2254:
2265:
2272:
3666:
120:
4398:
5633:
5627:
103:
3671:
405:. Known material includes the holotype, MLL Pv-005, a posterior dorsal vertebra; the paratype MLL-Pv-002, a posterior cervical vertebra; MLL-Pv-010, a partial skeleton including cervical vertebrae, dorsal vertebrae, ribs, a caudal vertebra, and incomplete scapulae; and other isolated vertebrae. Other dicraeosaurid fossils have been found in the Mulichinco Formation, including a femur and a natural cranial
3128:
5622:
3120:
5616:
4390:
2260:
3140:
3134:
4406:
785:
445:, and unlike other dicraeosaurids in which forking of the neural spines only extended into the middle dorsal vertebrae. The posterior dorsal vertebrae show an autapomorphic pair of deep fossae near the bases of the neural spines. The scapula has a ridge on the medial surface near the acromion, which is not present in other dicraeosaurids.
441:
foramen is present on the proximal end of the cervical ribs, which probably is a nutrient foramen and not a pneumatic structure. The anterior dorsal vertebrae show an autapomorphic ridge on the anterior centrodiapophyseal lamina. Even the posterior dorsal vertebrae had forked neural spines, as in its close relative
440:
contain pneumatic chambers. However, as in other dicraeosaurids, the dorsal centra lack the pneumatic foramina that are characteristic of most sauropods. The cervical centra have ventral keels that are forked anteriorly and posteriorly, unlike the simple midline keels typical of other diplodocoids. A
964:
Windholz, Guillermo J; Coria, Rodolfo A.; Bellardini, Flavio; Baiano, Mattia A.; Pino, Diego; Ortega, Francisco; Currie, Philip J. (2022-12-21). "On a dicraeosaurid specimen from the
Mulichinco Formation (Valanginian, Neuquén Basin) of Argentina and phylogenetic relationships of the South American
872:
Coria, Rodolfo A.; Currie, Philip J.; Ortega, Francisco; Baiano, Mattia A. (2019-11-25). "An Early
Cretaceous, medium-sized carcharodontosaurid theropod (Dinosauria, Saurischia) from the Mulichinco Formation (upper Valanginian), Neuquén Province, Patagonia, Argentina".
731:. Windholz et al. argued that, while phylogenetic analyses do not currently provide clear resolution for its affinities, on chronological and biogeographic grounds it is most likely to belong to a clade uniting the South American dicraeosaurids.
1041:
Gallina, Pablo A.; ApesteguĂa, Sebastián; Carballido, JosĂ© L.; Garderes, Juan P. (2022). "Southernmost spiny backs and whiplash tails: flagellicaudatans from South
America". In Otero, Alejandro; Carballido, José L.; Pol, Diego (eds.).
1111:
Windholz, Guillermo J.; Carballido, José L.; Coria, Rodolfo A.; Zurriaguz, Virginia L.; Rauhut, Oliver W. M. (2022-11-22). "How pneumatic were the presacral vertebrae of dicraeosaurid (Sauropoda: Diplodocoidea) dinosaurs?".
742:
lived in what is now
Argentina during the Valanginian age of the Cretaceous, and is a member of the Bajadan faunal assemblage. It coexisted with an indeterminate species of diplodocid and the small, early carcharodontosaur
1067:
Windholz, Guillermo J.; Coria, Rodolfo A.; Zurriaguz, Virginia L. (2019-11-29). "Vertebral pneumatic structures in the Early
Cretaceous sauropod dinosaur Pilmatueia faundezi from northwestern Patagonia, Argentina".
992:
Paulina
Carabajal, A.; Coria, R.A.; Currie, P.J.; Koppelhus, E.B. (2017-12-12). "A natural cranial endocast with possible dicraeosaurid (Sauropoda, Diplodocoidea) affinities from the Lower Cretaceous of Patagonia".
370:, at a site called Pilmatué. Fossil excavations at Pilmatué began in 2009, and the discovery of dicraeosaurid remains at Pilmatué was first announced in 2012, at a paleontology conference in Buenos Aires. In 2019,
825:
Rodolfo A. Coria; Guillermo J. Windholz; Francisco Ortega; Philip J. Currie (2018). "A new dicraeosaurid sauropod from the Lower
Cretaceous (Mulichinco Formation, Valanginian, Neuquén Basin) of Argentina".
705:
within
Dicraeosauridae, following an analysis by Windholtz et al. that recovered their favored hypothesis of South American dicraeosaurid monophyly, with South American dicraeosaurids highlighted in red.
378:. The genus name refers to the Pilmatué locality, and the species epithet recognizes Ramón Faúndez, manager of the Museo Municipal de Las Lajas, who supported the excavation project. The discovery of
382:
in the
Valanginian, an age of the Early Cretaceous with poorly known dinosaur faunas, helped fill a gap in the dicraeosaurid fossil record between Jurassic dicraeosaurids and the later
1222:
Whitlock, John A.; Wilson
Mantilla, Jeffrey A. (2020-12-23). "The Late Jurassic sauropod dinosaur 'Morosaurus' agilis Marsh, 1889 reexamined and reinterpreted as a dicraeosaurid".
5699:
926:
Coria, R. A.; Ortega, F.; Succar, C.; Currie, P.; Koppelhus, E. (2012). "First record of a dicraeosaurid sauropod from the Lower Cretaceous (Valanginian) of Neuquén Basin".
424:, like other dicraeosaurids, had tall, deeply forked neural spines on its cervical and dorsal vertebrae. The cervical neural spines were low compared to those of
1356:
715:
is a dicraeosaurid sauropod. Its phylogenetic position within Dicraeosauridae is uncertain, with some analyses finding it to be closely related to
5719:
1051:
342:
vertebrae compared to other dicraeosaurids, which were otherwise characterized by a reduction in pneumaticity relative to other sauropods.
5724:
1283:"First Lower Cretaceous record of Podocarpaceae wood associated with dinosaur remains from Patagonia, Neuquén Province, Argentina"
2264:
2253:
2271:
436:. Unlike most dicraeosaurids, which have reduced vertebral pneumaticity compared to other sauropods, the cervical vertebrae of
3665:
5709:
5704:
1349:
4397:
119:
769:
The article was released online on 29 August 2018, but formally published in the January 2019 issue of the journal.
5632:
5626:
5714:
432:. As in most dicraeosaurids, the neural spines were inclined forward, unlike the backswept neural spines seen in
5669:
3670:
1444:
1342:
102:
1281:
Gnaedinger, Silvia; Coria, Rodolfo A.; Koppelhus, Eva; CasadĂo, Silvio; Tunik, Maisa; Currie, Philip (2017).
5694:
3127:
3139:
3119:
5659:
5621:
5350:
1632:
339:
5615:
4389:
3133:
4405:
2259:
1294:
1231:
1160:
835:
398:
262:
2335:
363:
5390:
2462:
1282:
1263:
1247:
1204:
1093:
908:
851:
114:
5526:
5067:
3814:
3455:
3184:
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1320:
1255:
1196:
1178:
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1047:
1020:
974:
900:
547:
5397:
5242:
5182:
4824:
4716:
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4295:
4234:
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1310:
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1002:
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882:
843:
790:
315:
30:
1147:
Gallina, Pablo A.; ApesteguĂa, Sebastián; Canale, Juan I.; Haluza, Alejandro (2019-02-04).
5664:
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2071:
2064:
2056:
2043:
1960:
1506:
1493:
1456:
1429:
1404:
1365:
719:
and other analyses finding it to be a more basal dicraeosaurid, outside the clade uniting
461:
311:
235:
193:
1298:
1235:
1164:
839:
5541:
5456:
5338:
5283:
5233:
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2806:
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2592:
2499:
2398:
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2194:
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2003:
1974:
1967:
1913:
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1602:
1191:
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2159:
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2100:
1996:
1933:
1924:
1806:
1680:
1648:
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1559:
1538:
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912:
855:
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371:
255:
222:
65:
1208:
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5359:
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5218:
5204:
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5167:
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5085:
5074:
5025:
5006:
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4877:
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4803:
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4738:
4674:
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4455:
4437:
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4352:
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4220:
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4078:
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4014:
3962:
3948:
3934:
3919:
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3763:
3756:
3575:
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3541:
3523:
3448:
3434:
3341:
3298:
3291:
3233:
3226:
2989:
2904:
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2662:
2634:
2627:
2606:
2578:
2564:
2484:
2428:
2407:
2093:
1981:
1885:
1841:
1834:
1786:
1758:
1719:
1545:
745:
612:
571:
330:
270:
1306:
1243:
1149:"A new long-spined dinosaur from Patagonia sheds light on sauropod defense system"
1006:
886:
847:
1125:
5577:
5485:
5471:
5367:
5256:
5211:
5108:
5101:
5094:
5060:
4992:
4970:
4891:
4782:
4477:
4284:
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4107:
4049:
4028:
4021:
3993:
3926:
3898:
3863:
3856:
3842:
3741:
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3627:
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3095:
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2934:
2927:
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2762:
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2641:
2585:
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2321:
2187:
2145:
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2111:
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2031:
2024:
1940:
1906:
1892:
1735:
1673:
1609:
1586:
1552:
1524:
347:
304:
40:
1173:
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5405:
5374:
5136:
4963:
4810:
4745:
4709:
4695:
4630:
4601:
4564:
4527:
4470:
4056:
4000:
3955:
3905:
3877:
3835:
3771:
3719:
3701:
3688:
3612:
3605:
3598:
3584:
3484:
3477:
3397:
3309:
3284:
3014:
2964:
2948:
2912:
2813:
2689:
2613:
2527:
2470:
2421:
2173:
1989:
1878:
1742:
1701:
1666:
1395:
780:
499:
350:, an age of the Cretaceous period for which dinosaur faunas are poorly known.
177:
85:
50:
1324:
1259:
1251:
1182:
1133:
1089:
1024:
978:
904:
5441:
5323:
5129:
4608:
4586:
4535:
4256:
4213:
4199:
3941:
3634:
3620:
3505:
3219:
3067:
3060:
2791:
2740:
2303:
2290:
2205:
2180:
2138:
2017:
1857:
1624:
1566:
475:
402:
367:
328:
was probably closely related to other South American dicraeosaurids such as
209:
131:
90:
1200:
5449:
5115:
4925:
4128:
3969:
3828:
3081:
2571:
2414:
2124:
1899:
1871:
1779:
1694:
1687:
1594:
1531:
1413:
1389:
406:
307:
164:
151:
80:
75:
60:
55:
45:
895:
4485:
2889:
2733:
2620:
2078:
1315:
1015:
95:
70:
1081:
101:
5014:
4192:
2010:
1383:
141:
1334:
5316:
1820:
5648:
4422:
3686:
3155:
2288:
1491:
1376:
1338:
2978:
1046:. Cham: Springer International Publishing. pp. 209–236.
16:
Sauropod dinosaur genus from the early Cretaceous Period
277:
5518:
5423:
5349:
5302:
5266:
5232:
5150:
5084:
5024:
4942:
4909:
4835:
4435:
4351:
4294:
4166:
3979:
3888:
3812:
3781:
3699:
3574:
3531:
3522:
3424:
3415:
3352:
3317:
3308:
3205:
3194:
3168:
3045:
3013:
2997:
2988:
2879:
2723:
2672:
2541:
2510:
2442:
2397:
2301:
2204:
2110:
2042:
1950:
1923:
1855:
1796:
1756:
1718:
1646:
1576:
1504:
1455:
1428:
959:
957:
955:
953:
951:
949:
947:
945:
943:
941:
374:and colleagues named the new genus and species
409:, and it is possible that they also belong to
1350:
1036:
1034:
8:
965:dicraeosaurids (Sauropoda, Diplodocoidea)".
5700:Early Cretaceous dinosaurs of South America
867:
865:
820:
818:
816:
814:
812:
810:
808:
806:
804:
802:
5645:
5515:
5308:
5263:
5030:
4948:
4917:
4841:
4447:
4443:
4432:
4419:
4300:
3985:
3820:
3711:
3707:
3696:
3683:
3528:
3421:
3314:
3211:
3202:
3176:
3165:
3152:
3042:
2994:
2549:
2454:
2450:
2313:
2309:
2298:
2285:
2116:
2048:
1863:
1793:
1764:
1658:
1654:
1516:
1512:
1501:
1488:
1434:
1425:
1373:
1357:
1343:
1335:
20:
1314:
1190:
1172:
1114:Biological Journal of the Linnean Society
1014:
894:
1044:South American Sauropodomorph Dinosaurs
798:
762:
7:
587:
563:
539:
515:
491:
467:
457:
314:that lived in Argentina during the
1224:Journal of Vertebrate Paleontology
14:
5650:Topics in sauropodomorph research
5631:
5625:
5620:
5614:
4404:
4396:
4388:
3669:
3664:
3138:
3132:
3126:
3118:
2270:
2263:
2258:
2252:
783:
118:
753:were present in the ecosystem.
397:fossils have been found in the
318:. Its type and only species is
1:
5720:Fossil taxa described in 2019
1307:10.1016/j.cretres.2017.06.014
1244:10.1080/02724634.2020.1780600
1007:10.1016/j.cretres.2017.12.001
887:10.1016/j.cretres.2019.104319
848:10.1016/j.cretres.2018.08.019
5741:
5725:Early Cretaceous sauropods
1174:10.1038/s41598-018-37943-3
1126:10.1093/biolinnean/blac131
5655:
5644:
5612:
5514:
5311:
5033:
4951:
4920:
4844:
4450:
4446:
4431:
4418:
4385:
4303:
3988:
3823:
3714:
3710:
3695:
3682:
3662:
3214:
3179:
3164:
3151:
3115:
3041:
2552:
2457:
2453:
2316:
2312:
2297:
2284:
2250:
2119:
2051:
1866:
1767:
1661:
1657:
1519:
1515:
1500:
1487:
1437:
1424:
1372:
701:Phylogenetic position of
626:
609:
592:
585:
568:
561:
544:
537:
520:
513:
496:
489:
472:
465:
276:
269:
115:Scientific classification
113:
23:
5670:List of sauropod species
310:belonging to the family
967:Comptes Rendus Palevol
107:
5660:Sauropod neck posture
5351:Opisthocoelicaudiinae
105:
5710:Fossils of Argentina
5705:Cretaceous Argentina
399:Mulichinco Formation
354:Discovery and naming
5519:Dubious titanosaurs
2336:Chinshakiangosaurus
1299:2017CrRes..78..228G
1287:Cretaceous Research
1236:2020JVPal..40E0600W
1165:2019NatSR...9.1392G
995:Cretaceous Research
875:Cretaceous Research
840:2019CrRes..93...33C
828:Cretaceous Research
411:Pilmatueia faundezi
395:Pilmatueia faundezi
376:Pilmatueia faundezi
362:were discovered in
360:Pilmatueia faundezi
321:Pilmatueia faundezi
282:Pilmatueia faundezi
106:Pilmatueia faundezi
5391:Opisthocoelicaudia
2463:Archaeodontosaurus
1153:Scientific Reports
108:
5682:
5681:
5678:
5677:
5640:
5639:
5610:
5609:
5606:
5605:
5602:
5601:
5527:Bruhathkayosaurus
5510:
5509:
5506:
5505:
5502:
5501:
5228:
5227:
5146:
5145:
5068:Panamericansaurus
4905:
4904:
4901:
4900:
4414:
4413:
4383:
4382:
4379:
4378:
4375:
4374:
4371:
4370:
4347:
4346:
3815:Titanosauriformes
3808:
3807:
3678:
3677:
3660:
3659:
3656:
3655:
3652:
3651:
3648:
3647:
3644:
3643:
3456:Brachytrachelopan
3411:
3410:
3407:
3406:
3185:Haplocanthosaurus
3147:
3146:
3113:
3112:
3109:
3108:
3105:
3104:
3046:Dubious sauropods
3037:
3036:
3033:
3032:
3029:
3028:
2537:
2536:
2438:
2437:
2373:Protognathosaurus
2280:
2279:
2248:
2247:
2244:
2243:
2240:
2239:
2236:
2235:
2232:
2231:
2228:
2227:
2224:
2223:
1851:
1850:
1752:
1751:
1642:
1641:
1483:
1482:
1479:
1478:
1475:
1474:
1082:10.1111/let.12363
1053:978-3-030-95958-6
725:Brachytrachelopan
710:
709:
695:
694:
686:
685:
677:
676:
668:
667:
659:
658:
650:
649:
641:
640:
548:Brachytrachelopan
296:
295:
265:
5732:
5646:
5635:
5629:
5624:
5618:
5516:
5398:Pellegrinisaurus
5309:
5264:
5243:Bustingorrytitan
5183:Futalognkosaurus
5031:
4949:
4918:
4842:
4825:Diamantinasauria
4717:Pellegrinisaurus
4623:Jiangshanosaurus
4514:Baotianmansaurus
4448:
4444:
4433:
4420:
4409:
4408:
4401:
4400:
4393:
4392:
4316:Diamantinasaurus
4301:
4296:Diamantinasauria
4235:Jiangshanosaurus
3986:
3821:
3712:
3708:
3697:
3684:
3673:
3668:
3529:
3422:
3354:Rebbachisaurinae
3315:
3212:
3207:Rebbachisauridae
3203:
3177:
3166:
3153:
3142:
3136:
3131:
3130:
3123:
3122:
3054:Bothriospondylus
3043:
2995:
2771:Eomamenchisaurus
2725:Mamenchisauridae
2705:Lapparentosaurus
2649:Spinophorosaurus
2550:
2455:
2451:
2314:
2310:
2299:
2286:
2275:
2274:
2267:
2262:
2256:
2117:
2049:
1952:Massospondylidae
1864:
1794:
1765:
1729:Jaklapallisaurus
1709:Thecodontosaurus
1659:
1655:
1517:
1513:
1502:
1489:
1435:
1426:
1419:
1418:
1374:
1359:
1352:
1345:
1336:
1329:
1328:
1318:
1278:
1272:
1271:
1219:
1213:
1212:
1194:
1176:
1144:
1138:
1137:
1108:
1102:
1101:
1064:
1058:
1057:
1038:
1029:
1028:
1018:
989:
983:
982:
973:(45): 991–1019.
961:
936:
935:
923:
917:
916:
898:
869:
860:
859:
822:
793:
791:Dinosaurs portal
788:
787:
786:
770:
767:
588:
564:
540:
516:
492:
468:
458:
453:
452:
390:Fossil specimens
364:Neuquén Province
316:Early Cretaceous
279:
254:
247:
234:
221:
208:
192:
123:
122:
100:
37:
31:Early Cretaceous
29:Temporal range:
21:
5740:
5739:
5735:
5734:
5733:
5731:
5730:
5729:
5685:
5684:
5683:
5674:
5665:Sauropod hiatus
5651:
5636:
5598:
5535:Campylodoniscus
5498:
5419:
5345:
5298:
5268:Nemegtosauridae
5262:
5224:
5161:Argentinosaurus
5142:
5080:
5020:
4938:
4914:
4897:
4857:Atsinganosaurus
4837:Lirainosaurinae
4831:
4818:Xianshanosaurus
4703:Normanniasaurus
4689:Narambuenatitan
4572:Dongyangosaurus
4440:
4427:
4410:
4403:
4395:
4387:
4367:
4343:
4323:Sarmientosaurus
4290:
4243:Phuwiangosaurus
4162:
3975:
3890:Brachiosauridae
3884:
3817:
3804:
3777:
3704:
3691:
3674:
3640:
3570:
3518:
3426:Dicraeosauridae
3417:Flagellicaudata
3403:
3348:
3304:
3278:Nopcsaspondylus
3270:Maraapunisaurus
3199:
3197:Diplodocimorpha
3190:
3173:
3160:
3143:
3125:
3117:
3101:
3025:
3009:
2984:
2957:Tehuelchesaurus
2875:
2835:Tienshanosaurus
2719:
2668:
2656:Tehuelchesaurus
2546:
2533:
2512:Vulcanodontidae
2506:
2447:
2434:
2393:
2343:Gongxianosaurus
2306:
2293:
2276:
2269:
2220:
2200:
2106:
2065:Jingshanosaurus
2057:Chuxiongosaurus
2044:Sauropodiformes
2038:
1961:Adeopapposaurus
1946:
1919:
1860:
1847:
1792:
1761:
1748:
1714:
1651:
1638:
1572:
1509:
1507:Sauropodomorpha
1496:
1494:Sauropodomorpha
1471:
1457:Sauropodomorpha
1451:
1445:Avemetatarsalia
1430:Avemetatarsalia
1420:
1405:Avemetatarsalia
1379:
1378:
1368:
1366:Sauropodomorpha
1363:
1333:
1332:
1280:
1279:
1275:
1230:(6): e1780600.
1221:
1220:
1216:
1146:
1145:
1141:
1110:
1109:
1105:
1066:
1065:
1061:
1054:
1040:
1039:
1032:
991:
990:
986:
963:
962:
939:
925:
924:
920:
871:
870:
863:
824:
823:
800:
789:
784:
782:
779:
774:
773:
768:
764:
759:
737:
696:
687:
678:
669:
660:
651:
642:
462:Dicraeosauridae
451:
419:
392:
356:
338:had relatively
312:Dicraeosauridae
292:
285:
253:
245:
236:Dicraeosauridae
232:
219:
206:
194:Sauropodomorpha
190:
117:
109:
99:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
35:
34:
27:
17:
12:
11:
5:
5738:
5736:
5728:
5727:
5722:
5717:
5712:
5707:
5702:
5697:
5695:Dicraeosaurids
5687:
5686:
5680:
5679:
5676:
5675:
5673:
5672:
5667:
5662:
5656:
5653:
5652:
5649:
5642:
5641:
5638:
5637:
5613:
5611:
5608:
5607:
5604:
5603:
5600:
5599:
5597:
5596:
5589:
5581:
5574:
5566:
5559:
5552:
5545:
5542:Clasmodosaurus
5538:
5531:
5522:
5520:
5512:
5511:
5508:
5507:
5504:
5503:
5500:
5499:
5497:
5496:
5489:
5482:
5475:
5468:
5461:
5457:Maxakalisaurus
5453:
5446:
5438:
5429:
5427:
5421:
5420:
5418:
5417:
5409:
5402:
5394:
5387:
5379:
5371:
5364:
5355:
5353:
5347:
5346:
5344:
5343:
5339:Quaesitosaurus
5335:
5328:
5320:
5312:
5306:
5300:
5299:
5297:
5296:
5288:
5284:Quaesitosaurus
5280:
5272:
5270:
5261:
5260:
5253:
5246:
5238:
5236:
5234:Saltasauroidea
5230:
5229:
5226:
5225:
5223:
5222:
5215:
5208:
5201:
5194:
5186:
5179:
5171:
5164:
5156:
5154:
5148:
5147:
5144:
5143:
5141:
5140:
5133:
5126:
5119:
5112:
5105:
5098:
5090:
5088:
5082:
5081:
5079:
5078:
5071:
5064:
5057:
5050:
5046:Maxakalisaurus
5042:
5039:Adamantisaurus
5034:
5028:
5022:
5021:
5019:
5018:
5011:
5003:
4996:
4989:
4985:Epachthosaurus
4981:
4974:
4967:
4960:
4957:Antarctosaurus
4952:
4946:
4940:
4939:
4937:
4936:
4929:
4921:
4915:
4912:Eutitanosauria
4910:
4907:
4906:
4903:
4902:
4899:
4898:
4896:
4895:
4888:
4885:Mansourasaurus
4881:
4874:
4867:
4860:
4853:
4845:
4839:
4833:
4832:
4830:
4829:
4821:
4814:
4807:
4800:
4793:
4786:
4779:
4772:
4764:
4760:Tastavinsaurus
4756:
4749:
4742:
4735:
4728:
4725:Petrobrasaurus
4721:
4713:
4706:
4699:
4692:
4685:
4678:
4671:
4667:Malarguesaurus
4663:
4656:
4648:
4641:
4634:
4627:
4619:
4612:
4605:
4598:
4594:Epachthosaurus
4590:
4583:
4575:
4568:
4561:
4554:
4547:
4540:
4532:
4524:
4517:
4510:
4507:Austroposeidon
4503:
4496:
4489:
4482:
4474:
4467:
4459:
4451:
4441:
4436:
4429:
4428:
4423:
4416:
4415:
4412:
4411:
4386:
4384:
4381:
4380:
4377:
4376:
4373:
4372:
4369:
4368:
4366:
4365:
4364:
4363:
4357:
4355:
4349:
4348:
4345:
4344:
4342:
4341:
4333:
4326:
4319:
4312:
4304:
4298:
4292:
4291:
4289:
4288:
4281:
4274:
4271:Tangvayosaurus
4267:
4260:
4253:
4246:
4239:
4231:
4224:
4217:
4210:
4203:
4196:
4189:
4181:
4172:
4170:
4164:
4163:
4161:
4160:
4153:
4150:Tastavinsaurus
4146:
4139:
4132:
4125:
4122:Pukyongosaurus
4118:
4111:
4104:
4101:Liubangosaurus
4097:
4089:
4086:Liaoningotitan
4082:
4075:
4067:
4060:
4053:
4046:
4039:
4032:
4025:
4018:
4011:
4008:Astrophocaudia
4004:
3997:
3989:
3983:
3981:Somphospondyli
3977:
3976:
3974:
3973:
3966:
3959:
3952:
3945:
3938:
3931:
3923:
3916:
3909:
3902:
3894:
3892:
3886:
3885:
3883:
3882:
3874:
3867:
3860:
3853:
3846:
3839:
3832:
3824:
3818:
3813:
3810:
3809:
3806:
3805:
3803:
3802:
3799:Lourinhasaurus
3795:
3787:
3785:
3783:Camarasauridae
3779:
3778:
3776:
3775:
3768:
3760:
3753:
3749:Dashanpusaurus
3745:
3738:
3730:
3723:
3715:
3705:
3700:
3693:
3692:
3687:
3680:
3679:
3676:
3675:
3663:
3661:
3658:
3657:
3654:
3653:
3650:
3649:
3646:
3645:
3642:
3641:
3639:
3638:
3631:
3624:
3617:
3609:
3602:
3595:
3592:Dinheirosaurus
3588:
3580:
3578:
3572:
3571:
3569:
3568:
3561:
3553:
3546:
3537:
3535:
3526:
3520:
3519:
3517:
3516:
3509:
3502:
3495:
3488:
3481:
3474:
3466:
3459:
3452:
3445:
3438:
3430:
3428:
3419:
3413:
3412:
3409:
3408:
3405:
3404:
3402:
3401:
3394:
3391:Rebbachisaurus
3387:
3380:
3373:
3366:
3358:
3356:
3350:
3349:
3347:
3346:
3338:
3331:
3323:
3321:
3312:
3306:
3305:
3303:
3302:
3295:
3288:
3281:
3274:
3266:
3263:Lavocatisaurus
3259:
3252:
3244:
3237:
3230:
3223:
3215:
3209:
3200:
3195:
3192:
3191:
3189:
3188:
3180:
3174:
3169:
3162:
3161:
3156:
3149:
3148:
3145:
3144:
3116:
3114:
3111:
3110:
3107:
3106:
3103:
3102:
3100:
3099:
3092:
3089:Qinlingosaurus
3085:
3078:
3071:
3064:
3057:
3049:
3047:
3039:
3038:
3035:
3034:
3031:
3030:
3027:
3026:
3024:
3023:
3022:(see below ↓ )
3019:
3017:
3011:
3010:
3008:
3007:
3006:(see below ↓ )
3003:
3001:
2992:
2986:
2985:
2983:
2982:
2975:
2968:
2961:
2953:
2945:
2938:
2931:
2924:
2917:
2909:
2901:
2893:
2885:
2883:
2877:
2876:
2874:
2873:
2866:
2859:
2852:
2845:
2838:
2831:
2828:Rhomaleopakhus
2824:
2817:
2810:
2807:Mamenchisaurus
2803:
2795:
2788:
2781:
2774:
2767:
2759:
2756:Chuanjiesaurus
2752:
2748:Cetiosauriscus
2744:
2737:
2729:
2727:
2721:
2720:
2718:
2717:
2709:
2701:
2693:
2686:
2678:
2676:
2670:
2669:
2667:
2666:
2659:
2652:
2645:
2638:
2631:
2624:
2617:
2610:
2603:
2600:Dashanpusaurus
2596:
2593:Cetiosauriscus
2589:
2582:
2575:
2568:
2561:
2553:
2547:
2542:
2539:
2538:
2535:
2534:
2532:
2531:
2524:
2516:
2514:
2508:
2507:
2505:
2504:
2500:Zizhongosaurus
2496:
2488:
2481:
2474:
2467:
2458:
2448:
2443:
2440:
2439:
2436:
2435:
2433:
2432:
2425:
2418:
2411:
2403:
2401:
2399:Lessemsauridae
2395:
2394:
2392:
2391:
2387:Tuebingosaurus
2383:
2376:
2369:
2361:
2353:
2346:
2339:
2332:
2325:
2317:
2307:
2302:
2295:
2294:
2289:
2282:
2281:
2278:
2277:
2251:
2249:
2246:
2245:
2242:
2241:
2238:
2237:
2234:
2233:
2230:
2229:
2226:
2225:
2222:
2221:
2219:
2218:
2217:
2216:
2210:
2208:
2202:
2201:
2199:
2198:
2195:Sefapanosaurus
2191:
2184:
2177:
2170:
2167:Melanorosaurus
2163:
2156:
2149:
2142:
2135:
2128:
2120:
2114:
2108:
2107:
2105:
2104:
2097:
2090:
2082:
2075:
2068:
2061:
2052:
2046:
2040:
2039:
2037:
2036:
2028:
2021:
2014:
2007:
2004:Massospondylus
2000:
1993:
1986:
1978:
1975:Glacialisaurus
1971:
1968:Coloradisaurus
1964:
1956:
1954:
1948:
1947:
1945:
1944:
1937:
1929:
1927:
1921:
1920:
1918:
1917:
1914:Tuebingosaurus
1910:
1903:
1896:
1889:
1882:
1875:
1867:
1861:
1856:
1853:
1852:
1849:
1848:
1846:
1845:
1838:
1831:
1828:Pachysauriscus
1824:
1817:
1814:Gresslyosaurus
1810:
1802:
1800:
1798:Plateosauridae
1791:
1790:
1783:
1776:
1773:Plateosauravus
1768:
1762:
1757:
1754:
1753:
1750:
1749:
1747:
1746:
1739:
1732:
1724:
1722:
1716:
1715:
1713:
1712:
1705:
1698:
1691:
1684:
1677:
1670:
1662:
1652:
1647:
1644:
1643:
1640:
1639:
1637:
1636:
1629:
1621:
1613:
1606:
1603:Chromogisaurus
1599:
1591:
1582:
1580:
1574:
1573:
1571:
1570:
1563:
1556:
1549:
1542:
1535:
1528:
1520:
1510:
1505:
1498:
1497:
1492:
1485:
1484:
1481:
1480:
1477:
1476:
1473:
1472:
1470:
1469:
1468:
1467:
1461:
1459:
1453:
1452:
1450:
1449:
1448:
1447:
1438:
1432:
1422:
1421:
1417:
1416:
1407:
1398:
1392:
1386:
1377:
1370:
1369:
1364:
1362:
1361:
1354:
1347:
1339:
1331:
1330:
1273:
1214:
1139:
1103:
1076:(3): 369–381.
1059:
1052:
1030:
984:
937:
918:
861:
797:
796:
795:
794:
778:
775:
772:
771:
761:
760:
758:
755:
751:Podocarp trees
736:
733:
708:
707:
698:
697:
693:
692:
689:
688:
684:
683:
680:
679:
675:
674:
671:
670:
666:
665:
662:
661:
657:
656:
653:
652:
648:
647:
644:
643:
639:
638:
635:
634:
625:
622:
621:
618:
617:
608:
605:
604:
601:
600:
591:
586:
584:
581:
580:
577:
576:
567:
562:
560:
557:
556:
553:
552:
543:
538:
536:
533:
532:
529:
528:
519:
514:
512:
509:
508:
505:
504:
495:
490:
488:
485:
484:
481:
480:
471:
466:
464:
456:
450:
449:Classification
447:
418:
415:
391:
388:
355:
352:
294:
293:
286:
274:
273:
267:
266:
243:
239:
238:
230:
226:
225:
217:
213:
212:
204:
197:
196:
188:
181:
180:
175:
168:
167:
162:
155:
154:
149:
145:
144:
139:
135:
134:
129:
125:
124:
111:
110:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
39:
38:
28:
15:
13:
10:
9:
6:
4:
3:
2:
5737:
5726:
5723:
5721:
5718:
5716:
5715:Neuquén Basin
5713:
5711:
5708:
5706:
5703:
5701:
5698:
5696:
5693:
5692:
5690:
5671:
5668:
5666:
5663:
5661:
5658:
5657:
5654:
5647:
5643:
5634:
5630:
5628:
5623:
5617:
5595:
5594:
5590:
5587:
5586:
5585:Mongolosaurus
5582:
5580:
5579:
5575:
5572:
5571:
5570:Macrurosaurus
5567:
5565:
5564:
5560:
5558:
5557:
5553:
5551:
5550:
5549:Hypselosaurus
5546:
5544:
5543:
5539:
5537:
5536:
5532:
5529:
5528:
5524:
5523:
5521:
5517:
5513:
5495:
5494:
5490:
5488:
5487:
5483:
5481:
5480:
5476:
5474:
5473:
5469:
5467:
5466:
5465:Neuquensaurus
5462:
5459:
5458:
5454:
5452:
5451:
5447:
5444:
5443:
5439:
5436:
5435:
5431:
5430:
5428:
5426:
5425:Saltasaurinae
5422:
5415:
5414:
5413:Zhuchengtitan
5410:
5408:
5407:
5403:
5400:
5399:
5395:
5393:
5392:
5388:
5385:
5384:
5383:Dreadnoughtus
5380:
5377:
5376:
5372:
5370:
5369:
5365:
5362:
5361:
5357:
5356:
5354:
5352:
5348:
5341:
5340:
5336:
5334:
5333:
5332:Qingxiusaurus
5329:
5326:
5325:
5321:
5319:
5318:
5314:
5313:
5310:
5307:
5305:
5304:Saltasauridae
5301:
5294:
5293:
5289:
5286:
5285:
5281:
5279:
5278:
5277:Nemegtosaurus
5274:
5273:
5271:
5269:
5265:
5259:
5258:
5254:
5252:
5251:
5247:
5245:
5244:
5240:
5239:
5237:
5235:
5231:
5221:
5220:
5216:
5214:
5213:
5209:
5207:
5206:
5202:
5200:
5199:
5198:Mendozasaurus
5195:
5192:
5191:
5187:
5185:
5184:
5180:
5177:
5176:
5175:Dzharatitanis
5172:
5170:
5169:
5165:
5163:
5162:
5158:
5157:
5155:
5153:
5149:
5139:
5138:
5134:
5132:
5131:
5127:
5125:
5124:
5123:Gondwanatitan
5120:
5118:
5117:
5113:
5111:
5110:
5106:
5104:
5103:
5099:
5097:
5096:
5092:
5091:
5089:
5087:
5083:
5077:
5076:
5072:
5070:
5069:
5065:
5063:
5062:
5058:
5056:
5055:
5054:Muyelensaurus
5051:
5048:
5047:
5043:
5041:
5040:
5036:
5035:
5032:
5029:
5027:
5023:
5017:
5016:
5012:
5009:
5008:
5004:
5002:
5001:
4997:
4995:
4994:
4990:
4987:
4986:
4982:
4980:
4979:
4978:Chucarosaurus
4975:
4973:
4972:
4968:
4966:
4965:
4961:
4959:
4958:
4954:
4953:
4950:
4947:
4945:
4944:Colossosauria
4941:
4935:
4934:
4933:Menucocelsior
4930:
4928:
4927:
4923:
4922:
4919:
4916:
4913:
4908:
4894:
4893:
4889:
4887:
4886:
4882:
4880:
4879:
4875:
4873:
4872:
4871:Lirainosaurus
4868:
4866:
4865:
4861:
4859:
4858:
4854:
4852:
4851:
4847:
4846:
4843:
4840:
4838:
4834:
4827:
4826:
4822:
4820:
4819:
4815:
4813:
4812:
4808:
4806:
4805:
4801:
4799:
4798:
4794:
4792:
4791:
4790:Trigonosaurus
4787:
4785:
4784:
4780:
4778:
4777:
4773:
4770:
4769:
4765:
4762:
4761:
4757:
4755:
4754:
4753:Ruyangosaurus
4750:
4748:
4747:
4743:
4741:
4740:
4736:
4734:
4733:
4732:Pitekunsaurus
4729:
4727:
4726:
4722:
4719:
4718:
4714:
4712:
4711:
4707:
4705:
4704:
4700:
4698:
4697:
4693:
4691:
4690:
4686:
4684:
4683:
4682:Mnyamawamtuka
4679:
4677:
4676:
4672:
4669:
4668:
4664:
4662:
4661:
4660:Magyarosaurus
4657:
4654:
4653:
4652:Ligabuesaurus
4649:
4647:
4646:
4645:Laplatasaurus
4642:
4640:
4639:
4638:Karongasaurus
4635:
4633:
4632:
4628:
4625:
4624:
4620:
4618:
4617:
4613:
4611:
4610:
4606:
4604:
4603:
4599:
4596:
4595:
4591:
4589:
4588:
4584:
4581:
4580:
4579:Dreadnoughtus
4576:
4574:
4573:
4569:
4567:
4566:
4562:
4560:
4559:
4555:
4553:
4552:
4548:
4546:
4545:
4544:Borealosaurus
4541:
4538:
4537:
4533:
4530:
4529:
4525:
4523:
4522:
4521:Barrosasaurus
4518:
4516:
4515:
4511:
4509:
4508:
4504:
4502:
4501:
4497:
4495:
4494:
4490:
4488:
4487:
4483:
4480:
4479:
4475:
4473:
4472:
4468:
4465:
4464:
4463:Aegyptosaurus
4460:
4458:
4457:
4453:
4452:
4449:
4445:
4442:
4439:
4434:
4430:
4426:
4421:
4417:
4407:
4402:
4399:
4391:
4361:
4360:
4359:
4358:
4356:
4354:
4350:
4339:
4338:
4334:
4332:
4331:
4330:Savannasaurus
4327:
4325:
4324:
4320:
4318:
4317:
4313:
4311:
4310:
4309:Australotitan
4306:
4305:
4302:
4299:
4297:
4293:
4287:
4286:
4282:
4280:
4279:
4275:
4273:
4272:
4268:
4266:
4265:
4261:
4259:
4258:
4254:
4252:
4251:
4247:
4245:
4244:
4240:
4237:
4236:
4232:
4230:
4229:
4225:
4223:
4222:
4218:
4216:
4215:
4211:
4209:
4208:
4204:
4202:
4201:
4197:
4195:
4194:
4190:
4187:
4186:
4182:
4179:
4178:
4177:Australodocus
4174:
4173:
4171:
4169:
4168:Euhelopodidae
4165:
4159:
4158:
4157:Triunfosaurus
4154:
4152:
4151:
4147:
4145:
4144:
4140:
4138:
4137:
4136:Sauroposeidon
4133:
4131:
4130:
4126:
4124:
4123:
4119:
4117:
4116:
4115:Padillasaurus
4112:
4110:
4109:
4105:
4103:
4102:
4098:
4095:
4094:
4093:Ligabuesaurus
4090:
4088:
4087:
4083:
4081:
4080:
4076:
4073:
4072:
4068:
4066:
4065:
4061:
4059:
4058:
4054:
4052:
4051:
4047:
4045:
4044:
4040:
4038:
4037:
4036:Chubutisaurus
4033:
4031:
4030:
4026:
4024:
4023:
4019:
4017:
4016:
4012:
4010:
4009:
4005:
4003:
4002:
3998:
3996:
3995:
3991:
3990:
3987:
3984:
3982:
3978:
3972:
3971:
3967:
3965:
3964:
3960:
3958:
3957:
3953:
3951:
3950:
3946:
3944:
3943:
3939:
3937:
3936:
3932:
3929:
3928:
3924:
3922:
3921:
3917:
3915:
3914:
3913:Brachiosaurus
3910:
3908:
3907:
3903:
3901:
3900:
3896:
3895:
3893:
3891:
3887:
3880:
3879:
3875:
3873:
3872:
3868:
3866:
3865:
3861:
3859:
3858:
3854:
3852:
3851:
3850:Fushanosaurus
3847:
3845:
3844:
3840:
3838:
3837:
3833:
3831:
3830:
3826:
3825:
3822:
3819:
3816:
3811:
3801:
3800:
3796:
3794:
3793:
3789:
3788:
3786:
3784:
3780:
3774:
3773:
3769:
3766:
3765:
3761:
3759:
3758:
3754:
3751:
3750:
3746:
3744:
3743:
3739:
3736:
3735:
3734:Bashunosaurus
3731:
3729:
3728:
3724:
3722:
3721:
3717:
3716:
3713:
3709:
3706:
3703:
3698:
3694:
3690:
3685:
3681:
3672:
3667:
3637:
3636:
3632:
3630:
3629:
3625:
3623:
3622:
3618:
3615:
3614:
3610:
3608:
3607:
3603:
3601:
3600:
3596:
3594:
3593:
3589:
3587:
3586:
3582:
3581:
3579:
3577:
3573:
3567:
3566:
3562:
3559:
3558:
3557:Atlantosaurus
3554:
3552:
3551:
3547:
3544:
3543:
3539:
3538:
3536:
3534:
3533:Apatosaurinae
3530:
3527:
3525:
3521:
3515:
3514:
3510:
3508:
3507:
3503:
3501:
3500:
3499:Smitanosaurus
3496:
3494:
3493:
3489:
3487:
3486:
3482:
3480:
3479:
3475:
3472:
3471:
3470:Dyslocosaurus
3467:
3465:
3464:
3463:Dicraeosaurus
3460:
3458:
3457:
3453:
3451:
3450:
3446:
3444:
3443:
3442:Amargatitanis
3439:
3437:
3436:
3432:
3431:
3429:
3427:
3423:
3420:
3418:
3414:
3400:
3399:
3395:
3393:
3392:
3388:
3386:
3385:
3381:
3379:
3378:
3377:Katepensaurus
3374:
3372:
3371:
3370:Itapeuasaurus
3367:
3365:
3364:
3363:Demandasaurus
3360:
3359:
3357:
3355:
3351:
3344:
3343:
3339:
3337:
3336:
3332:
3330:
3329:
3328:Cathartesaura
3325:
3324:
3322:
3320:
3319:Limaysaurinae
3316:
3313:
3311:
3307:
3301:
3300:
3296:
3294:
3293:
3289:
3287:
3286:
3282:
3280:
3279:
3275:
3272:
3271:
3267:
3265:
3264:
3260:
3258:
3257:
3256:Histriasaurus
3253:
3250:
3249:
3248:Dzharatitanis
3245:
3243:
3242:
3241:Comahuesaurus
3238:
3236:
3235:
3231:
3229:
3228:
3224:
3222:
3221:
3217:
3216:
3213:
3210:
3208:
3204:
3201:
3198:
3193:
3187:
3186:
3182:
3181:
3178:
3175:
3172:
3171:Diplodocoidea
3167:
3163:
3159:
3158:Diplodocoidea
3154:
3150:
3141:
3137:
3135:
3129:
3121:
3098:
3097:
3093:
3091:
3090:
3086:
3084:
3083:
3079:
3077:
3076:
3075:Gigantosaurus
3072:
3070:
3069:
3065:
3063:
3062:
3058:
3056:
3055:
3051:
3050:
3048:
3044:
3040:
3021:
3020:
3018:
3016:
3012:
3005:
3004:
3002:
3000:
2999:Diplodocoidea
2996:
2993:
2991:
2987:
2981:
2980:
2976:
2974:
2973:
2969:
2967:
2966:
2962:
2959:
2958:
2954:
2951:
2950:
2946:
2944:
2943:
2942:Narindasaurus
2939:
2937:
2936:
2932:
2930:
2929:
2925:
2923:
2922:
2921:Losillasaurus
2918:
2915:
2914:
2910:
2907:
2906:
2902:
2899:
2898:
2894:
2892:
2891:
2887:
2886:
2884:
2882:
2878:
2872:
2871:
2870:Zigongosaurus
2867:
2865:
2864:
2863:Yuanmousaurus
2860:
2858:
2857:
2856:Xinjiangtitan
2853:
2851:
2850:
2849:Wamweracaudia
2846:
2844:
2843:
2842:Tonganosaurus
2839:
2837:
2836:
2832:
2830:
2829:
2825:
2823:
2822:
2818:
2816:
2815:
2811:
2809:
2808:
2804:
2801:
2800:
2799:Klamelisaurus
2796:
2794:
2793:
2789:
2787:
2786:
2782:
2780:
2779:
2778:Huangshanlong
2775:
2773:
2772:
2768:
2765:
2764:
2760:
2758:
2757:
2753:
2750:
2749:
2745:
2743:
2742:
2738:
2736:
2735:
2731:
2730:
2728:
2726:
2722:
2715:
2714:
2710:
2707:
2706:
2702:
2699:
2698:
2697:Ferganasaurus
2694:
2692:
2691:
2687:
2685:
2684:
2680:
2679:
2677:
2675:
2674:Cetiosauridae
2671:
2665:
2664:
2660:
2658:
2657:
2653:
2651:
2650:
2646:
2644:
2643:
2639:
2637:
2636:
2632:
2630:
2629:
2625:
2623:
2622:
2618:
2616:
2615:
2611:
2609:
2608:
2604:
2602:
2601:
2597:
2595:
2594:
2590:
2588:
2587:
2583:
2581:
2580:
2576:
2574:
2573:
2569:
2567:
2566:
2562:
2560:
2559:
2555:
2554:
2551:
2548:
2545:
2540:
2530:
2529:
2525:
2523:
2522:
2521:Tazoudasaurus
2518:
2517:
2515:
2513:
2509:
2502:
2501:
2497:
2494:
2493:
2489:
2487:
2486:
2482:
2480:
2479:
2478:Ohmdenosaurus
2475:
2473:
2472:
2468:
2465:
2464:
2460:
2459:
2456:
2452:
2449:
2446:
2441:
2431:
2430:
2426:
2424:
2423:
2419:
2417:
2416:
2412:
2410:
2409:
2405:
2404:
2402:
2400:
2396:
2389:
2388:
2384:
2382:
2381:
2380:Schleitheimia
2377:
2375:
2374:
2370:
2367:
2366:
2362:
2359:
2358:
2357:Leonerasaurus
2354:
2352:
2351:
2347:
2345:
2344:
2340:
2338:
2337:
2333:
2331:
2330:
2329:Blikanasaurus
2326:
2324:
2323:
2319:
2318:
2315:
2311:
2308:
2305:
2300:
2296:
2292:
2287:
2283:
2273:
2268:
2266:
2261:
2255:
2214:
2213:
2212:
2211:
2209:
2207:
2203:
2197:
2196:
2192:
2190:
2189:
2185:
2183:
2182:
2178:
2176:
2175:
2171:
2169:
2168:
2164:
2162:
2161:
2160:Leonerasaurus
2157:
2155:
2154:
2153:Lamplughsaura
2150:
2148:
2147:
2143:
2141:
2140:
2136:
2134:
2133:
2129:
2127:
2126:
2122:
2121:
2118:
2115:
2113:
2109:
2103:
2102:
2101:Yunnanosaurus
2098:
2096:
2095:
2091:
2088:
2087:
2083:
2081:
2080:
2076:
2074:
2073:
2069:
2067:
2066:
2062:
2059:
2058:
2054:
2053:
2050:
2047:
2045:
2041:
2034:
2033:
2029:
2027:
2026:
2022:
2020:
2019:
2015:
2013:
2012:
2008:
2006:
2005:
2001:
1999:
1998:
1997:Lufengosaurus
1994:
1992:
1991:
1987:
1984:
1983:
1979:
1977:
1976:
1972:
1970:
1969:
1965:
1963:
1962:
1958:
1957:
1955:
1953:
1949:
1943:
1942:
1938:
1936:
1935:
1934:Eucnemesaurus
1931:
1930:
1928:
1926:
1925:Riojasauridae
1922:
1916:
1915:
1911:
1909:
1908:
1904:
1902:
1901:
1897:
1895:
1894:
1890:
1888:
1887:
1883:
1881:
1880:
1876:
1874:
1873:
1869:
1868:
1865:
1862:
1859:
1854:
1844:
1843:
1839:
1837:
1836:
1832:
1830:
1829:
1825:
1823:
1822:
1818:
1816:
1815:
1811:
1809:
1808:
1807:Euskelosaurus
1804:
1803:
1801:
1799:
1795:
1789:
1788:
1784:
1782:
1781:
1777:
1775:
1774:
1770:
1769:
1766:
1763:
1760:
1755:
1745:
1744:
1740:
1738:
1737:
1733:
1731:
1730:
1726:
1725:
1723:
1721:
1717:
1711:
1710:
1706:
1704:
1703:
1699:
1697:
1696:
1692:
1690:
1689:
1685:
1683:
1682:
1681:Bagualosaurus
1678:
1676:
1675:
1671:
1669:
1668:
1664:
1663:
1660:
1656:
1653:
1650:
1649:Bagualosauria
1645:
1635:
1634:
1630:
1627:
1626:
1622:
1619:
1618:
1617:Pampadromaeus
1614:
1612:
1611:
1607:
1605:
1604:
1600:
1597:
1596:
1592:
1589:
1588:
1584:
1583:
1581:
1579:
1578:Saturnaliidae
1575:
1569:
1568:
1564:
1562:
1561:
1560:Pampadromaeus
1557:
1555:
1554:
1550:
1548:
1547:
1543:
1541:
1540:
1539:Gigantoscelus
1536:
1534:
1533:
1529:
1527:
1526:
1522:
1521:
1518:
1514:
1511:
1508:
1503:
1499:
1495:
1490:
1486:
1465:
1464:
1463:
1462:
1460:
1458:
1454:
1446:
1442:
1441:
1440:
1439:
1436:
1433:
1431:
1427:
1423:
1415:
1411:
1408:
1406:
1402:
1399:
1397:
1393:
1391:
1387:
1385:
1381:
1380:
1375:
1371:
1367:
1360:
1355:
1353:
1348:
1346:
1341:
1340:
1337:
1326:
1322:
1317:
1312:
1308:
1304:
1300:
1296:
1292:
1288:
1284:
1277:
1274:
1269:
1265:
1261:
1257:
1253:
1249:
1245:
1241:
1237:
1233:
1229:
1225:
1218:
1215:
1210:
1206:
1202:
1198:
1193:
1188:
1184:
1180:
1175:
1170:
1166:
1162:
1158:
1154:
1150:
1143:
1140:
1135:
1131:
1127:
1123:
1119:
1115:
1107:
1104:
1099:
1095:
1091:
1087:
1083:
1079:
1075:
1071:
1063:
1060:
1055:
1049:
1045:
1037:
1035:
1031:
1026:
1022:
1017:
1012:
1008:
1004:
1000:
996:
988:
985:
980:
976:
972:
968:
960:
958:
956:
954:
952:
950:
948:
946:
944:
942:
938:
933:
929:
922:
919:
914:
910:
906:
902:
897:
892:
888:
884:
880:
876:
868:
866:
862:
857:
853:
849:
845:
841:
837:
833:
829:
821:
819:
817:
815:
813:
811:
809:
807:
805:
803:
799:
792:
781:
776:
766:
763:
756:
754:
752:
748:
747:
741:
734:
732:
730:
726:
722:
721:Dicraeosaurus
718:
714:
704:
700:
699:
691:
690:
682:
681:
673:
672:
664:
663:
655:
654:
646:
645:
637:
636:
633:
632:
631:
624:
623:
620:
619:
616:
615:
614:
607:
606:
603:
602:
599:
598:
597:
596:Amargatitanis
590:
589:
583:
582:
579:
578:
575:
574:
573:
566:
565:
559:
558:
555:
554:
551:
550:
549:
542:
541:
535:
534:
531:
530:
527:
526:
525:
524:Dicraeosaurus
518:
517:
511:
510:
507:
506:
503:
502:
501:
494:
493:
487:
486:
483:
482:
479:
478:
477:
470:
469:
463:
460:
459:
455:
454:
448:
446:
444:
439:
435:
431:
427:
423:
416:
414:
412:
408:
404:
400:
396:
389:
387:
385:
381:
377:
373:
372:Rodolfo Coria
369:
365:
361:
353:
351:
349:
346:dates to the
345:
341:
337:
333:
332:
327:
323:
322:
317:
313:
309:
306:
302:
301:
290:
284:
283:
275:
272:
268:
264:
260:
257:
252:
251:
244:
241:
240:
237:
231:
228:
227:
224:
223:Diplodocoidea
218:
216:Superfamily:
215:
214:
211:
205:
202:
199:
198:
195:
189:
186:
183:
182:
179:
176:
173:
170:
169:
166:
163:
160:
157:
156:
153:
150:
147:
146:
143:
140:
137:
136:
133:
130:
127:
126:
121:
116:
112:
104:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
32:
26:
22:
19:
5619:
5593:Titanosaurus
5591:
5583:
5576:
5568:
5563:Loricosaurus
5561:
5556:Iuticosaurus
5554:
5547:
5540:
5533:
5525:
5493:Yamanasaurus
5491:
5484:
5477:
5470:
5463:
5455:
5448:
5440:
5434:Abditosaurus
5432:
5411:
5404:
5396:
5389:
5381:
5373:
5366:
5360:Abditosaurus
5358:
5337:
5330:
5322:
5315:
5292:Tapuiasaurus
5290:
5282:
5275:
5255:
5250:Titanomachya
5248:
5241:
5219:Puertasaurus
5217:
5210:
5205:Notocolossus
5203:
5196:
5190:Jiangxititan
5188:
5181:
5173:
5168:Drusilasaura
5166:
5159:
5152:Lognkosauria
5135:
5128:
5121:
5114:
5107:
5100:
5093:
5086:Aeolosaurini
5075:Rinconsaurus
5073:
5066:
5059:
5052:
5044:
5037:
5026:Rinconsauria
5013:
5007:Tengrisaurus
5005:
5000:Quetecsaurus
4998:
4991:
4983:
4976:
4969:
4962:
4955:
4931:
4924:
4890:
4883:
4878:Lohuecotitan
4876:
4869:
4864:Garrigatitan
4862:
4855:
4850:Ampelosaurus
4848:
4823:
4816:
4809:
4804:Uberabatitan
4802:
4797:Sonidosaurus
4795:
4788:
4781:
4774:
4768:Tengrisaurus
4766:
4758:
4751:
4744:
4739:Rapetosaurus
4737:
4730:
4723:
4715:
4708:
4701:
4694:
4687:
4680:
4675:Malawisaurus
4673:
4665:
4658:
4650:
4643:
4636:
4629:
4621:
4616:Huabeisaurus
4614:
4607:
4600:
4592:
4585:
4577:
4570:
4563:
4558:Choconsaurus
4556:
4551:Brasilotitan
4549:
4542:
4534:
4526:
4519:
4512:
4505:
4500:Atacamatitan
4498:
4493:Argyrosaurus
4491:
4484:
4476:
4469:
4461:
4456:Abdarainurus
4454:
4438:Titanosauria
4425:Titanosauria
4394:
4353:Titanosauria
4337:Wintonotitan
4335:
4328:
4321:
4314:
4307:
4283:
4278:Yongjinglong
4276:
4269:
4264:Tambatitanis
4262:
4255:
4248:
4241:
4233:
4228:Huanghetitan
4226:
4221:Huabeisaurus
4219:
4212:
4207:Gannansaurus
4205:
4198:
4191:
4185:Chiayusaurus
4183:
4175:
4155:
4148:
4141:
4134:
4127:
4120:
4113:
4106:
4099:
4091:
4084:
4079:Jiutaisaurus
4077:
4071:Jiangxititan
4069:
4064:Garumbatitan
4062:
4055:
4048:
4043:Dongbeititan
4041:
4034:
4027:
4020:
4015:Austrosaurus
4013:
4006:
3999:
3992:
3968:
3963:Venenosaurus
3961:
3954:
3949:Sonorasaurus
3947:
3940:
3935:Giraffatitan
3933:
3925:
3920:Cedarosaurus
3918:
3911:
3904:
3897:
3876:
3871:Pelorosaurus
3869:
3862:
3855:
3848:
3841:
3834:
3827:
3797:
3792:Camarasaurus
3790:
3770:
3764:Haestasaurus
3762:
3757:Europasaurus
3755:
3747:
3740:
3732:
3725:
3718:
3633:
3626:
3619:
3611:
3604:
3597:
3590:
3583:
3576:Diplodocinae
3565:Brontosaurus
3563:
3555:
3548:
3542:Amphicoelias
3540:
3524:Diplodocidae
3511:
3504:
3497:
3491:
3490:
3483:
3476:
3468:
3461:
3454:
3449:Bajadasaurus
3447:
3440:
3435:Amargasaurus
3433:
3396:
3389:
3382:
3375:
3368:
3361:
3342:Rayososaurus
3340:
3333:
3326:
3299:Zapalasaurus
3297:
3292:Xenoposeidon
3290:
3283:
3276:
3268:
3261:
3254:
3246:
3239:
3234:Campananeyen
3232:
3227:Amazonsaurus
3225:
3218:
3183:
3124:
3094:
3087:
3080:
3073:
3066:
3059:
3052:
2990:Neosauropoda
2977:
2970:
2963:
2955:
2947:
2940:
2933:
2926:
2919:
2911:
2905:Haestasaurus
2903:
2895:
2888:
2868:
2861:
2854:
2847:
2840:
2833:
2826:
2819:
2812:
2805:
2797:
2790:
2783:
2776:
2769:
2761:
2754:
2746:
2739:
2732:
2713:Patagosaurus
2711:
2703:
2695:
2688:
2681:
2663:Volkheimeria
2661:
2654:
2647:
2640:
2635:Perijasaurus
2633:
2628:Nebulasaurus
2626:
2619:
2612:
2607:Dystrophaeus
2605:
2598:
2591:
2584:
2579:Barapasaurus
2577:
2570:
2565:Asiatosaurus
2563:
2556:
2526:
2519:
2498:
2490:
2485:Rhoetosaurus
2483:
2476:
2469:
2461:
2429:Lessemsaurus
2427:
2420:
2413:
2408:Antetonitrus
2406:
2385:
2378:
2371:
2363:
2355:
2348:
2341:
2334:
2327:
2320:
2257:
2193:
2186:
2179:
2172:
2165:
2158:
2151:
2144:
2137:
2130:
2123:
2099:
2094:Yizhousaurus
2092:
2084:
2077:
2070:
2063:
2055:
2030:
2023:
2016:
2009:
2002:
1995:
1988:
1982:Ignavusaurus
1980:
1973:
1966:
1959:
1939:
1932:
1912:
1905:
1898:
1891:
1886:Ignavusaurus
1884:
1877:
1870:
1842:Yimenosaurus
1840:
1835:Plateosaurus
1833:
1826:
1819:
1812:
1805:
1787:Xixiposaurus
1785:
1778:
1771:
1759:Plateosauria
1741:
1734:
1727:
1720:Unaysauridae
1707:
1700:
1693:
1686:
1679:
1672:
1665:
1631:
1623:
1615:
1608:
1601:
1593:
1585:
1565:
1558:
1551:
1546:Guaibasaurus
1544:
1537:
1530:
1523:
1409:
1400:
1290:
1286:
1276:
1227:
1223:
1217:
1156:
1152:
1142:
1117:
1113:
1106:
1073:
1069:
1062:
1043:
998:
994:
987:
970:
966:
931:
927:
921:
896:11336/122794
878:
874:
831:
827:
765:
746:Lajasvenator
744:
739:
738:
735:Paleoecology
729:Amargasaurus
728:
724:
720:
717:Amargasaurus
716:
712:
711:
702:
629:
628:
627:
613:Bajadasaurus
611:
610:
594:
593:
572:Amargasaurus
570:
569:
546:
545:
522:
521:
498:
497:
474:
473:
443:Amargasaurus
442:
437:
434:Amargasaurus
433:
430:Bajadasaurus
429:
426:Amargasaurus
425:
421:
420:
410:
394:
393:
384:Amargasaurus
383:
379:
375:
359:
357:
343:
335:
331:Amargasaurus
329:
325:
320:
319:
299:
298:
297:
288:
281:
280:
271:Type species
258:
249:
248:
200:
184:
171:
158:
24:
18:
5578:Microcoelus
5486:Saltasaurus
5472:Paralititan
5368:Alamosaurus
5257:Udelartitan
5212:Patagotitan
5109:Bravasaurus
5102:Arrudatitan
5095:Aeolosaurus
5061:Overosaurus
4993:Jainosaurus
4971:Bonitasaura
4892:Paludititan
4783:Traukutitan
4478:Angolatitan
4285:Yunmenglong
4250:Qiaowanlong
4143:Sibirotitan
4108:Oceanotitan
4050:Europatitan
4029:Brontomerus
4022:Brohisaurus
3994:Angolatitan
3927:Galvesaurus
3899:Abydosaurus
3864:Ornithopsis
3857:Fusuisaurus
3843:Eucamerotus
3742:Daanosaurus
3727:Aragosaurus
3628:Supersaurus
3550:Apatosaurus
3513:Tharosaurus
3384:Nigersaurus
3335:Limaysaurus
3096:Ultrasaurus
2972:Turiasaurus
2935:Moabosaurus
2928:Mierasaurus
2897:Bellusaurus
2881:Turiasauria
2821:Qijianglong
2785:Hudiesaurus
2763:Datousaurus
2683:Cetiosaurus
2642:Shunosaurus
2586:Bellusaurus
2558:Algoasaurus
2544:Eusauropoda
2492:Sanpasaurus
2445:Gravisauria
2365:Pulanesaura
2350:Isanosaurus
2322:Amygdalodon
2188:Pulanesaura
2146:Irisosaurus
2132:Anchisaurus
2112:Anchisauria
2086:Xingxiulong
2032:Xingxiulong
2025:Sarahsaurus
1941:Riojasaurus
1907:Sarahsaurus
1893:Kholumolumo
1736:Macrocollum
1674:Asylosaurus
1610:Nhandumirim
1587:Buriolestes
1553:Mbiresaurus
1525:Buriolestes
1316:11336/30408
1293:: 228–239.
1159:(1): 1392.
1120:: 103–120.
1016:11336/86084
1001:: 437–441.
928:Ameghiniana
417:Description
358:Fossils of
348:Valanginian
305:diplodocoid
36:Valanginian
5689:Categories
5479:Rocasaurus
5406:Qunkasaura
5375:Baurutitan
5137:Shingopana
4964:Baalsaurus
4811:Volgatitan
4746:Rukwatitan
4710:Nullotitan
4696:Ninjatitan
4631:Kaijutitan
4602:Gandititan
4565:Daxiatitan
4528:Baurutitan
4471:Andesaurus
4362:see below↓
4057:Fukuititan
4001:Arkharavia
3956:Soriatitan
3906:Atlasaurus
3878:Rugocaudia
3836:Duriatitan
3772:Yuzhoulong
3720:Abrosaurus
3702:Macronaria
3689:Macronaria
3613:Kaatedocus
3606:Galeamopus
3599:Diplodocus
3585:Barosaurus
3492:Pilmatueia
3485:Lingwulong
3478:Kaatedocus
3398:Tataouinea
3310:Khebbashia
3285:Sidersaura
3015:Macronaria
2965:Tendaguria
2949:Oplosaurus
2913:Janenschia
2814:Omeisaurus
2690:Chebsaurus
2614:Janenschia
2528:Vulcanodon
2471:Kotasaurus
2422:Ledumahadi
2215:see below↓
2174:Meroktenos
1990:Leyesaurus
1879:Gyposaurus
1743:Unaysaurus
1702:Pantydraco
1667:Arcusaurus
1633:Saturnalia
1466:see below↓
1414:Dinosauria
1396:Sauropsida
934:(4R): R44.
881:: 104319.
777:References
740:Pilmatueia
713:Pilmatueia
703:Pilmatueia
630:Pilmatueia
500:Lingwulong
438:Pilmatueia
422:Pilmatueia
380:Pilmatueia
344:Pilmatueia
336:Pilmatueia
326:Pilmatueia
300:Pilmatueia
250:Pilmatueia
178:Saurischia
165:Dinosauria
25:Pilmatueia
5442:Bonatitan
5324:Isisaurus
5130:Punatitan
4609:Hamititan
4587:Elaltitan
4536:Bonatitan
4257:Silutitan
4214:Gobititan
4200:Euhelopus
3942:Lusotitan
3635:Tornieria
3621:Leinkupal
3506:Suuwassea
3220:Agustinia
3068:Dinodocus
3061:Cardiodon
2792:Jingiella
2741:Anhuilong
2304:Sauropoda
2291:Sauropoda
2206:Sauropoda
2181:Mussaurus
2139:Camelotia
2018:Pradhania
1858:Massopoda
1625:Panphagia
1567:Panphagia
1382:Kingdom:
1325:0195-6671
1268:234424022
1260:0272-4634
1252:1937-2809
1183:2045-2322
1134:0024-4066
1098:212766423
1090:0024-1164
1025:0195-6671
979:1631-0683
913:214475057
905:0195-6671
856:135017018
834:: 33–48.
757:Footnotes
476:Suuwassea
403:Argentina
368:Argentina
340:pneumatic
210:Sauropoda
138:Kingdom:
132:Eukaryota
5450:Ibirania
5116:Caieiria
4926:Inawentu
4129:Ruixinia
3970:Vouivria
3829:Astrodon
3082:Neosodon
2572:Bagualia
2415:Ingentia
2125:Aardonyx
2072:Qianlong
1900:Musankwa
1872:Gryponyx
1780:Ruehleia
1695:Nambalia
1688:Efraasia
1595:Eoraptor
1532:Eoraptor
1390:Chordata
1388:Phylum:
1384:Animalia
1209:59603627
1201:30718633
407:endocast
308:sauropod
229:Family:
152:Chordata
148:Phylum:
142:Animalia
128:Domain:
4486:Arackar
2890:Amanzia
2734:Analong
2621:Jobaria
2079:Seitaad
1394:Class:
1295:Bibcode
1232:Bibcode
1192:6362061
1161:Bibcode
1070:Lethaia
836:Bibcode
242:Genus:
5015:Vahiny
4776:Tiamat
4193:Erketu
2011:Ngwevu
1323:
1266:
1258:
1250:
1207:
1199:
1189:
1181:
1132:
1096:
1088:
1050:
1023:
977:
911:
903:
854:
727:, and
291:, 2019
289:et al.
287:Coria
259:et al.
1410:Clade
1401:Clade
1264:S2CID
1248:eISSN
1205:S2CID
1094:S2CID
909:S2CID
852:S2CID
303:is a
256:Coria
201:Clade
185:Clade
172:Clade
159:Clade
5317:Igai
1821:Issi
1443:see
1321:ISSN
1256:ISSN
1197:PMID
1179:ISSN
1130:ISSN
1086:ISSN
1048:ISBN
1021:ISSN
975:ISSN
901:ISSN
428:and
263:2019
41:Preęž’
2979:Zby
1311:hdl
1303:doi
1240:doi
1187:PMC
1169:doi
1122:doi
1118:138
1078:doi
1011:hdl
1003:doi
891:hdl
883:doi
879:111
844:doi
401:of
5691::
1412::
1403::
1319:.
1309:.
1301:.
1291:78
1289:.
1285:.
1262:.
1254:.
1246:.
1238:.
1228:40
1226:.
1203:.
1195:.
1185:.
1177:.
1167:.
1155:.
1151:.
1128:.
1116:.
1092:.
1084:.
1074:53
1072:.
1033:^
1019:.
1009:.
999:84
997:.
971:21
969:.
940:^
932:49
930:.
907:.
899:.
889:.
877:.
864:^
850:.
842:.
832:93
830:.
801:^
749:.
723:,
413:.
386:.
366:,
334:.
324:.
261:,
203::
187::
174::
161::
91:Pg
33:,
5588:?
5573:?
5530:?
5460:?
5445:?
5437:?
5416:?
5401:?
5386:?
5378:?
5363:?
5342:?
5327:?
5295:?
5287:?
5193:?
5178:?
5049:?
5010:?
4988:?
4828:?
4771:?
4763:?
4720:?
4670:?
4655:?
4626:?
4597:?
4582:?
4539:?
4531:?
4481:?
4466:?
4340:?
4238:?
4188:?
4180:?
4096:?
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3930:?
3881:?
3767:?
3752:?
3737:?
3616:?
3560:?
3545:?
3473:?
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2908:?
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2802:?
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2035:?
1985:?
1628:?
1620:?
1598:?
1590:?
1358:e
1351:t
1344:v
1327:.
1313::
1305::
1297::
1270:.
1242::
1234::
1211:.
1171::
1163::
1157:9
1136:.
1124::
1100:.
1080::
1056:.
1027:.
1013::
1005::
981:.
915:.
893::
885::
858:.
846::
838::
278:â€
246:â€
233:â€
220:â€
207:â€
191:â€
96:N
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