2.0.co;2
5587:
William Berryman Scott (1941). "Part V: Perissodactyla".
5523:
Horace Elmer Wood (1934). "Revision of the Hyrachyidae".
4967:"Molecular phylogeny and evolution of the Perissodactyla"
4577:
Zookeeping: An Introduction to the Science and Technology
4384:"Strangles (Distemper) in Horses - MSD Veterinary Manual"
3783:). At that time, this category also included the tapirs (
613:
604:
589:
4910:
The History of Life: A View from the Southern Hemisphere
3725:. For the same reason, however, new animals such as the
1199:. Elephants and hyraxes are now considered to belong to
5994:
Proceedings of the Royal Society B: Biological Sciences
5463:
5461:
4438:
Dyce, Sack and Wensing's Textbook of Veterinary Anatomy
4359:
Dyce, Sack and Wensing's Textbook of Veterinary Anatomy
3787:), more precisely the lowland or South American tapir (
2936:(a basal family of Tapiromorpha; from the Eocene epoch)
5033:"Withers height, body mass and shape of proboscideans"
3024:(sometimes recognized as a subfamily of deperetellids)
5538:
Horace Elmer Wood (1937). "Perissodactyl suborders".
4836:
4834:
3037:(sometimes recognized as a subfamily of lophialetids)
1211:. Molecular genetic findings suggest that the cloven
607:
592:
583:
4679:
4677:
2711:
one of the largest land mammals to have ever existed
616:
610:
601:
586:
577:
527:
519:
511:
503:
477:
469:
449:
439:
430:
422:
412:
404:
387:
362:
354:
7655:
7565:
7549:
7521:
7497:
7473:
7440:
7423:
7250:
7233:
7070:
7009:
7000:
6939:
6910:
6889:
6832:
6803:
6782:
6773:
6748:
6739:
6344:
Proceedings of the National Academy of Sciences USA
5724:. Johns Hopkins University Press. pp. 242–267.
4785:
Proceedings of the National Academy of Sciences USA
4472:. NSW Government - Department of Primary Industries
598:
62:. Unsourced material may be challenged and removed.
7114:(lorises, lemurs, tarsiers, monkeys, apes, humans)
5525:Bulletin of the American Museum of Natural History
5510:Bulletin of the American Museum of Natural History
3924:had become extinct by the end of the 19th century.
2829:Several mammal groups traditionally classified as
798:and other mammals can be seen in the shape of the
6611:Textbook of Paleozoology, Volume III: vertebrates
4440:(5th ed.). Missouri: Elsevier. p. 512.
4361:(5th ed.). Missouri: Elsevier. p. 511.
1247:Internal taxonomy of the Euungulata after Welker
6546:International Union for Conservation of Nature.
5818:Journal of the Palaeontological Society of India
5763:Journal of the Palaeontological Society of India
5744:Journal of the Palaeontological Society of India
5329:
5327:
5182:
5180:
5178:
4722:"Summary of Laurasiatheria (Mammalia) Phylogeny"
3862:animals and referred to all of them together as
2062:: horses and allies, seven species in one genus
1434:Internal relationships of extant Perissodactyla
701:chambers as artiodactyls, with the exception of
6566:Systematic Zoology. Part 2: Vortex or craniotes
5503:
5501:
5499:
4625:(2nd ed.). Wiley-Blackwell. p. 1015.
5567:
5565:
5563:
5561:
5290:Tsubamoto, T.; Tun, S.T.; et al. (2003).
4138:"Glossary. American Museum of Natural History"
3949:International Union for Conservation of Nature
2920:(might be a basal perissodactyl grade instead)
7153:
6683:
6125:Pieter Missiaen; Philip D. Gingerich (2012).
5401:
5399:
3831:(1769–1832) introduced the term "pachyderm" (
1021:, where the food is digested by microbes. No
8:
6486:. Oxford University Press. pp. 530–537.
4965:Cynthia C. Steiner; Oliver A. Ryder (2011).
4651:International Environment Library Consortium
3799:("beasts"). He combined the rhinos with the
2432:: rhinoceroses, five species in four genera
6264:. London: Chapman & Hall. p. 343.
1042:Unlike artiodactyls, perissodactyls lack a
7643:
7562:
7437:
7409:
7247:
7219:
7160:
7146:
7138:
7006:
6907:
6800:
6779:
6745:
6690:
6676:
6668:
6285:Mikael Fortelius; John Kappelmann (1993).
5709:. Oxford University Press. pp. 24–36.
5576:. Oxford University Press. pp. 13–23.
4592:"Understand Normal Mare and Foal Behavior"
4496:"The carotid rete and artiodactyl success"
3069:Internal classification of Perissodactyla
1329: (South American ungulates,
1280: (Even-toed ungulates and cetaceans)
955:mould) of the guttural pouch (also called
685:, or positioned posteriorly. By contrast,
225:
135:
6522:
6373:
6363:
6294:Zoological Journal of the Linnean Society
6214:Hallesches Jahrbuch für Geowissenschaften
6152:
6142:
6109:
6098:Zoological Journal of the Linnean Society
6013:
5964:
5954:
5866:
5856:
5643:Zoological Journal of the Linnean Society
5441:
5431:
5351:
5340:Zoological Journal of the Linnean Society
5307:
5204:
5158:
5148:
5099:
5089:
5048:
5015:
4982:
4971:Zoological Journal of the Linnean Society
4875:
4814:
4804:
4747:
4737:
4720:Jingyang Hu; Zhang Yaping; Li Yu (2012).
4653:. San Diego Zoo Wildlife Alliance Library
4527:
4326:
2855:have traditionally been placed among the
122:Learn how and when to remove this message
7021:(Hedgehogs, shrews, moles and relatives)
6624:Don E. Wilson, DeeAnn M. Reeder (eds.):
5000:"A new species of tapir from the Amazon"
4621:Angus, McKinnon; Noah, Heninger (2011).
3947:Present threat levels, according to the
3795:and put both genera in the group of the
3067:
1432:
1245:
6598:Oxford University Press, Oxford, 2005.
6287:"The Largest land mammal ever imagined"
4590:Williams, Jennifer (15 December 2008).
4494:Mitchell, G; Lust, A (23 August 2008).
4069:
3064:Higher classification of perissodactyls
6464:
6453:
5683:
5672:
4156:"The Largest Beasts to Walk the Earth"
2666:genus first formally described in 1804
6596:The Origin & Evolution of Mammals
4932:Molecular Phylogenetics and Evolution
4687:Molecular Phylogenetics and Evolution
4352:
4350:
4348:
4346:
3885:(1831–1899) came up with the concept
1407:Cretaceous–Paleogene extinction event
884:Rhinos have one or two horns made of
649:. The order includes about 17 living
7:
7903:682f936e-e4c4-4c38-b32e-d24c54f384c0
7802:2d58add6-3cfc-406c-b2ed-76db036bdfbd
6431:Johann Friedrich Blumenbach (1779).
4040:; however, the northern subspecies,
2779:. Chalicotherioidea died out in the
2613:Dicerorhinus sumatrensis sumatrensis
2376:: tapirs, five species in one genus
731:, who also coined the order's name.
645: 'finger, toe') is an order of
60:adding citations to reliable sources
6229:The Ecology of Browsing and Grazing
5722:The Beginning of the Age of Mammals
3683:from the Chalicotheriidae line, or
3410:
3386:
3332:
3320:
3265:
3253:
3229:
3165:
3111:
3099:
3087:
3075:
2622:Dicerorhinus sumatrensis harrissoni
1945:
1920:
1868:
1861:
1778:
1753:
1728:
1703:
1692:
1604:
1579:
1527:
1520:
1495:
1470:
1459:
1452:
1440:
1331: especially
1298:
1269:
1257:
564:is the largest living perissodactyl
6503:Proceedings of the Royal Society B
6414:Introduction à l'Anatomie Comparée
6306:10.1111/j.1096-3642.1993.tb02560.x
5896:Journal of Vertebrate Paleontology
5779:Journal of Vertebrate Paleontology
5655:10.1111/j.1096-3642.1984.tb00545.x
5470:Journal of Vertebrate Paleontology
5353:10.1111/j.1096-3642.2001.tb02270.x
4256:10.1111/j.1439-0264.1975.tb00637.x
3852:Henri Marie Ducrotay de Blainville
3707:, in the Middle and Upper Eocene.
2791:were hippo-like, aquatic animals.
1422:molecular genetic analysis in the
1083:around 3 million years ago in the
25:
7962:Extant Ypresian first appearances
7053:(Horses, rhinoceroses and tapirs)
5193:Evolution: Education and Outreach
4293:Lorente, Malena (2 August 2019).
3944:was once eradicated in the wild.
3858:over to the tapirs and rhinos as
3616:or near its collision with Asia.
2634:Dicerorhinus sumatrensis lasiotis
7627:
7626:
6111:10.1046/j.1096-3642.2002.00005.x
4984:10.1111/j.1096-3642.2011.00752.x
4739:10.3724/sp.j.1141.2012.e05-06e65
3803:, a group now consisting of the
2966:(basal grade of chalicotheroids)
2739:) also developed in the Eocene.
2583:Rhinoceros sondaicus annamiticus
1231:(a common taxon of carnivorans,
573:
267:
36:
6484:The Evolution of Perissodactyls
5707:The Evolution of Perissodactyls
5574:The Evolution of Perissodactyls
3895:is now considered a synonym of
3771:(1707–1778) classified horses (
1430:, about 47 million years ago.
853:Odd-toed ungulates have a long
47:needs additional citations for
4238:Hussain, S. (September 1975).
4202:Allen, Jeremiah (March 2010).
2749:, which became extinct in the
2576:Rhinoceros sondaicus sondaicus
2517:South-western black rhinoceros
2508:South-central black rhinoceros
2457:North-eastern black rhinoceros
932:, a small outpocketing of the
1:
6644:Walker's Mammals of the World
6581:Walker's Mammals of the World
6131:Acta Palaeontologica Polonica
5908:10.1080/02724634.2020.1761370
5622:Leonard B. Radinsky (1964). "
5277:10.1016/j.geobios.2003.07.005
5037:Acta Palaeontologica Polonica
4208:. Other Places Travel Guide.
4005:Three species are considered
3864:onguligrades à doigts impairs
3847:, meaning "one-hoof animal".
3763:In 1758, in his seminal work
2611:Western Sumatran rhinoceros,
2580:Vietnamese Javan rhinoceros,
2574:Indonesian Javan rhinoceros,
2521:Diceros bicornis occidentalis
697:, rather than in one or more
5956:10.1371/journal.pone.0157556
5482:10.1080/02724634.2011.579669
5433:10.1371/journal.pone.0109232
5234:10.1080/08912963.2014.915823
5150:10.1371/journal.pone.0157556
5091:10.1371/journal.pone.0109232
4116:10.1080/08912963.2014.915823
3955:Four species are considered
3612:suggests an origin in India
2876:Superfamily Brontotherioidea
2627:Northern Sumatran rhinoceros
2596:Rhinoceros sondaicus inermis
1129:Reproduction and development
1032:surface-area-to-volume ratio
861:between the front and cheek
745:, a hornless rhino from the
6627:Mammal Species of the World
6433:Handbook of Natural History
6260:Benton, Michael J. (1997).
6237:10.1007/978-3-540-72422-3_2
6092:David J. Froehlich (2002).
4055:is not considered at risk (
4042:Ceratotherium simum cottoni
2618:Eastern Sumatran rhinoceros
2550:Ceratotherium simum cottoni
2473:Diceros bicornis chobiensis
2130:Equus africanus somaliensis
1309: (Odd-toed ungulates)
1079:after the formation of the
914:of odd-toed ungulates is:
384:(horses, asses, and zebras)
7998:
7977:Taxa named by Richard Owen
7090:(Rabbits, hares and pikas)
7045:(Dogs, cats and relatives)
6951:(Tenrecs and golden moles)
2833:, long-understood to be a
2724:and the more basal family
2482:Diceros bicornis ladoensis
2152:Equus africanus atlanticus
1156:
895:, which have a bony core.
7624:
7412:
7408:
7222:
7218:
7180:
6876:(Kangaroos and relatives)
6707:
5628:American Museum Novitates
5334:Luke T. Holbrook (2001).
5206:10.1007/s12052-009-0136-1
4048:) is close to extinction.
4046:northern white rhinoceros
3432:
3415:
3408:
3391:
3384:
3354:
3337:
3330:
3318:
3287:
3270:
3263:
3251:
3234:
3227:
3187:
3170:
3163:
3133:
3116:
3109:
3097:
3085:
2546:Northern white rhinoceros
2541:Ceratotherium simum simum
2537:Southern white rhinoceros
2503:Diceros bicornis michaeli
2494:Diceros bicornis longipes
2452:Diceros bicornis bicornis
2445:Southern black rhinoceros
2342:Hartmann's mountain zebra
2121:Equus africanus africanus
1968:
1950:
1943:
1925:
1918:
1891:
1873:
1866:
1859:
1801:
1783:
1776:
1758:
1751:
1733:
1726:
1708:
1701:
1690:
1627:
1609:
1602:
1584:
1577:
1550:
1532:
1525:
1518:
1500:
1493:
1475:
1468:
1457:
1450:
1320:
1303:
1296:
1274:
1267:
817:are reduced in horses. A
351:
346:
264:Scientific classification
262:
233:
224:
138:
6868:(Bilbies and bandicoots)
6642:Ronald M. Nowak (1999),
6262:Vertebrate Palaeontology
5720:Kenneth D. Rose (2006).
5552:10.1093/jmammal/18.1.106
5296:Paleontological Research
3912:Interactions with humans
3874:and introduced the name
3813:Mathurin Jacques Brisson
2607:Dicerorhinus sumatrensis
2499:Eastern black rhinoceros
2487:Western black rhinoceros
1930:Dicerorhinus sumatrensis
1125:(tapirs, other rhinos).
1077:arrived in South America
638: 'odd' and
6760:(Platypus and echidnas)
6446:Georges Cuvier (1798).
6365:10.1073/pnas.0903672106
6171:Journal of Paleontology
6063:10.1126/science.1213859
5017:10.1644/12-MAMM-A-169.1
4806:10.1073/pnas.0603797103
4182:Encyclopædia Britannica
3793:Hippopotamus terrestris
3542:Leonard Burton Radinsky
2812:Palaeoloxodon namadicus
2674:Live reconstruction of
2589:Indian Javan rhinoceros
2478:Uganda black rhinoceros
2464:Diceros bicornis brucii
2272:Equus quagga burchellii
2211:Equus hemionus hemippus
2172:Equus hemionus hemionus
2091:Equus ferus przewalskii
1223:. More distant are the
989:Streptococcus equi equi
981:internal carotid artery
875:temporomandibular joint
27:Order of hoofed mammals
7595:South American tapir (
6991:(Dugongs and manatees)
6930:(Anteaters and sloths)
6515:10.1098/rspb.2006.0229
6463:Cite journal requires
6399:Magasin Encyclopédique
6006:10.1098/rspb.2015.3026
5682:Cite journal requires
5050:10.4202/app.00136.2014
4944:10.1006/mpev.2000.0903
4699:10.1006/mpev.1996.0391
4512:10.1098/rsbl.2008.0138
4436:Singh, Baljit (2018).
4357:Singh, Baljit (2018).
3988:Central American tapir
3925:
3760:
3650:
2712:
2689:
2667:
2512:Diceros bicornis minor
2469:Chobe black rhinoceros
2346:Equus zebra hartmannae
2308:Equus quagga crawshayi
2140:Equus africanus asinus
1159:List of perissodactyls
1141:
1063:
957:Guttural Pouch Mycosis
891:, unlike the horns of
850:
780:
565:
7924:Paleobiology Database
7485:Sumatran rhinoceros (
6860:(Quolls and dunnarts)
6216:. Suppl. 23: 161–173.
6144:10.4202/app.2010.0093
5837:Nature Communications
5812:Kishor Kumar (2005).
5031:Larramendi A (2016).
4415:MSD Veterinary Manual
4388:MSD Veterinary Manual
3957:critically endangered
3919:
3883:Othniel Charles Marsh
3758:
3643:
2980:Suborder Ceratomorpha
2701:Scale diagram of the
2700:
2686:Chalicotherium grande
2673:
2655:
2366:Suborder Ceratomorpha
2299:Equus quagga chapmani
2290:Equus quagga borensis
2234:Equus kiang holdereri
2220:or Tibetan wild ass,
2190:Equus hemionus onager
2161:or Asiatic wild ass,
1136:
1057:
848:
778:
559:
234:Clockwise from left:
7797:Fauna Europaea (new)
7292:Przewalski's horse (
7172:(Odd-toed ungulates)
5540:Journal of Mammalogy
5004:Journal of Mammalogy
4144:on 20 November 2021.
4015:South American tapir
3557:Evolutionary history
2906:Superfamily Equoidea
2889:Suborder Hippomorpha
2871:Order Perissodactyla
2570:Rhinoceros sondaicus
2561:Rhinoceros unicornis
2317:Equus quagga selousi
2240:Equus kiang polyodon
2181:Equus hemionus kulan
2101:Equus ferus caballus
2052:Suborder Hippomorpha
2047:Order Perissodactyla
1973:Rhinoceros unicornis
1955:Rhinoceros sondaicus
1193:convergent evolution
1038:Lack of carotid rete
324:Perissodactylamorpha
248:Rhinoceros unicornis
56:improve this article
7452:Indian rhinoceros (
7175:species by suborder
6509:(1614): 1159–1166.
6435:. pp. 168–448.
6356:2009PNAS..10621754O
6350:(51): 21754–21759.
6055:2012Sci...335..959S
5947:2016PLoSO..1157556G
5849:2014NatCo...5.5570R
5424:2014PLoSO...9j9232C
5309:10.2517/prpsj.7.219
5187:Donald, R. (2009).
5141:2016PLoSO..1157556G
5082:2014PLoSO...9j9232C
5010:(6): 1331 to 1345.
4868:10.1038/nature14249
4860:2015Natur.522...81W
4797:2006PNAS..103.9929N
4726:Zoological Research
4623:Equine Reproduction
3965:Sumatran rhinoceros
3825:even-toed ungulates
3703:to emerge were the
3538:Edward Drinker Cope
3070:
2801:(formerly known as
2648:Prehistoric members
2603:Sumatran rhinoceros
2532:Ceratotherium simum
2333:Cape mountain zebra
2281:Equus quagga boehmi
2263:Equus quagga quagga
2199:Equus hemionus khur
1878:Ceratotherium simum
1435:
1252:
1097:habitat destruction
945:Veterinary practice
918:0-3 . 0-1 . 2-4 . 3
893:even-toed ungulates
673:), Rhinocerotidae (
653:divided into three
7533:Black rhinoceros (
7509:White rhinoceros (
7461:Javan rhinoceros (
7323:African wild ass (
6844:(Monito del monte)
6325:Tapir Conservation
6000:(1833): 20153026.
5858:10.1038/ncomms6570
5703:Radinskya yupingae
5382:10.1111/pala.12471
5222:Historical Biology
4556:www.britannica.com
4299:Journal of Anatomy
4104:Historical Biology
3942:Przewalski's horse
3926:
3761:
3759:Richard Owen, 1856
3651:
3599:as well as to the
3568:adaptive radiation
3526:Titanotheriomorpha
3068:
2928:Clade Tapiromorpha
2713:
2690:
2668:
2384:Tapirus terrestris
2168:Mongolian wild ass
2134:Domesticated ass (
2087:Przewalski's horse
1806:Tapirus terrestris
1433:
1379:protein sequencing
1246:
1142:
1104:Lifestyle and diet
1064:
977:Hypoglossal Nerves
851:
781:
677:), and Tapiridae (
566:
256:Tapirus terrestris
7982:Panperissodactyla
7949:
7948:
7911:Open Tree of Life
7649:Taxon identifiers
7640:
7639:
7620:
7619:
7616:
7615:
7559:
7545:
7544:
7434:
7404:
7403:
7400:
7399:
7294:E. f. przewalskii
7267:
7244:
7173:
7135:
7134:
7131:
7130:
7127:
7126:
7123:
7122:
7115:
7107:
7099:
7091:
7083:
7062:
7054:
7046:
7038:
7030:
7022:
6992:
6984:
6976:
6968:
6960:
6959:(Elephant shrews)
6952:
6931:
6923:
6904:
6885:
6884:
6877:
6869:
6861:
6853:
6852:(Marsupial moles)
6850:Notoryctemorphia
6845:
6824:
6816:
6813:Paucituberculata
6797:
6769:
6768:
6761:
6653:978-0-8018-5789-8
6579:Ronald M. Nowak:
6246:978-3-540-72421-6
6210:Hyrachyus minimus
6206:Hyrachyus minimus
6049:(6071): 959–962.
4791:(26): 9929–9934.
4632:978-0-8138-1971-6
4596:equusmagazine.com
4311:10.1111/joa.13065
4011:Indian rhinoceros
3789:Tapirus terrestus
3735:Isthmus of Panama
3581:rhinos. Finds of
3505:
3504:
3500:
3499:
3491:
3490:
3482:
3481:
3473:
3472:
3464:
3463:
3455:
3454:
3446:
3445:
3368:
3367:
3302:
3301:
3211:
3210:
3202:
3201:
3147:
3146:
2957:Chalicotherioidea
2863:and the northern
2835:wastebasket taxon
2757:Chalicotherioidea
2557:Indian rhinoceros
2393:Tapirus pinchaque
2337:Equus zebra zebra
2228:Equus kiang kiang
2082:Equus ferus ferus
2043:
2042:
2038:
2037:
2029:
2028:
2020:
2019:
2011:
2010:
2002:
2001:
1993:
1992:
1984:
1983:
1907:
1906:
1844:
1843:
1835:
1834:
1826:
1825:
1817:
1816:
1788:Tapirus pinchaque
1679:
1678:
1670:
1669:
1661:
1660:
1652:
1651:
1643:
1642:
1566:
1565:
1403:Panperissodactyla
1371:
1370:
1366:
1365:
1357:
1356:
1348:
1347:
1292:Panperissodactyla
1235:, and pangolins,
1227:(Chiroptera) and
1081:Isthmus of Panama
1060:Indian rhinoceros
921:1-3 . 1 . 2-4 . 3
857:with an extended
554:
553:
523:Paraceratheriidae
426:Chalicotherioidea
342:
244:Indian rhinoceros
132:
131:
124:
106:
16:(Redirected from
7989:
7942:
7941:
7932:
7931:
7919:
7918:
7906:
7905:
7896:
7895:
7883:
7882:
7880:NHMSYS0000377074
7870:
7869:
7857:
7856:
7844:
7843:
7831:
7830:
7818:
7817:
7805:
7804:
7792:
7791:
7779:
7778:
7766:
7765:
7753:
7752:
7740:
7739:
7727:
7726:
7717:
7716:
7704:
7703:
7691:
7690:
7689:
7676:
7675:
7674:
7644:
7630:
7629:
7586:Mountain tapir (
7563:
7558:
7557:
7553:
7438:
7433:
7432:
7428:
7410:
7379:Mountain zebra (
7303:Domestic horse (
7266:
7265:
7257:
7248:
7243:
7242:
7238:
7220:
7171:
7162:
7155:
7148:
7139:
7113:
7105:
7097:
7089:
7081:
7072:Euarchontoglires
7060:
7052:
7044:
7036:
7028:
7020:
7007:
6990:
6982:
6974:
6966:
6958:
6950:
6929:
6921:
6908:
6903:
6902:
6894:
6875:
6867:
6866:Peramelemorphia
6859:
6851:
6843:
6822:
6821:Didelphimorphia
6815:(Shrew opossums)
6814:
6801:
6796:
6795:
6787:
6780:
6759:
6750:Australosphenida
6746:
6692:
6685:
6678:
6669:
6664:
6594:Thomas S. Kemp:
6552:
6551:
6543:
6537:
6536:
6526:
6494:
6488:
6487:
6479:
6473:
6472:
6466:
6461:
6459:
6451:
6443:
6437:
6436:
6428:
6422:
6421:
6409:
6403:
6402:
6394:
6388:
6387:
6377:
6367:
6335:
6329:
6328:
6316:
6310:
6309:
6291:
6282:
6276:
6275:
6257:
6251:
6250:
6224:
6218:
6217:
6201:
6195:
6194:
6183:10.1666/10-087.1
6165:
6159:
6158:
6156:
6146:
6122:
6116:
6115:
6113:
6089:
6083:
6082:
6034:
6028:
6027:
6017:
5985:
5979:
5978:
5968:
5958:
5926:
5920:
5919:
5887:
5881:
5880:
5870:
5860:
5828:
5822:
5821:
5820:: 101–113, 2005.
5809:
5803:
5802:
5773:
5767:
5766:
5754:
5748:
5747:
5741:
5732:
5726:
5725:
5717:
5711:
5710:
5698:
5692:
5691:
5685:
5680:
5678:
5670:
5665:
5659:
5658:
5649:(1–2): 229–244.
5638:
5632:
5631:
5619:
5613:
5612:
5584:
5578:
5577:
5569:
5556:
5555:
5535:
5529:
5528:
5520:
5514:
5513:
5505:
5494:
5493:
5465:
5456:
5455:
5445:
5435:
5403:
5394:
5393:
5364:
5358:
5357:
5355:
5331:
5322:
5321:
5311:
5287:
5281:
5280:
5252:
5246:
5245:
5217:
5211:
5210:
5208:
5184:
5173:
5172:
5162:
5152:
5120:
5114:
5113:
5103:
5093:
5061:
5055:
5054:
5052:
5028:
5022:
5021:
5019:
4995:
4989:
4988:
4986:
4977:(4): 1289–1303.
4962:
4956:
4955:
4929:
4920:
4914:
4913:
4904:
4898:
4897:
4879:
4838:
4829:
4828:
4818:
4808:
4776:
4770:
4769:
4751:
4741:
4717:
4711:
4710:
4681:
4672:
4669:
4663:
4662:
4660:
4658:
4643:
4637:
4636:
4618:
4612:
4611:
4609:
4607:
4598:. Archived from
4587:
4581:
4580:
4573:
4567:
4566:
4564:
4562:
4548:
4542:
4541:
4531:
4491:
4482:
4481:
4479:
4477:
4467:
4458:
4452:
4451:
4433:
4427:
4426:
4424:
4422:
4406:
4400:
4399:
4397:
4395:
4379:
4373:
4372:
4354:
4341:
4340:
4330:
4305:(6): 1024–1035.
4290:
4284:
4283:
4235:
4229:
4226:
4220:
4219:
4199:
4193:
4192:
4190:
4188:
4174:
4168:
4167:
4166:on 15 July 2015.
4152:
4146:
4145:
4140:. Archived from
4134:
4128:
4127:
4099:
4093:
4092:
4090:
4088:
4074:
4034:white rhinoceros
3978:Six species are
3973:African wild ass
3969:black rhinoceros
3961:Javan rhinoceros
3775:) together with
3751:Research history
3533:Chalicotheriidae
3531:Originally, the
3411:
3387:
3333:
3321:
3291:Chalicotheriidae
3266:
3254:
3230:
3166:
3159:Brontotherioidea
3112:
3100:
3088:
3076:
3071:
2971:Chalicotheriidae
2761:Chalicotheriidae
2566:Javan rhinoceros
2528:White rhinoceros
2440:Diceros bicornis
2436:Black rhinoceros
2420:Tapirus kabomani
2304:Crawshay's zebra
2268:Burchell's zebra
2238:Southern kiang,
2177:Turkmenian kulan
2108:African wild ass
1946:
1921:
1896:Diceros bicornis
1869:
1862:
1779:
1763:Tapirus kabomani
1754:
1729:
1704:
1693:
1605:
1580:
1528:
1521:
1496:
1471:
1460:
1453:
1441:
1436:
1299:
1270:
1258:
1253:
1215:(containing the
965:Glossopharyngeal
963:(Nerves IX-XII:
951:(infection with
936:that drains the
925:
923:
922:
919:
877:is high and the
626:
625:
622:
621:
618:
615:
612:
609:
606:
603:
600:
597:
594:
591:
588:
585:
582:
579:
562:white rhinoceros
529:
521:
513:
505:
479:
471:
451:
441:
434:Chalicotheriidae
432:
424:
414:
406:
389:
364:
356:
337:
272:
271:
229:
214:Latest Paleocene
212:
149:
142:Temporal range:
136:
127:
120:
116:
113:
107:
105:
71:"Perissodactyla"
64:
40:
32:
21:
7997:
7996:
7992:
7991:
7990:
7988:
7987:
7986:
7952:
7951:
7950:
7945:
7937:
7935:
7927:
7922:
7914:
7909:
7901:
7899:
7891:
7886:
7878:
7873:
7865:
7860:
7852:
7847:
7839:
7834:
7826:
7821:
7813:
7808:
7800:
7795:
7787:
7782:
7774:
7769:
7761:
7756:
7748:
7743:
7735:
7730:
7722:
7720:
7712:
7707:
7699:
7694:
7685:
7684:
7679:
7670:
7669:
7664:
7651:
7641:
7636:
7612:
7604:Malayan tapir (
7577:Baird's tapir (
7555:
7554:
7541:
7517:
7493:
7469:
7430:
7429:
7427:
7419:
7396:
7388:Grévy's zebra (
7259:
7258:
7256:
7240:
7239:
7237:
7229:
7214:
7176:
7170:Perissodactyla
7166:
7136:
7119:
7066:
7051:Perissodactyla
6996:
6935:
6896:
6895:
6893:
6881:
6858:Dasyuromorphia
6842:Microbiotheria
6834:Australidelphia
6828:
6789:
6788:
6786:
6765:
6735:
6703:
6696:
6654:
6641:
6561:
6559:Further reading
6556:
6555:
6545:
6544:
6540:
6496:
6495:
6491:
6481:
6480:
6476:
6462:
6452:
6445:
6444:
6440:
6430:
6429:
6425:
6411:
6410:
6406:
6396:
6395:
6391:
6337:
6336:
6332:
6318:
6317:
6313:
6289:
6284:
6283:
6279:
6272:
6259:
6258:
6254:
6247:
6226:
6225:
6221:
6203:
6202:
6198:
6167:
6166:
6162:
6154:1854/LU-3178691
6124:
6123:
6119:
6091:
6090:
6086:
6036:
6035:
6031:
5987:
5986:
5982:
5941:(7): e0157556.
5928:
5927:
5923:
5889:
5888:
5884:
5868:1854/LU-5972511
5830:
5829:
5825:
5811:
5810:
5806:
5775:
5774:
5770:
5756:
5755:
5751:
5739:
5734:
5733:
5729:
5719:
5718:
5714:
5700:
5699:
5695:
5681:
5671:
5667:
5666:
5662:
5640:
5639:
5635:
5621:
5620:
5616:
5601:10.2307/1005518
5586:
5585:
5581:
5571:
5570:
5559:
5537:
5536:
5532:
5522:
5521:
5517:
5507:
5506:
5497:
5467:
5466:
5459:
5418:(10): e109232.
5405:
5404:
5397:
5366:
5365:
5361:
5333:
5332:
5325:
5289:
5288:
5284:
5254:
5253:
5249:
5219:
5218:
5214:
5186:
5185:
5176:
5135:(7): e0157556.
5122:
5121:
5117:
5076:(10): e109232.
5063:
5062:
5058:
5030:
5029:
5025:
4997:
4996:
4992:
4964:
4963:
4959:
4927:
4922:
4921:
4917:
4906:
4905:
4901:
4854:(7554): 81–84.
4840:
4839:
4832:
4778:
4777:
4773:
4719:
4718:
4714:
4683:
4682:
4675:
4670:
4666:
4656:
4654:
4645:
4644:
4640:
4633:
4620:
4619:
4615:
4605:
4603:
4589:
4588:
4584:
4575:
4574:
4570:
4560:
4558:
4550:
4549:
4545:
4500:Biology Letters
4493:
4492:
4485:
4475:
4473:
4465:
4461:Freeman, Paul.
4460:
4459:
4455:
4448:
4447:978-0-323442640
4435:
4434:
4430:
4420:
4418:
4409:Lascola, Kara.
4408:
4407:
4403:
4393:
4391:
4381:
4380:
4376:
4369:
4368:978-0-323442640
4356:
4355:
4344:
4292:
4291:
4287:
4237:
4236:
4232:
4227:
4223:
4216:
4201:
4200:
4196:
4186:
4184:
4176:
4175:
4171:
4154:
4153:
4149:
4136:
4135:
4131:
4101:
4100:
4096:
4086:
4084:
4076:
4075:
4071:
4066:
4038:near-threatened
3940:is extinct and
3914:
3765:Systema Naturae
3753:
3718:Paraceratherium
3638:
3628:soon after the
3564:
3559:
3522:Rhinocerotoidea
3501:
3492:
3483:
3474:
3465:
3456:
3447:
3380:Rhinocerotoidea
3369:
3303:
3212:
3203:
3191:Brontotheriidae
3174:Lambdotheriidae
3148:
3120:Palaeotheriidae
3066:
3055:Phenacodontidae
3048:Anthracobunidae
2987:Rhinocerotoidea
2918:Palaeotheriidae
2901:Pachynolophidae
2882:Brontotheriidae
2853:Anthracobunidae
2798:Paraceratherium
2741:Palaeotheriidae
2726:Lambdotheriidae
2716:Brontotheriidae
2707:Paraceratherium
2703:paraceratheriid
2656:Restoration of
2650:
2411:Tapirus indicus
2402:Tapirus bairdii
2380:Brazilian tapir
2295:Chapman's zebra
2232:Eastern kiang,
2226:Western kiang,
2204:Syrian wild ass
2195:Indian wild ass
2126:Somali wild ass
2117:Nubian wild ass
2112:Equus africanus
2039:
2030:
2021:
2012:
2003:
1994:
1985:
1908:
1845:
1836:
1827:
1818:
1738:Tapirus bairdii
1713:Tapirus indicus
1680:
1671:
1662:
1653:
1644:
1567:
1419:
1367:
1358:
1349:
1330:
1197:common ancestry
1166:
1161:
1155:
1139:Brazilian tapir
1131:
1106:
1052:
1040:
1027:small intestine
1017:in an enlarged
1007:
916:
915:
843:
841:Skull and teeth
773:
742:Paraceratherium
737:
576:
572:
494:Rhinocerotoidea
436:(chalicotheres)
391:Palaeotheriidae
366:Brontotheriidae
336:
266:
252:Brazilian tapir
220:
211:
210:
205:
200:
195:
190:
185:
180:
175:
170:
165:
160:
155:
144:
143:
140:
128:
117:
111:
108:
65:
63:
53:
41:
28:
23:
22:
15:
12:
11:
5:
7995:
7993:
7985:
7984:
7979:
7974:
7969:
7967:Perissodactyla
7964:
7954:
7953:
7947:
7946:
7944:
7943:
7933:
7920:
7907:
7897:
7884:
7871:
7858:
7845:
7832:
7819:
7806:
7793:
7784:Fauna Europaea
7780:
7767:
7754:
7741:
7728:
7718:
7714:Perissodactyla
7705:
7701:Perissodactyla
7692:
7687:Perissodactyla
7677:
7661:
7659:
7657:Perissodactyla
7653:
7652:
7647:
7638:
7637:
7625:
7622:
7621:
7618:
7617:
7614:
7613:
7611:
7610:
7601:
7592:
7583:
7573:
7571:
7560:
7547:
7546:
7543:
7542:
7540:
7539:
7529:
7527:
7519:
7518:
7516:
7515:
7505:
7503:
7495:
7494:
7492:
7491:
7487:D. sumatrensis
7481:
7479:
7471:
7470:
7468:
7467:
7458:
7448:
7446:
7435:
7431:(Rhinoceroses)
7425:Rhinocerotidae
7421:
7420:
7413:
7406:
7405:
7402:
7401:
7398:
7397:
7395:
7394:
7385:
7376:
7370:Plains zebra (
7357:
7356:
7347:
7338:
7329:
7310:
7309:
7300:
7299:
7298:
7270:
7268:
7245:
7241:(Horse family)
7231:
7230:
7223:
7216:
7215:
7213:
7212:
7210:Laurasiatheria
7206:
7200:
7194:
7188:
7181:
7178:
7177:
7167:
7165:
7164:
7157:
7150:
7142:
7133:
7132:
7129:
7128:
7125:
7124:
7121:
7120:
7118:
7117:
7109:
7101:
7093:
7085:
7076:
7074:
7068:
7067:
7065:
7064:
7056:
7048:
7040:
7032:
7024:
7015:
7013:
7011:Laurasiatheria
7004:
6998:
6997:
6995:
6994:
6986:
6978:
6970:
6965:Tubulidentata
6962:
6957:Macroscelidea
6954:
6945:
6943:
6937:
6936:
6934:
6933:
6925:
6916:
6914:
6905:
6887:
6886:
6883:
6882:
6880:
6879:
6874:Diprotodontia
6871:
6863:
6855:
6847:
6838:
6836:
6830:
6829:
6827:
6826:
6818:
6809:
6807:
6798:
6777:
6771:
6770:
6767:
6766:
6764:
6763:
6754:
6752:
6743:
6737:
6736:
6734:
6733:
6727:
6721:
6715:
6708:
6705:
6704:
6697:
6695:
6694:
6687:
6680:
6672:
6666:
6665:
6652:
6639:
6622:
6607:
6592:
6577:
6560:
6557:
6554:
6553:
6548:"IUCN redlist"
6538:
6489:
6474:
6465:|journal=
6438:
6423:
6404:
6389:
6330:
6311:
6277:
6270:
6252:
6245:
6219:
6196:
6177:(4): 665–677.
6160:
6117:
6104:(2): 141–256.
6084:
6029:
5980:
5921:
5882:
5823:
5804:
5768:
5749:
5727:
5712:
5693:
5684:|journal=
5660:
5633:
5614:
5595:(5): 747–964.
5591:. New Series.
5579:
5557:
5530:
5515:
5495:
5476:(4): 895–901.
5457:
5395:
5376:(4): 547–564.
5359:
5323:
5282:
5247:
5212:
5199:(2): 289–302.
5174:
5115:
5056:
5023:
4990:
4957:
4915:
4899:
4830:
4771:
4712:
4693:(2): 195–200.
4673:
4664:
4638:
4631:
4613:
4602:on 13 May 2017
4582:
4568:
4543:
4506:(4): 415–418.
4483:
4453:
4446:
4428:
4401:
4382:Rush, Bonnie.
4374:
4367:
4342:
4285:
4250:(3): 193–222.
4230:
4221:
4214:
4194:
4169:
4147:
4129:
4094:
4082:paleobiodb.org
4068:
4067:
4065:
4062:
4061:
4060:
4049:
4022:
4019:mountain zebra
4003:
3984:mountain tapir
3976:
3913:
3910:
3905:Perissodactyla
3901:Perissodactyla
3897:Perissodactyla
3876:Perissodactyla
3829:Georges Cuvier
3752:
3749:
3655:phenacodontids
3637:
3634:
3622:meridiungulate
3576:from the late
3563:
3560:
3558:
3555:
3551:Lophiodontidae
3503:
3502:
3498:
3497:
3494:
3493:
3489:
3488:
3485:
3484:
3480:
3479:
3476:
3475:
3471:
3470:
3467:
3466:
3462:
3461:
3458:
3457:
3453:
3452:
3449:
3448:
3444:
3443:
3440:
3439:
3436:Rhinocerotidae
3431:
3428:
3427:
3424:
3423:
3419:Hyracodontidae
3414:
3409:
3407:
3404:
3403:
3400:
3399:
3390:
3385:
3383:
3375:
3374:
3371:
3370:
3366:
3365:
3362:
3361:
3353:
3350:
3349:
3346:
3345:
3336:
3331:
3329:
3319:
3317:
3309:
3308:
3305:
3304:
3300:
3299:
3296:
3295:
3286:
3283:
3282:
3279:
3278:
3274:Lophiodontidae
3269:
3264:
3262:
3252:
3250:
3247:
3246:
3243:
3242:
3238:Isectolophidae
3233:
3228:
3226:
3218:
3217:
3214:
3213:
3209:
3208:
3205:
3204:
3200:
3199:
3196:
3195:
3186:
3183:
3182:
3179:
3178:
3169:
3164:
3162:
3154:
3153:
3150:
3149:
3145:
3144:
3141:
3140:
3132:
3129:
3128:
3125:
3124:
3115:
3110:
3108:
3098:
3096:
3086:
3084:
3081:Perissodactyla
3074:
3065:
3062:
3061:
3060:
3059:
3058:
3051:
3044:
3043:
3042:
3041:
3040:
3039:
3038:
3031:
3025:
3018:
3016:Deperetellidae
3004:
3003:
3002:
3000:Hyracodontidae
2996:
2977:
2976:
2975:
2974:
2973:
2967:
2952:
2950:Lophiodontidae
2937:
2934:Isectolophidae
2925:
2924:
2923:
2922:
2921:
2914:
2903:
2897:
2895:Hyopsodontidae
2886:
2885:
2884:
2868:
2846:
2843:phenacodontids
2827:
2816:
2807:Indricotherium
2803:Baluchitherium
2793:Hyracodontidae
2784:
2770:Chalicotherium
2765:Lophiodontidae
2754:
2733:
2649:
2646:
2645:
2644:
2643:
2642:
2641:
2640:
2639:
2638:
2637:
2636:
2624:
2615:
2600:
2599:
2598:
2586:
2578:
2563:
2554:
2553:
2552:
2543:
2525:
2524:
2523:
2514:
2505:
2496:
2484:
2475:
2466:
2454:
2429:Rhinocerotidae
2424:
2423:
2422:
2416:Kabomani tapir
2413:
2404:
2395:
2389:Mountain tapir
2386:
2363:
2362:
2361:
2360:
2359:
2350:
2349:
2348:
2339:
2324:Mountain zebra
2321:
2320:
2319:
2310:
2301:
2292:
2286:Maneless zebra
2283:
2274:
2265:
2244:
2243:
2242:
2236:
2230:
2215:
2214:
2213:
2201:
2192:
2186:Persian onager
2183:
2174:
2163:Equus hemionus
2156:
2155:
2154:
2145:Atlas wild ass
2142:
2132:
2123:
2105:
2104:
2103:
2093:
2084:
2041:
2040:
2036:
2035:
2032:
2031:
2027:
2026:
2023:
2022:
2018:
2017:
2014:
2013:
2009:
2008:
2005:
2004:
2000:
1999:
1996:
1995:
1991:
1990:
1987:
1986:
1982:
1981:
1978:
1977:
1967:
1964:
1963:
1960:
1959:
1949:
1944:
1942:
1939:
1938:
1935:
1934:
1924:
1919:
1917:
1914:
1913:
1910:
1909:
1905:
1904:
1901:
1900:
1890:
1887:
1886:
1883:
1882:
1872:
1867:
1865:
1860:
1858:
1856:Rhinocerotidae
1851:
1850:
1847:
1846:
1842:
1841:
1838:
1837:
1833:
1832:
1829:
1828:
1824:
1823:
1820:
1819:
1815:
1814:
1811:
1810:
1800:
1797:
1796:
1793:
1792:
1782:
1777:
1775:
1772:
1771:
1768:
1767:
1757:
1752:
1750:
1747:
1746:
1743:
1742:
1732:
1727:
1725:
1722:
1721:
1718:
1717:
1707:
1702:
1700:
1691:
1689:
1686:
1685:
1682:
1681:
1677:
1676:
1673:
1672:
1668:
1667:
1664:
1663:
1659:
1658:
1655:
1654:
1650:
1649:
1646:
1645:
1641:
1640:
1637:
1636:
1626:
1623:
1622:
1619:
1618:
1608:
1603:
1601:
1598:
1597:
1594:
1593:
1583:
1578:
1576:
1573:
1572:
1569:
1568:
1564:
1563:
1560:
1559:
1549:
1546:
1545:
1542:
1541:
1537:Equus hemionus
1531:
1526:
1524:
1519:
1517:
1514:
1513:
1510:
1509:
1499:
1494:
1492:
1489:
1488:
1485:
1484:
1474:
1469:
1467:
1458:
1456:
1451:
1449:
1446:Perissodactyla
1439:
1418:
1417:Modern members
1415:
1375:Meridiungulata
1369:
1368:
1364:
1363:
1360:
1359:
1355:
1354:
1351:
1350:
1346:
1345:
1342:
1341:
1327:Meridiungulata
1319:
1316:
1315:
1312:
1311:
1307:Perissodactyla
1302:
1297:
1295:
1287:
1286:
1283:
1282:
1273:
1268:
1266:
1256:
1205:Laurasiatheria
1182:carotid artery
1165:
1164:Outer taxonomy
1162:
1154:
1151:
1130:
1127:
1105:
1102:
1051:
1048:
1039:
1036:
1006:
1003:
961:cranial nerves
930:guttural pouch
912:dental formula
842:
839:
819:common feature
772:
769:
736:
733:
722:paraceratheres
569:Perissodactyla
552:
551:
550:
549:
548:
547:
546:
545:
544:
543:
538:(rhinoceroses
536:Rhinocerotidae
533:
531:Eggysodontidae
525:
517:
515:Hyracodontidae
509:
496:(rhinoceroses
491:
490:
489:
483:
475:
473:Deperetellidae
457:
456:
455:
453:Lophiodontidae
447:
446:
445:
437:
410:
408:Isectolophidae
397:
396:
395:
394:
393:
385:
369:
368:(brontotheres)
360:
358:Hyopsodontidae
349:
348:
344:
343:
334:Perissodactyla
331:
327:
326:
321:
314:
313:
308:
304:
303:
298:
294:
293:
288:
284:
283:
278:
274:
273:
260:
259:
231:
230:
222:
221:
206:
201:
196:
191:
186:
181:
176:
171:
166:
161:
156:
151:
150:
141:
139:Perissodactyla
130:
129:
44:
42:
35:
26:
24:
18:Perissodactyls
14:
13:
10:
9:
6:
4:
3:
2:
7994:
7983:
7980:
7978:
7975:
7973:
7972:Mammal orders
7970:
7968:
7965:
7963:
7960:
7959:
7957:
7940:
7934:
7930:
7925:
7921:
7917:
7912:
7908:
7904:
7898:
7894:
7889:
7885:
7881:
7876:
7872:
7868:
7863:
7859:
7855:
7850:
7846:
7842:
7837:
7833:
7829:
7824:
7820:
7816:
7811:
7807:
7803:
7798:
7794:
7790:
7785:
7781:
7777:
7772:
7768:
7764:
7759:
7755:
7751:
7746:
7742:
7738:
7733:
7729:
7725:
7719:
7715:
7710:
7706:
7702:
7697:
7693:
7688:
7682:
7678:
7673:
7667:
7663:
7662:
7660:
7658:
7654:
7650:
7645:
7635:
7634:
7623:
7609:
7607:
7602:
7600:
7598:
7597:T. terrestris
7593:
7591:
7589:
7584:
7582:
7580:
7575:
7574:
7572:
7570:
7569:
7564:
7561:
7552:
7548:
7538:
7536:
7531:
7530:
7528:
7526:
7525:
7520:
7514:
7512:
7507:
7506:
7504:
7502:
7501:
7500:Ceratotherium
7496:
7490:
7488:
7483:
7482:
7480:
7478:
7477:
7472:
7466:
7464:
7459:
7457:
7455:
7450:
7449:
7447:
7445:
7444:
7439:
7436:
7426:
7422:
7418:
7417:
7411:
7407:
7393:
7391:
7386:
7384:
7382:
7377:
7375:
7373:
7367:
7366:
7365:
7359:
7358:
7355:
7353:
7348:
7346:
7344:
7339:
7337:
7335:
7330:
7328:
7326:
7320:
7319:
7318:
7312:
7311:
7308:
7306:
7301:
7297:
7295:
7290:
7289:
7288:
7286:
7280:
7279:
7278:
7272:
7271:
7269:
7263:
7255:
7254:
7249:
7246:
7236:
7232:
7228:
7227:
7221:
7217:
7211:
7207:
7205:
7201:
7199:
7195:
7193:
7189:
7187:
7183:
7182:
7179:
7174:
7163:
7158:
7156:
7151:
7149:
7144:
7143:
7140:
7116:
7110:
7108:
7102:
7100:
7094:
7092:
7086:
7084:
7078:
7077:
7075:
7073:
7069:
7063:
7059:Artiodactyla
7057:
7055:
7049:
7047:
7041:
7039:
7033:
7031:
7025:
7023:
7019:Eulipotyphla
7017:
7016:
7014:
7012:
7008:
7005:
7003:
7002:Boreoeutheria
6999:
6993:
6987:
6985:
6979:
6977:
6971:
6969:
6963:
6961:
6955:
6953:
6949:Afrosoricida
6947:
6946:
6944:
6942:
6938:
6932:
6926:
6924:
6918:
6917:
6915:
6913:
6909:
6906:
6900:
6892:
6888:
6878:
6872:
6870:
6864:
6862:
6856:
6854:
6848:
6846:
6840:
6839:
6837:
6835:
6831:
6825:
6819:
6817:
6811:
6810:
6808:
6806:
6802:
6799:
6793:
6785:
6781:
6778:
6776:
6772:
6762:
6756:
6755:
6753:
6751:
6747:
6744:
6742:
6738:
6732:
6728:
6726:
6722:
6720:
6716:
6714:
6710:
6709:
6706:
6701:
6693:
6688:
6686:
6681:
6679:
6674:
6673:
6670:
6663:
6659:
6655:
6649:
6645:
6640:
6637:
6636:0-8018-8221-4
6633:
6629:
6628:
6623:
6620:
6619:3-334-00223-3
6616:
6612:
6608:
6605:
6604:0-19-850761-5
6601:
6597:
6593:
6590:
6589:0-8018-5789-9
6586:
6582:
6578:
6575:
6574:3-8274-0307-3
6571:
6567:
6563:
6562:
6558:
6549:
6542:
6539:
6534:
6530:
6525:
6520:
6516:
6512:
6508:
6504:
6500:
6493:
6490:
6485:
6478:
6475:
6470:
6457:
6449:
6442:
6439:
6434:
6427:
6424:
6419:
6415:
6408:
6405:
6400:
6393:
6390:
6385:
6381:
6376:
6371:
6366:
6361:
6357:
6353:
6349:
6345:
6341:
6334:
6331:
6326:
6322:
6315:
6312:
6307:
6303:
6299:
6295:
6288:
6281:
6278:
6273:
6271:0-412-73810-4
6267:
6263:
6256:
6253:
6248:
6242:
6238:
6234:
6230:
6223:
6220:
6215:
6211:
6207:
6200:
6197:
6192:
6188:
6184:
6180:
6176:
6172:
6164:
6161:
6155:
6150:
6145:
6140:
6136:
6132:
6128:
6121:
6118:
6112:
6107:
6103:
6099:
6095:
6088:
6085:
6080:
6076:
6072:
6068:
6064:
6060:
6056:
6052:
6048:
6044:
6040:
6033:
6030:
6025:
6021:
6016:
6011:
6007:
6003:
5999:
5995:
5991:
5984:
5981:
5976:
5972:
5967:
5962:
5957:
5952:
5948:
5944:
5940:
5936:
5932:
5925:
5922:
5917:
5913:
5909:
5905:
5901:
5897:
5893:
5886:
5883:
5878:
5874:
5869:
5864:
5859:
5854:
5850:
5846:
5842:
5838:
5834:
5827:
5824:
5819:
5815:
5808:
5805:
5800:
5796:
5792:
5788:
5784:
5780:
5772:
5769:
5764:
5760:
5753:
5750:
5745:
5738:
5731:
5728:
5723:
5716:
5713:
5708:
5704:
5697:
5694:
5689:
5676:
5664:
5661:
5656:
5652:
5648:
5644:
5637:
5634:
5629:
5625:
5618:
5615:
5610:
5606:
5602:
5598:
5594:
5590:
5583:
5580:
5575:
5568:
5566:
5564:
5562:
5558:
5553:
5549:
5545:
5541:
5534:
5531:
5526:
5519:
5516:
5511:
5504:
5502:
5500:
5496:
5491:
5487:
5483:
5479:
5475:
5471:
5464:
5462:
5458:
5453:
5449:
5444:
5439:
5434:
5429:
5425:
5421:
5417:
5413:
5409:
5402:
5400:
5396:
5391:
5387:
5383:
5379:
5375:
5371:
5370:Palaeontology
5363:
5360:
5354:
5349:
5345:
5341:
5337:
5330:
5328:
5324:
5319:
5315:
5310:
5305:
5302:(3): 219–43.
5301:
5297:
5293:
5286:
5283:
5278:
5274:
5270:
5266:
5262:
5261:Pachynolophus
5258:
5257:Pachynolophus
5251:
5248:
5243:
5239:
5235:
5231:
5227:
5223:
5216:
5213:
5207:
5202:
5198:
5194:
5190:
5183:
5181:
5179:
5175:
5170:
5166:
5161:
5156:
5151:
5146:
5142:
5138:
5134:
5130:
5126:
5119:
5116:
5111:
5107:
5102:
5097:
5092:
5087:
5083:
5079:
5075:
5071:
5067:
5060:
5057:
5051:
5046:
5042:
5038:
5034:
5027:
5024:
5018:
5013:
5009:
5005:
5001:
4994:
4991:
4985:
4980:
4976:
4972:
4968:
4961:
4958:
4953:
4949:
4945:
4941:
4937:
4933:
4926:
4919:
4916:
4911:
4903:
4900:
4895:
4891:
4887:
4883:
4878:
4873:
4869:
4865:
4861:
4857:
4853:
4849:
4845:
4837:
4835:
4831:
4826:
4822:
4817:
4812:
4807:
4802:
4798:
4794:
4790:
4786:
4782:
4775:
4772:
4767:
4763:
4759:
4755:
4750:
4745:
4740:
4735:
4731:
4727:
4723:
4716:
4713:
4708:
4704:
4700:
4696:
4692:
4688:
4680:
4678:
4674:
4668:
4665:
4652:
4648:
4642:
4639:
4634:
4628:
4624:
4617:
4614:
4601:
4597:
4593:
4586:
4583:
4578:
4572:
4569:
4557:
4553:
4547:
4544:
4539:
4535:
4530:
4525:
4521:
4517:
4513:
4509:
4505:
4501:
4497:
4490:
4488:
4484:
4471:
4464:
4457:
4454:
4449:
4443:
4439:
4432:
4429:
4416:
4412:
4405:
4402:
4389:
4385:
4378:
4375:
4370:
4364:
4360:
4353:
4351:
4349:
4347:
4343:
4338:
4334:
4329:
4324:
4320:
4316:
4312:
4308:
4304:
4300:
4296:
4289:
4286:
4281:
4277:
4273:
4269:
4265:
4261:
4257:
4253:
4249:
4245:
4241:
4234:
4231:
4225:
4222:
4217:
4215:9780982261965
4211:
4207:
4206:
4198:
4195:
4183:
4179:
4173:
4170:
4165:
4161:
4157:
4151:
4148:
4143:
4139:
4133:
4130:
4125:
4121:
4117:
4113:
4109:
4105:
4098:
4095:
4083:
4079:
4073:
4070:
4063:
4058:
4057:least concern
4054:
4050:
4047:
4043:
4039:
4035:
4031:
4027:
4023:
4020:
4016:
4012:
4008:
4004:
4001:
4000:Grévy's zebra
3997:
3993:
3992:Malayan tapir
3989:
3985:
3981:
3977:
3974:
3970:
3966:
3962:
3958:
3954:
3953:
3952:
3950:
3945:
3943:
3939:
3934:
3931:
3930:domestication
3923:
3918:
3911:
3909:
3906:
3902:
3898:
3894:
3890:
3889:
3884:
3879:
3877:
3873:
3872:Isle of Wight
3869:
3865:
3861:
3857:
3856:solidungulate
3853:
3848:
3846:
3842:
3838:
3834:
3830:
3826:
3822:
3818:
3814:
3810:
3806:
3802:
3798:
3794:
3790:
3786:
3782:
3778:
3774:
3770:
3766:
3757:
3750:
3748:
3746:
3745:
3744:Elasmotherium
3740:
3736:
3732:
3728:
3724:
3720:
3719:
3712:
3710:
3706:
3702:
3698:
3697:
3692:
3691:
3686:
3682:
3679:, as well as
3678:
3677:Ganderalophus
3674:
3670:
3666:
3665:
3660:
3659:hyopsodontids
3656:
3648:
3647:
3642:
3635:
3633:
3631:
3627:
3626:South America
3623:
3617:
3615:
3611:
3610:Cambaytherium
3607:
3606:Cambaytherium
3602:
3598:
3594:
3590:
3586:
3585:
3584:Cambaytherium
3579:
3575:
3574:
3569:
3561:
3556:
3554:
3552:
3548:
3543:
3539:
3534:
3529:
3527:
3523:
3519:
3515:
3510:
3496:
3495:
3487:
3486:
3478:
3477:
3469:
3468:
3460:
3459:
3451:
3450:
3442:
3441:
3438:
3437:
3430:
3429:
3426:
3425:
3422:
3420:
3413:
3412:
3406:
3405:
3402:
3401:
3398:
3396:
3395:Amynodontidae
3389:
3388:
3381:
3377:
3376:
3373:
3372:
3364:
3363:
3360:
3359:
3352:
3351:
3348:
3347:
3344:
3342:
3335:
3334:
3327:
3323:
3322:
3315:
3311:
3310:
3307:
3306:
3298:
3297:
3294:
3292:
3285:
3284:
3281:
3280:
3277:
3275:
3268:
3267:
3260:
3256:
3255:
3249:
3248:
3245:
3244:
3241:
3239:
3232:
3231:
3224:
3220:
3219:
3216:
3215:
3207:
3206:
3198:
3197:
3194:
3192:
3185:
3184:
3181:
3180:
3177:
3175:
3168:
3167:
3160:
3156:
3155:
3152:
3151:
3143:
3142:
3139:
3138:
3131:
3130:
3127:
3126:
3123:
3121:
3114:
3113:
3106:
3102:
3101:
3094:
3090:
3089:
3082:
3078:
3077:
3073:
3072:
3063:
3056:
3052:
3049:
3045:
3036:
3032:
3030:
3029:Lophialetidae
3026:
3023:
3019:
3017:
3013:
3012:
3011:
3010:
3005:
3001:
2997:
2995:
2994:Amynodontidae
2991:
2990:
2989:
2988:
2983:
2982:
2981:
2978:
2972:
2968:
2965:
2961:
2960:
2959:
2958:
2953:
2951:
2947:
2946:
2945:
2944:
2938:
2935:
2931:
2930:
2929:
2926:
2919:
2915:
2913:
2909:
2908:
2907:
2904:
2902:
2898:
2896:
2892:
2891:
2890:
2887:
2883:
2879:
2878:
2877:
2874:
2873:
2872:
2869:
2866:
2862:
2858:
2854:
2850:
2847:
2844:
2840:
2839:hyopsodontids
2836:
2832:
2828:
2825:
2820:
2817:
2814:
2813:
2808:
2804:
2800:
2799:
2794:
2790:
2789:Amynodontidae
2785:
2782:
2778:
2777:
2772:
2771:
2766:
2762:
2758:
2755:
2752:
2748:
2747:
2742:
2738:
2734:
2731:
2727:
2723:
2722:
2717:
2710:
2708:
2704:
2699:
2695:
2694:
2693:
2687:
2683:
2681:
2677:
2672:
2665:
2661:
2660:
2659:Palaeotherium
2654:
2647:
2635:
2632:
2628:
2625:
2623:
2619:
2616:
2614:
2610:
2609:
2608:
2604:
2601:
2597:
2594:
2590:
2587:
2585:
2584:
2579:
2577:
2573:
2572:
2571:
2567:
2564:
2562:
2558:
2555:
2551:
2547:
2544:
2542:
2538:
2535:
2534:
2533:
2529:
2526:
2522:
2518:
2515:
2513:
2509:
2506:
2504:
2500:
2497:
2495:
2492:
2488:
2485:
2483:
2479:
2476:
2474:
2470:
2467:
2465:
2462:
2458:
2455:
2453:
2450:
2446:
2443:
2442:
2441:
2437:
2434:
2433:
2431:
2430:
2425:
2421:
2417:
2414:
2412:
2408:
2407:Malayan tapir
2405:
2403:
2399:
2398:Baird's tapir
2396:
2394:
2390:
2387:
2385:
2381:
2378:
2377:
2375:
2374:
2369:
2368:
2367:
2364:
2358:
2354:
2353:Grévy's zebra
2351:
2347:
2343:
2340:
2338:
2334:
2331:
2330:
2329:
2325:
2322:
2318:
2314:
2313:Selous' zebra
2311:
2309:
2305:
2302:
2300:
2296:
2293:
2291:
2287:
2284:
2282:
2278:
2277:Grant's zebra
2275:
2273:
2269:
2266:
2264:
2261:
2257:
2254:
2253:
2252:
2248:
2245:
2241:
2237:
2235:
2231:
2229:
2225:
2224:
2223:
2219:
2216:
2212:
2209:
2205:
2202:
2200:
2196:
2193:
2191:
2187:
2184:
2182:
2178:
2175:
2173:
2169:
2166:
2165:
2164:
2160:
2157:
2153:
2150:
2146:
2143:
2141:
2137:
2133:
2131:
2127:
2124:
2122:
2118:
2115:
2114:
2113:
2109:
2106:
2102:
2098:
2094:
2092:
2088:
2085:
2083:
2080:
2076:
2073:
2072:
2071:
2067:
2064:
2063:
2061:
2060:
2055:
2054:
2053:
2050:
2049:
2048:
2045:
2044:
2034:
2033:
2025:
2024:
2016:
2015:
2007:
2006:
1998:
1997:
1989:
1988:
1980:
1979:
1976:
1975:
1974:
1966:
1965:
1962:
1961:
1958:
1957:
1956:
1948:
1947:
1941:
1940:
1937:
1936:
1933:
1932:
1931:
1923:
1922:
1916:
1915:
1912:
1911:
1903:
1902:
1899:
1898:
1897:
1889:
1888:
1885:
1884:
1881:
1880:
1879:
1871:
1870:
1864:
1863:
1857:
1853:
1852:
1849:
1848:
1840:
1839:
1831:
1830:
1822:
1821:
1813:
1812:
1809:
1808:
1807:
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1498:
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1408:
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1400:
1396:
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1300:
1293:
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1284:
1281:
1279:
1272:
1271:
1264:
1260:
1259:
1255:
1254:
1250:
1244:
1242:
1238:
1234:
1230:
1226:
1222:
1218:
1214:
1210:
1206:
1202:
1198:
1194:
1190:
1185:
1183:
1179:
1175:
1171:
1163:
1160:
1152:
1150:
1148:
1140:
1135:
1128:
1126:
1124:
1120:
1116:
1112:
1111:grass feeders
1103:
1101:
1098:
1094:
1090:
1089:Baird's tapir
1086:
1082:
1078:
1074:
1070:
1061:
1056:
1049:
1047:
1045:
1037:
1035:
1033:
1028:
1024:
1020:
1016:
1012:
1004:
1002:
999:
994:
990:
986:
982:
978:
974:
970:
966:
962:
958:
954:
950:
949:Aspergillosis
946:
943:
939:
935:
934:auditory tube
931:
926:
913:
909:
905:
901:
896:
894:
890:
887:
882:
881:is enlarged.
880:
876:
872:
871:lacrimal bone
868:
864:
860:
856:
847:
840:
838:
836:
832:
828:
824:
820:
816:
812:
807:
805:
801:
797:
793:
789:
785:
777:
770:
768:
767:are striped.
766:
762:
758:
754:
753:
748:
744:
743:
734:
732:
730:
725:
723:
719:
718:chalicotheres
715:
711:
706:
704:
700:
696:
692:
688:
684:
680:
676:
672:
668:
664:
660:
656:
652:
648:
644:
640:
637:
633:
630:
629:Ancient Greek
624:
570:
563:
558:
541:
540:sensu stricto
537:
534:
532:
526:
524:
518:
516:
510:
508:
507:Amynodontidae
502:
501:
499:
495:
492:
487:
484:
482:
476:
474:
468:
467:
466:
463:
462:
461:
458:
454:
448:
444:
438:
435:
429:
428:
427:
421:
420:
419:
418:
411:
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403:
402:
401:
398:
392:
386:
383:
380:
379:
378:
375:
374:
373:
370:
367:
361:
359:
353:
352:
350:
345:
340:
335:
332:
329:
328:
325:
322:
319:
316:
315:
312:
309:
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305:
302:
299:
296:
295:
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279:
276:
275:
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257:
253:
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245:
241:
237:
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228:
223:
219:
215:
209:
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199:
194:
189:
184:
179:
174:
169:
164:
159:
154:
148:
137:
134:
126:
123:
115:
104:
101:
97:
94:
90:
87:
83:
80:
76:
73: –
72:
68:
67:Find sources:
61:
57:
51:
50:
45:This article
43:
39:
34:
33:
30:
19:
7656:
7631:
7605:
7596:
7588:T. pinchaque
7587:
7578:
7566:
7534:
7522:
7510:
7498:
7486:
7476:Dicerorhinus
7474:
7463:R. sondaicus
7462:
7454:R. unicornis
7453:
7441:
7416:Ceratomorpha
7414:
7389:
7380:
7371:
7362:
7360:
7351:
7342:
7333:
7325:E. africanus
7324:
7315:
7313:
7304:
7293:
7284:
7283:Wild horse (
7275:
7273:
7251:
7224:
7169:
7098:(Treeshrews)
6981:Proboscidea
6922:(Armadillos)
6805:Ameridelphia
6758:Monotremata
6643:
6625:
6610:
6595:
6580:
6565:
6541:
6506:
6502:
6492:
6483:
6477:
6456:cite journal
6441:
6432:
6426:
6417:
6413:
6407:
6398:
6392:
6347:
6343:
6333:
6324:
6320:
6314:
6297:
6293:
6280:
6261:
6255:
6228:
6222:
6213:
6209:
6205:
6199:
6174:
6170:
6163:
6134:
6130:
6120:
6101:
6097:
6087:
6046:
6042:
6032:
5997:
5993:
5983:
5938:
5934:
5924:
5899:
5895:
5885:
5840:
5836:
5826:
5817:
5807:
5782:
5778:
5771:
5762:
5752:
5743:
5730:
5721:
5715:
5706:
5702:
5696:
5675:cite journal
5663:
5646:
5642:
5636:
5627:
5624:Paleomoropus
5623:
5617:
5592:
5588:
5582:
5573:
5543:
5539:
5533:
5524:
5518:
5509:
5473:
5469:
5415:
5411:
5373:
5369:
5362:
5343:
5339:
5299:
5295:
5285:
5268:
5264:
5260:
5256:
5250:
5225:
5221:
5215:
5196:
5192:
5132:
5128:
5118:
5073:
5069:
5059:
5040:
5036:
5026:
5007:
5003:
4993:
4974:
4970:
4960:
4938:(1): 34–44.
4935:
4931:
4918:
4909:
4902:
4851:
4847:
4788:
4784:
4774:
4732:(6): 65–74.
4729:
4725:
4715:
4690:
4686:
4667:
4655:. Retrieved
4650:
4641:
4622:
4616:
4604:. Retrieved
4600:the original
4595:
4585:
4576:
4571:
4559:. Retrieved
4555:
4546:
4503:
4499:
4476:25 September
4474:. Retrieved
4469:
4456:
4437:
4431:
4421:25 September
4419:. Retrieved
4414:
4404:
4394:25 September
4392:. Retrieved
4387:
4377:
4358:
4302:
4298:
4288:
4247:
4243:
4233:
4224:
4204:
4197:
4185:. Retrieved
4181:
4178:"Dawn horse"
4172:
4164:the original
4159:
4150:
4142:the original
4132:
4107:
4103:
4097:
4085:. Retrieved
4081:
4072:
4041:
4030:plains zebra
3946:
3935:
3927:
3904:
3900:
3896:
3892:
3886:
3880:
3875:
3868:Richard Owen
3863:
3860:multungulate
3859:
3855:
3849:
3844:
3840:
3833:Pachydermata
3821:proboscidean
3816:
3796:
3792:
3788:
3784:
3781:Hippopotamus
3780:
3772:
3764:
3762:
3742:
3723:Tethys Ocean
3716:
3713:
3708:
3705:brontotheres
3694:
3688:
3684:
3680:
3676:
3662:
3652:
3644:
3618:
3609:
3605:
3596:
3588:
3582:
3571:
3565:
3550:
3547:Tapiromorpha
3546:
3530:
3525:
3509:Richard Owen
3506:
3433:
3416:
3392:
3355:
3338:
3314:Ceratomorpha
3288:
3271:
3235:
3223:Tapiromorpha
3188:
3171:
3134:
3117:
3080:
3022:Rhodopagidae
3007:Superfamily
3006:
2985:Superfamily
2984:
2979:
2955:Superfamily
2954:
2940:
2927:
2912:Indolophidae
2905:
2888:
2875:
2870:
2861:Tethys Ocean
2810:
2806:
2802:
2796:
2774:
2768:
2744:
2719:
2705:
2691:
2685:
2678:
2676:chalicothere
2657:
2633:
2621:
2612:
2606:
2595:
2581:
2575:
2569:
2560:
2549:
2540:
2531:
2520:
2511:
2502:
2493:
2481:
2472:
2463:
2451:
2439:
2426:
2419:
2410:
2401:
2392:
2383:
2370:
2365:
2357:Equus grevyi
2356:
2345:
2336:
2327:
2316:
2307:
2298:
2289:
2280:
2271:
2262:
2251:Equus quagga
2250:
2247:Plains zebra
2239:
2233:
2227:
2221:
2210:
2198:
2189:
2180:
2171:
2162:
2151:
2139:
2129:
2120:
2111:
2100:
2090:
2081:
2069:
2056:
2051:
2046:
1971:
1969:
1953:
1951:
1928:
1926:
1894:
1892:
1876:
1874:
1804:
1802:
1786:
1784:
1761:
1759:
1736:
1734:
1711:
1709:
1632:Equus quagga
1630:
1628:
1614:Equus grevyi
1612:
1610:
1587:
1585:
1553:
1551:
1535:
1533:
1505:Equus asinus
1503:
1501:
1478:
1476:
1445:
1420:
1399:Notoungulata
1392:
1384:Macrauchenia
1382:
1372:
1333:Notoungulata
1321:
1306:
1304:
1278:Artiodactyla
1275:
1248:
1213:Artiodactyla
1195:rather than
1189:polyphyletic
1186:
1170:artiodactyls
1167:
1143:
1123:leaf feeders
1115:white rhinos
1107:
1065:
1050:Distribution
1044:carotid rete
1041:
1034:is smaller.
1008:
998:S. equi equi
997:
993:S. equi equi
992:
988:
952:
927:
897:
886:agglutinated
883:
852:
808:
786:
782:
750:
740:
738:
729:Richard Owen
726:
714:palaeotheres
710:brontotheres
707:
687:artiodactyls
675:rhinoceroses
642:
639:
635:
632:
568:
567:
539:
497:
460:Ceratomorpha
459:
415:
400:Tapiromorpha
399:
376:
371:
333:
317:
255:
247:
240:Equus quagga
239:
236:plains zebra
133:
118:
112:October 2021
109:
99:
92:
85:
78:
66:
54:Please help
49:verification
46:
29:
7823:iNaturalist
7681:Wikispecies
7535:D. bicornis
7364:Hippotigris
7343:E. hemionus
7305:E. caballus
7260:(including
7226:Hippomorpha
7208:Superorder
7202:Infraclass
7104:Dermoptera
7096:Scandentia
7088:Lagomorpha
7037:(Pangolins)
7027:Chiroptera
6983:(Elephants)
6973:Hyracoidea
6729:(unranked)
6609:AH Müller:
5785:: 219–225.
5271:(1): 1–16.
4877:11336/14769
4463:"Strangles"
4187:7 September
3841:Solidungula
3673:Balochistan
3601:Tethytheria
3589:Kalitherium
3341:Helaletidae
3093:Hippomorpha
2964:Eomoropidae
2849:Desmostylia
2831:condylarths
2824:Helaletidae
2781:Pleistocene
2664:palaeothere
2328:Equus zebra
2222:Equus kiang
2070:Equus ferus
1589:Equus zebra
1555:Equus kiang
1480:Equus ferus
1241:Pegasoferae
1023:gallbladder
953:Aspergillus
924:× 2 = 30-44
837:is absent.
627:, from
481:Helaletidae
443:Eomoropidae
372:Hippomorpha
7956:Categories
7606:T. indicus
7579:T. bairdii
7443:Rhinoceros
7043:Carnivora
7035:Pholidota
6967:(Aardvark)
6941:Afrotheria
6920:Cingulata
6901:inclusive)
6823:(Opossums)
6794:inclusive)
6784:Metatheria
6741:Yinotheria
6725:Vertebrata
6723:Subphylum
6401:: 164–190.
6300:: 85–101.
5765:: 101–110.
5746:: 101–113.
5527:: 181–295.
5512:: 252–258.
4749:1807/64815
4606:4 February
4064:References
4007:vulnerable
3996:wild horse
3980:endangered
3805:lagomorphs
3709:Megacerops
3685:Eotitanops
3681:Litolophus
3675:, such as
3664:Sifrhippus
3536:coined by
3518:Tapiroidea
3326:Tapiroidea
3259:Ancylopoda
3009:Tapiroidea
2943:Ancylopoda
2857:afrotheres
2837:, such as
2819:Tapiroidea
2735:Equoidea (
2721:Megacerops
2684:(formerly
2066:Wild horse
1389:Litopterna
1337:Litopterna
1263:Euungulata
1221:Euungulata
1201:Afrotheria
1157:See also:
1062:shown here
979:) and the
938:middle ear
867:premaxilla
823:astragalus
804:rhinoceros
763:and young
720:, and the
695:intestines
643:, dáktylos
636:, perissós
498:sensu lato
465:Tapiroidea
417:Ancylopoda
145:56–0
82:newspapers
7551:Tapiridae
7390:E. grevyi
7372:E. quagga
7361:Subgenus
7334:E. asinus
7314:Subgenus
7274:Subgenus
7112:Primates
7106:(Colugos)
7080:Rodentia
6975:(Hyraxes)
6912:Xenarthra
6899:Placental
6792:Marsupial
6191:129473193
6137:: 21–34.
5916:226263139
5902:: 1–147.
5390:214053761
4520:1744-9561
4470:Strangles
4319:0021-8782
4264:0340-2096
4160:Phenomena
3893:Mesaxonia
3888:Mesaxonia
3881:In 1884,
3850:In 1861,
3845:Solipèdes
3837:peccaries
3731:ruminants
3701:megafauna
3696:Hyrachyus
3636:Phylogeny
3630:K–T event
3597:Radinskya
3578:Paleocene
3573:Radinskya
3358:Tapiridae
3035:Eoletidae
2941:Suborder
2751:Oligocene
2373:Tapiridae
2095:Domestic
1698:Tapiridae
1424:Paleocene
1411:Paleogene
1397:from the
1387:from the
1237:Pholidota
1233:Carnivora
1217:cetaceans
1178:elephants
1147:precocial
1073:Oligocene
1011:ruminants
985:Strangles
973:Accessory
855:upper jaw
796:ungulates
788:Ungulates
757:epidermis
747:Oligocene
693:in their
691:cellulose
683:vestigial
647:ungulates
486:Tapiridae
347:Families
287:Kingdom:
281:Eukaryota
7867:14100001
7721:BioLib:
7666:Wikidata
7633:Category
7556:(Tapirs)
7511:C. simum
7381:E. zebra
7352:E. kiang
7341:Onager (
7332:Donkey (
7285:E. ferus
7204:Eutheria
7198:Mammalia
7192:Chordata
7186:Animalia
7184:Kingdom
6989:Sirenia
6891:Eutheria
6719:Chordata
6713:Animalia
6711:Kingdom
6662:98023686
6533:17329227
6450:: 1–710.
6420:: 1–540.
6384:20007379
6327:: 12–14.
6071:22363006
6024:27358361
5975:27384169
5935:PLOS ONE
5877:25410701
5843:: 5570.
5799:86206151
5490:84351811
5452:25295875
5412:PLOS ONE
5346:: 1–54.
5318:84061253
5242:84391276
5228:(8): 8.
5169:27384169
5129:PLOS ONE
5110:25295875
5070:PLOS ONE
4952:11286489
4886:25799987
4825:16785431
4766:30067369
4758:23266984
4538:18426746
4337:31373392
4280:34568531
4124:84391276
4110:(8): 8.
4032:and the
4017:and the
3971:and the
3951:(2012):
3817:le tapir
3769:Linnaeus
3739:Pliocene
3727:mammoths
3690:Eohippus
3646:Eohippus
3514:Equoidea
3105:Equoidea
2746:Eohippus
2680:Anisodon
1209:Laurasia
1153:Taxonomy
1137:A young
1085:Pliocene
900:incisors
879:mandible
859:diastema
835:clavicle
829:and the
827:scaphoid
752:Eohippus
655:families
641:δάκτυλος
634:περισσός
488:(tapirs)
377:Equoidea
311:Mammalia
301:Chordata
297:Phylum:
291:Animalia
277:Domain:
7568:Tapirus
7524:Diceros
7350:Kiang (
7235:Equidae
7190:Phylum
7168:Extant
6928:Pilosa
6731:Amniota
6717:Phylum
6698:Extant
6524:2189562
6375:2799835
6352:Bibcode
6321:Tapirus
6079:4603597
6051:Bibcode
6043:Science
6015:4936024
5966:4934866
5943:Bibcode
5845:Bibcode
5630:: 1–28.
5609:1005518
5546:: 106.
5443:4189980
5420:Bibcode
5265:Geobios
5160:4934866
5137:Bibcode
5101:4189980
5078:Bibcode
4894:4467386
4856:Bibcode
4816:1479866
4793:Bibcode
4707:9126561
4657:12 June
4529:2610139
4328:6875937
4272:1199544
4205:Namibia
4087:18 July
3809:rodents
3797:Belluae
3785:Tapirus
3737:in the
3669:Wyoming
3562:Origins
3382:
3328:
3316:
3261:
3225:
3161:
3137:Equidae
3107:
3095:
3083:
2865:Pacific
2776:Moropus
2737:equines
2427:Family
2371:Family
2059:Equidae
2057:Family
1465:Equidae
1448:
1394:Toxodon
1294:
1265:
1174:hyraxes
1119:equines
1093:tarpans
1071:to the
1015:stomach
904:canines
889:keratin
815:fibulae
800:humerus
735:Anatomy
699:stomach
659:Equidae
651:species
382:Equidae
330:Order:
307:Class:
218:present
96:scholar
7939:105959
7936:uBio:
7916:541948
7900:NZOR:
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