1178:
884:
1326:
1128:
765:
387:
118:
1297:
663:
320:
1218:
3469:
302:
957:
840:
361:
The Prince Creek
Formation aged from 80 to 61.7 million years ago. The Kikak-Tegoseak Quarry, where almost all of the dinosaur fossil are from, is located near the middle of the formation, and is about 70.6 to 69.1 million years ago. A lower section, the Kogosukruk Tongue, ages from 72 to 71 million
405:, and were, at the time of burial, on a muddy coastal plain. Leafy plants, roots and pollen are known from the formation, and they show that trampling by dinosaurs was common. It can be proven that during the Maastrichtian the Prince Creek Formation bordered a large body of water by the presence of
477:
towering above them. The mean temperature was 5 to 6 °C (41 to 43 °F), with the mean temperature during the cold months being 2 to 4 °C (36 to 39 °F) and the mean temperature during the warm months being 10 to 12 °C (50 to 54 °F). Mean annual precipitation was 500 to
4474:
Fiorillo, A. R.; Hasiotis, S. T.; Kobayashi, Y.; Breithaupt, B. H.; McCarthy, P. J. (2011). "Bird tracks from the Upper
Cretaceous Cantwell Formation of Denali National Park, Alaska, USA: a new perspective on ancient northern polar vertebrate biodiversity".
5275:
Flaig, P.P.; McCarthy, P.J.; Fiorillo, A.R. (2011). "A Tidally-Influenced, High-Latitude
Coastal-Plain: the Upper Cretaceous (Maastrichtian) Prince Creek Formation, North Slope, Alaska". In Stephanie K. Davidson; Sophie Leleu; Colin P. North (eds.).
5210:
Eberle, Jaelyn J.; Clemens, William A.; McCarthy, Paul J.; Fiorillo, Anthony R.; Erickson, Gregory M.; Druckenmiller, Patrick S. (2019-02-14). "Northernmost record of the
Metatheria: a new Late Cretaceous pediomyid from the North Slope of Alaska".
4014:
Weishampel, David B; et al. (2004). "Dinosaur distribution (Late
Cretaceous, North America)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 574-588.
3960:"Anatomy, Evolution, and Paleoenvironmental Interpretation of an Ancient Arctic Coastal Plain: Integrated Paleopedology and Palynology from the Upper Cretaceous (Maastrichtian) Prince Creek Formation, North Slope, Alaska, USA"
5303:
Flaig, P.P.; Fiorillo, A.R.; McCarthy, P.J. (2014). "Dinosaur-bearing hyperconcentrated flows of
Cretaceous Arctic Alaska—Recurring catastrophic event beds on a distal paleopolar coastal plain".
5415:
378:
and mollusks. In the middle of the formation is the
Coleville River Bluff, which extends from the Late Campanian to the Middle/Late Maastrichtian, in which pollen spores are common.
3011:
Very abundant in the early
Maastrichtian and Indeterminate level of the formation, becoming rarer until the Middle/Late Maastrichtian. Also found in the Schrader Bluff Formation.
4227:
Marincovich, L.; Brouwers, E. M.; Carter, L. D. (1985). "Early
Tertiary marine fossils from northern Alaska: Implications for Arctic Ocean paleogeography and faunal evolution".
5266:
Flaig, P.P. (2010). "Depositional
Environments of the Late Cretaceous (Maaastrichtian) Dinosaur-Bearing Prince Creek Formation: Colville River Region, North Slope, Alaska".
500:
Indeterminate tyrannosaurid remains are present, mostly in the form of teeth. The teeth are from the Kikak-Tegoseak Quarry, Liscomb Quarry, and Byers Bed, totaling 8 teeth.
5395:
5019:
Sharpe, Henry S.; Powers, Mark J.; Dyer, Aaron D.; Rhodes, Matthew M.; McIntosh, Annie P.; Garros, Christiana W.; Currie, Philip J.; Funston, Gregory F. (2024-04-16).
5375:
1167:
species, found in one of the highest latitudes of centrosaurine discoveries. A discovery in the Kikak-Tegoseak Quarry was identified in 2013 as a juvenile of
4964:
Brown, C.M.; Druckenmiller, P. (2011). "Basal ornithopod (Dinosauria: Ornithischia) teeth from the Prince Creek Formation (early Maastrichtian) of Alaska".
5380:
2426:
Indeterminate remains abundant in the Early Maastrichtian, and still numerous in the Late Maastrichtian. Also found in the Schrader Bluff Formation.
3933:
5360:
5339:
Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. 861 pp.
4414:(2000). "Theropod Teeth from the Prince Creek Formation (Cretaceous) of Northern Alaska, with Speculations on Arctic Dinosaur Paleoecology".
3498:
Common and abundant in the Early Maastrichtian, becoming rarer towards the Late Maastrichtian. Also found in the Schrader Bluff Formation.
1171:. This discovery shows that the crest started to develop in the front of the snout, then extending farther back until it reaches the eye.
478:
1,500 millimetres (20 to 59 in)/year. The paleolatitude of the formation at the time of deposition was around 80°–85°N, high in the
5139:
5060:"The First Definite Lambeosaurine Bone From the Liscomb Bonebed of the Upper Cretaceous Prince Creek Formation, Alaska, United States"
4832:(1987). "Plant Megafossils, Vertebrate Remains, and Paleoclimate of the Kogosukruk Tongue (Late Cretaceous), North Slope, Alaska". In
5344:
5293:
4020:
3979:
4658:(2009). "Description of two partial Troodon braincases from the Prince Creek Formation (Upper Cretaceous), North Slope Alaska".
4294:"Stratigraphy and Facies of Cretaceous Schrader Bluff and Prince Creek Formations in Colville River Bluffs, North Slope, Alaska"
5420:
4703:"A New Troodontid Theropod, Talos sampsoni gen. et sp. nov., from the Upper Cretaceous Western Interior Basin of North America"
4192:(2005). "The First Pachycephalosaurine (Dinosauria) from the Paleo-Arctic of Alaska and its Paleogeographic Implications".
5400:
1177:
264:
225:
218:
5134:. Anchorage, Alaska: U.S. Dept. of the Interior, Minerals Management Service, Alaska Outer Continental Shelf Region.
5004:
Re-examination of the cranial osteology of the Arctic Alaskan hadrosaurine with implications for its taxonomic status
3712:
Numerous in the Late Campanian, becoming abundant in the Maastrichtian. Also found in the Schrader Bluff Formation.
5365:
4323:
Druckenmiller, Patrick S.; Erickson, Gregory M.; Brinkman, Donald; Brown, Caleb M.; Eberle, Jaelyn J. (June 2021).
5058:
McCarthy, Paul J.; Tykoski, Ronald S.; Kobayashi, Yoshitsugu; Fiorillo, Anthony R.; Takasaki, Ryuji (2019-03-29).
5410:
5405:
5385:
4427:
4205:
883:
5370:
4701:
Zanno, Lindsay E.; Varricchio, David J.; O'Connor, Patrick M.; Titus, Alan L.; Knell, Michael J. (2011-09-19).
1884:
438:
169:
4248:
5003:
1537:
159:
4575:
Chiarenza, A. A.; Fiorillo, A. R.; Tykoski, R. S.; McCarthy, P. J.; Flaig, P. P.; Contreras, D. L. (2020).
1936:
4888:(Dinosauria: Ceratopsidae) Nasal Reveals Unexpected Complexity of Craniofacial Ontogeny and Integument in
4289:
4273:
3964:
New Frontiers in Paleopedology and Terrestrial Paleoclimatology: Paleosols and Soil Surface Analog Systems
3094:
4639:
3274:
5390:
4411:
3400:
1858:
4643:
4285:
2949:
2400:
5155:
Eberle, Jaelyn J.; Clemens, William A.; Erickson, Gregory M.; Druckenmiller, Patrick S. (2023-01-01).
4837:
4651:
4277:
3580:
2229:
5312:
5220:
5168:
5156:
5071:
5020:
4973:
4905:
4714:
4667:
4588:
4526:
4484:
4336:
4265:
4236:
4189:
4181:
4080:
3856:
3758:
3444:
3131:
2914:
2737:
2158:
1118:
131:
5278:
From River to Rock Record: The Preservation of Fluvial Sediments and their Subsequent Interpretation
4829:
4771:
Sullivan, R.M. (2006). "A taxonomic review of the Pachycephalosauridae (Dinosauria: Ornithischia)".
4058:
3892:
3508:
3239:
2878:
2808:
2641:
2606:
2570:
2534:
2123:
1972:
4825:
4281:
4269:
3821:
3364:
3203:
3167:
2985:
2772:
2498:
2365:
2329:
1898:
1740:
A conifer, one of two from the region. It dominated the forest community of the Kogosukruk Tongue.
338:
2843:
2265:
1718:
5328:
5244:
5192:
4849:
4833:
4683:
4655:
4550:
4431:
4209:
3989:
3722:
3056:
3021:
2676:
2463:
2194:
932:
902:
894:
4457:"3.33 Alaska, United States; 3. Prince Creek Formation," in Weishampel, et al. (2004). Page 587.
1325:
1127:
2008:
1789:
1750:
764:
386:
117:
5340:
5289:
5236:
5184:
5135:
5105:
5087:
5040:
4933:
4750:
4732:
4616:
4542:
4362:
4354:
4108:
4016:
3975:
1226:
850:
754:
705:
673:
5320:
5281:
5228:
5176:
5095:
5079:
5032:
4981:
4923:
4913:
4740:
4722:
4675:
4647:
4606:
4596:
4534:
4492:
4423:
4407:
4344:
4244:
4201:
4185:
4156:
4098:
4088:
4054:
3967:
3544:
1478:
1436:
1296:
1187:
1138:
662:
4861:
4001:
875:
812:
654:
350:
397:
During the time when the Prince Creek Formation was deposited, Earth was going through a
5316:
5224:
5172:
5075:
4977:
4909:
4718:
4671:
4592:
4530:
4488:
4340:
4240:
4084:
5100:
5059:
4928:
4883:
4745:
4702:
4611:
4576:
4103:
4066:
1305:
1217:
721:
257:
141:
353:. Dinosaur remains are among the fossils that have been recovered from the formation.
5354:
5332:
5196:
4687:
4554:
4514:
4435:
4213:
3959:
1247:
1092:
1061:
823:
623:
479:
467:
346:
325:
69:
31:
27:
5248:
3468:
4679:
4062:
3435:
1447:
390:
206:
5232:
5180:
5036:
4515:"An ornithomimosaurian from the Upper Cretaceous Prince Creek Formation of Alaska"
4918:
4727:
4601:
4538:
4513:
Watanabe, Akinobu; Erickson, Gregory M.; Druckenmiller, Patrick S. (2013-09-01).
4496:
4093:
1845:
1821:
1653:
1501:
1335:
1027:
829:
775:
688:
559:
44:
5083:
4293:
2355:
Common in the Early Maastrichtian. Also found in the Schrader Bluff Formation.
2148:
Common in the Early Maastrichtian. Also found in the Schrader Bluff Formation.
956:
413:
in nearby rock. Large amounts of plants material are represented by peridonoid
4349:
4324:
1776:
1639:
1581:
1560:
1013:
545:
474:
443:
398:
89:
54:
5240:
5188:
5091:
5044:
5006:
Ryuji Takasaki, Anthony R. Fiorillo, Ronald S. Tykoski, Yoshitsugu Kobayashi.
4736:
4577:"The first juvenile dromaeosaurid (Dinosauria: Theropoda) from Arctic Alaska"
4546:
4358:
279:
266:
240:
227:
1412:
817:
367:
366:. The youngest part of the formation is Ocean Point, which extends into the
363:
198:
184:
151:
Kikak-Tegoseak Quarry, Kogosukruk Tongue, Ocean Point, Coleville River Bluff
94:
38:
5285:
5109:
4937:
4754:
4620:
4366:
4112:
5324:
4161:
4140:
3971:
4896:
4071:
1549:
1525:
1468:
1283:
800:
593:
470:
414:
402:
375:
188:
84:
79:
64:
59:
49:
839:
4145:
from the North Slope (Prince Creek Formation: Maastrichtian) of Alaska"
1646:
1020:
943:
939:
796:
552:
99:
74:
1380:
447:
410:
406:
371:
342:
307:
4985:
1612:
986:
518:
5280:. Vol. 97. Society for Sedimentary Geology. pp. 233–264.
5268:
Unpublished Ph.D. Dissertations, the University of Alaska-Fairbanks
1687:
1617:
1158:
An abundance of skeletal remains, including an immature juvenile.
991:
926:
Dental remains, including teeth. Braincases have also been found.
523:
463:
455:
433:
422:
418:
385:
202:
1320:
The first confirmed lambeosaurine in the Prince Creek Formation.
482:, and would have likely experienced 120 days of winter darkness.
459:
451:
430:
426:
4882:
Fiorillo, A.R.; Tykoski, R.S. (2013). Farke, Andrew A. (ed.).
5132:
1992 Proceedings, International Conference on Arctic Margins
1355:
A single "hypsilophodontid" cheek tooth not attributable to
4877:
4875:
4873:
4871:
4428:
10.1671/0272-4634(2000)020[0675:ttftpc]2.0.co;2
4325:"Nesting at extreme polar latitudes by non-avian dinosaurs"
4206:
10.1666/0022-3360(2005)079[0997:tfpdft]2.0.co;2
950:. The most abundant theropod. As of 2011, a dubious genus.
5025:
Lambe, 1917 clarifies issues in ontogeny and biogeography"
5157:"A new tiny eutherian from the Late Cretaceous of Alaska"
4773:
New Mexico Museum of Natural History and Science Bulletin
4249:
10.1130/0091-7613(1985)13<770:ETMFFN>2.0.CO;2
795:
One partial skull including a bone near the front of the
4067:"A Diminutive New Tyrannosaur from the Top of the World"
122:
General location of the Prince Creek Formation, in red
5021:"Craniomandibular anatomy of a juvenile specimen of
3958:
Flaig, P.P.; McCarthy, P.J.; Fiorillo, A.R. (2013).
5125:
5123:
5121:
5119:
313:
295:
256:
217:
212:
194:
180:
175:
165:
155:
147:
137:
127:
21:
5416:Fossiliferous stratigraphic units of North America
752:A new species of dromaeosaurid closely related to
462:. Concluded on the large amounts of dinosaurs and
938:are approximately 50% larger than specimens from
473:lacking ground ice with dinosaurs dominating and
4570:
4568:
4566:
4564:
4766:
4764:
4469:
4467:
4465:
4463:
4141:"A new species of the centrosaurine ceratopsid
3953:
3951:
3949:
4453:
4451:
4449:
4447:
4445:
4959:
4957:
4955:
4953:
4951:
4949:
4947:
8:
4634:
4632:
4630:
3918:Also found in the Schrader Bluff Formation.
3882:Also found in the Schrader Bluff Formation.
3846:Also found in the Schrader Bluff Formation.
3811:Also found in the Schrader Bluff Formation.
3784:Also found in the Schrader Bluff Formation.
3748:Also found in the Schrader Bluff Formation.
3686:Also found in the Schrader Bluff Formation.
3659:Also found in the Schrader Bluff Formation.
3632:Also found in the Schrader Bluff Formation.
3605:Also found in the Schrader Bluff Formation.
3570:Also found in the Schrader Bluff Formation.
3534:Also found in the Schrader Bluff Formation.
3462:Also found in the Schrader Bluff Formation.
3425:Also found in the Schrader Bluff Formation.
3390:Also found in the Schrader Bluff Formation.
3354:Also found in the Schrader Bluff Formation.
3327:Also found in the Schrader Bluff Formation.
3300:Also found in the Schrader Bluff Formation.
3264:Also found in the Schrader Bluff Formation.
3229:Also found in the Schrader Bluff Formation.
3193:Also found in the Schrader Bluff Formation.
3157:Also found in the Schrader Bluff Formation.
3121:Also found in the Schrader Bluff Formation.
3083:Also found in the Schrader Bluff Formation.
3046:Also found in the Schrader Bluff Formation.
2975:Also found in the Schrader Bluff Formation.
2939:Also found in the Schrader Bluff Formation.
2904:Also found in the Schrader Bluff Formation.
2868:Also found in the Schrader Bluff Formation.
2833:Also found in the Schrader Bluff Formation.
2798:Also found in the Schrader Bluff Formation.
2762:Also found in the Schrader Bluff Formation.
2727:Also found in the Schrader Bluff Formation.
2701:Also found in the Schrader Bluff Formation.
2666:Also found in the Schrader Bluff Formation.
2631:Also found in the Schrader Bluff Formation.
2596:Also found in the Schrader Bluff Formation.
2560:Also found in the Schrader Bluff Formation.
2524:Also found in the Schrader Bluff Formation.
2488:Also found in the Schrader Bluff Formation.
2453:Also found in the Schrader Bluff Formation.
2390:Also found in the Schrader Bluff Formation.
2319:Also found in the Schrader Bluff Formation.
2290:Also found in the Schrader Bluff Formation.
2255:Also found in the Schrader Bluff Formation.
2219:Also found in the Schrader Bluff Formation.
2184:Also found in the Schrader Bluff Formation.
2113:Also found in the Schrader Bluff Formation.
2087:Also found in the Schrader Bluff Formation.
2061:Also found in the Schrader Bluff Formation.
2034:Also found in the Schrader Bluff Formation.
1998:Also found in the Schrader Bluff Formation.
1962:Also found in the Schrader Bluff Formation.
1925:Also found in the Schrader Bluff Formation.
1273:Disassociated parts from multiple juveniles
4134:
4132:
4130:
4128:
4126:
4124:
4122:
1278:Originally identified as a distinct genus (
4820:
4818:
4816:
4814:
4812:
4810:
4808:
4806:
1515:Most common in the Prince Creek Formation
1236:Remains of adult and juvenile individuals
466:, the Prince Creek Formation was likely a
116:
18:
5396:Sandstone formations of the United States
5099:
4927:
4917:
4804:
4802:
4800:
4798:
4796:
4794:
4792:
4790:
4788:
4786:
4744:
4726:
4610:
4600:
4348:
4301:U.S. Geological Survey Professional Paper
4160:
4102:
4092:
4402:
4400:
4398:
4396:
4260:
4258:
4139:Fiorillo, A.R.; Tykoski, R.S.T. (2012).
4049:
3934:List of dinosaur-bearing rock formations
1685:
1682:
1376:
1059:
1056:
591:
588:
4394:
4392:
4390:
4388:
4386:
4384:
4382:
4380:
4378:
4376:
4176:
4174:
4172:
4047:
4045:
4043:
4041:
4039:
4037:
4035:
4033:
4031:
4029:
3962:. In Driese, S.G.; Nordt, L.C. (eds.).
3945:
1112:A squamosal, and the back of the dome.
393:of the Liscomb Bonebed in their habitat
4857:
4847:
3997:
3987:
946:. Remains were previously assigned to
5014:
5012:
4999:
4997:
4995:
4508:
4506:
3116:Specimens of preserved pollen spores
7:
5376:Maastrichtian Stage of North America
4318:
4316:
4314:
5213:Journal of Systematic Palaeontology
5161:Journal of Systematic Palaeontology
5130:Thurston, D.K.; Fujita, K. (1994).
4477:Journal of Systematic Palaeontology
3966:. Vol. 104. pp. 179–230.
1669:Uncertain or tentative taxa are in
1043:Uncertain or tentative taxa are in
747:Small dentary tip from a juvenile.
575:Uncertain or tentative taxa are in
5029:Journal of Vertebrate Paleontology
4966:Canadian Journal of Earth Sciences
4660:Journal of Vertebrate Paleontology
4519:Journal of Vertebrate Paleontology
4416:Journal of Vertebrate Paleontology
14:
1629:Taxon falsely reported as present
1207:Remains previously attributed to
1003:Taxon falsely reported as present
535:Taxon falsely reported as present
3467:
1324:
1295:
1282:), recent studies have found it
1216:
1176:
1126:
955:
882:
838:
763:
742:Pediomys Point - Liscomb Quarry
661:
345:with strata dating to the Early
318:
300:
1634:Dubious taxon or junior synonym
1008:Dubious taxon or junior synonym
540:Dubious taxon or junior synonym
4680:10.1080/02724634.2009.10010370
1383:of the Prince Creek Formation
697:Footprints from a small bird.
1:
5381:Cretaceous–Paleogene boundary
5361:Geologic formations of Alaska
5233:10.1080/14772019.2018.1560369
5181:10.1080/14772019.2023.2232359
5037:10.1080/02724634.2024.2326644
4149:Acta Palaeontologica Polonica
1690:of the Prince Creek Formation
1064:of the Prince Creek Formation
4919:10.1371/journal.pone.0065802
4728:10.1371/journal.pone.0024487
4602:10.1371/journal.pone.0235078
4539:10.1080/02724634.2013.770750
4497:10.1080/14772019.2010.509356
4094:10.1371/journal.pone.0091287
3493:Bissacate gymnosperm pollen
258:Approximate paleocoordinates
3041:Samples of distinct pollen
2899:Samples of distinct pollen
2863:Samples of distinct pollen
2828:Samples of distinct pollen
2793:Samples of distinct pollen
2757:Samples of distinct pollen
2722:Samples of distinct pollen
2696:Samples of distinct pollen
2661:Samples of distinct pollen
2626:Samples of distinct pollen
2591:Samples of distinct pollen
1596:
970:
732:Saurornitholestinae indet.
502:
401:phase. The rocks in it are
5437:
5084:10.1038/s41598-019-41325-8
4886:Pachyrhinosaurus perotorum
4842:Geologic Studies in Alaska
3836:Coleville River Formation
3380:Coleville Bluff Formation
1169:Pachyrhinosaurus perotorum
4350:10.1016/j.cub.2021.05.041
3755:
3577:
3349:Fossilized pollen spores
3271:
3152:Preserved pollen samples
3006:Preserved pollen samples
2673:
2397:
2005:
1603:
1599:
1379:
977:
973:
509:
505:
439:Wodehouseia edmontonicola
362:years ago, in the latest
115:
26:
3295:Distinct pollen remains
1885:Schrader Bluff Formation
1601:
1121:from Alaska discovered.
975:
507:
170:Schrader Bluff Formation
4194:Journal of Paleontology
1811:An aquatic angiosperm.
1397:Stratigraphic position
1286:indistinguishable from
160:Sagavanirktok Formation
5421:Paleontology in Alaska
5286:10.2110/sepmsp.097.233
3908:Coleville River Bluff
3872:Coleville River Bluff
3801:Coleville River Bluff
3774:Coleville River Bluff
3738:Coleville River Bluff
3702:Coleville River Bluff
3676:Coleville River Bluff
3649:Coleville River Bluff
3622:Coleville River Bluff
3595:Coleville River Bluff
3560:Coleville River Bluff
3524:Coleville River Bluff
3488:Coleville River Bluff
3452:Coleville River Bluff
3415:Coleville River Bluff
3344:Coleville River Bluff
3317:Coleville River Bluff
3290:Coleville River Bluff
3254:Coleville River Bluff
3219:Coleville River Bluff
3183:Coleville River Bluff
3147:Coleville River Bluff
3111:Coleville River Bluff
3073:Coleville River Bluff
3036:Coleville River Bluff
3001:Coleville River Bluff
2965:Coleville River Bluff
2929:Coleville River Bluff
2894:Coleville River Bluff
2858:Coleville River Bluff
2823:Coleville River Bluff
2788:Coleville River Bluff
2752:Coleville River Bluff
2717:Coleville River Bluff
2691:Coleville River Bluff
2656:Coleville River Bluff
2621:Coleville River Bluff
2586:Coleville River Bluff
2550:Coleville River Bluff
2514:Coleville River Bluff
2478:Coleville River Bluff
2443:Coleville River Bluff
2416:Coleville River Bluff
2380:Coleville River Bluff
2345:Coleville River Bluff
2314:Septate fungal hypha.
2309:Coleville River Bluff
2280:Coleville River Bluff
2245:Coleville River Bluff
2209:Coleville River Bluff
2174:Coleville River Bluff
2138:Coleville River Bluff
2103:Coleville River Bluff
2077:Coleville River Bluff
2051:Coleville River Bluff
2024:Coleville River Bluff
1988:Coleville River Bluff
1952:Coleville River Bluff
1915:Coleville River Bluff
1873:Coleville River Bluff
1677:taxa are discredited.
1268:Kikak-Tegoseak Quarry
1153:Kikak-Tegoseak Quarry
1051:taxa are discredited.
912:Kikak-Tegoseak Quarry
790:Kikak-Tegoseak Quarry
641:Kikak-Tegoseak Quarry
583:taxa are discredited.
394:
374:, based on the age of
335:Prince Creek Formation
22:Prince Creek Formation
5325:10.2110/palo.2013.133
5023:Edmontosaurus regalis
4162:10.4202/app.2011.0033
3972:10.2110/sepmsp.104.14
3529:Pollen spore remains
1937:Polycingulatisporites
1779:, known from leaves.
799:and the front of the
717:Distal metatarsal IV
486:Vertebrate paleofauna
389:
4335:(16): 3469–3478.e5.
3095:Converrucosisporites
1574:Lower Maastrichtian
1512:Lower Maastrichtian
1489:Lower Maastrichtian
1461:Lower Maastrichtian
1429:Lower Maastrichtian
1163:The youngest of the
921:Magical Mystery Bar
370:, at the end of the
132:Geological formation
16:Geological formation
5401:Mudstone formations
5317:2014Palai..29..594F
5225:2019JSPal..17.1805E
5173:2023JSPal..2132359E
5076:2019NatSR...9.5384T
4978:2011CaJES..48.1342B
4910:2013PLoSO...865802F
4834:Hamilton, Thomas D.
4719:2011PLoSO...624487Z
4672:2009JVPal..29..178F
4593:2020PLoSO..1535078C
4531:2013JVPal..33.1169W
4489:2011JSPal...9...33F
4341:2021CBio...31E3469D
4241:1985Geo....13..770M
4085:2014PLoSO...991287F
3275:Cicatricosisporites
2923:E. procumbentformis
1119:pachycephalosaurine
815:closely related to
276: /
237: /
28:Stratigraphic range
5064:Scientific Reports
3565:Carbonized pollen
3401:Camarozonosporites
2203:M. pseudosenonicus
1883:Also found in the
1859:Pulcheripollenites
1837:Kogosukruk Tongue
1804:Kogosukruk Tongue
1765:Kogosukruk Tongue
1733:Kogosukruk Tongue
1624:Reclassified taxon
1577:Over 60 specimens
1107:Kogosukruk Tongue
998:Reclassified taxon
530:Reclassified taxon
450:, and pollen from
395:
203:carbonaceous shale
5366:Cretaceous Alaska
5219:(21): 1805–1824.
4844:. pp. 47–48.
4838:Galloway, John P.
4274:Houseknecht, D.W.
3925:
3924:
3420:Pollen specimens
3322:Preserved pollen
2950:Dictyophyllidites
2401:Laegivatosporites
1681:
1680:
1672:
1660:
1659:
1656:
1649:
1642:
1635:
1630:
1625:
1620:
1590:
1589:
1370:
1369:
1317:A supraoccipital
1227:Leptoceratopsidae
1055:
1054:
1046:
1034:
1033:
1030:
1023:
1016:
1009:
1004:
999:
994:
964:
963:
851:Saurornitholestes
755:Saurornitholestes
706:Ornithomimosauria
649:Fossilized teeth
632:D. cf albertensis
587:
586:
578:
566:
565:
562:
555:
548:
541:
536:
531:
526:
331:
330:
5428:
5411:Fluvial deposits
5406:Deltaic deposits
5386:Paleogene Alaska
5336:
5299:
5271:
5253:
5252:
5207:
5201:
5200:
5152:
5146:
5145:
5127:
5114:
5113:
5103:
5055:
5049:
5048:
5016:
5007:
5001:
4990:
4989:
4972:(9): 1342–1354.
4961:
4942:
4941:
4931:
4921:
4890:Pachyrhinosaurus
4879:
4866:
4865:
4859:
4855:
4853:
4845:
4822:
4781:
4780:
4768:
4759:
4758:
4748:
4730:
4698:
4692:
4691:
4636:
4625:
4624:
4614:
4604:
4572:
4559:
4558:
4525:(5): 1169–1175.
4510:
4501:
4500:
4471:
4458:
4455:
4440:
4439:
4404:
4371:
4370:
4352:
4320:
4309:
4308:
4298:
4282:Brizzolara, D.W.
4262:
4253:
4252:
4224:
4218:
4217:
4178:
4167:
4166:
4164:
4143:Pachyrhinosaurus
4136:
4117:
4116:
4106:
4096:
4051:
4024:
4012:
4006:
4005:
3999:
3995:
3993:
3985:
3955:
3581:Aquilapollenites
3471:
2685:H. scollardensis
2650:I. marylandensis
2230:Lycopodiacidites
1683:
1670:
1652:
1645:
1638:
1633:
1628:
1623:
1616:
1613:
1597:
1543:Upper Campanian
1479:Multituberculata
1437:multituberculate
1377:
1328:
1314:Liscomb Bonebed
1299:
1220:
1188:Thescelosaurinae
1180:
1165:Pachyrhinosaurus
1139:Pachyrhinosaurus
1130:
1057:
1044:
1026:
1019:
1012:
1007:
1002:
997:
990:
987:
971:
959:
886:
859:S. cf. langstoni
842:
767:
665:
589:
576:
558:
551:
544:
539:
534:
529:
522:
519:
503:
337:is a geological
324:
322:
321:
306:
304:
303:
291:
290:
288:
287:
286:
281:
277:
274:
273:
272:
269:
252:
251:
249:
248:
247:
242:
238:
235:
234:
233:
230:
120:
111:
41:
19:
5436:
5435:
5431:
5430:
5429:
5427:
5426:
5425:
5371:Campanian Stage
5351:
5350:
5311:(11): 594–611.
5302:
5296:
5274:
5265:
5262:
5257:
5256:
5209:
5208:
5204:
5154:
5153:
5149:
5142:
5129:
5128:
5117:
5057:
5056:
5052:
5018:
5017:
5010:
5002:
4993:
4986:10.1139/e11-017
4963:
4962:
4945:
4881:
4880:
4869:
4856:
4846:
4824:
4823:
4784:
4770:
4769:
4762:
4700:
4699:
4695:
4638:
4637:
4628:
4587:(7): e0235078.
4574:
4573:
4562:
4512:
4511:
4504:
4473:
4472:
4461:
4456:
4443:
4406:
4405:
4374:
4329:Current Biology
4322:
4321:
4312:
4296:
4290:Takahashi, K.I.
4264:
4263:
4256:
4226:
4225:
4221:
4200:(5): 997–1001.
4180:
4179:
4170:
4138:
4137:
4120:
4055:Fiorillo, A. R.
4053:
4052:
4027:
4013:
4009:
3996:
3986:
3982:
3957:
3956:
3947:
3942:
3930:
3857:Aequitriradites
3759:Anacololsidites
3696:A. amygdaloides
3259:Pollen samples
3224:Pollen remains
3188:Pollen samples
3132:Clavatisporites
3078:Pollen samples
2970:Pollen remains
2915:Erdtmannipollis
2738:Gleicheniidites
2555:Pollen remains
2519:Pollen remains
2483:Pollen remains
2472:K. trispissatus
2448:Pollen remains
2421:Pollen remains
2385:Pollen remains
2350:Pollen remains
2285:Pollen remains
2250:Pollen remains
2214:Pollen remains
2179:Pollen remains
2159:Mantonisporites
2143:Pollen remains
1668:
1611:
1595:
1571:Pediomys Point
1492:Isolated teeth
1464:Isolated teeth
1432:Isolated teeth
1375:
1284:ontogenetically
1267:
1266:Liscomb Bonebed
1042:
985:
969:
915:Liscomb Quarry
865:Old Bone Beach
714:Old Bone Beach
682:G. vegrandiunis
638:Liscomb Quarry
574:
517:
498:
493:
488:
436:, projectates,
384:
359:
351:Late Cretaceous
319:
317:
301:
299:
284:
282:
278:
275:
270:
267:
265:
263:
262:
245:
243:
239:
236:
231:
228:
226:
224:
223:
123:
110:
109:
108:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
37:70.6–69.1
36:
35:
34:
17:
12:
11:
5:
5434:
5432:
5424:
5423:
5418:
5413:
5408:
5403:
5398:
5393:
5388:
5383:
5378:
5373:
5368:
5363:
5353:
5352:
5349:
5348:
5337:
5300:
5294:
5272:
5261:
5258:
5255:
5254:
5202:
5147:
5141:978-1125448038
5140:
5115:
5050:
5008:
4991:
4943:
4867:
4858:|journal=
4782:
4760:
4693:
4666:(1): 178–187.
4652:Mccarthy, P.J.
4640:Fiorillo, A.R.
4626:
4560:
4502:
4459:
4441:
4422:(4): 675–682.
4412:Gangloff, R.A.
4408:Fiorillo, A.R.
4372:
4310:
4278:Stricker, G.D.
4254:
4219:
4186:Fiorillo, A.R.
4182:Gangloff, R.A.
4168:
4155:(3): 561–573.
4118:
4059:Tykoski, R. S.
4025:
4007:
3998:|journal=
3980:
3944:
3943:
3941:
3938:
3937:
3936:
3929:
3926:
3923:
3922:
3920:
3915:
3910:
3905:
3897:
3893:Aequitriradite
3887:
3886:
3884:
3879:
3874:
3869:
3861:
3851:
3850:
3848:
3843:
3838:
3833:
3826:
3816:
3815:
3813:
3808:
3803:
3798:
3789:
3788:
3786:
3781:
3776:
3771:
3763:
3753:
3752:
3750:
3745:
3740:
3735:
3727:
3717:
3716:
3714:
3709:
3704:
3699:
3691:
3690:
3688:
3683:
3678:
3673:
3664:
3663:
3661:
3656:
3651:
3646:
3637:
3636:
3634:
3629:
3624:
3619:
3610:
3609:
3607:
3602:
3597:
3592:
3585:
3575:
3574:
3572:
3567:
3562:
3557:
3549:
3539:
3538:
3536:
3531:
3526:
3521:
3513:
3509:Balmeisporites
3503:
3502:
3500:
3495:
3490:
3485:
3483:Indeterminate
3480:
3478:Indeterminate
3474:
3473:
3464:
3459:
3454:
3449:
3440:
3430:
3429:
3427:
3422:
3417:
3412:
3405:
3395:
3394:
3392:
3387:
3385:Pollen spores
3382:
3377:
3369:
3359:
3358:
3356:
3351:
3346:
3341:
3332:
3331:
3329:
3324:
3319:
3314:
3305:
3304:
3302:
3297:
3292:
3287:
3279:
3269:
3268:
3266:
3261:
3256:
3251:
3244:
3240:Cingulizonates
3234:
3233:
3231:
3226:
3221:
3216:
3208:
3198:
3197:
3195:
3190:
3185:
3180:
3172:
3162:
3161:
3159:
3154:
3149:
3144:
3136:
3126:
3125:
3123:
3118:
3113:
3108:
3099:
3088:
3087:
3085:
3080:
3075:
3070:
3062:
3051:
3050:
3048:
3043:
3038:
3033:
3026:
3016:
3015:
3013:
3008:
3003:
2998:
2990:
2980:
2979:
2977:
2972:
2967:
2962:
2954:
2944:
2943:
2941:
2936:
2934:Pollen spores
2931:
2926:
2919:
2909:
2908:
2906:
2901:
2896:
2891:
2883:
2879:Expressipollis
2873:
2872:
2870:
2865:
2860:
2855:
2848:
2838:
2837:
2835:
2830:
2825:
2820:
2813:
2809:Foraminisporis
2803:
2802:
2800:
2795:
2790:
2785:
2777:
2767:
2766:
2764:
2759:
2754:
2749:
2742:
2732:
2731:
2729:
2724:
2719:
2714:
2706:
2705:
2703:
2698:
2693:
2688:
2681:
2671:
2670:
2668:
2663:
2658:
2653:
2646:
2642:Impardecispora
2636:
2635:
2633:
2628:
2623:
2618:
2611:
2607:Inundatisporis
2601:
2600:
2598:
2593:
2588:
2583:
2575:
2571:Ischyosporites
2565:
2564:
2562:
2557:
2552:
2547:
2539:
2535:Ischyosporites
2529:
2528:
2526:
2521:
2516:
2511:
2503:
2493:
2492:
2490:
2485:
2480:
2475:
2468:
2458:
2457:
2455:
2450:
2445:
2440:
2431:
2430:
2428:
2423:
2418:
2413:
2405:
2395:
2394:
2392:
2387:
2382:
2377:
2370:
2360:
2359:
2357:
2352:
2347:
2342:
2334:
2324:
2323:
2321:
2316:
2311:
2306:
2304:Indeterminate
2301:
2299:Indeterminate
2295:
2294:
2292:
2287:
2282:
2277:
2270:
2260:
2259:
2257:
2252:
2247:
2242:
2234:
2224:
2223:
2221:
2216:
2211:
2206:
2199:
2189:
2188:
2186:
2181:
2176:
2171:
2163:
2153:
2152:
2150:
2145:
2140:
2135:
2128:
2124:Osmundacidites
2118:
2117:
2115:
2110:
2108:Pollen spores
2105:
2100:
2092:
2091:
2089:
2084:
2082:Pollen spores
2079:
2074:
2066:
2065:
2063:
2058:
2056:Pollen spores
2053:
2048:
2039:
2038:
2036:
2031:
2029:Pollen spores
2026:
2021:
2013:
2003:
2002:
2000:
1995:
1993:Pollen spores
1990:
1985:
1977:
1973:Podocarpidites
1967:
1966:
1964:
1959:
1957:Pollen spores
1954:
1949:
1941:
1930:
1929:
1927:
1922:
1920:Pollen spores
1917:
1912:
1903:
1892:
1891:
1889:
1880:
1878:Pollen spores
1875:
1870:
1863:
1853:
1852:
1850:
1841:
1839:
1834:
1826:
1816:
1815:
1813:
1808:
1806:
1801:
1794:
1784:
1783:
1781:
1772:
1767:
1762:
1759:H. quercifolia
1755:
1745:
1744:
1742:
1737:
1735:
1730:
1723:
1713:
1712:
1709:
1706:
1703:
1700:
1697:
1693:
1692:
1679:
1678:
1663:
1661:
1658:
1657:
1650:
1643:
1636:
1631:
1626:
1621:
1605:
1604:
1602:
1600:
1594:
1591:
1588:
1587:
1585:
1578:
1575:
1572:
1569:
1564:
1556:
1555:
1553:
1546:
1544:
1541:
1538:Colville River
1534:
1529:
1521:
1520:
1518:
1516:
1513:
1510:
1508:
1507:Indeterminate
1505:
1498:
1497:
1495:
1493:
1490:
1487:
1485:
1484:Indeterminate
1482:
1475:
1474:
1472:
1465:
1462:
1459:
1457:
1451:
1443:
1442:
1440:
1433:
1430:
1427:
1425:
1416:
1408:
1407:
1404:
1401:
1398:
1395:
1392:
1389:
1385:
1384:
1374:
1371:
1368:
1367:
1365:
1361:Thescelosaurus
1352:
1347:
1345:
1343:Indeterminate
1340:
1331:
1330:
1321:
1318:
1315:
1312:
1311:Indeterminate
1309:
1306:Lambeosaurinae
1302:
1301:
1292:
1275:
1270:
1263:
1252:
1242:
1241:
1239:
1237:
1234:
1232:
1231:Indeterminate
1229:
1223:
1222:
1213:
1209:Thescelosaurus
1204:
1199:
1197:
1195:Indeterminate
1192:
1183:
1182:
1173:
1160:
1155:
1150:
1143:
1133:
1132:
1123:
1114:
1109:
1104:
1097:
1087:
1086:
1083:
1080:
1077:
1074:
1071:
1067:
1066:
1062:Ornithischians
1053:
1052:
1037:
1035:
1032:
1031:
1024:
1017:
1010:
1005:
1000:
995:
979:
978:
976:
974:
968:
967:Ornithischians
965:
962:
961:
952:
928:
923:
909:
899:
889:
888:
879:
872:
867:
862:
855:
845:
844:
835:
805:
792:
787:
780:
770:
769:
760:
749:
744:
739:
737:Indeterminate
734:
728:
727:
725:
718:
715:
712:
711:Indeterminate
709:
702:
701:
699:
694:
692:
685:
678:
668:
667:
658:
651:
646:
635:
628:
618:
617:
614:
611:
608:
605:
602:
598:
597:
585:
584:
569:
567:
564:
563:
556:
549:
542:
537:
532:
527:
511:
510:
508:
506:
497:
494:
492:
489:
487:
484:
383:
380:
358:
355:
329:
328:
315:
311:
310:
297:
293:
292:
280:83.2°N 115.9°W
260:
254:
253:
241:70.0°N 151.5°W
221:
215:
214:
210:
209:
196:
192:
191:
182:
178:
177:
173:
172:
167:
163:
162:
157:
153:
152:
149:
145:
144:
142:Colville Group
139:
135:
134:
129:
125:
124:
121:
113:
112:
105:
104:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
42:
24:
23:
15:
13:
10:
9:
6:
4:
3:
2:
5433:
5422:
5419:
5417:
5414:
5412:
5409:
5407:
5404:
5402:
5399:
5397:
5394:
5392:
5389:
5387:
5384:
5382:
5379:
5377:
5374:
5372:
5369:
5367:
5364:
5362:
5359:
5358:
5356:
5346:
5345:0-520-24209-2
5342:
5338:
5334:
5330:
5326:
5322:
5318:
5314:
5310:
5306:
5301:
5297:
5295:9781565763074
5291:
5287:
5283:
5279:
5273:
5269:
5264:
5263:
5259:
5250:
5246:
5242:
5238:
5234:
5230:
5226:
5222:
5218:
5214:
5206:
5203:
5198:
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5186:
5182:
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5170:
5166:
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5158:
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5148:
5143:
5137:
5133:
5126:
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5120:
5116:
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5107:
5102:
5097:
5093:
5089:
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5077:
5073:
5069:
5065:
5061:
5054:
5051:
5046:
5042:
5038:
5034:
5030:
5026:
5024:
5015:
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5009:
5005:
5000:
4998:
4996:
4992:
4987:
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4975:
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4960:
4958:
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4954:
4952:
4950:
4948:
4944:
4939:
4935:
4930:
4925:
4920:
4915:
4911:
4907:
4904:(6): e65802.
4903:
4899:
4898:
4893:
4891:
4887:
4884:"An Immature
4878:
4876:
4874:
4872:
4868:
4863:
4851:
4843:
4839:
4835:
4831:
4830:Parrish, J.T.
4827:
4821:
4819:
4817:
4815:
4813:
4811:
4809:
4807:
4805:
4803:
4801:
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4761:
4756:
4752:
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4738:
4734:
4729:
4724:
4720:
4716:
4713:(9): e24487.
4712:
4708:
4704:
4697:
4694:
4689:
4685:
4681:
4677:
4673:
4669:
4665:
4661:
4657:
4653:
4649:
4645:
4644:Tykoski, R.S.
4641:
4635:
4633:
4631:
4627:
4622:
4618:
4613:
4608:
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4598:
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4509:
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4503:
4498:
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4460:
4454:
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4437:
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4425:
4421:
4417:
4413:
4409:
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4399:
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4389:
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4364:
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4330:
4326:
4319:
4317:
4315:
4311:
4306:
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4295:
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4283:
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4275:
4271:
4267:
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4250:
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4234:
4230:
4223:
4220:
4215:
4211:
4207:
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4177:
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4169:
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4144:
4135:
4133:
4131:
4129:
4127:
4125:
4123:
4119:
4114:
4110:
4105:
4100:
4095:
4090:
4086:
4082:
4079:(3): e91287.
4078:
4074:
4073:
4068:
4064:
4063:Dodson, Peter
4060:
4056:
4050:
4048:
4046:
4044:
4042:
4040:
4038:
4036:
4034:
4032:
4030:
4026:
4022:
4021:0-520-24209-2
4018:
4011:
4008:
4003:
3991:
3983:
3981:9781565763227
3977:
3973:
3969:
3965:
3961:
3954:
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3950:
3946:
3939:
3935:
3932:
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3927:
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3916:
3914:
3911:
3909:
3906:
3904:
3902:
3898:
3896:
3895:
3894:
3889:
3888:
3885:
3883:
3880:
3878:
3875:
3873:
3870:
3868:
3867:
3866:A. spinulosus
3862:
3860:
3859:
3858:
3853:
3852:
3849:
3847:
3844:
3842:
3839:
3837:
3834:
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3831:
3827:
3825:
3824:
3823:
3818:
3817:
3814:
3812:
3809:
3807:
3804:
3802:
3799:
3797:
3795:
3791:
3790:
3787:
3785:
3782:
3780:
3777:
3775:
3772:
3770:
3768:
3764:
3762:
3761:
3760:
3754:
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3744:
3741:
3739:
3736:
3734:
3732:
3728:
3726:
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3719:
3718:
3715:
3713:
3710:
3708:
3705:
3703:
3700:
3698:
3697:
3693:
3692:
3689:
3687:
3684:
3682:
3679:
3677:
3674:
3672:
3671:
3666:
3665:
3662:
3660:
3657:
3655:
3652:
3650:
3647:
3645:
3643:
3639:
3638:
3635:
3633:
3630:
3628:
3625:
3623:
3620:
3618:
3616:
3612:
3611:
3608:
3606:
3603:
3601:
3598:
3596:
3593:
3591:
3590:
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3584:
3583:
3582:
3576:
3573:
3571:
3568:
3566:
3563:
3561:
3558:
3556:
3555:
3550:
3548:
3547:
3546:
3541:
3540:
3537:
3535:
3532:
3530:
3527:
3525:
3522:
3520:
3518:
3514:
3512:
3511:
3510:
3505:
3504:
3501:
3499:
3496:
3494:
3491:
3489:
3486:
3484:
3481:
3479:
3476:
3475:
3472:
3470:
3465:
3463:
3460:
3458:
3455:
3453:
3450:
3448:
3447:
3446:
3441:
3439:
3438:
3437:
3432:
3431:
3428:
3426:
3423:
3421:
3418:
3416:
3413:
3411:
3410:
3406:
3404:
3403:
3402:
3397:
3396:
3393:
3391:
3388:
3386:
3383:
3381:
3378:
3376:
3374:
3370:
3368:
3367:
3366:
3365:Cibotiumspora
3361:
3360:
3357:
3355:
3352:
3350:
3347:
3345:
3342:
3340:
3339:
3338:C. dorogensis
3334:
3333:
3330:
3328:
3325:
3323:
3320:
3318:
3315:
3313:
3311:
3307:
3306:
3303:
3301:
3298:
3296:
3293:
3291:
3288:
3286:
3284:
3280:
3278:
3277:
3276:
3270:
3267:
3265:
3262:
3260:
3257:
3255:
3252:
3250:
3249:
3245:
3243:
3242:
3241:
3236:
3235:
3232:
3230:
3227:
3225:
3222:
3220:
3217:
3215:
3214:
3209:
3207:
3206:
3205:
3204:Cingutriletes
3200:
3199:
3196:
3194:
3191:
3189:
3186:
3184:
3181:
3179:
3177:
3173:
3171:
3170:
3169:
3168:Circulodinium
3164:
3163:
3160:
3158:
3155:
3153:
3150:
3148:
3145:
3143:
3141:
3137:
3135:
3134:
3133:
3128:
3127:
3124:
3122:
3119:
3117:
3114:
3112:
3109:
3107:
3105:
3100:
3098:
3097:
3096:
3090:
3089:
3086:
3084:
3081:
3079:
3076:
3074:
3071:
3069:
3068:
3067:C. apisulacea
3063:
3061:
3059:
3058:
3053:
3052:
3049:
3047:
3044:
3042:
3039:
3037:
3034:
3032:
3031:
3027:
3025:
3024:
3023:
3018:
3017:
3014:
3012:
3009:
3007:
3004:
3002:
2999:
2997:
2995:
2991:
2989:
2988:
2987:
2986:Deltoidospora
2982:
2981:
2978:
2976:
2973:
2971:
2968:
2966:
2963:
2961:
2959:
2955:
2953:
2952:
2951:
2946:
2945:
2942:
2940:
2937:
2935:
2932:
2930:
2927:
2925:
2924:
2920:
2918:
2917:
2916:
2911:
2910:
2907:
2905:
2902:
2900:
2897:
2895:
2892:
2890:
2889:
2884:
2882:
2881:
2880:
2875:
2874:
2871:
2869:
2866:
2864:
2861:
2859:
2856:
2854:
2853:
2849:
2847:
2846:
2845:
2840:
2839:
2836:
2834:
2831:
2829:
2826:
2824:
2821:
2819:
2818:
2814:
2812:
2811:
2810:
2805:
2804:
2801:
2799:
2796:
2794:
2791:
2789:
2786:
2784:
2782:
2778:
2776:
2775:
2774:
2773:Foveosporites
2769:
2768:
2765:
2763:
2760:
2758:
2755:
2753:
2750:
2748:
2747:
2743:
2741:
2740:
2739:
2734:
2733:
2730:
2728:
2725:
2723:
2720:
2718:
2715:
2713:
2712:
2708:
2707:
2704:
2702:
2699:
2697:
2694:
2692:
2689:
2687:
2686:
2682:
2680:
2679:
2678:
2672:
2669:
2667:
2664:
2662:
2659:
2657:
2654:
2652:
2651:
2647:
2645:
2644:
2643:
2638:
2637:
2634:
2632:
2629:
2627:
2624:
2622:
2619:
2617:
2616:
2612:
2610:
2609:
2608:
2603:
2602:
2599:
2597:
2594:
2592:
2589:
2587:
2584:
2582:
2580:
2576:
2574:
2573:
2572:
2567:
2566:
2563:
2561:
2558:
2556:
2553:
2551:
2548:
2546:
2544:
2540:
2538:
2537:
2536:
2531:
2530:
2527:
2525:
2522:
2520:
2517:
2515:
2512:
2510:
2508:
2504:
2502:
2501:
2500:
2499:Integricorpus
2495:
2494:
2491:
2489:
2486:
2484:
2481:
2479:
2476:
2474:
2473:
2469:
2467:
2466:
2465:
2460:
2459:
2456:
2454:
2451:
2449:
2446:
2444:
2441:
2439:
2437:
2433:
2432:
2429:
2427:
2424:
2422:
2419:
2417:
2414:
2412:
2410:
2406:
2404:
2403:
2402:
2396:
2393:
2391:
2388:
2386:
2383:
2381:
2378:
2376:
2375:
2371:
2369:
2368:
2367:
2366:Lairidordites
2362:
2361:
2358:
2356:
2353:
2351:
2348:
2346:
2343:
2341:
2339:
2335:
2333:
2332:
2331:
2330:Leosphaeridia
2326:
2325:
2322:
2320:
2317:
2315:
2312:
2310:
2307:
2305:
2302:
2300:
2297:
2296:
2293:
2291:
2288:
2286:
2283:
2281:
2278:
2276:
2275:
2274:L. dissolutum
2271:
2269:
2268:
2267:
2262:
2261:
2258:
2256:
2253:
2251:
2248:
2246:
2243:
2241:
2239:
2235:
2233:
2232:
2231:
2226:
2225:
2222:
2220:
2217:
2215:
2212:
2210:
2207:
2205:
2204:
2200:
2198:
2197:
2196:
2191:
2190:
2187:
2185:
2182:
2180:
2177:
2175:
2172:
2170:
2168:
2164:
2162:
2161:
2160:
2155:
2154:
2151:
2149:
2146:
2144:
2141:
2139:
2136:
2134:
2133:
2129:
2127:
2126:
2125:
2120:
2119:
2116:
2114:
2111:
2109:
2106:
2104:
2101:
2099:
2098:
2094:
2093:
2090:
2088:
2085:
2083:
2080:
2078:
2075:
2073:
2072:
2068:
2067:
2064:
2062:
2059:
2057:
2054:
2052:
2049:
2047:
2045:
2041:
2040:
2037:
2035:
2032:
2030:
2027:
2025:
2022:
2020:
2018:
2014:
2012:
2011:
2010:
2004:
2001:
1999:
1996:
1994:
1991:
1989:
1986:
1984:
1982:
1978:
1976:
1975:
1974:
1969:
1968:
1965:
1963:
1960:
1958:
1955:
1953:
1950:
1948:
1947:
1942:
1940:
1939:
1938:
1932:
1931:
1928:
1926:
1923:
1921:
1918:
1916:
1913:
1911:
1909:
1904:
1902:
1901:
1900:
1899:Proteacidites
1894:
1893:
1890:
1888:
1886:
1881:
1879:
1876:
1874:
1871:
1869:
1868:
1864:
1862:
1861:
1860:
1855:
1854:
1851:
1849:
1847:
1842:
1840:
1838:
1835:
1833:
1831:
1827:
1825:
1824:
1823:
1818:
1817:
1814:
1812:
1809:
1807:
1805:
1802:
1800:
1799:
1795:
1793:
1792:
1791:
1786:
1785:
1782:
1780:
1778:
1773:
1771:
1768:
1766:
1763:
1761:
1760:
1756:
1754:
1753:
1752:
1747:
1746:
1743:
1741:
1738:
1736:
1734:
1731:
1729:
1728:
1724:
1722:
1721:
1720:
1715:
1714:
1710:
1707:
1704:
1701:
1698:
1695:
1694:
1691:
1689:
1684:
1676:
1667:
1664:
1662:
1655:
1651:
1648:
1644:
1641:
1637:
1632:
1627:
1622:
1619:
1615:
1614:
1610:
1607:
1606:
1598:
1592:
1586:
1583:
1579:
1576:
1573:
1570:
1568:
1567:U. hutchisoni
1565:
1563:
1562:
1558:
1557:
1554:
1551:
1547:
1545:
1542:
1539:
1535:
1533:
1530:
1528:
1527:
1523:
1522:
1519:
1517:
1514:
1511:
1509:
1506:
1503:
1500:
1499:
1496:
1494:
1491:
1488:
1486:
1483:
1480:
1477:
1476:
1473:
1470:
1466:
1463:
1460:
1458:
1455:
1452:
1450:
1449:
1445:
1444:
1441:
1438:
1434:
1431:
1428:
1426:
1424:
1420:
1417:
1415:
1414:
1410:
1409:
1405:
1402:
1399:
1396:
1393:
1390:
1387:
1386:
1382:
1378:
1372:
1366:
1364:
1362:
1358:
1353:
1351:
1348:
1346:
1344:
1341:
1339:
1337:
1333:
1332:
1329:
1327:
1322:
1319:
1316:
1313:
1310:
1307:
1304:
1303:
1300:
1298:
1293:
1291:
1289:
1288:Edmontosaurus
1285:
1281:
1276:
1274:
1271:
1269:
1264:
1262:
1261:
1257:
1253:
1251:
1250:
1249:
1248:Edmontosaurus
1244:
1243:
1240:
1238:
1235:
1233:
1230:
1228:
1225:
1224:
1221:
1219:
1214:
1212:
1210:
1205:
1203:
1200:
1198:
1196:
1193:
1191:
1189:
1185:
1184:
1181:
1179:
1174:
1172:
1170:
1166:
1161:
1159:
1156:
1154:
1151:
1149:
1148:
1144:
1142:
1141:
1140:
1135:
1134:
1131:
1129:
1124:
1122:
1120:
1115:
1113:
1110:
1108:
1105:
1103:
1102:
1098:
1096:
1095:
1094:
1093:Alaskacephale
1089:
1088:
1084:
1081:
1078:
1075:
1072:
1069:
1068:
1065:
1063:
1058:
1050:
1041:
1038:
1036:
1029:
1025:
1022:
1018:
1015:
1011:
1006:
1001:
996:
993:
989:
988:
984:
981:
980:
972:
966:
960:
958:
953:
951:
949:
945:
941:
937:
935:
929:
927:
924:
922:
919:
916:
913:
910:
908:
907:
905:
900:
898:
897:
896:
891:
890:
887:
885:
880:
877:
873:
871:
868:
866:
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860:
856:
854:
853:
852:
847:
846:
843:
841:
836:
834:
832:
831:
826:
825:
824:Tyrannosaurus
820:
819:
814:
813:tyrannosaurid
810:
806:
804:
802:
798:
793:
791:
788:
786:
785:
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772:
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639:
636:
634:
633:
629:
627:
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625:
624:Dromaeosaurus
620:
619:
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612:
609:
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582:
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490:
485:
483:
481:
480:Arctic Circle
476:
472:
469:
465:
461:
457:
453:
449:
445:
441:
440:
435:
432:
428:
424:
423:fungal hyphae
420:
416:
412:
408:
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400:
392:
388:
381:
379:
377:
373:
369:
365:
356:
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349:stage of the
348:
347:Maastrichtian
344:
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66:
61:
56:
51:
46:
40:
33:
32:Maastrichtian
29:
25:
20:
5391:Danian Stage
5308:
5304:
5277:
5267:
5260:Bibliography
5216:
5212:
5205:
5164:
5160:
5150:
5131:
5067:
5063:
5053:
5028:
5022:
4969:
4965:
4901:
4895:
4889:
4885:
4841:
4826:Spicer, R.A.
4776:
4772:
4710:
4706:
4696:
4663:
4659:
4648:Currie, P.J.
4584:
4580:
4522:
4518:
4483:(1): 33–49.
4480:
4476:
4419:
4415:
4332:
4328:
4304:
4300:
4286:Ryherd, T.J.
4266:Flores, R.M.
4232:
4228:
4222:
4197:
4193:
4190:Norton, D.W.
4152:
4148:
4142:
4076:
4070:
4010:
3963:
3917:
3912:
3907:
3900:
3899:
3891:
3890:
3881:
3876:
3871:
3865:
3863:
3855:
3854:
3845:
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3835:
3829:
3828:
3820:
3819:
3810:
3805:
3800:
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3792:
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3778:
3773:
3766:
3765:
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3756:
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3742:
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3730:
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3721:
3720:
3711:
3706:
3701:
3695:
3694:
3685:
3680:
3675:
3669:
3667:
3658:
3653:
3648:
3641:
3640:
3631:
3626:
3621:
3614:
3613:
3604:
3599:
3594:
3589:A. trialatus
3588:
3587:
3579:
3578:
3569:
3564:
3559:
3553:
3551:
3543:
3542:
3533:
3528:
3523:
3516:
3515:
3507:
3506:
3497:
3492:
3487:
3482:
3477:
3466:
3461:
3456:
3451:
3443:
3442:
3436:Botryococcus
3434:
3433:
3424:
3419:
3414:
3408:
3407:
3399:
3398:
3389:
3384:
3379:
3372:
3371:
3363:
3362:
3353:
3348:
3343:
3337:
3335:
3326:
3321:
3316:
3309:
3308:
3299:
3294:
3289:
3282:
3281:
3273:
3272:
3263:
3258:
3253:
3247:
3246:
3238:
3237:
3228:
3223:
3218:
3213:C. congruens
3212:
3210:
3202:
3201:
3192:
3187:
3182:
3175:
3174:
3166:
3165:
3156:
3151:
3146:
3139:
3138:
3130:
3129:
3120:
3115:
3110:
3103:
3101:
3093:
3091:
3082:
3077:
3072:
3066:
3064:
3055:
3054:
3045:
3040:
3035:
3029:
3028:
3020:
3019:
3010:
3005:
3000:
2993:
2992:
2984:
2983:
2974:
2969:
2964:
2957:
2956:
2948:
2947:
2938:
2933:
2928:
2922:
2921:
2913:
2912:
2903:
2898:
2893:
2888:E. accuratus
2887:
2885:
2877:
2876:
2867:
2862:
2857:
2852:F. scabratus
2851:
2850:
2844:Fibulapollis
2842:
2841:
2832:
2827:
2822:
2817:F. undulosus
2816:
2815:
2807:
2806:
2797:
2792:
2787:
2780:
2779:
2771:
2770:
2761:
2756:
2751:
2746:G. senonicus
2745:
2744:
2736:
2735:
2726:
2721:
2716:
2710:
2709:
2700:
2695:
2690:
2684:
2683:
2675:
2674:
2665:
2660:
2655:
2649:
2648:
2640:
2639:
2630:
2625:
2620:
2615:I. tappaniae
2614:
2613:
2605:
2604:
2595:
2590:
2585:
2578:
2577:
2569:
2568:
2559:
2554:
2549:
2542:
2541:
2533:
2532:
2523:
2518:
2513:
2506:
2505:
2497:
2496:
2487:
2482:
2477:
2471:
2470:
2462:
2461:
2452:
2447:
2442:
2435:
2434:
2425:
2420:
2415:
2408:
2407:
2399:
2398:
2389:
2384:
2379:
2373:
2372:
2364:
2363:
2354:
2349:
2344:
2337:
2336:
2328:
2327:
2318:
2313:
2308:
2303:
2298:
2289:
2284:
2279:
2273:
2272:
2266:Lunatadinium
2264:
2263:
2254:
2249:
2244:
2237:
2236:
2228:
2227:
2218:
2213:
2208:
2202:
2201:
2193:
2192:
2183:
2178:
2173:
2166:
2165:
2157:
2156:
2147:
2142:
2137:
2132:O. wellmanii
2131:
2130:
2122:
2121:
2112:
2107:
2102:
2096:
2095:
2086:
2081:
2076:
2070:
2069:
2060:
2055:
2050:
2043:
2042:
2033:
2028:
2023:
2016:
2015:
2007:
2006:
1997:
1992:
1987:
1980:
1979:
1971:
1970:
1961:
1956:
1951:
1945:
1943:
1935:
1933:
1924:
1919:
1914:
1907:
1905:
1897:
1895:
1882:
1877:
1872:
1866:
1865:
1857:
1856:
1843:
1836:
1829:
1828:
1820:
1819:
1810:
1803:
1797:
1796:
1788:
1787:
1774:
1769:
1764:
1758:
1757:
1749:
1748:
1739:
1732:
1727:P. wigginsii
1726:
1725:
1719:Parataxodium
1717:
1716:
1686:
1674:
1665:
1608:
1566:
1559:
1531:
1524:
1453:
1448:Gypsonictops
1446:
1422:
1418:
1411:
1360:
1357:Parksosaurus
1356:
1354:
1349:
1342:
1334:
1323:
1294:
1287:
1279:
1277:
1272:
1265:
1259:
1255:
1254:
1246:
1245:
1215:
1208:
1206:
1201:
1194:
1186:
1175:
1168:
1164:
1162:
1157:
1152:
1147:P. perotorum
1146:
1145:
1137:
1136:
1125:
1116:
1111:
1106:
1101:A. gangloffi
1100:
1099:
1091:
1090:
1060:
1048:
1039:
982:
954:
947:
933:
930:
925:
920:
917:
914:
911:
903:
901:
893:
892:
881:
869:
864:
858:
857:
849:
848:
837:
828:
822:
816:
808:
807:
794:
789:
783:
782:
774:
773:
762:
753:
751:
746:
741:
736:
731:
722:ornithomimid
720:Possibly an
696:
687:
681:
680:
672:
671:
660:
648:
643:
640:
637:
631:
630:
622:
621:
592:
580:
571:
514:
499:
437:
396:
391:Hadrosaurids
360:
334:
332:
285:83.2; -115.9
246:70.0; -151.5
5070:(1): 5384.
4270:Myers, M.D.
4235:(11): 770.
3723:Annulispora
3670:A. dentatus
3554:A. cribrata
3409:C. ambigens
3248:C. bialatus
3057:Crassispora
3030:C. fragilis
3022:Cycadopites
2677:Hannisporis
2464:Kurtzipites
2340:"stellata"
2195:Mancicorpus
2097:O. arcticus
1946:P. reduncus
1846:sphenophyte
1822:Equisetites
1798:Q. angulata
1675:crossed out
1654:Morphotaxon
1582:metatherian
1502:Marsupialia
1336:Ornithopoda
1049:crossed out
1028:Morphotaxon
948:T. formosus
931:Remains of
876:dromaeosaur
830:Tarbosaurus
809:Nanuqsaurus
784:N. hoglundi
776:Nanuqsaurus
689:Denali Park
655:dromaeosaur
581:crossed out
560:Morphotaxon
475:angiosperms
283: /
244: /
219:Coordinates
5355:Categories
4779:: 347–365.
3940:References
3830:A. clavate
3445:B. braunii
2169:sp. indet
1867:P. krempii
1777:angiosperm
1705:Abundance
1671:small text
1640:Ichnotaxon
1561:Unnuakomys
1400:Abundance
1350:One tooth
1280:Ugrunaaluk
1117:The first
1079:Abundance
1045:small text
1014:Ichnotaxon
918:Byers Bed
644:Byers Bed
610:Abundance
577:small text
546:Ichnotaxon
446:bisaccate
444:hinterland
399:greenhouse
5333:128713816
5241:1477-2019
5197:260668330
5189:1477-2019
5092:2045-2322
5045:0272-4634
4860:ignored (
4850:cite book
4737:1932-6203
4688:197535475
4656:Flaig, P.
4555:130049294
4547:0272-4634
4436:130766946
4359:0960-9822
4214:130669713
4000:ignored (
3990:cite book
2711:H. amplus
2374:L. magnus
2071:O. parvus
2009:Ovoidites
1790:Quereuxia
1751:Hollickia
1702:Location
1609:Color key
1550:eutherian
1532:S. mikros
1469:eutherian
1413:Cimolodon
1394:Location
1076:Location
983:Color key
818:Lythronax
801:lower jaw
607:Location
594:Theropods
515:Color key
496:Theropods
491:Dinosaurs
415:dinocysts
376:ostracods
368:Paleogene
364:Campanian
339:formation
199:siltstone
185:Sandstone
176:Lithology
156:Underlies
148:Sub-units
5249:92613824
5110:30926823
4938:23840371
4897:PLoS ONE
4840:(eds.).
4755:21949721
4707:PLOS ONE
4621:32639990
4581:PLOS ONE
4367:34171301
4292:(2007).
4113:24621577
4072:PLoS ONE
4061:(2014).
3928:See also
1699:Species
1580:A small
1526:Sikuomys
1467:A small
1435:A small
1391:Species
1073:Species
674:Gruipeda
604:Species
471:woodland
403:alluvial
271:115°54′W
232:151°30′W
213:Location
207:ash-fall
189:mudstone
166:Overlies
30:: Early
5313:Bibcode
5305:PALAIOS
5221:Bibcode
5169:Bibcode
5101:6440964
5072:Bibcode
4974:Bibcode
4929:3686821
4906:Bibcode
4746:3176273
4715:Bibcode
4668:Bibcode
4612:7343144
4589:Bibcode
4527:Bibcode
4485:Bibcode
4337:Bibcode
4237:Bibcode
4229:Geology
4104:3951350
4081:Bibcode
4065:(ed.).
3913:Pollen
3877:Pollen
3841:Pollen
3806:Pollen
3779:Pollen
3743:Pollen
3707:Pollen
3681:Pollen
3654:Pollen
3627:Pollen
3600:Pollen
3457:Pollen
1770:Leaves
1711:Images
1647:Ootaxon
1548:A tiny
1504:indet.
1481:indet.
1423:nitidus
1406:Images
1381:Mammals
1373:Mammals
1338:indet.
1308:indet.
1260:regalis
1190:indet.
1085:Images
1021:Ootaxon
944:Montana
940:Alberta
895:Troodon
797:maxilla
708:indet.
616:Images
553:Ootaxon
382:Habitat
314:Country
268:83°12′N
229:70°00′N
181:Primary
138:Unit of
106:↓
5343:
5331:
5292:
5270:: 311.
5247:
5239:
5195:
5187:
5138:
5108:
5098:
5090:
5043:
4936:
4926:
4753:
4743:
4735:
4686:
4619:
4609:
4553:
4545:
4434:
4365:
4357:
4212:
4111:
4101:
4019:
3978:
3769:sp. 1
3545:Azonia
3285:sp. 1
2019:? sp.
1708:Notes
1696:Genus
1688:Plants
1593:Plants
1536:Lower
1403:Notes
1388:Genus
1202:Teeth
1082:Notes
1070:Genus
870:Teeth
827:, and
613:Notes
601:Genus
458:, and
456:shrubs
448:pollen
434:spores
411:pyrite
407:gypsum
372:Danian
343:Alaska
323:
308:Alaska
305:
296:Region
5329:S2CID
5245:S2CID
5193:S2CID
5167:(1).
4684:S2CID
4551:S2CID
4432:S2CID
4307:: 52.
4297:(PDF)
4210:S2CID
3822:Alete
3617:spp.
3312:spp.
2996:spp.
2411:spp.
1666:Notes
1618:Taxon
1040:Notes
992:Taxon
811:is a
572:Notes
524:Taxon
468:polar
464:flora
460:herbs
452:trees
419:algae
195:Other
5341:ISBN
5290:ISBN
5237:ISSN
5185:ISSN
5136:ISBN
5106:PMID
5088:ISSN
5041:ISSN
4934:PMID
4862:help
4751:PMID
4733:ISSN
4617:PMID
4543:ISSN
4363:PMID
4355:ISSN
4305:1748
4109:PMID
4017:ISBN
4002:help
3976:ISBN
3903:sp.
3864:cf.
3796:sp.
3733:sp.
3668:cf.
3644:sp.
3552:cf.
3519:sp.
3375:sp.
3336:cf.
3211:cf.
3178:sp.
3142:sp.
3106:sp.
3102:cf.
3092:cf.
3065:cf.
2960:sp.
2886:cf.
2783:sp.
2581:sp.
2545:sp.
2509:sp.
2438:sp.
2240:sp.
2046:sp.
1983:sp.
1944:cf.
1934:cf.
1910:sp.
1906:cf.
1896:cf.
1832:sp.
1456:sp.
1421:cf.
1258:cf.
942:and
431:moss
429:and
427:fern
409:and
333:The
128:Type
45:PreꞒ
5321:doi
5282:doi
5229:doi
5177:doi
5096:PMC
5080:doi
5033:doi
4982:doi
4924:PMC
4914:doi
4741:PMC
4723:doi
4676:doi
4607:PMC
4597:doi
4535:doi
4493:doi
4424:doi
4345:doi
4245:doi
4202:doi
4157:doi
4099:PMC
4089:doi
3968:doi
1775:An
1359:or
936:sp.
357:Age
341:in
326:USA
5357::
5327:.
5319:.
5309:29
5307:.
5288:.
5243:.
5235:.
5227:.
5217:17
5215:.
5191:.
5183:.
5175:.
5165:21
5163:.
5159:.
5118:^
5104:.
5094:.
5086:.
5078:.
5066:.
5062:.
5039:.
5031:.
5027:.
5011:^
4994:^
4980:.
4970:48
4968:.
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4932:.
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4912:.
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4870:^
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3974:.
3948:^
3901:A.
3794:A.
3767:A.
3731:A.
3642:A.
3615:A.
3517:B.
3373:C.
3310:C.
3283:C.
3176:C.
3140:C.
3104:C.
3060:?
2994:D.
2958:D.
2579:I.
2543:I.
2507:I.
2436:L.
2409:L.
2338:L.
2238:L.
2167:M.
2044:O.
2017:O.
1981:P.
1908:P.
1887:.
1848:.
1844:A
1830:E.
1673:;
1584:.
1552:.
1540:.
1471:.
1454:G.
1439:.
1419:C.
1363:.
1290:.
1256:E.
1211:.
1047:;
934:T.
906:sp
904:T.
878:.
874:A
833:.
821:,
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758:.
724:.
657:.
653:A
579:;
454:,
442:,
425:,
421:,
417:,
205:,
201:,
187:,
95:Pg
39:Ma
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