292:). They sometimes spend their lives in just one tree, and in very large trees with complex structure many generations can live in different parts of the same tree. Red tree voles have been shown to have a home-range averaging around 800 m, though this area is somewhat smaller in old forest (<80 years) and larger in young forest. Within this home-range they can have nests in up to 6 different trees, although 2 nests are average for females and 3 nests are average for males.
272:
75:
50:
31:
308:
birth and it is common to find two separate litters living in the same nest. During peak breeding periods (winter and spring), males disperse and travel longer distances from home in search for females that are sexually receptive. Females tend to stay closer to their nests, therefore having smaller home ranges.
295:
When eating
Douglas-fir needles, they carefully remove the fine resin ducts (which resemble coarse, straight hairs) along each edge of the needle, which then become a major component of their nests (see image below). As well as these resin ducts, their nests are composed of small Douglas-fir cuttings
307:
Red tree voles have small litters consisting of 1 to 4 offspring and a long gestation period of 28 days compared to other
Microtines. Juveniles depend on their mothers for 47 to 60 days before complete independence. Females exhibit post-partum mating where they can breed within a few days of giving
337:
has determined their status as vulnerable on the list of sensitive species on the Oregon coast. They are vulnerable to timber harvest due to their arboreal lifestyle and their low reproductive rates contribute to the concern of the species becoming further threatened. They are an important food
267:
Red tree voles are about 6-8 in long, including the tail. When they are young, they exhibit a dull brown coat and develop a more reddish color with age. They are nocturnal and very difficult to see, but they can be detected by finding piles or wads of these resin ducts on the ground.
324:
are also another primary predator of the red tree vole. Females are easier to detect by predators due to their maternal nests. The time they spend outside of their nests searching for food to bring back for their young also puts them at a higher risk of predation.
550:
Miller, Mark P.; Forsman, Eric D.; Swingle, James K.; Miller, Stephanie A.; Haig, Susan M. (2010). "Size-Associated
Morphological Variation in the Red Tree Vole (Arborimus longicaudus)".
338:
source for the threatened
Northern Spotted Owl and have been listed as a "survey and manage species" under the Northwest Forest Plan (NWFP) in order to help manage the area's ecosystem.
3136:
296:
and their own scat. These nests are often built on or in large or epicormic branches, broken tops, cavities and split trunks, and are always within the live crown.
3281:
3214:
3301:
3110:
275:
The characteristic pile of discarded resin ducts (which run along the outside edges of
Douglas-fir needles) produced by a red tree vole when eating.
3149:
658:
Dunk, Jeffrey R.; Hawley, Jennifer J.V.G (24 April 2009). "Red-tree vole habitat suitability modeling: Implications for conservation management".
585:
Swingle, James K.; Foreman, Eric D. (2009). "Home Range Areas and
Activity Patterns of Red Tree Voles (Arborimus longicaudus) in Western Oregon".
244:. It is found only in coastal forests of Oregon and northern California. They feed exclusively on the needles of conifers, primarily Douglas-fir (
404:
Mammal
Species of the World a Taxonomic and Geographic Reference (D. E. Wilson and D. M. Reeder eds.). Johns Hopkins University Press, Baltimore.
623:
Swingle, James K.; Foreman, Eric D.; Anthony, Robert G. (2010). "Survival, Mortality, and
Predators of Red Tree Voles (Arborimus longicaudus)".
711:
425:
3291:
1943:
453:
Annotated
Bibliography of the Red Tree Vole (Arborimus Longicaudus), Sonoma Tree Vole (A. Pomo), and White-footed Vole (A. Albipes)
3175:
1952:
3193:
3154:
2418:
2117:
505:
Swingle James. Daily activity patterns, survival, and movements of red tree voles (Arborimus longicaudus) in western Oregon.
2685:
284:
Red tree voles live almost exclusively in
Douglas-fir trees, though they have occasionally been found in Sitka spruce and
2243:
1934:
1611:
1405:
3188:
3296:
3219:
2364:
1734:
1684:
74:
3075:
2934:
2658:
2615:
2465:
1970:
1916:
1804:
1794:
704:
2755:
2737:
2694:
2667:
2382:
2041:
2400:
2649:
2579:
1925:
1387:
815:
3037:
2588:
2552:
2234:
2225:
2358:
1203:
1161:
3253:
3012:
2409:
2108:
334:
179:
1459:
1275:
1019:
924:
3097:
2975:
2966:
2810:
2764:
2570:
2391:
2373:
2308:
2299:
1761:
1752:
1693:
1423:
1414:
1354:
1212:
915:
697:
519:
317:
1441:
1378:
1001:
942:
806:
470:
379:
39:
3206:
2479:
485:
3286:
2050:
2032:
1825:
1743:
1369:
1360:
1293:
1010:
824:
675:
640:
602:
567:
285:
69:
3258:
2441:
1524:
3141:
3240:
3084:
2916:
2216:
1571:
1553:
1302:
1194:
421:
221:
196:
3245:
2889:
2871:
2485:
2456:
2427:
2132:
2059:
1907:
1657:
1488:
1432:
1396:
1084:
861:
757:
667:
632:
594:
559:
374:
304:
Both sexes of red tree vole build nests, though the maternal nests are considerably larger.
3167:
3227:
2606:
2326:
1885:
1639:
1544:
1450:
906:
689:
54:
2773:
2561:
1867:
1834:
1770:
888:
2993:
2207:
2141:
2023:
1961:
1624:
1331:
1221:
1176:
1126:
1051:
933:
271:
3275:
3162:
2630:
2597:
1858:
1562:
1506:
1248:
1102:
969:
365:
356:
59:
679:
644:
606:
571:
456:. U.S. Department of Agriculture Forest Service, Pacific Northwest Research Station.
3003:
2949:
2703:
2447:
2267:
2198:
2181:
1816:
1675:
1666:
1630:
1591:
1497:
1340:
1316:
1257:
1230:
1093:
1075:
852:
3089:
333:
Their habitats in old forests have become a concern for the species. Logging and
671:
451:
415:
3201:
3180:
3123:
3069:
2282:
1988:
1605:
1515:
1322:
1284:
1266:
1239:
1185:
1060:
1034:
960:
897:
870:
721:
146:
3060:
224:, found in the US states of Oregon and California. They were formerly known as
2746:
2533:
2518:
2509:
1785:
1702:
1479:
1167:
951:
779:
769:
241:
136:
2899:
2862:
2852:
2791:
2718:
2676:
2099:
1648:
1117:
984:
879:
842:
156:
86:
30:
320:
is one of the red tree voles' primary predators. Recent studies show that
3232:
3054:
2801:
2500:
2336:
2077:
2006:
1141:
789:
751:
739:
636:
598:
106:
3115:
2834:
2825:
2167:
2159:
2085:
1893:
1849:
1585:
1538:
1473:
563:
3128:
2258:
1717:
763:
745:
733:
417:
Mammals of the Pacific Northwest: From the Coast to the High Cascades
237:
217:
126:
116:
96:
3031:
321:
270:
248:), though they occasionally eat the needles of western hemlock (
3102:
3035:
693:
400:
Musser, G. G. and M. D. Carleton. 2005. Superfamily Muroidea.
618:
616:
3044:
3001:
2992:
2947:
2897:
2888:
2860:
2851:
2823:
2799:
2790:
2716:
2628:
2531:
2498:
2334:
2325:
2280:
2256:
2179:
2158:
2130:
2097:
2076:
2004:
1987:
1905:
1884:
1847:
1814:
1783:
1715:
1139:
1115:
1073:
1032:
982:
850:
841:
787:
778:
545:
543:
380:10.2305/IUCN.UK.2018-1.RLTS.T42615A22389366.en
228:and have also been called the red tree mouse.
705:
8:
465:
463:
450:Forsman, James K.; Swingle, Eric D. (2016).
3032:
2998:
2894:
2857:
2796:
2331:
2176:
2094:
2001:
1902:
847:
784:
712:
698:
690:
520:"Field Guide to Red Tree Vole Nests, USFS"
518:Lesmeister, Damon; Swingle, James (2017).
48:
29:
20:
378:
347:
440:Maser, C. 1998; Wooster and Town. 2002
3282:IUCN Red List near threatened species
7:
1944:Ogilvie Mountains collared lemming (
513:
511:
484:Hayes, John P. (27 December 1996).
366:IUCN Red List of Threatened Species
14:
420:. Oregon State University Press.
493:American Society of Mammalogists
73:
3302:Taxa named by Frederick W. True
1953:Richardson's collared lemming (
2244:North American brown lemming (
1:
2118:Przewalski's steppe lemming (
660:Forest Ecology and Management
495:. Mammalian Species (532): 1.
2686:Imaizumi's red-backed vole (
2365:White-tailed mountain vole (
720:Extant species of subfamily
672:10.1016/j.foreco.2009.04.041
2935:California red tree mouse (
2659:Tien Shan red-backed vole (
2466:Lake Baikal mountain vole (
1971:Unalaska collared lemming (
1935:Nelson's collared lemming (
1917:Northern collared lemming (
1612:Schelkovnikov's pine vole (
1406:Liechtenstein's pine vole (
3318:
2756:Hokkaido red-backed vole (
2738:Japanese red-backed vole (
2695:Northern red-backed vole (
2668:Southern red-backed Vole (
2419:StoliÄŤka's mountain vole (
2383:Gobi Altai mountain vole (
2042:Transcaucasian mole vole (
3292:Mammals described in 1890
2650:Western red-backed vole (
1926:Ungava collared lemming (
1735:Forrest's mountain vole (
1388:Mediterranean pine vole (
816:Southwestern water vole (
728:
185:
178:
70:Scientific classification
68:
46:
37:
28:
23:
2616:Ward's red-backed vole (
2589:Yunnan red-backed vole (
2553:Kachin red-backed vole (
2235:Siberian brown lemming (
2226:Wrangel Island lemming (
1795:Duke of Bedford's vole (
3013:Long-clawed mole vole (
2410:Mongolian silver vole (
2401:Royle's mountain vole (
2109:Yellow steppe lemming (
552:Northwestern Naturalist
355:Scheuering, E. (2018).
236:The red tree vole is a
2976:Eastern heather vole (
2967:Western heather vole (
2811:Round-tailed muskrat (
2765:Grey red-backed vole (
2571:Southwest China vole (
2392:Central Kashmir vole (
2374:Silver mountain vole (
2309:Southern bog lemming (
2300:Northern bog lemming (
1762:Sikkim mountain vole (
1753:Linzhi mountain vole (
1415:Lusitanian pine vole (
486:"Aborimus Longicaudus"
280:Nesting and home range
276:
226:Phenacomys longicaudus
3254:Paleobiology Database
3076:Arborimus longicaudus
3046:Arborimus longicaudus
1379:Daghestan pine vole (
1042:(Voles from Pakistan)
1002:Caucasian snow vole (
943:Japanese grass vole (
807:European water vole (
414:Maser, Chris (1998).
373:: e.T42615A22389366.
359:Arborimus longicaudus
335:habitat fragmentation
274:
246:Pseudotsuga menziesii
213:Arborimus longicaudus
189:Arborimus longicaudus
2051:Northern mole vole (
2033:Southern mole vole (
1826:Narrow-headed vole (
1744:Chinese scrub vole (
1460:Thomas's pine vole (
1370:Calabria pine vole (
1361:Bavarian pine vole (
1294:European pine vole (
1276:Schidlovsky's vole (
1020:Robert's snow vole (
1011:European snow vole (
925:Middendorff's vole (
825:Montane water vole (
637:10.3955/046.084.0305
599:10.3955/046.083.0310
329:Conservation efforts
322:short-tailed weasels
318:northern spotted owl
260:), and Bishop pine (
2917:White-footed vole (
2580:Père David's vole (
2217:Beringian lemming (
1694:Zempoaltépec vole (
1572:Jalapan pine vole (
1424:Major's pine vole (
1390:M. duodecimcostatus
1303:Transcaspian vole (
916:Maximowicz's vole (
171:A. longicaudus
40:Conservation status
3015:P. schaposchnikowi
2872:Balkan snow vole (
2486:Flat-headed vole (
2457:Large-eared vole (
2428:Tuva silver vole (
2060:Zaisan mole vole (
1658:Long-tailed vole (
1489:Gray-tailed vole (
1442:Savi's pine vole (
1433:Alpine pine vole (
1204:DoÄźramaci's vole (
564:10.1898/nwn09-36.1
290:Tsuga heterophylla
277:
250:Tsuga heterophylla
3297:Mammals of Oregon
3269:
3268:
3241:Open Tree of Life
3038:Taxon identifiers
3029:
3028:
3025:
3024:
2988:
2987:
2960:
2910:
2884:
2883:
2847:
2846:
2786:
2785:
2731:
2726:(other Red-backed
2643:
2607:Yulungshan vole (
2546:
2349:
2321:
2320:
2293:
2192:
2173:
2154:
2153:
2091:
2072:
2071:
2017:
1998:
1983:
1982:
1899:
1880:
1879:
1805:P. liangshanensis
1728:
1640:California vole (
1614:M. schelkovnikovi
1545:Guatemalan vole (
1525:Townsend's vole (
1518:M. pennsylvanicus
1451:Tatra pine vole (
1408:M. liechtensteini
1152:
1045:
995:
907:Lacustrine vole (
837:
836:
800:
625:Northwest Science
587:Northwest Science
527:PNW Station, USFS
427:978-0-87071-438-2
252:), Sitka spruce (
222:Pacific Northwest
204:
203:
63:
16:Species of rodent
3309:
3262:
3261:
3249:
3248:
3236:
3235:
3223:
3222:
3210:
3209:
3197:
3196:
3184:
3183:
3171:
3170:
3158:
3157:
3145:
3144:
3132:
3131:
3119:
3118:
3106:
3105:
3093:
3092:
3080:
3079:
3078:
3065:
3064:
3063:
3033:
2999:
2959:
2958:
2954:
2909:
2908:
2904:
2895:
2890:Pliophenacomyini
2858:
2797:
2730:
2729:
2723:
2642:
2641:
2638:(some Red-backed
2635:
2545:
2544:
2538:
2348:
2347:
2341:
2332:
2292:
2291:
2287:
2246:L. trimucronatus
2208:Norway lemming (
2191:
2190:
2186:
2177:
2172:
2171:
2163:
2142:Steppe lemming (
2095:
2090:
2089:
2081:
2035:E. fuscocapillus
2024:Alai mole vole (
2016:
2015:
2011:
2002:
1997:
1996:
1992:
1962:Arctic lemming (
1919:D. groenlandicus
1903:
1898:
1897:
1889:
1727:
1726:
1725:(Mountain voles)
1722:
1705:M. xanthognathus
1651:M. chrotorrhinus
1554:TarabundĂ vole (
1547:M. guatemalensis
1381:M. daghestanicus
1332:Bucharian vole (
1305:M. transcaspicus
1222:Tien Shan vole (
1213:GĂĽnther's vole (
1195:Cabrera's vole (
1177:Anatolian vole (
1151:
1150:
1146:
1127:Sagebrush vole (
1044:
1043:
1039:
994:
993:
989:
972:M. sachalinensis
934:Mongolian vole (
848:
799:
798:
794:
785:
758:Euarchontoglires
714:
707:
700:
691:
684:
683:
655:
649:
648:
620:
611:
610:
582:
576:
575:
547:
538:
537:
535:
533:
524:
515:
506:
503:
497:
496:
490:
481:
475:
474:
467:
458:
457:
447:
441:
438:
432:
431:
411:
405:
398:
392:
391:
389:
387:
382:
352:
254:Picea sitchensis
191:
78:
77:
57:
52:
51:
33:
21:
3317:
3316:
3312:
3311:
3310:
3308:
3307:
3306:
3272:
3271:
3270:
3265:
3257:
3252:
3244:
3239:
3231:
3228:Observation.org
3226:
3218:
3213:
3205:
3200:
3192:
3187:
3179:
3174:
3166:
3161:
3153:
3148:
3140:
3135:
3127:
3122:
3114:
3109:
3101:
3096:
3088:
3083:
3074:
3073:
3068:
3059:
3058:
3053:
3040:
3030:
3021:
2984:
2957:(Heather voles)
2956:
2955:
2953:
2943:
2926:Red tree vole (
2906:
2905:
2903:
2880:
2843:
2819:
2782:
2727:
2725:
2724:
2722:
2712:
2652:C. californicus
2639:
2637:
2636:
2634:
2624:
2598:Chaotung vole (
2582:E. melanogaster
2542:
2540:
2539:
2537:
2527:
2494:
2421:A. stoliczkanus
2345:
2343:
2342:
2340:
2327:Clethrionomyini
2317:
2289:
2288:
2286:
2276:
2270:M. schisticolor
2252:
2189:(True lemmings)
2188:
2187:
2185:
2165:
2164:
2162:
2150:
2126:
2083:
2082:
2080:
2068:
2013:
2012:
2010:
1994:
1993:
1991:
1979:
1973:D. unalascensis
1946:D. nunatakensis
1891:
1890:
1888:
1886:Dicrostonychini
1876:
1859:Szechuan vole (
1843:
1810:
1779:
1724:
1723:
1721:
1711:
1642:M. californicus
1563:Woodland Vole (
1507:Creeping vole (
1397:Felten's vole (
1372:M. brachycercus
1296:M. subterraneus
1278:M. schidlovskii
1249:Southern vole (
1148:
1147:
1145:
1135:
1111:
1103:Mandarin vole (
1085:Brandt's vole (
1069:
1041:
1040:
1038:
1028:
991:
990:
988:
978:
970:Sakhalin vole (
927:M. middendorffi
918:M. maximowiczii
862:Clarke's vole (
833:
796:
795:
793:
774:
724:
718:
688:
687:
657:
656:
652:
622:
621:
614:
584:
583:
579:
549:
548:
541:
531:
529:
522:
517:
516:
509:
504:
500:
488:
483:
482:
478:
469:
468:
461:
449:
448:
444:
439:
435:
428:
413:
412:
408:
399:
395:
385:
383:
354:
353:
349:
344:
331:
314:
302:
286:western hemlock
282:
234:
200:
193:
187:
174:
72:
64:
55:Near Threatened
53:
49:
42:
17:
12:
11:
5:
3315:
3313:
3305:
3304:
3299:
3294:
3289:
3284:
3274:
3273:
3267:
3266:
3264:
3263:
3250:
3237:
3224:
3211:
3198:
3185:
3172:
3159:
3146:
3133:
3120:
3107:
3094:
3081:
3066:
3050:
3048:
3042:
3041:
3036:
3027:
3026:
3023:
3022:
3020:
3019:
3009:
3007:
2996:
2994:Prometheomyini
2990:
2989:
2986:
2985:
2983:
2982:
2973:
2969:P. intermedius
2963:
2961:
2945:
2944:
2942:
2941:
2932:
2928:A. longicaudus
2923:
2913:
2911:
2892:
2886:
2885:
2882:
2881:
2879:
2878:
2868:
2866:
2855:
2849:
2848:
2845:
2844:
2842:
2841:
2831:
2829:
2821:
2820:
2818:
2817:
2807:
2805:
2794:
2788:
2787:
2784:
2783:
2781:
2780:
2774:Smith's vole (
2771:
2762:
2753:
2744:
2734:
2732:
2714:
2713:
2711:
2710:
2704:Shansei vole (
2701:
2692:
2683:
2674:
2665:
2656:
2646:
2644:
2626:
2625:
2623:
2622:
2613:
2604:
2595:
2586:
2577:
2568:
2562:Pratt's vole (
2559:
2549:
2547:
2529:
2528:
2526:
2525:
2516:
2506:
2504:
2496:
2495:
2493:
2492:
2473:
2472:
2468:A. olchonensis
2463:
2454:
2448:Lemming vole (
2435:
2434:
2425:
2416:
2407:
2398:
2389:
2380:
2371:
2352:
2350:
2329:
2323:
2322:
2319:
2318:
2316:
2315:
2306:
2296:
2294:
2290:(Bog lemmings)
2278:
2277:
2275:
2274:
2268:Wood lemming (
2264:
2262:
2254:
2253:
2251:
2250:
2241:
2232:
2223:
2214:
2205:
2199:Amur lemming (
2195:
2193:
2174:
2156:
2155:
2152:
2151:
2149:
2148:
2138:
2136:
2128:
2127:
2125:
2124:
2120:E. przewalskii
2115:
2105:
2103:
2092:
2074:
2073:
2070:
2069:
2067:
2066:
2057:
2048:
2039:
2030:
2020:
2018:
1999:
1985:
1984:
1981:
1980:
1978:
1977:
1968:
1959:
1955:D. richardsoni
1950:
1941:
1932:
1923:
1913:
1911:
1900:
1882:
1881:
1878:
1877:
1875:
1874:
1868:Marie's vole (
1865:
1855:
1853:
1845:
1844:
1842:
1841:
1835:Radde's vole (
1832:
1822:
1820:
1812:
1811:
1809:
1808:
1801:
1791:
1789:
1781:
1780:
1778:
1777:
1771:Blyth's vole (
1768:
1759:
1755:N. linzhiensis
1750:
1741:
1731:
1729:
1713:
1712:
1710:
1709:
1700:
1691:
1687:M. richardsoni
1682:
1676:Singing vole (
1673:
1667:Mexican vole (
1664:
1660:M. longicaudus
1655:
1646:
1637:
1633:M. abbreviatus
1631:Insular vole (
1625:incertae sedis
1619:
1618:
1599:
1598:
1594:M. ochrogaster
1592:Prairie vole (
1579:
1578:
1569:
1560:
1551:
1532:
1531:
1522:
1513:
1504:
1498:Montane vole (
1495:
1486:
1467:
1466:
1457:
1448:
1439:
1430:
1421:
1417:M. lusitanicus
1412:
1403:
1394:
1385:
1376:
1367:
1348:
1347:
1341:Juniper vole (
1338:
1329:
1310:
1309:
1300:
1291:
1282:
1273:
1269:M. qazvinensis
1264:
1258:Paradox vole (
1255:
1246:
1242:M. kermanensis
1237:
1231:Persian vole (
1228:
1219:
1210:
1201:
1192:
1183:
1174:
1155:
1153:
1137:
1136:
1134:
1133:
1123:
1121:
1113:
1112:
1110:
1109:
1100:
1094:Plateau vole (
1091:
1081:
1079:
1071:
1070:
1068:
1067:
1058:
1048:
1046:
1030:
1029:
1027:
1026:
1017:
1008:
998:
996:
980:
979:
977:
976:
967:
958:
949:
940:
931:
922:
913:
909:M. limnophilus
904:
895:
889:Gerbe's vole (
886:
877:
873:M. evoronensis
868:
858:
856:
845:
839:
838:
835:
834:
832:
831:
822:
813:
803:
801:
782:
776:
775:
773:
772:
766:
760:
754:
748:
742:
736:
729:
726:
725:
719:
717:
716:
709:
702:
694:
686:
685:
666:(5): 626–634.
650:
631:(3): 255–265.
612:
593:(3): 273–286.
577:
539:
507:
498:
476:
459:
442:
433:
426:
406:
393:
346:
345:
343:
340:
330:
327:
313:
310:
301:
298:
281:
278:
262:Pinus muricata
256:), grand fir (
240:in the family
233:
230:
202:
201:
194:
183:
182:
176:
175:
168:
166:
162:
161:
154:
150:
149:
144:
140:
139:
134:
130:
129:
124:
120:
119:
114:
110:
109:
104:
100:
99:
94:
90:
89:
84:
80:
79:
66:
65:
47:
44:
43:
38:
35:
34:
26:
25:
24:Red tree vole
15:
13:
10:
9:
6:
4:
3:
2:
3314:
3303:
3300:
3298:
3295:
3293:
3290:
3288:
3285:
3283:
3280:
3279:
3277:
3260:
3255:
3251:
3247:
3242:
3238:
3234:
3229:
3225:
3221:
3216:
3212:
3208:
3203:
3199:
3195:
3190:
3186:
3182:
3177:
3173:
3169:
3164:
3160:
3156:
3151:
3147:
3143:
3138:
3134:
3130:
3125:
3121:
3117:
3112:
3108:
3104:
3099:
3095:
3091:
3086:
3082:
3077:
3071:
3067:
3062:
3056:
3052:
3051:
3049:
3047:
3043:
3039:
3034:
3018:
3016:
3011:
3010:
3008:
3006:
3005:
3000:
2997:
2995:
2991:
2981:
2979:
2974:
2972:
2970:
2965:
2964:
2962:
2952:
2951:
2946:
2940:
2938:
2933:
2931:
2929:
2924:
2922:
2920:
2915:
2914:
2912:
2902:
2901:
2896:
2893:
2891:
2887:
2877:
2875:
2870:
2869:
2867:
2865:
2864:
2859:
2856:
2854:
2850:
2840:
2838:
2837:O. zibethicus
2833:
2832:
2830:
2828:
2827:
2822:
2816:
2814:
2809:
2808:
2806:
2804:
2803:
2798:
2795:
2793:
2789:
2779:
2777:
2772:
2770:
2768:
2763:
2761:
2759:
2754:
2752:
2750:
2745:
2743:
2741:
2736:
2735:
2733:
2721:
2720:
2715:
2709:
2707:
2702:
2700:
2698:
2693:
2691:
2689:
2684:
2682:
2680:
2675:
2673:
2671:
2666:
2664:
2662:
2657:
2655:
2653:
2648:
2647:
2645:
2633:
2632:
2631:Clethrionomys
2627:
2621:
2619:
2614:
2612:
2610:
2605:
2603:
2601:
2596:
2594:
2592:
2587:
2585:
2583:
2578:
2576:
2574:
2569:
2567:
2565:
2560:
2558:
2556:
2551:
2550:
2548:
2536:
2535:
2530:
2524:
2522:
2517:
2515:
2513:
2508:
2507:
2505:
2503:
2502:
2497:
2491:
2489:
2483:
2482:
2481:
2475:
2474:
2471:
2469:
2464:
2462:
2460:
2455:
2453:
2451:
2445:
2444:
2443:
2437:
2436:
2433:
2431:
2426:
2424:
2422:
2417:
2415:
2413:
2408:
2406:
2404:
2399:
2397:
2395:
2390:
2388:
2386:
2381:
2379:
2377:
2376:A. argentatus
2372:
2370:
2368:
2362:
2361:
2360:
2354:
2353:
2351:
2346:Central Asia)
2339:
2338:
2333:
2330:
2328:
2324:
2314:
2312:
2307:
2305:
2303:
2298:
2297:
2295:
2285:
2284:
2279:
2273:
2271:
2266:
2265:
2263:
2261:
2260:
2255:
2249:
2247:
2242:
2240:
2238:
2233:
2231:
2229:
2224:
2222:
2220:
2215:
2213:
2211:
2206:
2204:
2202:
2197:
2196:
2194:
2184:
2183:
2178:
2175:
2169:
2161:
2157:
2147:
2145:
2140:
2139:
2137:
2135:
2134:
2129:
2123:
2121:
2116:
2114:
2112:
2107:
2106:
2104:
2102:
2101:
2096:
2093:
2087:
2079:
2075:
2065:
2063:
2058:
2056:
2054:
2049:
2047:
2045:
2040:
2038:
2036:
2031:
2029:
2027:
2022:
2021:
2019:
2009:
2008:
2003:
2000:
1990:
1986:
1976:
1974:
1969:
1967:
1965:
1960:
1958:
1956:
1951:
1949:
1947:
1942:
1940:
1938:
1933:
1931:
1929:
1924:
1922:
1920:
1915:
1914:
1912:
1910:
1909:
1904:
1901:
1895:
1887:
1883:
1873:
1871:
1866:
1864:
1862:
1857:
1856:
1854:
1852:
1851:
1846:
1840:
1838:
1833:
1831:
1829:
1824:
1823:
1821:
1819:
1818:
1813:
1807:
1806:
1802:
1800:
1798:
1793:
1792:
1790:
1788:
1787:
1782:
1776:
1774:
1769:
1767:
1765:
1764:N. sikimensis
1760:
1758:
1756:
1751:
1749:
1747:
1742:
1740:
1738:
1733:
1732:
1730:
1720:
1719:
1714:
1708:
1706:
1701:
1699:
1697:
1692:
1690:
1688:
1683:
1681:
1679:
1674:
1672:
1670:
1665:
1663:
1661:
1656:
1654:
1652:
1647:
1645:
1643:
1638:
1636:
1634:
1628:
1627:
1626:
1621:
1620:
1617:
1615:
1609:
1608:
1607:
1601:
1600:
1597:
1595:
1589:
1588:
1587:
1581:
1580:
1577:
1575:
1570:
1568:
1566:
1561:
1559:
1557:
1556:M. oaxacensis
1552:
1550:
1548:
1542:
1541:
1540:
1534:
1533:
1530:
1528:
1527:M. townsendii
1523:
1521:
1519:
1516:Meadow vole (
1514:
1512:
1510:
1505:
1503:
1501:
1496:
1494:
1492:
1491:M. canicaudus
1487:
1485:
1483:
1477:
1476:
1475:
1469:
1468:
1465:
1463:
1458:
1456:
1454:
1449:
1447:
1445:
1440:
1438:
1436:
1431:
1429:
1427:
1422:
1420:
1418:
1413:
1411:
1409:
1404:
1402:
1400:
1395:
1393:
1391:
1386:
1384:
1382:
1377:
1375:
1373:
1368:
1366:
1364:
1358:
1357:
1356:
1350:
1349:
1346:
1344:
1339:
1337:
1335:
1334:B. bucharicus
1330:
1328:
1326:
1323:Afghan vole (
1320:
1319:
1318:
1312:
1311:
1308:
1306:
1301:
1299:
1297:
1292:
1290:
1288:
1285:Social vole (
1283:
1281:
1279:
1274:
1272:
1270:
1267:Qazvin vole (
1265:
1263:
1261:
1256:
1254:
1252:
1247:
1245:
1243:
1240:Kerman vole (
1238:
1236:
1234:
1229:
1227:
1225:
1220:
1218:
1216:
1211:
1209:
1207:
1206:M. dogramacii
1202:
1200:
1198:
1193:
1191:
1189:
1186:Common vole (
1184:
1182:
1180:
1179:M. anatolicus
1175:
1173:
1171:
1165:
1164:
1163:
1157:
1156:
1154:
1144:
1143:
1138:
1132:
1130:
1125:
1124:
1122:
1120:
1119:
1114:
1108:
1106:
1105:L. mandarinus
1101:
1099:
1097:
1092:
1090:
1088:
1083:
1082:
1080:
1078:
1077:
1072:
1066:
1064:
1061:Murree vole (
1059:
1057:
1055:
1052:True's vole (
1050:
1049:
1047:
1037:
1036:
1031:
1025:
1023:
1018:
1016:
1014:
1009:
1007:
1005:
1000:
999:
997:
987:
986:
981:
975:
973:
968:
966:
964:
961:Tundra vole (
959:
957:
955:
954:M. mujanensis
950:
948:
946:
945:M. montebelli
941:
939:
937:
936:M. mongolicus
932:
930:
928:
923:
921:
919:
914:
912:
910:
905:
903:
901:
898:Taiwan vole (
896:
894:
892:
887:
885:
883:
878:
876:
874:
871:Evorsk vole (
869:
867:
865:
860:
859:
857:
855:
854:
849:
846:
844:
840:
830:
828:
823:
821:
819:
814:
812:
810:
805:
804:
802:
797:(Water voles)
792:
791:
786:
783:
781:
777:
771:
767:
765:
761:
759:
755:
753:
749:
747:
743:
741:
737:
735:
731:
730:
727:
723:
715:
710:
708:
703:
701:
696:
695:
692:
681:
677:
673:
669:
665:
661:
654:
651:
646:
642:
638:
634:
630:
626:
619:
617:
613:
608:
604:
600:
596:
592:
588:
581:
578:
573:
569:
565:
561:
557:
553:
546:
544:
540:
528:
521:
514:
512:
508:
502:
499:
494:
487:
480:
477:
472:
466:
464:
460:
455:
454:
446:
443:
437:
434:
429:
423:
419:
418:
410:
407:
403:
397:
394:
381:
376:
372:
368:
367:
362:
360:
351:
348:
341:
339:
336:
328:
326:
323:
319:
311:
309:
305:
299:
297:
293:
291:
287:
279:
273:
269:
265:
263:
259:
258:Abies grandis
255:
251:
247:
243:
239:
231:
229:
227:
223:
219:
215:
214:
209:
208:red tree vole
198:
192:
190:
184:
181:
180:Binomial name
177:
173:
172:
167:
164:
163:
160:
159:
155:
152:
151:
148:
145:
142:
141:
138:
135:
132:
131:
128:
125:
122:
121:
118:
115:
112:
111:
108:
105:
102:
101:
98:
95:
92:
91:
88:
85:
82:
81:
76:
71:
67:
61:
56:
45:
41:
36:
32:
27:
22:
19:
3045:
3014:
3004:Prometheomys
3002:
2977:
2968:
2948:
2936:
2927:
2925:
2918:
2907:(Tree voles)
2898:
2874:D. bogdanovi
2873:
2861:
2836:
2824:
2812:
2800:
2775:
2767:C. rufocanus
2766:
2757:
2748:
2747:Royal vole (
2740:C. andersoni
2739:
2717:
2706:C. shanseius
2705:
2696:
2688:C. imaizumii
2687:
2679:C. glareolus
2678:
2669:
2661:C. centralis
2660:
2651:
2629:
2617:
2608:
2599:
2590:
2581:
2572:
2564:E. chinensis
2563:
2554:
2532:
2520:
2519:Kolan vole (
2511:
2510:Ganzu vole (
2499:
2488:A. strelzowi
2487:
2480:Platycranius
2478:
2476:
2467:
2458:
2449:
2440:
2438:
2430:A. tuvinicus
2429:
2420:
2412:A. semicanus
2411:
2402:
2393:
2385:A. barakshin
2384:
2375:
2367:A. albicauda
2366:
2357:
2355:
2335:
2310:
2301:
2281:
2269:
2257:
2245:
2237:L. sibiricus
2236:
2227:
2218:
2209:
2201:L. amurensis
2200:
2180:
2143:
2131:
2119:
2110:
2098:
2061:
2052:
2044:E. lutescens
2043:
2034:
2025:
2014:(Mole voles)
2005:
1995:(mole voles)
1972:
1964:D. torquatus
1963:
1954:
1945:
1936:
1928:D. hudsonius
1927:
1918:
1906:
1869:
1861:V. millicens
1860:
1848:
1836:
1827:
1817:Stenocranius
1815:
1803:
1796:
1784:
1772:
1763:
1754:
1745:
1736:
1716:
1704:
1703:Taiga vole (
1695:
1686:
1685:Water vole (
1677:
1669:M. mexicanus
1668:
1659:
1650:
1641:
1632:
1623:
1622:
1613:
1604:
1602:
1593:
1584:
1582:
1574:M. quasiater
1573:
1565:M. pinetorum
1564:
1555:
1546:
1537:
1535:
1526:
1517:
1508:
1499:
1490:
1481:
1480:Beach Vole (
1472:
1470:
1461:
1452:
1443:
1435:M. multiplex
1434:
1425:
1416:
1407:
1398:
1389:
1380:
1371:
1363:M. bavaricus
1362:
1353:
1351:
1343:B. juldaschi
1342:
1333:
1324:
1317:Blanfordimys
1315:
1313:
1304:
1295:
1286:
1277:
1268:
1260:M. paradoxus
1259:
1250:
1241:
1232:
1223:
1215:M. guentheri
1214:
1205:
1196:
1187:
1178:
1169:
1168:Field vole (
1160:
1158:
1140:
1128:
1116:
1104:
1095:
1086:
1076:Lasiopodomys
1074:
1062:
1053:
1033:
1021:
1012:
1003:
992:(Snow voles)
983:
971:
963:M. oeconomus
962:
953:
952:Muisk vole (
944:
935:
926:
917:
908:
899:
890:
881:
872:
863:
853:Alexandromys
851:
826:
817:
809:A. amphibius
808:
788:
756:Superorder:
750:Infraclass:
663:
659:
653:
628:
624:
590:
586:
580:
558:(1): 63–73.
555:
551:
530:. Retrieved
526:
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492:
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370:
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188:
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18:
3202:NatureServe
3124:iNaturalist
3070:Wikispecies
2677:Bank vole (
2609:E. proditor
2555:E. cachinus
2541:(Voles from
2459:A. macrotis
2450:A. lemminus
2344:(Voles from
2302:S. borealis
2219:L. nigripes
2053:E. talpinus
1989:Ellobiusini
1908:Dicrostonyx
1828:S. gregalis
1797:P. bedfordi
1773:N. leucurus
1737:N. forresti
1696:M. umbrosus
1649:Rock vole (
1606:Hyrcanicola
1500:M. montanus
1453:M. tatricus
1325:B. afghanus
1287:M. socialis
1197:M. cabrerae
1170:M. agrestis
1129:L. curtatus
1087:L. brandtii
1054:H. fertilis
1035:Hyperacrius
900:M. kikuchii
880:Reed vole (
827:A. scherman
722:Arvicolinae
386:15 November
232:Description
147:Arvicolinae
143:Subfamily:
3276:Categories
2950:Phenacomys
2919:A. albipes
2776:C. smithii
2749:C. regulus
2697:C. rutilus
2670:C. gapperi
2591:E. miletus
2543:East Asia)
2534:Eothenomys
2442:Aschizomys
2394:A. montosa
2311:S. cooperi
2283:Synaptomys
2144:L. lagurus
2062:E. tancrei
2026:E. alaicus
1937:D. nelsoni
1892:(Collared
1870:V. musseri
1786:Proedromys
1509:M. oregoni
1482:M. breweri
1462:M. thomasi
1399:M. felteni
1188:M. arvalis
1022:C. roberti
1013:C. nivalis
864:M. clarkei
818:A. sapidus
780:Arvicolini
770:Cricetidae
532:5 February
342:References
242:Cricetidae
197:F. W. True
137:Cricetidae
3287:Arborimus
2978:P. ungava
2900:Arborimus
2863:Dinaromys
2853:Pliomyini
2835:Muskrat (
2813:N. alleni
2792:Ondatrini
2719:Craseomys
2600:E. olitor
2573:E. custos
2477:Subgenus
2439:Subgenus
2403:A. roylei
2356:Subgenus
2228:L. paulus
2210:L. lemmus
2111:E. luteus
2100:Eolagurus
1837:S. raddei
1678:M. miurus
1603:Subgenus
1583:Subgenus
1536:Subgenus
1471:Subgenus
1426:M. majori
1355:Terricola
1352:Subgenus
1314:Subgenus
1224:M. ilaeus
1159:Subgenus
1118:Lemmiscus
1096:L. fuscus
1063:H. wynnei
985:Chionomys
891:M. gerbei
882:M. fortis
843:Microtini
732:Kingdom:
312:Predation
165:Species:
158:Arborimus
93:Kingdom:
87:Eukaryota
3207:2.101630
3194:13000173
3142:11265946
3061:Q1761767
3055:Wikidata
2802:Neofiber
2618:E. wardi
2501:Caryomys
2359:Alticola
2337:Alticola
2168:Lemmings
2086:lemmings
2084:(Steppe
2078:Lagurini
2007:Ellobius
1894:lemmings
1746:N. irene
1444:M. savii
1251:M. levis
1233:M. irani
1162:Microtus
1142:Microtus
790:Arvicola
768:Family:
764:Rodentia
752:Eutheria
746:Mammalia
740:Chordata
738:Phylum:
734:Animalia
680:85266361
645:85851862
607:84217641
572:86157754
133:Family:
127:Rodentia
117:Mammalia
107:Chordata
103:Phylum:
97:Animalia
83:Domain:
60:IUCN 3.1
3181:1002195
3116:2437931
2937:A. pomo
2826:Ondatra
2521:C. inez
2160:Lemmini
2133:Lagurus
1850:Volemys
1586:Pedomys
1539:Pitymys
1474:Mynomes
1149:(Voles)
762:Order:
744:Class:
220:of the
216:) is a
199:, 1890)
153:Genus:
123:Order:
113:Class:
58: (
3259:161652
3246:663525
3220:251274
3155:180353
3103:328444
2758:C. rex
2728:voles)
2640:voles)
2512:C. eva
2259:Myopus
2182:Lemmus
1718:Neodon
1004:C. gud
678:
643:
605:
570:
471:"Blog"
424:
238:rodent
218:rodent
3233:81827
3168:42615
3137:IRMNG
3129:45719
676:S2CID
641:S2CID
603:S2CID
568:S2CID
523:(PDF)
489:(PDF)
3215:NCBI
3163:IUCN
3150:ITIS
3111:GBIF
3090:G6MG
534:2022
422:ISBN
388:2021
371:2018
316:The
206:The
3189:MSW
3176:MDD
3098:EoL
3085:CoL
668:doi
664:258
633:doi
595:doi
560:doi
375:doi
264:).
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