117:
960:
361:. In a 1985 paper he gives a brief comment on the genus. He stated without going into specific details, that while the fossil is superficially similar to those of Cathartidae members, however it possesses notable features which are different and as such he regarded the position of the genus as problematic. In their 2005 description of a Peruvian fossil, Marcelo Stucchi and Steven Emslie noted the disputed nature of
397:, and the width across the condyles is 14.9 millimetres (0.59 in). The diameter at the smallest area of the preserved bone shaft is 9.2 millimetres (0.36 in).
319:
quarry, possibly a fossilized watering hole. The quarry worked sedimentary rocks that are positioned approximately 25 feet (7.6 m) above the contact between the
305:
565:
2296:
389:
as being "not much larger than the Black
Vulture". The single known fossil is a partial upper leg bone, specifically the lower section of the right
2311:
2232:
2306:
2316:
558:
959:
494:
2291:
365:, but maintained the placement in Cathartidae and noted the genus as the oldest member of the family in the Americas.
522:
449:
116:
551:
2301:
2200:
357:
identifies the genus as a vulture. The fossil was reexamined in the early 1980s by avian paleontologist
2258:
219:
1054:
1044:
1533:
846:
664:
2286:
543:
450:"A new condor (Ciconiiformes, Vulturidae) from the Late Miocene/Early Pliocene Pisco Formation, Peru"
1255:
782:
1542:
1245:
799:
1931:
1894:
1870:
1327:
1195:
472:
393:. As preserved, the largest diameter of the bone is 17.9 millimetres (0.70 in) around the
111:
2263:
1287:
1235:
1175:
819:
745:
1860:
1367:
1307:
1215:
1205:
1185:
1113:
856:
836:
2245:
1627:
1297:
1266:
896:
876:
394:
332:
324:
309:
261:
238:
178:
2250:
1911:
1429:
1123:
631:
616:
464:
1985:
992:
977:
935:
678:
575:
358:
1491:
1021:
643:
579:
336:
312:
and given the number 1078. In 1923 Philip
Reinheinter collected the specimen from the
168:
420:
2280:
2170:
2121:
2051:
2033:
1803:
1755:
1519:
1463:
1456:
1436:
1165:
1071:
1064:
1032:
886:
378:
63:
476:
2163:
2149:
2094:
2071:
1999:
1949:
1938:
1921:
1769:
1669:
1662:
1648:
1579:
1413:
1317:
1225:
1082:
903:
701:
320:
331:. The fossil was first studied by the American paleontologist and ornithologist
2135:
1965:
1823:
1783:
1762:
1739:
1641:
1634:
1620:
1402:
1381:
1155:
1144:
1012:
755:
733:
715:
390:
313:
265:
38:
2223:
2142:
2087:
2060:
2026:
1992:
1976:
1958:
1884:
1848:
1690:
1683:
1599:
1553:
1505:
1477:
1470:
1357:
1338:
1103:
1001:
722:
694:
610:
328:
83:
48:
32:
2128:
2010:
1837:
1776:
1748:
1717:
1676:
1590:
1512:
1484:
1447:
1422:
1388:
1133:
920:
826:
772:
708:
687:
344:
283:
269:
128:
88:
289:
known, though its placement in the family
Cathartidae has been questioned.
2217:
2156:
2105:
2080:
2040:
1877:
1814:
1794:
1697:
1606:
1569:
1498:
1347:
809:
789:
765:
598:
530:
Transactions of the
Nebraska Academy of Sciences and Affiliated Societies
301:
273:
148:
78:
73:
58:
53:
43:
2237:
2114:
1904:
1708:
1655:
1277:
286:
254:
93:
68:
2019:
1830:
1724:
1395:
910:
592:
138:
2194:
1613:
604:
257:
468:
1374:
385:. This is contradicted by James Ducey's 1992 paper which lists
353:
158:
2198:
975:
629:
547:
339:
of the new genus and species was published in the journal
377:
was described as being one third larger than the modern
428:
Proceedings of the
Colorado Museum of Natural History
347:
explanations for the genus and species names, though
341:
Proceedings of the
Colorado Museum of Natural History
1927:
1917:
1900:
1890:
1866:
1856:
1575:
1565:
1530:
1363:
1353:
1323:
1313:
1303:
1293:
1283:
1273:
1261:
1251:
1241:
1231:
1221:
1211:
1201:
1191:
1181:
1171:
1161:
1151:
1139:
1129:
1119:
1109:
1099:
1060:
1050:
1040:
989:
916:
892:
882:
872:
862:
852:
842:
832:
815:
805:
795:
778:
761:
751:
741:
675:
660:
300:
is known from a single fragmentary fossil bone, the
2207:
2104:
2070:
2050:
2009:
1975:
1948:
1847:
1813:
1793:
1738:
1707:
1589:
1552:
1529:
1446:
1412:
1337:
1090:
1081:
1031:
1011:
988:
934:
732:
674:
651:
642:
351:is Latin and Greek for a phantom or apparition and
276:. The genus contains a single described species,
343:. In the description, Wetmore did not give any
304:housed in the paleontology collections of the
559:
443:
441:
421:"Fossil Birds from the Oligocene of Colorado"
414:
412:
410:
8:
488:
486:
2195:
1558:
1087:
985:
972:
648:
639:
626:
566:
552:
544:
20:
406:
502:. Vol. Avian Biology. p. 191
523:"Fossil Birds of the Nebraska Region"
7:
448:Stucchi, M.; Emslie, S.D. (2005).
306:Colorado Museum of Natural History
14:
373:When first described by Wetmore,
327:, thus dating the fossils to the
2297:Paleogene birds of North America
958:
115:
2312:Taxa named by Alexander Wetmore
1:
2307:Fossil taxa described in 1927
282:which is possibly the oldest
2333:
496:The Fossil Record of Birds
293:History and classification
1561:
984:
971:
956:
638:
625:
587:
225:
218:
112:Scientific classification
110:
23:
582:and their extinct allies
2317:Prehistoric bird genera
16:Extinct genus of birds
2259:Paleobiology Database
493:Olson, S.L. (1985).
419:Wetmore, A. (1927).
2209:Phasmagyps patritus
279:Phasmagyps patritus
232:Phasmagyps patritus
2292:New World vultures
521:Ducey, J. (1992).
2274:
2273:
2246:Open Tree of Life
2201:Taxon identifiers
2192:
2191:
2188:
2187:
2184:
2183:
2180:
2179:
1734:
1733:
1628:Cryptoleucopteryx
1055:Amphiserpentarius
967:
966:
954:
953:
930:
929:
333:Alexander Wetmore
325:Chadron Formation
310:Boulder, Colorado
268:, known from one
262:New World vulture
246:
245:
199:
2324:
2267:
2266:
2254:
2253:
2241:
2240:
2228:
2227:
2226:
2196:
1929:
1919:
1902:
1892:
1868:
1858:
1577:
1567:
1559:
1534:Archaehieraxinae
1532:
1430:Erythrotriorchis
1365:
1355:
1325:
1315:
1305:
1295:
1285:
1275:
1263:
1253:
1243:
1233:
1223:
1213:
1203:
1193:
1183:
1173:
1163:
1153:
1141:
1131:
1124:Apatosagittarius
1121:
1111:
1101:
1088:
1062:
1052:
1042:
991:
986:
973:
962:
918:
894:
884:
874:
864:
854:
844:
834:
817:
807:
797:
780:
763:
753:
743:
677:
665:Parasarcoramphus
662:
649:
640:
632:Accipitrimorphae
627:
617:Accipitrimorphae
568:
561:
554:
545:
538:
537:
527:
518:
512:
511:
509:
507:
501:
490:
481:
480:
454:
445:
436:
435:
425:
416:
383:Coragyps atratus
337:type description
329:Lower Chadronian
272:fossil found in
234:
230:
211:P. patritus
197:
190:
120:
119:
105:
35:
29:Temporal range:
21:
2332:
2331:
2327:
2326:
2325:
2323:
2322:
2321:
2302:Oligocene birds
2277:
2276:
2275:
2270:
2262:
2257:
2249:
2244:
2236:
2231:
2222:
2221:
2216:
2203:
2193:
2176:
2100:
2066:
2046:
2005:
1971:
1944:
1843:
1809:
1789:
1730:
1703:
1585:
1548:
1525:
1442:
1408:
1333:
1077:
1027:
1007:
993:Horusornithidae
980:
978:Accipitriformes
963:
950:
936:Accipitriformes
926:
728:
679:Teratornithidae
670:
634:
621:
583:
572:
542:
541:
525:
520:
519:
515:
505:
503:
499:
492:
491:
484:
452:
447:
446:
439:
423:
418:
417:
408:
403:
371:
359:Storrs L. Olson
295:
242:
236:
228:
227:
214:
196:
188:
114:
106:
104:
103:
102:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
31:
30:
27:
17:
12:
11:
5:
2330:
2328:
2320:
2319:
2314:
2309:
2304:
2299:
2294:
2289:
2279:
2278:
2272:
2271:
2269:
2268:
2255:
2242:
2229:
2213:
2211:
2205:
2204:
2199:
2190:
2189:
2186:
2185:
2182:
2181:
2178:
2177:
2175:
2174:
2167:
2160:
2153:
2146:
2139:
2132:
2125:
2118:
2110:
2108:
2102:
2101:
2099:
2098:
2091:
2084:
2076:
2074:
2068:
2067:
2065:
2064:
2056:
2054:
2048:
2047:
2045:
2044:
2037:
2030:
2023:
2015:
2013:
2007:
2006:
2004:
2003:
1996:
1989:
1981:
1979:
1973:
1972:
1970:
1969:
1962:
1954:
1952:
1946:
1945:
1943:
1942:
1935:
1925:
1915:
1908:
1898:
1888:
1881:
1874:
1864:
1853:
1851:
1845:
1844:
1842:
1841:
1834:
1827:
1819:
1817:
1811:
1810:
1808:
1807:
1799:
1797:
1791:
1790:
1788:
1787:
1780:
1773:
1766:
1759:
1752:
1744:
1742:
1736:
1735:
1732:
1731:
1729:
1728:
1721:
1713:
1711:
1705:
1704:
1702:
1701:
1694:
1687:
1680:
1673:
1666:
1659:
1652:
1645:
1638:
1631:
1624:
1617:
1610:
1603:
1595:
1593:
1587:
1586:
1584:
1583:
1573:
1562:
1556:
1550:
1549:
1547:
1546:
1538:
1536:
1527:
1526:
1524:
1523:
1516:
1509:
1502:
1495:
1492:Lophotriorchis
1488:
1481:
1474:
1467:
1460:
1452:
1450:
1444:
1443:
1441:
1440:
1433:
1426:
1418:
1416:
1410:
1409:
1407:
1406:
1399:
1392:
1385:
1378:
1371:
1361:
1351:
1343:
1341:
1335:
1334:
1332:
1331:
1321:
1311:
1301:
1291:
1281:
1271:
1259:
1249:
1239:
1229:
1219:
1209:
1199:
1189:
1179:
1169:
1159:
1149:
1137:
1127:
1117:
1107:
1096:
1094:
1092:incertae sedis
1085:
1079:
1078:
1076:
1075:
1068:
1058:
1048:
1037:
1035:
1029:
1028:
1026:
1025:
1017:
1015:
1009:
1008:
1006:
1005:
997:
995:
982:
981:
976:
969:
968:
965:
964:
957:
955:
952:
951:
949:
948:
947:
946:
940:
938:
932:
931:
928:
927:
925:
924:
914:
907:
900:
890:
880:
870:
860:
850:
840:
830:
823:
813:
803:
793:
786:
776:
769:
759:
749:
738:
736:
730:
729:
727:
726:
719:
712:
705:
698:
691:
683:
681:
672:
671:
669:
668:
657:
655:
653:incertae sedis
646:
644:Cathartiformes
636:
635:
630:
623:
622:
620:
619:
613:
607:
601:
595:
588:
585:
584:
573:
571:
570:
563:
556:
548:
540:
539:
513:
482:
437:
405:
404:
402:
399:
370:
367:
294:
291:
264:in the family
244:
243:
237:
223:
222:
216:
215:
207:
205:
201:
200:
186:
182:
181:
176:
172:
171:
169:Cathartiformes
166:
162:
161:
156:
152:
151:
146:
142:
141:
136:
132:
131:
126:
122:
121:
108:
107:
99:
98:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
37:
36:
28:
15:
13:
10:
9:
6:
4:
3:
2:
2329:
2318:
2315:
2313:
2310:
2308:
2305:
2303:
2300:
2298:
2295:
2293:
2290:
2288:
2285:
2284:
2282:
2265:
2260:
2256:
2252:
2247:
2243:
2239:
2234:
2230:
2225:
2219:
2215:
2214:
2212:
2210:
2206:
2202:
2197:
2173:
2172:
2168:
2166:
2165:
2161:
2159:
2158:
2154:
2152:
2151:
2147:
2145:
2144:
2140:
2138:
2137:
2133:
2131:
2130:
2126:
2124:
2123:
2122:Chondrohierax
2119:
2117:
2116:
2112:
2111:
2109:
2107:
2103:
2097:
2096:
2092:
2090:
2089:
2085:
2083:
2082:
2078:
2077:
2075:
2073:
2069:
2063:
2062:
2058:
2057:
2055:
2053:
2052:Lophospizinae
2049:
2043:
2042:
2038:
2036:
2035:
2034:Macheiramphus
2031:
2029:
2028:
2024:
2022:
2021:
2017:
2016:
2014:
2012:
2008:
2002:
2001:
1997:
1995:
1994:
1990:
1988:
1987:
1983:
1982:
1980:
1978:
1974:
1968:
1967:
1963:
1961:
1960:
1956:
1955:
1953:
1951:
1947:
1941:
1940:
1936:
1934:
1933:
1926:
1924:
1923:
1916:
1914:
1913:
1909:
1907:
1906:
1899:
1897:
1896:
1889:
1887:
1886:
1882:
1880:
1879:
1875:
1873:
1872:
1865:
1863:
1862:
1855:
1854:
1852:
1850:
1846:
1840:
1839:
1835:
1833:
1832:
1828:
1826:
1825:
1821:
1820:
1818:
1816:
1812:
1806:
1805:
1801:
1800:
1798:
1796:
1792:
1786:
1785:
1781:
1779:
1778:
1774:
1772:
1771:
1767:
1765:
1764:
1760:
1758:
1757:
1756:Dryotriorchis
1753:
1751:
1750:
1746:
1745:
1743:
1741:
1737:
1727:
1726:
1722:
1720:
1719:
1715:
1714:
1712:
1710:
1706:
1700:
1699:
1695:
1693:
1692:
1688:
1686:
1685:
1681:
1679:
1678:
1674:
1672:
1671:
1667:
1665:
1664:
1660:
1658:
1657:
1653:
1651:
1650:
1646:
1644:
1643:
1639:
1637:
1636:
1632:
1630:
1629:
1625:
1623:
1622:
1618:
1616:
1615:
1611:
1609:
1608:
1604:
1602:
1601:
1597:
1596:
1594:
1592:
1588:
1582:
1581:
1574:
1572:
1571:
1564:
1563:
1560:
1557:
1555:
1551:
1545:
1544:
1540:
1539:
1537:
1535:
1528:
1522:
1521:
1520:Stephanoaetus
1517:
1515:
1514:
1510:
1508:
1507:
1503:
1501:
1500:
1496:
1494:
1493:
1489:
1487:
1486:
1482:
1480:
1479:
1475:
1473:
1472:
1468:
1466:
1465:
1461:
1459:
1458:
1454:
1453:
1451:
1449:
1445:
1439:
1438:
1437:Megatriorchis
1434:
1432:
1431:
1427:
1425:
1424:
1420:
1419:
1417:
1415:
1411:
1405:
1404:
1400:
1398:
1397:
1393:
1391:
1390:
1386:
1384:
1383:
1379:
1377:
1376:
1372:
1370:
1369:
1362:
1360:
1359:
1352:
1350:
1349:
1345:
1344:
1342:
1340:
1336:
1330:
1329:
1322:
1320:
1319:
1312:
1310:
1309:
1302:
1300:
1299:
1292:
1290:
1289:
1282:
1280:
1279:
1272:
1269:
1268:
1260:
1258:
1257:
1256:Palaeoplancus
1250:
1248:
1247:
1240:
1238:
1237:
1230:
1228:
1227:
1220:
1218:
1217:
1210:
1208:
1207:
1200:
1198:
1197:
1190:
1188:
1187:
1180:
1178:
1177:
1170:
1168:
1167:
1166:Gigantohierax
1160:
1158:
1157:
1150:
1147:
1146:
1138:
1136:
1135:
1128:
1126:
1125:
1118:
1116:
1115:
1108:
1106:
1105:
1098:
1097:
1095:
1093:
1089:
1086:
1084:
1080:
1074:
1073:
1069:
1067:
1066:
1065:Pelargopappus
1059:
1057:
1056:
1049:
1047:
1046:
1039:
1038:
1036:
1034:
1033:Sagittariidae
1030:
1024:
1023:
1019:
1018:
1016:
1014:
1010:
1004:
1003:
999:
998:
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983:
979:
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961:
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933:
923:
922:
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887:Pleistovultur
881:
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839:
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783:Cathartidarum
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748:
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395:outer condyle
392:
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379:Black vulture
376:
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220:Binomial name
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198:Wetmore, 1927
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60:
55:
50:
45:
40:
34:
26:
22:
19:
2208:
2169:
2164:Lophoictinia
2162:
2155:
2150:Henicopernis
2148:
2141:
2134:
2127:
2120:
2113:
2095:Urotriorchis
2093:
2086:
2079:
2072:Melieraxinae
2059:
2039:
2032:
2025:
2018:
2000:Microspizias
1998:
1991:
1984:
1964:
1957:
1950:Haliaeetinae
1939:Polyboroides
1937:
1930:
1922:Neophrontops
1920:
1910:
1903:
1893:
1883:
1876:
1869:
1859:
1836:
1829:
1822:
1802:
1782:
1775:
1770:Pithecophaga
1768:
1761:
1754:
1747:
1723:
1716:
1696:
1689:
1682:
1675:
1670:Morphnarchus
1668:
1663:Leucopternis
1661:
1654:
1649:Helicolestes
1647:
1640:
1633:
1626:
1619:
1612:
1605:
1598:
1580:Garganoaetus
1578:
1568:
1543:Archaehierax
1541:
1518:
1511:
1504:
1497:
1490:
1483:
1476:
1469:
1462:
1455:
1435:
1428:
1421:
1414:Accipitrinae
1401:
1394:
1387:
1380:
1373:
1366:
1356:
1346:
1326:
1318:Titanohierax
1316:
1306:
1296:
1286:
1276:
1265:
1254:
1246:Palaeocircus
1244:
1234:
1226:Palaeohierax
1224:
1214:
1204:
1194:
1184:
1174:
1164:
1154:
1143:
1132:
1122:
1112:
1102:
1091:
1083:Accipitridae
1070:
1063:
1053:
1043:
1020:
1000:
919:
909:
904:Sarcoramphus
902:
895:
885:
875:
866:
865:
855:
845:
835:
825:
818:
808:
800:Diatropornis
798:
788:
781:
771:
764:
754:
744:
721:
714:
707:
702:Cathartornis
700:
693:
686:
663:
652:
533:
529:
516:
504:. Retrieved
495:
469:10.1650/7475
460:
456:
431:
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386:
382:
374:
372:
362:
352:
348:
345:etymological
340:
321:Pierre Shale
316:
297:
296:
278:
277:
249:
248:
247:
231:
226:
210:
209:
192:
191:
24:
18:
2287:Cathartidae
2136:Eutriorchis
1966:Icthyophaga
1932:Palaeoborus
1895:Mioneophron
1871:Arikarornis
1824:Chelictinia
1784:Terathopius
1763:Eutriorchis
1740:Circaetinae
1642:Geranospiza
1635:Geranoaetus
1621:Buteogallus
1403:Trigonoceps
1382:Necrosyrtes
1328:Vinchinavis
1196:Mioaegypius
1156:Dynatoaetus
1145:Cruschedula
1072:Sagittarius
1013:Pandionidae
945:See below β
756:Brasilogyps
734:Cathartidae
716:Taubatornis
463:: 107β113.
391:tibiotarsus
369:Description
335:; his 1927
314:Weld County
266:Cathartidae
179:Cathartidae
2281:Categories
2143:Hamirostra
2088:Micronisus
2061:Lophospiza
2027:Harpyopsis
1993:Kaupifalco
1977:Harpaginae
1959:Haliaeetus
1885:Gypohierax
1849:Gypaetinae
1691:Rostrhamus
1684:Pseudastur
1600:Busarellus
1554:Buteoninae
1506:Polemaetus
1478:Ictinaetus
1471:Hieraaetus
1358:Cryptogyps
1339:Aegypiinae
1288:Proictinia
1236:Palaeastur
1176:Gobihierax
1104:Amplibuteo
1045:Amanuensis
1002:Horusornis
867:Phasmagyps
820:Geronogyps
746:Aizenogyps
723:Teratornis
695:Argentavis
611:Telluraves
574:Genera of
506:3 December
457:The Condor
434:(2): 1β14.
401:References
387:Phasmagyps
375:Phasmagyps
363:Phasmagyps
298:Phasmagyps
250:Phasmagyps
193:Phasmagyps
33:Chadronian
25:Phasmagyps
2129:Elanoides
2011:Harpiinae
1861:Anchigyps
1838:Gampsonyx
1777:Spilornis
1749:Circaetus
1718:Haliastur
1677:Parabuteo
1591:Buteonini
1513:Spizaetus
1485:Lophaetus
1448:Aquilinae
1423:Accipiter
1389:Sarcogyps
1368:Gansugyps
1308:Qiluornis
1216:Necrastur
1206:Miohierax
1186:Milvoides
1134:Aviraptor
1114:Aquilavus
921:Wingegyps
857:Pampagyps
837:Hadrogyps
827:Gymnogyps
773:Cathartes
709:Oscaravis
688:Aiolornis
591:Kingdom:
317:Trigonias
284:New World
270:Oligocene
204:Species:
135:Kingdom:
129:Eukaryota
2224:Q7181027
2218:Wikidata
2157:Leptodon
2106:Perninae
2081:Melierax
2041:Morphnus
1986:Harpagus
1912:Neophron
1878:Gypaetus
1815:Elaninae
1795:Circinae
1698:Rupornis
1607:Butastur
1570:Bermuteo
1499:Nisaetus
1348:Aegypius
1298:Promilio
1267:Palaetus
897:Pliogyps
877:Perugyps
810:Dryornis
790:Coragyps
766:Breagyps
599:Chordata
597:Phylum:
593:Animalia
536:: 83β96.
477:85805971
323:and the
302:holotype
274:Colorado
253:, is an
175:Family:
149:Chordata
145:Phylum:
139:Animalia
125:Domain:
2251:6151127
2238:8564008
2115:Aviceda
1905:Neogyps
1709:Milvini
1656:Ictinia
1278:Pengana
1022:Pandion
615:Clade:
609:Clade:
603:Class:
580:condors
287:vulture
255:extinct
239:Wetmore
229:†
208:†
185:Genus:
165:Order:
155:Class:
100:↓
2264:172901
2171:Pernis
2020:Harpia
1831:Elanus
1804:Circus
1725:Milvus
1464:Clanga
1457:Aquila
1396:Torgos
911:Vultur
847:Kuntur
576:eagles
475:
349:phasma
241:, 1927
1614:Buteo
526:(PDF)
500:(PDF)
473:S2CID
453:(PDF)
424:(PDF)
258:genus
2233:GBIF
1375:Gyps
605:Aves
508:2012
354:Gyps
159:Aves
39:Preκ
534:XIX
465:doi
461:107
308:in
260:of
2283::
2261::
2248::
2235::
2220::
1148:"?
578:,
532:.
528:.
485:^
471:.
459:.
455:.
440:^
430:.
426:.
409:^
381:,
89:Pg
1928:β
1918:β
1901:β
1891:β
1867:β
1857:β
1576:β
1566:β
1531:β
1364:β
1354:β
1324:β
1314:β
1304:β
1294:β
1284:β
1274:β
1270:"
1264:"
1262:β
1252:β
1242:β
1232:β
1222:β
1212:β
1202:β
1192:β
1182:β
1172:β
1162:β
1152:β
1142:"
1140:β
1130:β
1120:β
1110:β
1100:β
1061:β
1051:β
1041:β
990:β
917:β
893:β
883:β
873:β
863:β
853:β
843:β
833:β
816:β
806:β
796:β
779:β
762:β
752:β
742:β
676:β
661:β
567:e
560:t
553:v
510:.
479:.
467::
432:7
189:β
94:N
84:K
79:J
74:T
69:P
64:C
59:D
54:S
49:O
44:κ
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