133:
111:
1300:
401:
is known from a dozen specimens, all found in or near the type locality. Among them are several well-preserved elements, including a well-preserved skull, partial skulls and braincases, several vertebrae including an almost complete sacrum, a left innominate, ribs, and partial limb elements.
385:
Baloch tribe which assisted
Gingerich and his team during their field work. "Qaisra" is also etymologically close to the royal title used in Persian and many Indo-European languages (e.g. Kaiser, Czar, Caesar). The species is named for Muhammad Arif, former paleontologist at the
437:, the most complete articulated skeleton of a protocetid, and they were similar in size: the latter had an estimated body weight of 620 kg (1,370 lb), compared to 590 kg (1,300 lb) for the former.
688:
Gingerich, Philip D; Haq, U; Khan, H; Zalmout, S (2001). "Eocene stratigraphy and archaeocete whales (Mammalia, Cetacea) of Drug Lahar in the eastern
Sulaiman range, Balochistan (Pakistan)".
1393:
1380:
1452:
739:
1457:
464:
The four sacral vertebrae are only partially fused: the first two are solidly fused, the third is locked in place by rib-like processes (a
1462:
584:
387:
551:
indicate protocetids represent a wide range of specializations, although which is ancestral to later whales is unclear.
132:
732:
540:
566:
345:
1320:
314:
1447:
1419:
590:
572:
1367:
1303:
725:
543:
in retaining pleurapophyseal connections between all sacral vertebrae. The sacral morphology of
127:
1424:
1406:
1398:
863:
705:
472:
448:
110:
1411:
1358:
697:
900:
1216:
1117:
920:
465:
1441:
982:
971:
943:
872:
852:
829:
818:
468:
68:
1250:
1225:
1149:
1126:
1025:
1011:
1002:
957:
950:
911:
879:
532:
514:
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499:
447:
located relatively anteriorly (above C) and a relatively broad frontal shield. The
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220:
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1343:
1352:
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1232:
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1177:
1142:
1135:
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936:
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493:
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471:), while the fourth has a caudal (tail-like) morphology including two ventral
457:
432:
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410:
88:
53:
360:
347:
329:
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1273:
1205:
1102:
1053:
1046:
929:
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144:
93:
37:
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1337:
1170:
767:
701:
382:
308:
293:
164:
83:
78:
63:
58:
48:
1385:
1239:
1156:
519:
390:
who contributed significantly to archaeocete paleontology in
Pakistan.
194:
98:
73:
779:
761:
690:
Contributions from the Museum of
Paleontology, University of Michigan
289:
174:
154:
30:
1314:
531:
has a sacrum where non of the vertebrae have fused centra, which is
1372:
444:
1318:
721:
671:
659:
644:
629:
617:
605:
265:
240:
443:
has a generalized protocetid skull with the external
255:
1327:
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1215:
1125:
1116:
1001:
981:
919:
910:
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828:
794:
640:
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655:
653:
475:. The fusion between S1 and S2 distinguished
733:
8:
1315:
1122:
916:
740:
726:
718:
109:
20:
586:Ander Dabh Shumali (Eocene of Pakistan)
559:
7:
601:
599:
14:
674:, Diagnosis, p. 296, 303, Fig. 23
1299:
1298:
131:
479:from other protocetids such as
1:
1453:Fossil taxa described in 2001
388:Geological Survey of Pakistan
381:The genus is named after the
118:Drawing of basic outline of
1458:Prehistoric cetacean genera
1479:
1294:
756:
430:is almost as complete as
252:
247:
128:Scientific classification
126:
117:
108:
23:
535:to be a protocetid, but
1463:Extinct mammals of Asia
662:, Discussion, pp. 307–8
593:. Retrieved July 2013.
575:. Retrieved July 2013.
299:48.6 to 40.4
1420:Paleobiology Database
672:Gingerich et al. 2001
660:Gingerich et al. 2001
645:Gingerich et al. 2001
630:Gingerich et al. 2001
618:Gingerich et al. 2001
606:Gingerich et al. 2001
591:Paleobiology Database
573:Paleobiology Database
266:Gingerich et al. 2001
241:Gingerich et al. 2001
555:Notes and references
513:' vertebrae are not
647:, Diagnosis, p. 296
620:, Etymology, p. 298
608:, Etymology, p. 296
357: /
342:, paleocoordinates
326: /
632:, Hypodigm, p. 297
523:. In contrast to
451:is more narrow in
1435:
1434:
1407:Open Tree of Life
1321:Taxon identifiers
1312:
1311:
1290:
1289:
1112:
1111:
864:Remingtonocetidae
473:chevron processes
428:Qaisracetus arifi
301:million years ago
274:
273:
268:
243:
1470:
1428:
1427:
1415:
1414:
1402:
1401:
1389:
1388:
1376:
1375:
1363:
1362:
1361:
1348:
1347:
1346:
1316:
1302:
1301:
1123:
917:
751:genera by family
742:
735:
728:
719:
713:
675:
669:
663:
657:
648:
642:
633:
627:
621:
615:
609:
603:
594:
582:
576:
564:
420:but larger than
408:is smaller than
372:
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368:
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354:
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341:
340:
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264:
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219:
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136:
135:
113:
103:
40:
29:Temporal range:
21:
16:Genus of mammals
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1471:
1469:
1468:
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1438:
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1423:
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1392:
1384:
1379:
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1366:
1357:
1356:
1351:
1342:
1341:
1336:
1323:
1313:
1308:
1286:
1245:
1211:
1108:
997:
977:
906:
901:Remingtonocetus
858:
824:
790:
752:
746:
716:
696:(11): 269–319.
687:
683:
678:
670:
666:
658:
651:
643:
636:
628:
624:
616:
612:
604:
597:
583:
579:
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561:
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515:dense and thick
466:pleurapophyseal
396:
379:
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351:
348:
346:
344:
343:
334:
332:
328:
325:
320:
317:
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313:
312:
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288:known from the
263:
238:
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130:
104:
102:
101:
96:
91:
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66:
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51:
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36:48.6–40.4
35:
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27:
17:
12:
11:
5:
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1474:
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1403:
1390:
1377:
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1319:
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1309:
1307:
1306:
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1263:
1255:
1253:
1247:
1246:
1244:
1243:
1236:
1229:
1221:
1219:
1217:Basilosaurinae
1213:
1212:
1210:
1209:
1202:
1195:
1188:
1181:
1174:
1167:
1160:
1153:
1146:
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1129:
1120:
1118:Basilosauridae
1114:
1113:
1110:
1109:
1107:
1106:
1099:
1092:
1085:
1078:
1071:
1064:
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1005:
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985:
979:
978:
976:
975:
968:
961:
954:
947:
940:
933:
925:
923:
921:Georgiacetinae
914:
908:
907:
905:
904:
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890:
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868:
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664:
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558:
556:
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395:
392:
378:
375:
281:is an extinct
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1128:
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1016:
1014:
1013:
1009:
1008:
1006:
1004:
1000:
994:
993:
989:
988:
986:
984:
983:Makaracetinae
980:
974:
973:
972:Pontobasileus
969:
967:
966:
962:
960:
959:
955:
953:
952:
948:
946:
945:
944:Carolinacetus
941:
939:
938:
934:
932:
931:
927:
926:
924:
922:
918:
915:
913:
909:
903:
902:
898:
896:
895:
891:
889:
888:
884:
882:
881:
877:
875:
874:
873:Andrewsiphius
870:
869:
867:
865:
861:
855:
854:
853:Himalayacetus
850:
848:
847:
843:
841:
840:
836:
835:
833:
831:
830:Ambulocetidae
827:
821:
820:
819:Ichthyolestes
816:
814:
813:
809:
807:
806:
802:
801:
799:
797:
793:
787:
783:
781:
777:
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769:
765:
763:
759:
758:
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750:
743:
738:
736:
731:
729:
724:
723:
720:
711:
707:
703:
702:2027.42/48661
699:
695:
691:
686:
685:
680:
673:
668:
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661:
656:
654:
650:
646:
641:
639:
635:
631:
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623:
619:
614:
611:
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592:
588:
587:
581:
578:
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569:
563:
560:
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552:
550:
546:
542:
538:
534:
530:
526:
522:
521:
516:
512:
508:
507:
502:
501:
496:
495:
490:
489:
484:
483:
478:
474:
470:
469:synchondrosis
467:
462:
460:
459:
454:
450:
446:
442:
438:
436:
434:
429:
425:
424:
419:
418:
413:
412:
407:
403:
400:
393:
391:
389:
384:
376:
374:
370:
361:12.4°N 65.7°E
339:
330:31.0°N 70.2°E
310:
306:
300:
295:
291:
287:
284:
280:
279:
267:
262:
261:
254:
253:
251:
246:
242:
237:
236:
229:
226:
225:
222:
216:
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107:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
31:Middle Eocene
26:
22:
19:
1448:Protocetidae
1328:
1279:
1272:
1265:
1258:
1251:Pachycetinae
1238:
1231:
1226:Basilosaurus
1224:
1204:
1197:
1190:
1183:
1176:
1169:
1162:
1155:
1150:Cynthiacetus
1148:
1141:
1134:
1127:Dorudontinae
1101:
1094:
1087:
1081:
1080:
1073:
1066:
1059:
1052:
1045:
1038:
1031:
1026:Crenatocetus
1024:
1017:
1012:Aegyptocetus
1010:
1003:Protocetinae
990:
970:
963:
958:Natchitochia
956:
951:Georgiacetus
949:
942:
935:
928:
912:Protocetidae
899:
892:
885:
880:Attockicetus
878:
871:
851:
844:
837:
817:
810:
803:
786:Artiodactyla
693:
689:
681:Bibliography
667:
625:
613:
585:
580:
567:
562:
548:
544:
536:
528:
524:
518:
510:
506:Georgiacetus
504:
500:Natchitochia
498:
492:
486:
480:
476:
463:
456:
452:
440:
439:
431:
427:
421:
415:
409:
405:
404:
398:
397:
380:
277:
276:
275:
259:
258:
234:
233:
221:Protocetinae
208:Protocetidae
191:Infraorder:
185:Artiodactyla
119:
24:
18:
1359:Qaisracetus
1353:Wikispecies
1329:Qaisracetus
1281:Platyosphys
1233:Basiloterus
1199:Supayacetus
1178:Pontogeneus
1143:Chrysocetus
1136:Ancalecetus
1082:Qaisracetus
1068:Phiomicetus
1061:Peregocetus
992:Makaracetus
894:Kutchicetus
839:Ambulocetus
796:Pakicetidae
749:Archaeocete
568:Qaisracetus
549:Qaisracetus
525:Qaisracetus
511:Qaisracetus
477:Qaisracetus
453:Qaisracetus
441:Qaisracetus
406:Qaisracetus
399:Qaisracetus
394:Description
364: /
333: /
305:Baluchistan
286:early whale
278:Qaisracetus
235:Qaisracetus
214:Subfamily:
120:Qaisracetus
25:Qaisracetus
1442:Categories
1267:Pachycetus
1260:Antaecetus
1192:Stromerius
1185:Saghacetus
1164:Masracetus
1096:Takracetus
1089:Rodhocetus
1075:Protocetus
1040:Gaviacetus
1033:Dhedacetus
1019:Artiocetus
965:Pappocetus
937:Babiacetus
887:Dalanistes
846:Gandakasia
774:Vertebrata
772:Subphylum
545:Rodhocetus
537:Rodhocetus
529:Rodhocetus
494:Gaviacetus
488:Rodhocetus
482:Protocetus
458:Takracetus
433:Rodhocetus
417:Babiacetus
411:Pappocetus
366:12.4; 65.7
335:31.0; 70.2
283:protocetid
1344:Q16758029
1274:Perucetus
1206:Zygorhiza
1103:Togocetus
1054:Maiacetus
1047:Indocetus
930:Aegicetus
812:Nalacetus
805:Pakicetus
541:primitive
423:Indocetus
377:Etymology
151:Kingdom:
145:Eukaryota
1338:Wikidata
1304:Category
1171:Ocucajea
780:Mammalia
768:Chordata
762:Animalia
760:Kingdom
710:50061585
517:like in
455:than in
435:kasranii
383:Qaisrani
309:Pakistan
294:Lutetian
260:Q. arifi
248:Species
201:Family:
175:Mammalia
165:Chordata
161:Phylum:
155:Animalia
141:Domain:
1412:4942541
1399:1192853
1386:4832597
1373:4526555
1240:Eocetus
1157:Dorudon
766:Phylum
589:in the
571:in the
533:derived
520:Eocetus
449:rostrum
352:65°42′E
349:12°24′N
321:70°12′E
318:31°00′N
227:Genus:
195:Cetacea
181:Order:
171:Class:
784:Order
778:Class
708:
503:, and
290:Eocene
1425:53242
1394:IRMNG
445:nares
303:) of
1381:GBIF
706:OCLC
547:and
414:and
44:PreꞒ
1368:EoL
698:hdl
539:is
509:.
426:.
373:).
1444::
1422::
1409::
1396::
1383::
1370::
1355::
1340::
704:.
694:30
692:.
652:^
637:^
598:^
527:,
497:,
491:,
485:,
461:.
307:,
296:,
122:.
94:Pg
38:Ma
33:,
741:e
734:t
727:v
712:.
700::
311:(
292:(
256:†
231:†
218:†
205:†
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:Ꞓ
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
