53:
33:
541:
Neilson, M.E.; Stepien C.A. (2009). "Escape from the Ponto-Caspian: evolution and biogeography of an endemic goby species flock (Benthophilinae: Gobiidae: Teleostei)".
742:
768:
716:
755:
579:
594:
Vasil'eva, E.D., Schwarzhans, W.W., Medvedev, D.A. & Vasil'ev, V.P. (2016): Cryptic
Species of Ponto-Caspian Bighead Goby of the Genus
807:
840:
760:
434:
367:
349:
690:
773:
438:
375:
353:
52:
262:
194:
occur in the brackish-water Black and
Caspian seas themselves. It was considered to be part of the broader goby
256:
329:
845:
335:
320:
305:
291:
248:
643:
471:
419:
398:
299:
276:
695:
517:
371:
794:
362:
270:
343:
812:
486:
453:
242:
207:
195:
835:
490:
457:
447:
428:
404:
383:
47:
799:
781:
747:
703:
575:
550:
389:
786:
480:
465:
314:
285:
413:
145:
681:
612:
198:
171:
94:
829:
513:
708:
137:
32:
729:
675:
183:
175:
104:
666:
617:
sp. nov. (Actinopterygii: Perciformes: Gobiidae) from the
Caspian Sea basin"
225:
179:
64:
554:
660:
569:
526:
220:
212:
202:
114:
84:
721:
191:
187:
734:
168:
74:
637:
164:
641:
611:
Vasil'eva, E.D.; Mousavi-Sabet, H.; Vasil'ev, V.P. (2015).
237:
There are currently 16 recognized species in this genus:
568:
J. S. Nelson; T. C. Grande; M. V. H. Wilson (2016).
650:
205:to the same region, although the 5th edition of
8:
638:
31:
20:
505:
600:Journal of Ichthyology, 56 (1): 1-18.
543:Molecular Phylogenetics and Evolution
211:does not list any subfamilies in the
7:
574:(5th ed.). Wiley. p. 752.
14:
624:Acta Ichthyologica et Piscatoria
51:
358:(Eastern Caspian bighead goby)
1:
862:
143:
136:
48:Scientific classification
46:
39:
30:
23:
516:; Pauly, Daniel (eds.).
330:Ponticola gymnotrachelus
257:Ponticola cephalargoides
530:. January 2016 version.
399:Ponticola platyrostris
325:(Caspian bighead goby)
300:Ponticola eurycephalus
841:Fish of the Black Sea
795:Paleobiology Database
271:Ponticola constructor
571:Fishes of the World
243:Ponticola bathybius
208:Fishes of the World
41:Ponticola bathybius
615:Ponticola iranicus
518:"Species in genus
448:Ponticola rizensis
429:Ponticola rhodioni
384:Ponticola kessleri
363:Ponticola iranicus
823:
822:
782:Open Tree of Life
644:Taxon identifiers
581:978-1-118-34233-6
494:
475:
461:
442:
423:
409:(Flat-snout goby)
408:
393:
379:
357:
339:
324:
309:
295:
280:
266:
252:
219:was described as
156:
155:
132:
853:
816:
815:
803:
802:
790:
789:
777:
776:
764:
763:
751:
750:
738:
737:
725:
724:
712:
711:
699:
698:
686:
685:
684:
671:
670:
669:
639:
632:
631:
621:
608:
602:
592:
586:
585:
565:
559:
558:
538:
532:
531:
510:
484:
481:Ponticola turani
469:
466:Ponticola syrman
451:
432:
417:
402:
387:
366:
347:
344:Ponticola iljini
333:
318:
315:Ponticola gorlap
303:
289:
286:Ponticola cyrius
281:(Caucasian goby)
274:
260:
246:
130:
56:
55:
35:
21:
861:
860:
856:
855:
854:
852:
851:
850:
826:
825:
824:
819:
811:
806:
798:
793:
785:
780:
772:
767:
759:
754:
746:
741:
733:
728:
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715:
707:
702:
694:
689:
680:
679:
674:
665:
664:
659:
646:
636:
635:
619:
610:
609:
605:
593:
589:
582:
567:
566:
562:
540:
539:
535:
512:
511:
507:
502:
443:(Riverine goby)
414:Ponticola ratan
394:(Big-head goby)
310:(Mushroom goby)
235:
152:
149:
129:
50:
17:
16:Genus of fishes
12:
11:
5:
859:
857:
849:
848:
846:Benthophilinae
843:
838:
828:
827:
821:
820:
818:
817:
804:
791:
778:
765:
752:
739:
726:
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687:
672:
656:
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634:
633:
603:
587:
580:
560:
533:
514:Froese, Rainer
504:
503:
501:
498:
497:
496:
495:(Turan's goby)
477:
462:
444:
425:
410:
395:
380:
359:
340:
326:
311:
296:
282:
267:
253:
234:
231:
215:. Originally,
199:Benthophilinae
154:
153:
151:Nordmann, 1840
150:
141:
140:
134:
133:
122:
118:
117:
112:
108:
107:
102:
98:
97:
95:Actinopterygii
92:
88:
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82:
78:
77:
72:
68:
67:
62:
58:
57:
44:
43:
37:
36:
28:
27:
15:
13:
10:
9:
6:
4:
3:
2:
858:
847:
844:
842:
839:
837:
834:
833:
831:
814:
809:
805:
801:
796:
792:
788:
783:
779:
775:
770:
766:
762:
757:
753:
749:
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740:
736:
731:
727:
723:
718:
714:
710:
705:
701:
697:
692:
688:
683:
677:
673:
668:
662:
658:
657:
655:
653:
649:
645:
640:
630:(2): 189–197.
629:
625:
618:
616:
607:
604:
601:
597:
591:
588:
583:
577:
573:
572:
564:
561:
556:
552:
549:(1): 84–102.
548:
544:
537:
534:
529:
528:
523:
521:
515:
509:
506:
499:
492:
488:
483:
482:
478:
476:(Syrman goby)
473:
468:
467:
463:
459:
455:
450:
449:
445:
440:
436:
431:
430:
426:
421:
416:
415:
411:
406:
401:
400:
396:
391:
386:
385:
381:
377:
373:
372:Mousavi-Sabet
369:
365:
364:
360:
355:
351:
346:
345:
341:
337:
332:
331:
327:
322:
317:
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45:
42:
38:
34:
29:
26:
22:
19:
651:
627:
623:
614:
606:
599:
598:(Gobiidae).
595:
590:
570:
563:
546:
542:
536:
525:
519:
508:
479:
464:
446:
427:
424:(Ratan goby)
412:
397:
382:
361:
342:
328:
313:
298:
284:
269:
255:
241:
236:
224:
216:
206:
176:fresh waters
159:
158:
157:
146:Gobius ratan
144:
138:Type species
125:
124:
40:
24:
18:
730:iNaturalist
676:Wikispecies
184:Caspian Sea
131:Iljin, 1927
105:Gobiiformes
830:Categories
500:References
186:region in
174:mostly to
836:Ponticola
682:Ponticola
652:Ponticola
596:Ponticola
520:Ponticola
435:Vasil'eva
368:Vasil'eva
350:Vasil'eva
226:Neogobius
217:Ponticola
196:subfamily
180:Black Sea
160:Ponticola
126:Ponticola
71:Kingdom:
65:Eukaryota
25:Ponticola
661:Wikidata
555:19475709
527:FishBase
472:Nordmann
439:Vasil'ev
420:Nordmann
376:Vasil'ev
354:Vasil'ev
277:Nordmann
221:subgenus
213:Gobiidae
115:Gobiidae
111:Family:
85:Chordata
81:Phylum:
75:Animalia
61:Domain:
748:1305530
722:2379222
667:Q753690
493:, 2008)
487:Kovačić
474:, 1840)
460:, 2008)
454:Kovačić
441:, 1994)
422:, 1840)
407:, 1814)
392:, 1861)
390:Günther
356:, 1996)
338:, 1857)
336:Kessler
323:, 1949)
308:, 1874)
306:Kessler
294:, 1874)
292:Kessler
279:, 1840)
265:, 1976)
263:Pinchuk
251:, 1877)
249:Kessler
233:Species
203:endemic
201:, also
192:species
190:. Some
188:Eurasia
178:of the
121:Genus:
101:Order:
91:Class:
813:297547
800:332808
787:165347
774:637986
761:172202
696:393674
578:
553:
489:&
456:&
437:&
405:Pallas
378:, 2015
374:&
352:&
172:native
169:gobies
808:WoRMS
743:IRMNG
735:91238
709:64279
620:(PDF)
491:Engin
458:Engin
321:Iljin
165:genus
163:is a
769:NCBI
756:ITIS
717:GBIF
691:BOLD
576:ISBN
551:PMID
704:CoL
223:of
167:of
832::
810::
797::
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771::
758::
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732::
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678::
663::
628:45
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547:52
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370:,
229:.
182:-
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485:(
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388:(
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247:(
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
