619:
1595:
Ethiopian wolf territories are largely stable, but packs can expand whenever the opportunity arises, such as when another pack disappears. The size of each territory correlates with the abundance of rodents, the number of wolves in a pack, and the survival of pups. Ethiopian wolves rest together in the open at night, and congregate for greetings and border patrols at dawn, noon, and evening. They may shelter from rain under overhanging rocks and behind boulders. The species never sleeps in dens, and only uses them for nursing pups. When patrolling their territories, Ethiopian wolves regularly scent-mark, and interact aggressively and vocally with other packs. Such confrontations typically end with the retreat of the smaller group.
1649:
1091:
1047:
1111:
1018:
989:
933:
904:
962:
866:
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1571:
123:
1685:. Solitary wolves hunt for rodents in the midst of the monkeys, ignoring juvenile monkeys, though these are similar in size to some of their prey. The monkeys, in turn, tolerate and largely ignore the wolves, although they take flight if they observe feral dogs, which sometimes prey on them. Within the troops, the wolves enjoy much higher success in capturing rodents than usual, perhaps because the monkeys' activities flush out the rodents, or because the presence of numerous larger animals makes it harder for rodents to spot a threat.
728:, Gojjam, and in every northern Afroalpine region where agriculture is well developed and human pressure acute. This revelation stressed the importance of the Bale Mountains wolf populations for the species' long-term survival, as well as the need to protect other surviving populations. A decade after the rabies outbreak, the Bale populations had fully recovered to pre-epizootic levels, prompting the species' downlisting to endangered in 2004, though it still remains the world's rarest canid, and Africa's most endangered
634:
175:
1665:... they are most amusing to watch, when hunting. The rats, which are brown, with short tails, live in big colonies and dart from burrow to burrow, while the cuberow stands motionless till one of them shows, when he makes a pounce for it. If he is unsuccessful, he seems to lose his temper, and starts digging violently; but this is only lost labour, as the ground is honeycombed with holes, and every rat is yards away before he has thrown up a pawful.
1797:
2292:
1616:
coat is gradually replaced with the adult colouration. By the age of five weeks, the pups feed on a combination of milk and solid food, and become completely weaned off milk at the age of 10 weeks to six months. All members of the pack contribute to protecting and feeding the pups, with subordinate females sometimes assisting the dominant female by suckling them. Full growth and sexual maturity are attained at the age of two years.
1497:
150:
2366:) are usually agonistic, with Ethiopian wolves dominating African wolves if the latter enter their territories, and vice versa. Although African golden wolves are inefficient rodent hunters and thus not in direct competition with Ethiopian wolves, it is likely that heavy human persecution prevents the former from attaining numbers large enough to completely displace the latter.
1461:
1194:
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521:
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in single figures for several years. Habitat loss in the
Ethiopian highlands is directly linked to agricultural expansion into Afroalpine areas. In the northern highlands, human density is among the highest in Africa, with 300 people per km in some localities, with almost all areas below 3,700 m having been converted into
2424:
The overall aim of the EWCP is to protect the wolf's
Afroalpine habitat in Bale, and establish additional conservation areas in Menz and Wollo. The EWCP carries out education campaigns for people outside the wolf's range to improve dog husbandry and manage diseases within and around the park, as well
2278:
farmers, herders, woodcutters, and many others. In Bale there are numerous villages in and around the area, comprising over 8,500 households with more than 12,500 dogs. In 2007, the estimate of households within wolf habitat numbered 1,756. Because of the high number of dogs, the risk of infection in
2243:
fields. Suitable areas of land below this limit are under some level of protection, such as Guassa-Menz and the
Denkoro Reserve, or within the southern highlands, such as the Arsi and Bale Mountains. The most vulnerable wolf populations to habitat loss are those within relatively low-lying Afroalpine
2238:
During the 1990s, wolf populations in Gosh Meda and
Guguftu became extinct. In both cases, the extent of Afroalpine habitat above the limit of agriculture had been reduced to less than 20 km. The EWCP team confirmed the extinction of a wolf population in Mt. Guna in 2011, whose numbers had been
1590:
of six wolves are more common. Packs are formed by dispersing males and a few females, which with the exception of the breeding female, are reproductively suppressed. Each pack has a well-established hierarchy, with dominance and subordination displays being common. Upon dying, a breeding female can
1549:
to rusty red, with dense whitish to pale ginger underfur. The fur of the throat, chest and underparts is white, with a distinct white band occurring around the sides of the neck. There is a sharp boundary between the red coat and white marks. The ears are thickly furred on the edges, though naked on
1213:
Due to the high density of rodents in their new
Afroalpine habitat, the ancestors of the Ethiopian wolf gradually developed into specialised rodent hunters. This specialisation is reflected in the animal's skull morphology, with its very elongated head, long jaw, and widely spaced teeth. During this
2323:
were more available. The extinction of wolves in Mt. Choqa was likely due to persecution. Although people living close to wolves in modern times believe that wolf populations are recovering, negative attitudes towards the species persist due to livestock predation. Wolves were largely unmolested by
2273:
In the Simien
Mountains National Park, human and livestock populations are increasing by 2% annually, with further road construction allowing easy access to peasants into wolf home ranges; 3,171 people in 582 households were found to be living in the park and 1,477 outside the park in October 2005.
1594:
These packs live in communal territories, which encompass 6 km (2.3 sq mi) of land on average. In areas with little food, the species lives in pairs, sometimes accompanied by pups, and defends larger territories averaging 13.4 km (5.2 sq mi). In the absence of disease,
1558:
during the wet season (August–October), and there is no evident seasonal variation in coat colour, though the contrast between the red coat and white markings increases with age and social rank. Females tend to have paler coats than males. During the breeding season, the female's coat turns yellow,
3282:
Gopalakrishnan, Shyam; Sinding, Mikkel-Holger S.; Ramos-Madrigal, Jazmín; Niemann, Jonas; Samaniego
Castruita, Jose A.; Vieira, Filipe G.; Carøe, Christian; Montero, Marc de Manuel; Kuderna, Lukas; Serres, Aitor; González-Basallote, Víctor Manuel; Liu, Yan-Hu; Wang, Guo-Dong; Marques-Bonet, Tomas;
2408:
in 1983, with the Bale
Mountains Research Project being established shortly after. This was followed by a detailed, four-year field study, which prompted the IUCN/SSC Canid Specialist Group to produce an action plan in 1997. The plan called for the education of people in wolf-inhabited areas, wolf
1615:
is 60–62 days, with pups being born between
October and December. Pups are born toothless and with their eyes closed, and are covered in a charcoal-grey coat with a buff patch on the chest and abdomen. Litters consist of two to six pups, which emerge from their den after three weeks, when the dark
3219:
Koepfli, Klaus-Peter; Pollinger, John; Godinho, Raquel; Robinson, Jacqueline; Lea, Amanda; Hendricks, Sarah; Schweizer, Rena M.; Thalmann, Olaf; Silva, Pedro; Fan, Zhenxin; Yurchenko, Andrey A.; Dobrynin, Pavel; Makunin, Alexey; Cahill, James A.; Shapiro, Beth; Álvares, Francisco; Brito, José C.;
1728:
around 3,200 to 4,500 m, with some wolves inhabiting the Bale
Mountains being present in montane grasslands at 3,000 m. Although specimens were collected in Gojjam and northwestern Shoa at 2,500 m in the early 20th century, no recent records exist of the species occurring below 3,000 m. In modern
1550:
the inside. The naked borders of the lips, the gums and palate are black. The lips, a small spot on the cheeks and an ascending crescent below the eyes are white. The thickly furred tail is white underneath, and has a black tip, though, unlike most other canids, there is no dark patch marking the
3043:
Vonholdt, B. M.; Pollinger, J. P.; Lohmueller, K. E.; Han, E.; Parker, H. G.; Quignon, P.; Degenhardt, J. D.; Boyko, A. R.; Earl, D. A.; Auton, A.; Reynolds, A.; Bryc, K.; Brisbin, A.; Knowles, J. C.; Mosher, D. S.; Spady, T. C.; Elkahloun, A.; Geffen, E.; Pilot, M.; Jedrzejewski, W.; Greco, C.;
2400:
Steps taken to ensure the survival of the Ethiopian wolf include dog vaccination campaigns in Bale, Menz, and Simien, sterilization programs for wolf-dog hybrids in Bale, rabies vaccination of wolves in parts of Bale, community and school education programs in Bale and Wollo, contributing to the
1315:
to both. There are two genetically distinct African golden wolf populations that exist in northwestern and eastern Africa. This suggests that Ethiopian wolves – or a close and extinct relative – once had a much larger range within Africa to admix with other canids. There is evidence of gene flow
2336:. It was once hunted by sportsmen, though this is now illegal. Vehicle collisions killed at least four wolves in the Sanetti Plateau since 1988, while two others were left with permanent limps. Similar accidents are a risk in areas where roads cut across wolf habitats, such as in Menz and Arsi.
2302:
Although wolves in Bale have learned to use cattle to conceal their presence when hunting for rodents, the level of grazing in the area can adversely affect the vegetation available for the wolves' prey. Although no declines in wolf populations related to overgrazing have occurred, high grazing
2204:
The Ethiopian wolf has been considered rare since it was first recorded scientifically. The species likely has always been confined to Afroalpine habitats, so it was never widespread. In historical times, all of the Ethiopian wolf's threats are both directly and indirectly human-induced, as the
296:
2344:
Management plans for hybridization with dogs involve sterilization of known hybrids. Incidences of Ethiopian wolf-dog hybridization have been recorded in Bale's Web Valley. At least four hybrids were identified and sterilized in the area. Although hybridization has not been detected elsewhere,
1270:
In 2015, a study of mitochondrial genome sequences and whole genome nuclear sequences of African and Eurasian canids indicated that extant wolf-like canids have colonised Africa from Eurasia at least five times throughout the Pliocene and Pleistocene, which is consistent with fossil evidence
1542:, males are larger than females, having 20% greater body mass. Adults measure 841–1,012 mm (33.1–39.8 in) in body length, and 530–620 mm (21–24 in) in height. Adult males weigh 14.2–19.3 kg (31–43 lb), while females weigh 11.2–14.15 kg (24.7–31.2 lb).
1271:
suggesting that much of African canid fauna diversity resulted from the immigration of Eurasian ancestors, likely coincident with Plio-Pleistocene climatic oscillations between arid and humid conditions. According to a phylogeny derived from nuclear sequences, the Eurasian golden jackal (
2221:
Rabies outbreaks, stemming from infected dogs, have killed many Ethiopian wolves over the 1990s and 2000s. Two well-documented outbreaks in Bale, one in 1991 and another in 2008–2009, resulted in the die-off or disappearance of 75% of known animals. Both incidents prompted reactive
2264:
or reduced fitness exists, the extremely small wolf population sizes, particularly those north of the Rift Valley, raise concerns among conservationists. Elsewhere, the Bale populations are fairly continuous, while those in Simien can still interbreed through habitat corridors.
1591:
be replaced by a resident daughter, though this increases the risk of inbreeding. Such a risk is sometimes circumvented by multiple paternity and extra-pack matings. The dispersal of wolves from their packs is largely restricted by the scarcity of unoccupied habitat.
671:
The Ethiopian wolf was recognised as requiring protection in 1938, and received it in 1974. The first in-depth studies on the species occurred in the 1980s with the onset of the American-sponsored Bale Mountains Research Project. Ethiopian wolf populations in the
3283:
Mirarab, Siavash; Fernandes, Carlos; Gaubert, Philippe; Koepfli, Klaus-Peter; Budd, Jane; Rueness, Eli Knispel; Heide-Jørgensen, Mads Peter; Petersen, Bent; Sicheritz-Ponten, Thomas; Bachmann, Lutz; Wiig, Øystein; Hansen, Anders J.; Gilbert, M. Thomas P. (2018).
1627:
Most females disperse from their natal pack at about two years of age, and some become "floaters" that may successfully immigrate into existing packs. Breeding pairs are most often unrelated to each other, suggesting that female-biased dispersal reduces
2260:, which is likely to increase with current rates of human expansion. The dangers posed by fragmentation include increased contact with humans, dogs, and livestock, and further risk of isolation and inbreeding in wolf populations. Although no evidence of
2521:
Martínez-Navarro, Bienvenido; Gossa, Tegenu; Carotenuto, Francesco; Bartolini-Lucenti, Saverio; Palmqvist, Paul; Asrat, Asfawossen; Figueirido, Borja; Rook, Lorenzo; Niespolo, Elizabeth M.; Renne, Paul R.; Herzlinger, Gadi; Hovers, Erella (2023-05-16).
2332:. However, they are perceived as threats to livestock elsewhere, with cases of retaliatory killings occurring in the Arsi Mountains. The Ethiopian wolf has not been recorded to be exploited for its fur, though in one case, wolf hides were used as
2274:
Although the area of the park has since been expanded, further settlement stopped, and grazing restricted, effective enforcement may take years. As of 2011, about 30,000 people live in 30 villages around and two within the park, including 4,650
1656:
Unlike most social carnivores, the Ethiopian wolf tends to forage and feed on small prey alone. It is most active during the day, the time when rodents are themselves most active, though they have been observed to hunt in groups when targeting
2349:
or a reduction in fitness, though this does not appear to have taken place. Due to the female's strong preference to avoid inbreeding, hybridization could be the result of not finding any males who are not close relatives outside of dogs.
719:
since the early 20th century, and in the Bale Mountains in the late 1950s. The status of the Ethiopian wolf was reassessed in the late 1990s, following improvements in travel conditions into northern Ethiopia. The surveys taken revealed
3879:
A sporting trip through Abyssinia : a narrative of a nine months' journey from the plains of the Hawash to the snows of Simien, with a description of the game, from elephant to ibex, and notes on the manners and customs of the
2093:
The local ericaceous forests and grasslands are under the protection of the Borena Saiynt Regional Park from agriculture as low as 3,200 m. The EWCP and the FZS have been involved in educational programs and wolf monitoring in the
1736:
The Ethiopian wolf uses all Afroalpine habitats, but has a preference for open areas containing short herbaceous and grassland communities inhabited by rodents, which are most abundant along flat or gently sloping areas with poor
3101:
Gottelli, D.; Sillero-Zubiri, C.; Applebaum, G. D.; Roy, M. S.; Girman, D. J.; Garcia-Moreno, J.; Ostrander, E. A.; Wayne, R. K. (1994). "Molecular genetics of the most endangered canid: The Ethiopian wolf Canis simensis".
1316:
between the eastern population and the Ethiopian wolf, which has led to the eastern population being distinct from the northwestern population. The common ancestor of both African golden wolf populations was a genetically
1519:
and has comparatively longer legs. Its skull is very flat, with a long facial region accounting for 58% of the skull's total length. The ears are broad, pointed, and directed forward. The teeth, particularly the
1680:
herds, a tactic thought to aid in ambushing rodents out of their holes by using the cattle to hide their presence. Ethiopian wolves have also been observed forming temporary associations with troops of grazing
653:) by Europeans and Ze Etiyopia by its citizens, wrote that the animal's skin was never worn by natives, as it was popularly believed that the wearer would die should any wolf hairs enter an open wound, while
1168:
approximately 1.8 million years ago. The Ethiopian wolf has survived numerous climatic changes in its Ethiopian highland habitat, with its range repeatedly expanding and contracting with glacial cycles.
1275:) diverged from the wolf/coyote lineage 1.9 million years ago, and with mitochondrial genome sequences indicating the Ethiopian wolf diverged from this lineage slightly prior to that. Further studies on
2230:
is not necessarily fatal to wolves, though a recent increase in infection has occurred, with outbreaks of canine distemper having been detected in 2005–2006 in Bale and in 2010 across subpopulations.
1164:
and dated to between 1.6 and 1.4 million years ago. The authors of this study state that the ancestors of the Ethiopian wolf arrived in Africa from Eurasia at the same time as the ancestors of the
2378:
appendices, though it is afforded full official protection under Ethiopia's Wildlife Conservation Regulations of 1974, Schedule VI, with the killing of a wolf carrying a two-year jail sentence.
2213:, road construction, and livestock farming), diseases (primarily rabies and canine distemper), conflict with humans (poisoning, persecution, and road kills), and hybridisation with dogs.
3029:
2409:
population monitoring, and the stemming of rabies in dog populations. The Ethiopian Wolf Conservation Programme was formed in 1995 by Oxford University, with donors including the
2205:
wolf's highland habitat, with its high annual rainfall and rich fertile soils, is ideal for agricultural activities. Its proximate threats include habitat loss and fragmentation (
1469:
This canid was initially classed as a distinct subspecies on account of its bright red coat, though this characteristic is unreliable as a taxonomic distinction. However, its
2135:
The wolf's range is protected by community resource management, and the Guassa Community Conservation Area. Educational campaigns are undertaken in schools near wolf ranges.
3272:. Master of Science Thesis. Center for Ecological and Evolutionary Synthesis Department of Bioscience Faculty of Mathematics and Natural Science, University of Oslo, Norway
3988:
9689:
2425:
as monitoring wolves in Bale, south and north Wollo. The program seeks to vaccinate up to 5,000 dogs a year to reduce rabies and distemper in wolf-inhabited areas.
2397:. Areas of suitable wolf habitat have recently increased to 87%, as a result of boundary extensions in Simien and the creation of the Arsi Mountains National Park.
618:
4010:
2428:
In 2016, the Korean company Sooam Biotech was reported to be attempting to clone the Ethiopian wolf using dogs as surrogate mothers to help conserve the species.
9767:
1562:
Animals resulting from Ethiopian wolf-dog hybridisation tend to be more heavily built than pure wolves, and have shorter muzzles and different coat patterns.
1611:
involves the breeding male following the female closely. The breeding female only accepts the advances of the breeding male, or males from other packs. The
1768:, and short grasslands growing in shallow soils. In its northern range, the wolf's habitat is composed of plant communities characterised by a matrix of
2775:
9890:
9860:
9663:
3220:
Geffen, Eli; Leonard, Jennifer A.; Helgen, Kristofer M.; Johnson, Warren E.; o'Brien, Stephen J.; Van Valkenburgh, Blaire; Wayne, Robert K. (2015).
1545:
The Ethiopian wolf has short guard hairs and thick underfur, which provides protection at temperatures as low as −15 °C. Its overall colour is
9715:
3955:
3783:
1276:
400:
in size and build, and is distinguished by its long and narrow skull, and its red and white fur. Unlike most large canids, which are widespread,
1673:, with the level of effort varying from scratching lightly at the hole to totally destroying a set of burrows, leaving metre-high earth mounds.
3805:
692:
in 1994. The IUCN/SSC Canid Specialist Group advocated a three-front strategy of education, wolf population monitoring, and rabies control in
3923:
2708:
2678:
9905:
534:
426:
at altitudes of 3,000–4,500 m, with the overall adult population estimated at 360–440 individuals in 2011, more than half of them in the
9895:
4066:
1532:
are relatively small. The Ethiopian wolf has eight mammae, of which only six are functional. The front paws have five toes, including a
9900:
9880:
2904:
2390:
664:. Since then, it was scarcely heard of in Europe up until the early 20th century, when several skins were shipped to England by Major
1156:
The earliest known fossil of the Ethiopian wolf is known from the Melka Wakena paleoanthropological site-complex in the Southeastern
5436:
3152:
Dalton, R. (2001). The skull morphology of the Ethiopian wolf (Canis simensis). B.Sc.thesis. University of Edinburgh, Edinburgh, UK.
1648:
1586:
The Ethiopian wolf is a social animal, living in family groups containing up to 20 adults (individuals older than one year), though
401:
3910:
711:, the paucity of information stemming from that area indicated that the species was likely declining there, while reports from the
4031:
4018:
9741:
4006:
154:
3423:"The physiology of cooperative breeding in a rare social canid; sex, suppression and pseudopregnancy in female Ethiopian wolves"
2401:
running of national parks, and population monitoring and surveying. A 10-year national action plan was formed in February 2011.
5780:
1832:
9875:
9759:
2470:
2386:
1993:
9720:
680:, though the critical state of the species was revealed during the early 1990s after a combination of shooting and a severe
2456:
This is in reference to the Ethiopian wolf's reported habit of following mares and cows about to give birth to feed on the
7998:
2418:
2405:
1820:
9885:
2414:
2394:
2382:
2060:
673:
3781:
Foraging ecology and trophic niche overlap between sympatric African wolf and Ethiopian wolf in the Ethiopian Highlands
3044:
Randi, E.; Bannasch, D.; Wilton, A.; Shearman, J.; Musiani, M.; Cargill, M.; Jones, P. G.; Qian, Z.; Huang, W. (2010).
1222:, which caused the species' Afroalpine habitat to fragment, thus isolating Ethiopian wolf populations from each other.
9754:
9702:
5192:
3906:
1888:
1812:
1670:
1392:
1334:
1090:
1046:
7989:
2776:"The effect of the last glacial age on speciation and population genetic structure of the endangered Ethiopian wolf (
1698:
1110:
1017:
9454:
9110:
7476:
1852:
988:
932:
903:
442:
9772:
1570:
961:
865:
6315:
3832:
707:. Elsewhere, information on Ethiopian wolves remained scarce; although first described in 1835 as living in the
9373:
9268:
8007:
6297:
5992:
5041:
2257:
538:
122:
6025:
5968:
5789:
3752:
2345:
scientists are concerned that it could pose a threat to the wolf population's genetic integrity, resulting in
411:
with very specific habitat requirements. It is one of the world's rarest canids, and Africa's most endangered
9837:
2826:
174:
9035:
8016:
7747:
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7375:
6757:
6333:
5806:
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2144:
1824:
1730:
1296:
1215:
3572:"Wolves are better hunters when monkeys are around: An unexpected co-existence in the Ethiopian highlands."
1180:
may be an evolutionary relic of a gray wolf-like ancestor's past invasion of northern Africa from Eurasia.
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8700:
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2188:
1781:
470:
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9806:
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8655:
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8131:
7503:
6273:
6255:
6204:
6142:
6124:
6115:
6052:
6001:
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4267:
3586:"Solitary Ethiopian wolves increase predation success on rodents when among grazing gelada monkey herds"
2491:
2261:
1872:
1633:
1501:
1312:
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1176:
analysis showed a closer relationship to the gray wolf and the coyote than to other African canids, and
689:
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269:
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1240:
1202:
3753:"Field report : Association between an Arabian wolf and a domestic dog in central Saudi Arabia"
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leaves have occasionally been found in Ethiopian wolf stomachs. The sedge possibly is ingested for
1800:
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1508:
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Life in Abyssinia: Being Notes Collected During Three Years' Residence and Travels in that Country
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696:. The establishment of the Ethiopian Wolf Conservation Programme in Bale soon followed in 1995 by
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4157:
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3571:
3529:
3453:
3135:
3009:
2818:
2739:
2731:
1507:
The Ethiopian wolf is similar in size and build to North America's coyote; it is larger than the
478:
441:, on account of its small numbers and fragmented range. Threats include increasing pressure from
308:
169:
9811:
5319:
1788:
moorlands at 3,200–3,600 m in Simien may provide a refuge for wolves in highly disturbed areas.
9694:
3014:
2010:
As of 2011, no sightings have been made, despite intensive monitoring during two field visits.
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Venkataraman, V. V.; Kerby, J. T.; Nguyen, N.; Ashenafi, Z. T.; Fashing, P. J. (2015-03-27).
2524:"The earliest Ethiopian wolf: implications for the species evolution and its future survival"
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ScienceDaily., 22 June 2015. <www.sciencedaily.com/releases/2015/06/150622085224.htm>.
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3222:"Genome-wide Evidence Reveals that African and Eurasian Golden Jackals Are Distinct Species"
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1962:
1900:
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or for parasite control. The species may scavenge on carcasses, but is usually displaced by
1828:
1796:
1784:, all of which are favoured by the wolf's rodent prey. Although marginal in importance, the
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3046:"Genome-wide SNP and haplotype analyses reveal a rich history underlying dog domestication"
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9382:
9044:
8994:
8985:
8860:
8806:
8750:
8619:
8592:
8485:
8303:
8262:
8227:
7972:
7035:
6715:
6706:
6213:
5951:
5400:
4850:
4777:
4471:
4276:
4210:
4187:
4022:
3787:
3473:"Inbreeding is reduced by female-biased dispersal and mating behavior in Ethiopian wolves"
1755:
1621:
685:
419:
131:
6748:
4735:
3863:
2381:
The species is present in several protected areas, including three areas in South Wollo (
528:
3929:
3471:
Randall, DA; Pollinger, JP; Wayne, RK; Tallents, LA; Johnson, PJ; Macdonald, DW (2007).
3300:
3237:
3115:
3061:
2798:
2556:
2523:
1733:, which extends up to 3,700 m, has largely restricted the species to the highest peaks.
1661:
calves. Major Percy-Cotton described the hunting behaviour of Ethiopian wolves as thus:
1632:. Inbreeding is ordinarily avoided because it leads to a reduction in progeny fitness (
9865:
9606:
9481:
9400:
9202:
8949:
8842:
8815:
7868:
7485:
7085:
6632:
6169:
6151:
6082:
5693:
5684:
5286:
5247:
4201:
4113:
4015:
3635:
3317:
3284:
3123:
3078:
3045:
2333:
2256:
Some Ethiopian wolf populations, particularly those in North Wollo, show signs of high
2169:
1924:
1844:
1658:
1575:
1496:
1481:
1235:
by its larger size, relatively longer legs, distinct reddish coat, and white markings.
654:
646:
591:
427:
423:
1989:
encroachment into rodent habitat, disease, and competition/predation by golden wolves
1980:
The second-largest population, the most genetically diverse, is a tourist attraction.
9854:
9728:
9445:
9341:
9314:
8940:
8638:
8583:
8105:
8049:
7790:
7431:
6886:
6448:
5613:
5580:
5382:
5373:
5310:
5210:
5168:
5020:
4613:
3892:
3877:
3422:
3373:
2806:
2743:
2604:
2140:
2025:
An ORDA Biodiversity Conservation Project is active in the area, in conjunction with
1920:
1604:
1587:
1308:
957:
716:
458:
159:
73:
3457:
3139:
2822:
2595:
9632:
9472:
9418:
9119:
8887:
8833:
8610:
8509:
8064:
8040:
7859:
7850:
7670:
7637:
7560:
7521:
6931:
6913:
6775:
6647:
6106:
5507:
5470:
5355:
5337:
4750:
4696:
4622:
4300:
3966:
2897:
Canids: Foxes, Wolves, Jackals, and Dogs:Status Survey and Conservation Action Plan
2304:
2223:
2019:
1525:
1516:
1219:
1205:
has resulted in a skull similar in shape to that of jackals and the South American
928:
4002:
3533:
3441:
2490:
For a full set of supporting references refer to the note (a) in the phylotree at
649:. European writers traveling in Ethiopia during the mid-19th century (then called
9824:
9746:
9676:
9586:
9086:
8277:
8194:
8167:
7916:
7830:
7762:
7688:
7679:
7646:
7604:
7569:
7286:
7179:
7011:
6551:
6463:
6016:
5589:
5409:
5074:
4988:
4971:
4948:
4911:
4795:
4759:
4604:
4393:
4248:
2895:. In Sillero-Zubiri, Claudio; Hoffmann, Michael; Macdonald, David Whyte (eds.).
2210:
2158:
The third-largest population, in the second-largest Afroalpine area in Ethiopia
2087:
1985:
1916:
1764:
1214:
period, the species likely attained its highest abundance, and had a relatively
725:
474:
450:
405:
48:
3894:
Neue Wirbelthiere zu der Fauna von Abyssinien gehörig, entdeckt und beschrieben
2539:
2125:
A core population with ideal habitat, it is increasingly a tourist attraction.
2056:
Isolation, habitat degradation, human-wildlife conflict, and road encroachment
9295:
8541:
8472:
8459:
8426:
8212:
7950:
7710:
7258:
7235:
7121:
7112:
7061:
7026:
7002:
6766:
6665:
6614:
6560:
6542:
6515:
6422:
6372:
5771:
5744:
5672:
5655:
5418:
5391:
5237:
5141:
5125:
4961:
4841:
4681:
4517:
4384:
4176:
4144:
4128:
3392:
3308:
3246:
3221:
2457:
2280:
2001:
1912:
1851:
calves. In areas where the big-headed African mole-rat is absent, the smaller
1840:
1747:
1710:
1629:
1529:
1470:
1400:
1206:
657:
434:
93:
58:
9577:
3602:
3585:
2547:
2307:
and vegetation deterioration in Afroalpine areas such as Delanta and Simien.
2193:
Most of the species' habitat occurs within the Bale Mountains National Park.
1983:
Human disturbance, road traffic, extensive agriculture, habitat degradation,
1754:
and grasses, with low vegetation cover. Other favourable habitats consist of
637:
Mounted specimen (1902), one of the first post-1835 specimens to reach Europe
9226:
9161:
8774:
8714:
8500:
8321:
7936:
7928:
7877:
7727:
7613:
7595:
7309:
7143:
6976:
6818:
6784:
6596:
6288:
5562:
5032:
4508:
4076:
3489:
3472:
3374:"Scent-marking and territorial behaviour of Ethiopian wolves Canis simensis"
3188:"A review of the family Canidae, with a classification by numerical methods"
2478:
2022:, isolation, and possible competition with abundant golden wolf populations
1876:
1785:
1759:
1725:
1717:
1713:
1477:
1460:
1198:
764:
729:
700:, in conjunction with the Ethiopian Wildlife Conservation Authority (EWCA).
661:
412:
226:
186:
98:
42:
9707:
4044:
3525:
3504:
Charlesworth D, Willis JH (2009). "The genetics of inbreeding depression".
3449:
3326:
3255:
3087:
2814:
2673:. Vol. 1 (3rd ed.). Johns Hopkins University Press. p. 577.
2565:
2279:
local wolf populations is high. Furthermore, intentional and unintentional
3204:
3131:
2067:, which is to encompass about a third of the wolf's range in North Wollo.
1193:
17:
9785:
9571:
8923:
8386:
8203:
8179:
7540:
7390:
7336:
7272:
7267:
7220:
7152:
7070:
6940:
6689:
6680:
6578:
6533:
6413:
6246:
6070:
5983:
5892:
5882:
5861:
5821:
5708:
5640:
5548:
5539:
4859:
4655:
4408:
4291:
4258:
4107:
4095:
3841:
3840:. IUCN/SSC Canid Specialist Group. Oxford, United Kingdom. Archived from
2320:
2122:
Although diminishing from disease, the population is healthy and stable.
1864:
1848:
1738:
1637:
1555:
1521:
1259:
refuted the separate genus in favour of placing the species in the genus
1161:
454:
206:
88:
83:
68:
63:
53:
9642:
3270:
Can hybridization be detected between African wolf and sympatric canids?
3069:
9832:
9668:
9305:
9152:
9026:
8741:
8253:
7583:
7049:
6605:
6569:
6436:
6404:
6388:
5901:
5604:
5183:
4664:
4571:
4489:
4417:
4359:
4336:
4229:
3992:
2954:
2735:
2245:
2095:
1879:, with farmers often leaving herds in wolf-inhabited areas unattended.
1776:
1770:
1721:
1559:
becomes woolier, and the tail turns brownish, losing much of its hair.
1533:
844:
760:
385:
236:
103:
78:
9681:
9614:
9409:
9356:
9101:
8789:
8523:
7243:
6904:
6739:
6496:
5933:
4705:
4631:
4580:
4562:
4535:
4456:
4318:
4101:
4089:
4038:
3187:
2275:
2240:
2030:
2026:
1966:
1896:
1682:
1677:
1232:
899:
712:
681:
408:
397:
216:
196:
9548:
3517:
2727:
1919:
for several decades. Southeast of the Rift Valley, it occurs in the
1218:. This changed about 15,000 years ago with the onset of the current
703:
Soon after, a further wolf population was discovered in the Central
688:. In response, the IUCN reclassified the species from endangered to
1377:
9819:
9391:
9250:
9193:
9066:
8869:
8726:
8353:
8335:
7958:
7628:
6877:
6862:
5523:
4814:
4309:
2375:
2325:
2290:
2107:
2039:
2033:
governments, and supported by the Guna Highland Water enterprise.
1908:
1860:
1855:
is targeted. In the Simien Mountains, the Ethiopian wolf preys on
1836:
1795:
1697:
1647:
1569:
1546:
1538:
1495:
1408:
1244:
1227:
1192:
1186:
1013:
632:
617:
246:
3285:"Interspecific Gene Flow Shaped the Evolution of the Genus Canis"
2671:
Mammal Species of the World: A Taxonomic and Geographic Reference
8765:
7661:
6986:
6922:
4595:
4327:
3983:
3630:
3628:
2329:
2296:
2103:
1911:. It has recently become extinct in Gosh Meda in north Shoa and
1904:
1751:
1742:
1404:
1321:
861:
715:
plateau were a century out of date. Wolves were recorded in the
438:
9655:
9552:
8457:
7914:
7218:
6370:
5505:
4946:
4126:
4048:
3806:"Inside the cloning factory that creates 500 new animals a day"
3738:
3726:
3714:
3702:
3690:
3678:
3666:
3654:
3619:
3546:
3408:
3359:
3344:
3161:
2941:
2847:
2084:
After Simien, the second-largest area north of the Rift Valley
1887:
Six current Ethiopian wolf populations are known. North of the
1839:
and eggs. Ethiopian wolves have twice been observed to feed on
1473:
are consistently longer than those of the nominate subspecies.
1231:
species present in Africa, and is readily distinguishable from
6895:
6504:
4832:
3912:
The Ethiopian Wolf: Status Survey and Conservation Action Plan
2589:
2587:
2585:
2583:
2581:
2579:
2577:
2575:
2474:
839:
693:
404:
feeders, the Ethiopian wolf is a highly specialised feeder of
2161:
Habitat degradation, expanding agriculture, and road traffic
2007:
An isolated patch, estimated at less than 20 km by 2004
1811:
In the Bale Mountains, the Ethiopian wolf's primary prey are
1745:. Prime wolf habitat in the Bale Mountains consists of short
489:
Alternative English names for the Ethiopian wolf include the
2404:
The species' critical situation was first publicised by the
1536:, while the hind paws have four. As is typical in the genus
684:
epidemic decimated most packs studied in the Web Valley and
2774:
Gottelli, D; Marino, J; Sillero-Zubiri, C; Funk, S (2004).
1669:
The technique described above is commonly used in hunting
641:
The species was first scientifically described in 1835 by
2283:
are frequent in the ericaceous moorlands wolves inhabit.
2184:
The largest population, with the highest density of prey
2315:
Direct killings of wolves were more frequent during the
1709:
The Ethiopian wolf is restricted to isolated pockets of
1197:
Ethiopian wolf skull: Despite its close relation to the
676:
were negatively affected by the political unrest of the
4007:
IUCN Species Survival Commission#Canid Specialist Group
3030:
The Variation of Animals and Plants Under Domestication
2926:
2899:. IUCN-The World Conservation Union. pp. 167–174.
2389:, and Borena Sayint Regional Park), one in north Shoa (
2324:
humans in Bale, as they were not considered threats to
2115:
As of 2010, an estimated 40% have been missing since a
2090:, ploughing, persecution, and local negative attitudes
2075:
Patches connected by corridors, totalling 243 km
1971:
Patches connected by corridors, totalling 273 km
3372:
Sillero-Zubiri, Claudio; Macdonald, David W. (1998).
1279:
found instances of Ethiopian wolves hybridizing with
627:
Neue Wirbelthiere zu der Fauna von Abyssinien gehörig
2883:
2881:
392:. In southeastern Ethiopia, it is also known as the
9561:
9354:
9303:
9294:
9248:
9224:
9191:
9150:
9099:
9065:
9024:
8983:
8966:
8921:
8895:
8886:
8787:
8763:
8739:
8713:
8636:
8581:
8539:
8522:
8498:
8470:
8425:
8384:
8351:
8334:
8275:
8225:
8192:
8166:
8103:
8062:
8029:
7970:
7949:
7927:
7829:
7788:
7760:
7726:
7709:
7659:
7626:
7582:
7539:
7520:
7429:
7388:
7334:
7308:
7284:
7256:
7234:
7178:
7151:
7142:
7083:
7048:
7024:
7000:
6985:
6974:
6875:
6816:
6801:
6728:
6678:
6646:
6631:
6494:
6461:
6435:
6402:
6386:
6286:
6244:
6104:
6069:
6014:
5981:
5931:
5890:
5881:
5860:
5819:
5769:
5743:
5706:
5671:
5654:
5638:
5602:
5561:
5537:
5522:
5469:
5284:
5245:
5236:
5181:
5140:
5122:
5072:
5019:
4970:
4959:
4910:
4812:
4748:
4722:
4694:
4653:
4644:
4593:
4506:
4469:
4443:
4406:
4382:
4373:
4357:
4289:
4256:
4247:
4227:
4185:
4174:
4142:
3897:(in German). Frankfurt am Main : S. Schmerber.
3751:Chris Barichievy; Shayne Clugston; Robert Sheldon.
3186:Clutton-Brock, J.; Corbet, G.G.; Hills, M. (1976).
2132:encroachment into rodent habitat, and road traffic
1807:), one of the Ethiopian wolf's primary prey animals
1243:originally classified the species under a separate
3834:Strategic Planning for Ethiopian Wolf Conservation
2164:Protected within the Arsi Mountains Regional Park
1676:Wolves in Bale have been observed to forage among
3989:Conservation of Ethiopian wolves (Canis simensis)
2051:Possibly declining, otherwise stable, as of 1998
1607:usually takes place between August and November.
2660:
2658:
2656:
1652:Southern Ethiopian wolf feeding, Bale Mountains.
1311:, and with the Ethiopian wolf being genetically
2045:Patchily distributed in an area of 140 km
1996:, and has been monitored regularly since 2003.
1663:
1255:considered it to be "only an exaggerated fox".
625:by Friedrich Carl Vogel (from Eduard Rüppell's
4011:International Union for Conservation of Nature
2469:This was later proven incorrect in 2010, when
4060:
3558:
3173:
2997:
2619:10.2305/IUCN.UK.2011-1.RLTS.T3748A10051312.en
2063:and its associates are working to create the
1640:expression of deleterious recessive alleles.
1332:As of 2005, two subspecies are recognised by
8:
3984:Ethiopian Wolf Conservation Programme (EWCP)
1263:, upon noting cranial similarities with the
473:, which seeks to protect the wolves through
2955:"Ethiopia | International Wolf Center"
2295:Ethiopian wolves resting alongside grazing
1720:inhabited by Afroalpine rodents. Its ideal
1528:measure 14–22 mm in length, while the
9549:
9300:
9079:
8980:
8901:
8892:
8719:
8536:
8478:
8467:
8454:
8348:
8172:
7967:
7924:
7911:
7843:
7740:
7723:
7588:
7553:
7536:
7314:
7281:
7231:
7215:
7148:
7054:
6997:
6982:
6813:
6658:
6643:
6441:
6399:
6383:
6367:
6075:
5887:
5878:
5749:
5677:
5668:
5651:
5573:
5534:
5519:
5515:
5502:
5466:
5242:
5152:
5137:
5119:
5025:
4967:
4956:
4943:
4728:
4650:
4449:
4379:
4370:
4253:
4244:
4240:
4224:
4182:
4171:
4150:
4139:
4123:
4067:
4053:
4045:
2769:
2767:
2669:. In Wilson, D. E.; Reader, D. M. (eds.).
2516:
2514:
2512:
2510:
1929:
1341:
545:
294:
148:
121:
31:
3831:IUCN; SSC Canid Specialist Group (2011).
3601:
3488:
3421:van Kesteren, Freya; et al. (2013).
3404:
3402:
3316:
3245:
3203:
3077:
2707:Sillero-Zubiri, C.; Gottelli, D. (1994).
2617:
2555:
1299:was used to compare members of the genus
3615:
3613:
3340:
3338:
3336:
2982:
2702:
2700:
2698:
2696:
2694:
2692:
2690:
2374:The Ethiopian wolf is not listed on the
2065:Abuna Yoseph Community Conservation Area
1624:have been observed in Ethiopian wolves.
3355:
3353:
2993:
2991:
2978:
2976:
2967:
2937:
2935:
2888:Sillero-Zubiri, C.; Marino, J. (2004).
2639:
2594:Marino, J.; Sillero-Zubiri, C. (2011).
2506:
2449:
2244:ranges, such as those in Aboi Gara and
1827:. Other secondary prey species include
539:question marks, boxes, or other symbols
2843:
2841:
2839:
2492:Evolution of the wolf#Wolf-like canids
2354:Competition with African golden wolves
2152:54 wolves in 9 packs, as of 2007-2010
2226:in 2003 and 2008–2009, respectively.
2119:outbreak was detected in local dogs.
7:
9838:FE981C90-6153-4BC5-89C2-F3A999EF5A11
3739:IUCN/SSC Canid Specialist Group 2011
3727:IUCN/SSC Canid Specialist Group 2011
3715:IUCN/SSC Canid Specialist Group 2011
3703:IUCN/SSC Canid Specialist Group 2011
3691:IUCN/SSC Canid Specialist Group 2011
3679:IUCN/SSC Canid Specialist Group 2011
3667:IUCN/SSC Canid Specialist Group 2011
3655:IUCN/SSC Canid Specialist Group 2011
3620:IUCN/SSC Canid Specialist Group 2011
3547:IUCN/SSC Canid Specialist Group 2011
3409:IUCN/SSC Canid Specialist Group 2011
3360:IUCN/SSC Canid Specialist Group 2011
3345:IUCN/SSC Canid Specialist Group 2011
3162:IUCN/SSC Canid Specialist Group 2011
2942:IUCN/SSC Canid Specialist Group 2011
2848:IUCN/SSC Canid Specialist Group 2011
541: instead of Ethiopic characters.
27:Canine native to Ethiopian Highlands
5437:South African small-spotted genet (
3918:. IUCN/SSC Canid Specialist Group.
2927:Sillero-Zubiri & MacDonald 1997
2605:IUCN Red List of Threatened Species
2269:Encroachment within protected areas
2187:Disease (rabies and distemper) and
2181:Declining, but stable in long term
1524:, are small and widely spaced. The
1079:
1071:
832:
825:
818:
810:
802:
794:
786:
778:
770:
756:
3124:10.1111/j.1365-294X.1994.tb00070.x
3018:. Vol. II. John Murray. pp. 12–13.
2391:Guassa Community Conservation Area
1875:. It typically poses no threat to
1225:The Ethiopian wolf is one of five
25:
5781:Broad-striped Malagasy mongoose (
3804:Zastrow, Mark (8 February 2016).
3636:"Canis simensis (Ethiopian wolf)"
2311:Human persecution and disturbance
2303:intensities are known to lead to
1582:Social and territorial behaviours
1324:and 28% Ethiopian wolf ancestry.
750:with timing in millions of years
668:during his travels in Abyssinia.
9891:Fauna of the Ethiopian Highlands
9861:IUCN Red List endangered species
2807:10.1111/j.1365-294x.2004.02226.x
1833:yellow-spotted brush-furred rats
1459:
1376:
1109:
1089:
1045:
1016:
987:
960:
931:
902:
864:
843:
519:
465:. Its conservation is headed by
433:The Ethiopian wolf is listed as
173:
4003:Ethiopian Wolf (Canis simensis)
2178:About 250 adults and subadults
1992:The entire range is within the
1915:, and has not been reported in
1702:Northern Ethiopian wolf in the
1574:Southern Ethiopian wolf in the
1343:
645:, who provided a skull for the
2387:Simien Mountains National Park
2112:A single patch of 112 km
1994:Simien Mountains National Park
1821:black-clawed brush-furred rats
1419:(Erlanger & Neumann, 1900)
660:that the species gave rise to
1:
5193:Malabar large-spotted civet (
3883:. London : Rowland Ward.
3442:10.1016/j.physbeh.2013.08.016
3033:. Vol. I. Orange Judd. p. 48.
2419:Wildlife Conservation Network
2406:Wildlife Conservation Society
1931:Summary of current status of
1847:calves. It will also prey on
422:is limited to seven isolated
9455:Indonesian mountain weasel (
9111:North American river otter (
7999:Humboldt's hog-nosed skunk (
5053:Macrogalidia musschenbroekii
3960:National Geographic Magazine
2415:Frankfurt Zoological Society
2395:Arsi Mountains National Park
2383:Bale Mountains National Park
2061:Frankfurt Zoological Society
1891:, the species occurs in the
1813:big-headed African mole-rats
1671:big-headed African mole-rats
1599:Reproduction and development
1160:. It is the right half of a
674:Bale Mountains National Park
9906:Ethiopian montane moorlands
6316:Somalian slender mongoose (
3868:. London, R. Ward, limited.
3192:Bull. Br. Mus. (Nat. Hist.)
2128:Human disturbance, rabies,
1335:Mammal Species of the World
443:expanding human populations
9922:
9896:Endangered fauna of Africa
9269:Asian small-clawed otter (
8008:American hog-nosed skunk (
7990:Molina's hog-nosed skunk (
6298:Angolan slender mongoose (
5993:Ethiopian dwarf mongoose (
5042:Small-toothed palm civet (
3865:The Game Animals of Africa
2665:Wozencraft, C. W. (2005).
2540:10.1038/s42003-023-04908-w
1945:Population size estimates
1853:Northeast African mole-rat
1815:, though it also feeds on
1805:Tachyoryctes macrocephalus
1758:grasslands, high-altitude
321:Erlanger and Neumann, 1900
9901:Mammals described in 1840
9881:Endemic fauna of Ethiopia
9174:
9082:
9036:Saharan striped polecat (
9007:
8904:
8722:
8481:
8466:
8453:
8175:
8017:Striped hog-nosed skunk (
7923:
7910:
7846:
7748:Mediterranean monk seal (
7743:
7591:
7556:
7477:Juan Fernández fur seal (
7441:South American fur seal (
7412:
7376:South American sea lion (
7371:
7317:
7230:
7214:
7057:
6849:
6758:South American gray fox (
6702:
6661:
6444:
6382:
6366:
6334:Common slender mongoose (
6200:
6078:
5964:
5914:
5807:Narrow-striped mongoose (
5802:
5752:
5680:
5576:
5518:
5514:
5501:
5465:
5329:Crested servaline genet (
5155:
5118:
5028:
4955:
4942:
4731:
4677:
4452:
4243:
4223:
4170:
4153:
4138:
4122:
4084:
3430:Physiology & Behavior
3393:10.1017/s0952836998007134
3309:10.1016/j.cub.2018.08.041
3247:10.1016/j.cub.2015.06.060
1303:. The study supports the
1104:
1084:
1077:
1069:
1040:
1011:
984:
955:
926:
897:
859:
837:
830:
823:
816:
808:
800:
792:
784:
776:
768:
314:
307:
302:
293:
275:
268:
170:Scientific classification
168:
146:
137:
129:
120:
34:
9392:Stoat/Beringian ermine (
9260:African clawless otter (
9012:African striped weasel (
8825:Yellow-throated marten (
8701:Formosan ferret-badger (
8405:Western mountain coati (
8396:Eastern mountain coati (
8313:Western lowland olingo (
8295:Eastern lowland olingo (
8123:Southern spotted skunk (
7817:Southern elephant seal (
7808:Northern elephant seal (
7598:Pagophilus groenlandicus
6375:("dog-like" carnivorans)
6188:Stripe-necked mongoose (
5809:Mungotictis decemlineata
4684:Herpailurus yagouaroundi
4131:("cat-like" carnivorans)
3967:Ethiopian wolves on high
3027:Darwin, Charles (1868).
2477:'s sole ancestor is the
2473:studies showed that the
2252:Population fragmentation
1307:being distinct from the
555:Linguistic group or area
9437:Yellow-bellied weasel (
8692:Burmese ferret-badger (
8674:Chinese ferret-badger (
8665:Bornean ferret badger (
8656:Vietnam ferret-badger (
8141:Eastern spotted skunk (
8132:Western spotted skunk (
7862:Leptonychotes weddellii
7504:Subantarctic fur seal (
7450:Australasian fur seal (
6274:Black-footed mongoose (
6256:Bushy-tailed mongoose (
6205:White-tailed mongoose (
6143:Indian brown mongoose (
6125:Short-tailed mongoose (
6116:Small Indian mongoose (
6053:Flat-headed kusimanse (
6026:Alexander's kusimanse (
6002:Common dwarf mongoose (
5969:Pousargues's mongoose (
5831:Brown-tailed mongoose (
5790:Grandidier's mongoose (
5044:Arctogalidia trivirgata
4268:Sunda clouded leopard (
3903:Sillero-Zubiri, Claudio
3874:Powell-Cotton, P. H. G.
2340:Hybridisation with dogs
2207:subsistence agriculture
1731:subsistence agriculture
1724:extends from above the
1476:Southeast Rift Valley:
1452:Southern Ethiopian wolf
1363:Northern Ethiopian wolf
1297:whole genome sequencing
1216:continuous distribution
396:. It is similar to the
9278:Congo clawless otter (
9181:Hydrictis maculicollis
9179:Spotted-necked otter (
9138:Southern river otter (
9089:Pteronura brasiliensis
8852:European pine marten (
8363:South American coati (
8091:Palawan stink badger (
7417:New Zealand sea lion (
6482:Japanese raccoon dog (
6179:Crab-eating mongoose (
6134:Indian grey mongoose (
5757:Ring-tailed vontsira (
5568:(bone-crushing hyenas)
5263:Central African oyan (
4824:Chinese mountain cat (
3792:The Rufford Foundation
3603:10.1093/jmammal/gyu013
2890:"6.4 – Ethiopian wolf"
2528:Communications Biology
2347:outbreeding depression
2299:
2189:agricultural expansion
2072:South Wollo highlands
1899:, in the northern and
1808:
1706:
1667:
1653:
1578:
1504:
1210:
736:Taxonomy and evolution
638:
630:
527:This section contains
471:Conservation Programme
418:The species's current
130:Ethiopian wolf on the
9876:Carnivorans of Africa
9807:Paleobiology Database
9518:Back-striped weasel (
9464:Black-footed ferret (
9236:Smooth-coated otter (
9014:Poecilogale albinucha
8683:Javan ferret-badger (
8569:Sumatran hog badger (
8551:Northern hog badger (
8245:Crab-eating raccoon (
8150:Pygmy spotted skunk (
7871:Lobodon carcinophagus
7607:Histriophoca fasciata
7400:Australian sea lion (
7346:California sea lion (
7095:American black bear (
6837:Black-backed jackal (
6828:Side-striped jackal (
6668:Chrysocyon brachyurus
6092:Long-nosed mongoose (
5365:Rusty-spotted genet (
5202:Large-spotted civet (
5051:Sulawesi palm civet (
4980:Owston's palm civet (
4554:Andean mountain cat (
3999:Department of Zoology
3971:BBC Wildlife Magazine
3490:10.1093/beheco/arm010
3205:10.5962/bhl.part.6922
2360:African golden wolves
2294:
2262:inbreeding depression
1883:Range and populations
1873:African golden wolves
1857:Abyssinian grass rats
1799:
1701:
1651:
1636:) due largely to the
1634:inbreeding depression
1573:
1502:Louis Agassiz Fuertes
1499:
1287:Admixture with other
1281:African golden wolves
1196:
690:critically endangered
636:
621:
547:Indigenous names for
303:Ethiopian wolf range
9333:Long-tailed weasel (
8911:Lyncodon patagonicus
8560:Greater hog badger (
8082:Sunda stink badger (
7772:Hawaiian monk seal (
7495:Guadalupe fur seal (
7468:Antarctic fur seal (
7459:Galápagos fur seal (
7355:Galápagos sea lion (
6473:Common raccoon dog (
6325:Cape gray mongoose (
6265:Jackson's mongoose (
6225:Paracynictis selousi
6216:Cynictis penicillata
5840:Durrell's vontsira (
5428:Giant forest genet (
5220:Large Indian civet (
5160:Small Indian civet (
4769:Sunda leopard cat (
4545:Southern tiger cat (
4481:African golden cat (
4158:African palm civet (
4027:Born Free Foundation
3997:University of Oxford
3641:Animal Diversity Web
3590:Journal of Mammalogy
3295:(21): 3441–3449.e5.
3268:Bahlk, S.H. (2015).
2612:: e.T3748A10051312.
2411:Born Free Foundation
1618:Cooperative breeding
1513:side-striped jackals
1348:Trinomial authority
1257:Juliet Clutton-Brock
1241:Glover Morrill Allen
1203:convergent evolution
561:Literal translation
455:disease transference
9886:Mammals of Ethiopia
9376:M. aistoodonnivalis
9212:Hairy-nosed otter (
9129:Neotropical otter (
8909:Patagonian weasel (
8372:White-nosed coati (
7572:Cystophora cristata
7563:Erignathus barbatus
7296:Northern fur seal (
7073:Helarctos malayanus
6709:Atelocynus microtis
6416:U. cinereoargenteus
6307:Egyptian mongoose (
6234:Rhynchogale melleri
6232:Meller's mongoose (
6223:Selous's mongoose (
6207:Ichneumia albicauda
6161:Collared mongoose (
6035:Angolan kusimanse (
5919:Liberian mongoose (
5272:West African oyan (
5171:Civettictis civetta
5102:Golden palm civet (
5060:Masked palm civet (
5035:Arctictis binturong
5009:Hemigalus derbyanus
5007:Banded palm civet (
4998:Hose's palm civet (
4787:Rusty-spotted cat (
4041:, images and videos
4016:Supporting the EWCP
3965:Williams, S. 2004.
3935:on 26 December 2011
3907:MacDonald, David W.
3301:2018CBio...28E3441G
3238:2015CBio...25.2158K
3116:1994MolEc...3..301G
3070:10.1038/nature08837
3062:2010Natur.464..898V
2929:, p. 5 & 8
2799:2004MolEc..13.2275G
2642:, Vol. Säugetiere:
2438:Black-backed jackal
2317:Ethiopian Civil War
2155:Probably declining
2078:16–19 (as of 2000)
2048:19–23 (as of 2000)
1935:
1835:, and occasionally
1801:Big-headed mole rat
1509:black-backed jackal
1500:Painting (1926) by
1372:Nominate subspecies
1305:African golden wolf
1265:side-striped jackal
1158:Ethiopian Highlands
1107:Black-backed jackal
1087:Side-striped jackal
678:Ethiopian Civil War
666:Percy Powell-Cotton
551:
447:habitat degradation
390:Ethiopian Highlands
372:), also called the
140:Conservation status
9491:European polecat (
9324:Colombian weasel (
8974:(African polecats)
8602:Caucasian badger (
8530:(Eurasian badgers)
7980:(hog-nosed skunks)
7717:("southern seals")
7616:Halichoerus grypus
7547:("northern seals")
7419:Phocarctos hookeri
7324:Eumetopias jubatus
7322:Steller sea lion (
7104:Asian black bear (
6854:African wild dog (
6808:(wolf-like canids)
6085:Atilax paludinosus
6044:Common kusimanse (
5971:Dologale dybowskii
5943:Gambian mongoose (
5727:Western falanouc (
5718:Eastern falanouc (
5687:Cryptoprocta ferox
5592:Parahyaena brunnea
5347:Johnston's genet (
5302:Abyssinian genet (
5254:(African linsangs)
5162:Viverricula indica
5093:Brown palm civet (
5084:Asian palm civet (
4991:Cynogale bennettii
4982:Chrotogale owstoni
4878:European wildcat (
4869:Black-footed cat (
4459:Leptailurus serval
4427:Asian golden cat (
4193:(Asiatic linsangs)
4021:2009-02-12 at the
3956:Wolves of Ethiopia
3786:2016-10-06 at the
3570:Dartmouth College.
3559:Powell-Cotton 1902
3477:Behavioral Ecology
3381:Journal of Zoology
3174:Powell-Cotton 1902
3010:Parkyns, Mansfield
3000:, pp. 206–207
2998:Powell-Cotton 1902
2393:), and one in the
2300:
2018:Small population,
1930:
1903:highlands, and in
1809:
1707:
1654:
1644:Hunting behaviours
1579:
1505:
1407:, north and south
1320:canid of 72% grey
1211:
639:
631:
614:Historical account
546:
479:community outreach
469:'s Ethiopian Wolf
9848:
9847:
9794:Open Tree of Life
9555:Taxon identifiers
9546:
9545:
9542:
9541:
9538:
9537:
9534:
9533:
9530:
9529:
9509:Siberian weasel (
9500:American ermine (
9428:Japanese weasel (
9383:Mountain weasel (
9367:
9290:
9289:
9076:
9061:
9060:
9057:
9056:
9045:Striped polecat (
8995:Marbled polecat (
8977:
8962:
8961:
8934:
8882:
8881:
8861:Japanese marten (
8807:American marten (
8800:
8658:M. cucphuongensis
8649:
8632:
8631:
8620:European badger (
8593:Japanese badger (
8533:
8486:American badger (
8449:
8448:
8445:
8444:
8421:
8420:
8417:
8416:
8345:
8304:Northern olingo (
8288:
8263:Cozumel raccoon (
8238:
8162:
8161:
8116:
8075:
7983:
7964:
7906:
7905:
7902:
7901:
7898:
7897:
7894:
7893:
7890:
7889:
7880:Ommatophoca rossi
7853:Hydrurga leptonyx
7840:
7837:(Antarctic seals)
7810:M. angustirostris
7801:
7784:
7783:
7750:Monachus monachus
7737:
7720:
7705:
7704:
7701:
7700:
7550:
7533:
7516:
7515:
7378:Otaria flavescens
7367:
7366:
7278:
7246:Odobenus rosmarus
7210:
7209:
7206:
7205:
7202:
7201:
7198:
7197:
7161:
7138:
7137:
7134:
7133:
7036:Spectacled bear (
6994:
6970:
6969:
6966:
6965:
6962:
6961:
6810:
6797:
6796:
6716:Crab-eating fox (
6707:Short-eared dog (
6655:
6640:
6627:
6626:
6509:
6451:Otocyon megalotis
6395:
6362:
6361:
6358:
6357:
6354:
6353:
6350:
6349:
6346:
6345:
6214:Yellow mongoose (
6065:
6064:
5952:Banded mongoose (
5921:Liberiictus kuhni
5875:
5856:
5855:
5852:
5851:
5739:
5738:
5665:
5634:
5633:
5630:
5629:
5626:
5625:
5570:
5531:
5497:
5496:
5493:
5492:
5489:
5488:
5461:
5460:
5457:
5456:
5453:
5452:
5449:
5448:
5401:Servaline genet (
5320:Bourlon's genet (
5295:
5256:
5232:
5231:
5114:
5113:
5086:P. hermaphroditus
4938:
4937:
4934:
4933:
4930:
4929:
4906:
4905:
4902:
4901:
4898:
4897:
4894:
4893:
4890:
4889:
4851:African wildcat (
4808:
4807:
4778:Flat-headed cat (
4718:
4717:
4502:
4501:
4439:
4438:
4353:
4352:
4349:
4348:
4277:Clouded leopard (
4236:
4211:Spotted linsang (
4195:
4160:Nandinia binotata
3954:Morell, V. 2006.
3925:978-2-8317-0407-4
3860:Lydekker, Richard
3104:Molecular Ecology
3056:(7290): 898–902.
2787:Molecular Ecology
2716:Mammalian Species
2680:978-0-8018-8221-0
2667:"Order Carnivora"
2197:
2196:
1974:102 (as of 2010)
1869:free-ranging dogs
1552:supracaudal gland
1489:
1488:
1446:
1438:
1430:
1420:
1174:mitochondrial DNA
1154:
1153:
1149:
1148:
1140:
1139:
1131:
1130:
1122:
1121:
1058:
1057:
1029:
1028:
1000:
999:
973:
972:
944:
943:
915:
914:
886:
885:
877:
876:
744:Phylogenetic tree
722:local extinctions
698:Oxford University
611:
610:
535:rendering support
467:Oxford University
463:free-ranging dogs
360:
359:
354:
346:
338:
330:
322:
163:
16:(Redirected from
9913:
9841:
9840:
9828:
9827:
9815:
9814:
9802:
9801:
9789:
9788:
9776:
9775:
9763:
9762:
9750:
9749:
9737:
9736:
9724:
9723:
9711:
9710:
9698:
9697:
9685:
9684:
9672:
9671:
9659:
9658:
9646:
9645:
9636:
9635:
9623:
9622:
9610:
9609:
9597:
9596:
9595:
9582:
9581:
9580:
9550:
9482:Malayan weasel (
9401:Steppe polecat (
9374:Sichuan weasel (
9366:
9365:
9361:
9301:
9238:L. perspicillata
9203:Eurasian otter (
9080:
9075:
9074:
9070:
8981:
8976:
8975:
8971:
8950:Greater grison (
8933:
8932:
8928:
8902:
8893:
8843:Nilgiri marten (
8816:Pacific marten (
8799:
8798:
8794:
8720:
8703:M. subaurantiaca
8648:
8647:
8646:(ferret-badgers)
8643:
8537:
8532:
8531:
8527:
8479:
8468:
8455:
8349:
8344:
8343:
8339:
8287:
8286:
8282:
8237:
8236:
8232:
8173:
8115:
8114:
8113:(spotted skunks)
8110:
8074:
8073:
8069:
7982:
7981:
7977:
7968:
7963:
7962:
7954:
7925:
7912:
7869:Crabeater seal (
7844:
7839:
7838:
7834:
7800:
7799:
7798:(elephant seals)
7795:
7774:N. schauinslandi
7741:
7736:
7735:
7731:
7724:
7719:
7718:
7714:
7589:
7554:
7549:
7548:
7544:
7537:
7532:
7531:
7525:
7486:Brown fur seal (
7461:A. galapagoensis
7348:Z. californianus
7315:
7282:
7277:
7276:
7262:
7232:
7224:
7216:
7160:
7159:
7155:
7149:
7064:Melursus ursinus
7055:
6998:
6993:
6990:
6983:
6950:Ethiopian wolf (
6814:
6809:
6806:
6659:
6654:
6651:
6644:
6639:
6636:
6525:Blanford's fox (
6508:
6501:
6442:
6400:
6394:
6391:
6384:
6376:
6368:
6170:Ruddy mongoose (
6152:Javan mongoose (
6083:Marsh mongoose (
6076:
6055:C. platycephalus
5888:
5879:
5874:
5871:
5869:
5750:
5694:Malagasy civet (
5678:
5669:
5664:
5659:
5652:
5647:
5574:
5569:
5566:
5535:
5530:
5527:
5520:
5516:
5503:
5467:
5294:
5291:
5255:
5252:
5243:
5153:
5149:
5138:
5133:
5120:
5026:
4968:
4957:
4944:
4738:Otocolobus manul
4729:
4651:
4527:Geoffroy's cat (
4450:
4380:
4371:
4366:
4254:
4245:
4241:
4235:
4232:
4225:
4202:Banded linsang (
4194:
4191:
4183:
4172:
4151:
4140:
4132:
4124:
4069:
4062:
4055:
4046:
4034:(Canis simensis)
3944:
3942:
3940:
3934:
3928:. Archived from
3917:
3898:
3884:
3869:
3855:
3853:
3852:
3846:
3839:
3818:
3817:
3815:
3813:
3801:
3795:
3779:Gutema, T. M.,
3777:
3771:
3770:
3768:
3766:
3757:
3748:
3742:
3736:
3730:
3724:
3718:
3717:, pp. 27–28
3712:
3706:
3705:, pp. 26–27
3700:
3694:
3693:, pp. 22–26
3688:
3682:
3676:
3670:
3669:, pp. 40–46
3664:
3658:
3652:
3646:
3645:
3632:
3623:
3622:, pp. 19–20
3617:
3608:
3607:
3605:
3581:
3575:
3568:
3562:
3556:
3550:
3544:
3538:
3537:
3501:
3495:
3494:
3492:
3468:
3462:
3461:
3427:
3418:
3412:
3406:
3397:
3396:
3378:
3369:
3363:
3357:
3348:
3342:
3331:
3330:
3320:
3279:
3273:
3266:
3260:
3259:
3249:
3216:
3210:
3209:
3207:
3183:
3177:
3171:
3165:
3159:
3153:
3150:
3144:
3143:
3098:
3092:
3091:
3081:
3040:
3034:
3025:
3019:
3007:
3001:
2995:
2986:
2980:
2971:
2965:
2959:
2958:
2951:
2945:
2939:
2930:
2924:
2918:
2917:
2915:
2913:
2894:
2885:
2876:
2875:
2873:
2871:
2861:"Ethiopian Wolf"
2857:
2851:
2845:
2834:
2833:
2831:
2825:. Archived from
2793:(8): 2275–2286.
2784:
2771:
2762:
2761:
2759:
2758:
2752:
2746:. Archived from
2713:
2709:"Canis simensis"
2704:
2685:
2684:
2662:
2651:
2637:
2631:
2630:
2628:
2626:
2621:
2591:
2570:
2569:
2559:
2518:
2494:
2488:
2482:
2467:
2461:
2454:
2358:Encounters with
2248:in North Wollo.
2228:Canine distemper
2117:canine distemper
1963:Simien Mountains
1936:
1893:Simien Mountains
1704:Simien Mountains
1613:gestation period
1463:
1444:
1436:
1428:
1426:
1418:
1397:Simien Mountains
1380:
1342:
1237:John Edward Gray
1166:African wild dog
1113:
1093:
1080:
1072:
1049:
1043:African wild dog
1020:
991:
964:
935:
906:
868:
848:
847:
833:
826:
819:
811:
803:
795:
787:
779:
771:
757:
748:wolf-like canids
740:
739:
709:Simien Mountains
552:
523:
522:
513:Indigenous names
503:Ethiopian jackal
352:
344:
336:
328:
320:
298:
281:
261:C. simensis
178:
177:
157:
152:
151:
125:
115:
45:
38:Temporal range:
32:
21:
9921:
9920:
9916:
9915:
9914:
9912:
9911:
9910:
9851:
9850:
9849:
9844:
9836:
9831:
9823:
9818:
9810:
9805:
9797:
9792:
9784:
9781:Observation.org
9779:
9771:
9766:
9758:
9753:
9745:
9740:
9732:
9727:
9719:
9714:
9706:
9701:
9693:
9688:
9680:
9675:
9667:
9662:
9654:
9649:
9641:
9639:
9631:
9626:
9618:
9613:
9605:
9600:
9591:
9590:
9585:
9576:
9575:
9570:
9557:
9547:
9526:
9502:M. richardsonii
9446:European mink (
9363:
9362:
9360:
9350:
9342:American mink (
9315:Amazon weasel (
9286:
9244:
9220:
9187:
9170:
9146:
9095:
9072:
9071:
9069:
9053:
9020:
9003:
8973:
8972:
8970:
8958:
8941:Lesser grison (
8930:
8929:
8927:
8917:
8878:
8796:
8795:
8793:
8783:
8759:
8735:
8709:
8645:
8644:
8642:
8628:
8577:
8529:
8528:
8526:
8518:
8494:
8475:
8462:
8441:
8413:
8380:
8341:
8340:
8338:
8330:
8284:
8283:
8281:
8271:
8234:
8233:
8231:
8221:
8188:
8158:
8125:S. angustifrons
8112:
8111:
8109:
8099:
8072:(stink badgers)
8071:
8070:
8068:
8058:
8050:Striped skunk (
8025:
8019:C. semistriatus
7979:
7978:
7976:
7956:
7955:
7953:
7945:
7939:Ailurus fulgens
7919:
7886:
7836:
7835:
7833:
7825:
7797:
7796:
7794:
7780:
7756:
7733:
7732:
7730:
7716:
7715:
7713:
7697:
7655:
7622:
7578:
7546:
7545:
7543:
7529:
7527:
7526:
7524:
7512:
7425:
7408:
7384:
7363:
7330:
7304:
7271:
7266:
7264:
7263:
7261:
7257:
7252:
7226:
7222:
7194:
7174:
7157:
7156:
7130:
7079:
7044:
7020:
6991:
6989:
6979:
6958:
6887:Golden jackal (
6871:
6845:
6807:
6805:
6793:
6724:
6718:Cerdocyon thous
6698:
6674:
6652:
6650:
6637:
6635:
6623:
6588:Rüppell's fox (
6502:
6500:
6490:
6475:N. procyonoides
6457:
6449:Bat-eared fox (
6431:
6396:
6392:
6378:
6374:
6342:
6327:H. pulverulenta
6282:
6240:
6196:
6163:U. semitorquata
6118:U. auropunctata
6100:
6061:
6010:
5977:
5960:
5927:
5910:
5872:
5870:
5865:
5864:
5848:
5815:
5798:
5765:
5759:Galidia elegans
5735:
5702:
5662:
5660:
5658:
5648:
5643:
5622:
5614:Spotted hyena (
5598:
5581:Striped hyena (
5567:
5565:
5557:
5528:
5526:
5510:
5485:
5445:
5383:Aquatic genet (
5374:Pardine genet (
5311:Angolan genet (
5292:
5290:
5280:
5265:P. richardsonii
5253:
5251:
5228:
5211:Malayan civet (
5177:
5169:African civet (
5145:
5144:
5134:
5129:
5128:
5124:
5110:
5068:
5015:
5000:Diplogale hosei
4964:
4951:
4926:
4886:
4804:
4744:
4724:
4714:
4690:
4673:
4646:
4640:
4614:Eurasian lynx (
4589:
4498:
4465:
4445:
4435:
4402:
4375:
4367:
4362:
4345:
4285:
4237:
4233:
4219:
4192:
4179:
4166:
4147:
4134:
4130:
4118:
4080:
4073:
4032:Ethiopian wolf
4023:Wayback Machine
3980:
3951:
3949:Further reading
3938:
3936:
3932:
3926:
3915:
3901:
3889:Rüppell, Eduard
3887:
3872:
3858:
3850:
3848:
3844:
3837:
3830:
3827:
3822:
3821:
3811:
3809:
3808:. New Scientist
3803:
3802:
3798:
3794:(November 2015)
3788:Wayback Machine
3778:
3774:
3764:
3762:
3755:
3750:
3749:
3745:
3737:
3733:
3725:
3721:
3713:
3709:
3701:
3697:
3689:
3685:
3677:
3673:
3665:
3661:
3653:
3649:
3634:
3633:
3626:
3618:
3611:
3583:
3582:
3578:
3569:
3565:
3557:
3553:
3545:
3541:
3518:10.1038/nrg2664
3506:Nat. Rev. Genet
3503:
3502:
3498:
3470:
3469:
3465:
3425:
3420:
3419:
3415:
3407:
3400:
3376:
3371:
3370:
3366:
3358:
3351:
3343:
3334:
3289:Current Biology
3281:
3280:
3276:
3267:
3263:
3232:(16): 2158–65.
3226:Current Biology
3218:
3217:
3213:
3185:
3184:
3180:
3172:
3168:
3160:
3156:
3151:
3147:
3100:
3099:
3095:
3042:
3041:
3037:
3026:
3022:
3008:
3004:
2996:
2989:
2981:
2974:
2966:
2962:
2957:. 28 July 2012.
2953:
2952:
2948:
2940:
2933:
2925:
2921:
2911:
2909:
2907:
2892:
2887:
2886:
2879:
2869:
2867:
2859:
2858:
2854:
2846:
2837:
2829:
2782:
2773:
2772:
2765:
2756:
2754:
2750:
2728:10.2307/3504136
2711:
2706:
2705:
2688:
2681:
2664:
2663:
2654:
2638:
2634:
2624:
2622:
2593:
2592:
2573:
2520:
2519:
2508:
2503:
2498:
2497:
2489:
2485:
2468:
2464:
2455:
2451:
2446:
2434:
2372:
2356:
2342:
2313:
2289:
2271:
2254:
2236:
2219:
2202:
2004:, South Gondar
1885:
1794:
1696:
1691:
1646:
1622:pseudopregnancy
1601:
1584:
1568:
1515:as well as the
1494:
1466:de Beaux, 1922
1458:
1454:
1445:(Heuglin, 1862)
1443:
1439:
1435:
1431:
1429:(Heuglin, 1862)
1427:
1424:C. s. semiensis
1422:
1421:
1417:
1415:C. s. crinensis
1375:
1369:
1365:
1330:
1293:
1187:Canis evolution
1150:
1141:
1132:
1123:
1059:
1030:
1001:
986:Ethiopian wolf
974:
945:
916:
887:
878:
842:
763: 3.5
738:
686:Sanetti Plateau
616:
606:
599:
585:
576:
558:Indigenous name
544:
543:
542:
533:Without proper
524:
520:
515:
507:Abyssinian wolf
487:
445:, resulting in
424:mountain ranges
289:
283:
277:
264:
172:
164:
153:
149:
142:
132:Sanetti Plateau
116:
114:
113:
112:
111:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
40:
39:
36:
28:
23:
22:
15:
12:
11:
5:
9919:
9917:
9909:
9908:
9903:
9898:
9893:
9888:
9883:
9878:
9873:
9868:
9863:
9853:
9852:
9846:
9845:
9843:
9842:
9829:
9816:
9803:
9790:
9777:
9764:
9751:
9738:
9725:
9712:
9699:
9686:
9673:
9660:
9647:
9637:
9624:
9620:canis-simensis
9611:
9607:Canis_simensis
9598:
9593:Canis simensis
9583:
9567:
9565:
9563:Canis simensis
9559:
9558:
9553:
9544:
9543:
9540:
9539:
9536:
9535:
9532:
9531:
9528:
9527:
9525:
9524:
9520:M. strigidorsa
9515:
9506:
9497:
9488:
9479:
9473:Least weasel (
9470:
9461:
9452:
9443:
9434:
9425:
9419:Haida ermine (
9416:
9407:
9403:M. eversmannii
9398:
9389:
9380:
9370:
9368:
9352:
9351:
9349:
9348:
9339:
9330:
9321:
9311:
9309:
9298:
9292:
9291:
9288:
9287:
9285:
9284:
9275:
9266:
9256:
9254:
9246:
9245:
9243:
9242:
9232:
9230:
9222:
9221:
9219:
9218:
9209:
9199:
9197:
9189:
9188:
9186:
9185:
9175:
9172:
9171:
9169:
9168:
9158:
9156:
9148:
9147:
9145:
9144:
9135:
9131:L. longicaudis
9126:
9120:Marine otter (
9117:
9107:
9105:
9097:
9096:
9094:
9093:
9083:
9077:
9063:
9062:
9059:
9058:
9055:
9054:
9052:
9051:
9042:
9032:
9030:
9022:
9021:
9019:
9018:
9008:
9005:
9004:
9002:
9001:
8991:
8989:
8978:
8964:
8963:
8960:
8959:
8957:
8956:
8947:
8937:
8935:
8919:
8918:
8916:
8915:
8905:
8899:
8890:
8884:
8883:
8880:
8879:
8877:
8876:
8867:
8858:
8849:
8840:
8834:Beech marten (
8831:
8822:
8813:
8803:
8801:
8785:
8784:
8782:
8781:
8771:
8769:
8761:
8760:
8758:
8757:
8747:
8745:
8737:
8736:
8734:
8733:
8723:
8717:
8711:
8710:
8708:
8707:
8698:
8689:
8680:
8671:
8662:
8652:
8650:
8634:
8633:
8630:
8629:
8627:
8626:
8617:
8611:Asian badger (
8608:
8599:
8589:
8587:
8579:
8578:
8576:
8575:
8566:
8557:
8553:A. albogularis
8547:
8545:
8534:
8520:
8519:
8517:
8516:
8510:Honey badger (
8506:
8504:
8496:
8495:
8493:
8492:
8482:
8476:
8471:
8464:
8463:
8458:
8451:
8450:
8447:
8446:
8443:
8442:
8440:
8439:
8438:
8437:
8431:
8429:
8423:
8422:
8419:
8418:
8415:
8414:
8412:
8411:
8402:
8392:
8390:
8382:
8381:
8379:
8378:
8369:
8359:
8357:
8346:
8332:
8331:
8329:
8328:
8319:
8310:
8301:
8291:
8289:
8273:
8272:
8270:
8269:
8260:
8251:
8247:P. cancrivorus
8241:
8239:
8223:
8222:
8220:
8219:
8215:B. sumichrasti
8210:
8200:
8198:
8190:
8189:
8187:
8186:
8176:
8170:
8164:
8163:
8160:
8159:
8157:
8156:
8147:
8138:
8129:
8119:
8117:
8101:
8100:
8098:
8097:
8088:
8078:
8076:
8060:
8059:
8057:
8056:
8047:
8041:Hooded skunk (
8037:
8035:
8027:
8026:
8024:
8023:
8014:
8005:
7996:
7986:
7984:
7965:
7947:
7946:
7944:
7943:
7933:
7931:
7921:
7920:
7915:
7908:
7907:
7904:
7903:
7900:
7899:
7896:
7895:
7892:
7891:
7888:
7887:
7885:
7884:
7875:
7866:
7860:Weddell seal (
7857:
7851:Leopard seal (
7847:
7841:
7827:
7826:
7824:
7823:
7814:
7804:
7802:
7786:
7785:
7782:
7781:
7779:
7778:
7768:
7766:
7758:
7757:
7755:
7754:
7744:
7738:
7721:
7707:
7706:
7703:
7702:
7699:
7698:
7696:
7695:
7686:
7677:
7671:Caspian seal (
7667:
7665:
7657:
7656:
7654:
7653:
7644:
7638:Spotted seal (
7634:
7632:
7624:
7623:
7621:
7620:
7611:
7602:
7592:
7586:
7580:
7579:
7577:
7576:
7567:
7561:Bearded seal (
7557:
7551:
7534:
7530:or true seals)
7528:(earless seals
7518:
7517:
7514:
7513:
7511:
7510:
7501:
7492:
7483:
7474:
7465:
7456:
7447:
7437:
7435:
7427:
7426:
7424:
7423:
7413:
7410:
7409:
7407:
7406:
7396:
7394:
7386:
7385:
7383:
7382:
7372:
7369:
7368:
7365:
7364:
7362:
7361:
7352:
7342:
7340:
7332:
7331:
7329:
7328:
7318:
7312:
7306:
7305:
7303:
7302:
7292:
7290:
7279:
7254:
7253:
7251:
7250:
7240:
7238:
7228:
7227:
7219:
7212:
7211:
7208:
7207:
7204:
7203:
7200:
7199:
7196:
7195:
7193:
7192:
7191:
7190:
7184:
7182:
7176:
7175:
7173:
7172:
7171:
7170:
7164:
7162:
7146:
7140:
7139:
7136:
7135:
7132:
7131:
7129:
7128:
7119:
7110:
7101:
7091:
7089:
7081:
7080:
7078:
7077:
7068:
7058:
7052:
7046:
7045:
7043:
7042:
7032:
7030:
7022:
7021:
7019:
7018:
7014:A. melanoleuca
7008:
7006:
6995:
6980:
6975:
6972:
6971:
6968:
6967:
6964:
6963:
6960:
6959:
6957:
6956:
6947:
6938:
6932:Eastern wolf (
6929:
6920:
6914:African wolf (
6911:
6902:
6896:Domestic dog (
6893:
6883:
6881:
6873:
6872:
6870:
6869:
6860:
6850:
6847:
6846:
6844:
6843:
6834:
6824:
6822:
6811:
6799:
6798:
6795:
6794:
6792:
6791:
6782:
6776:Sechuran fox (
6773:
6769:L. gymnocercus
6764:
6755:
6749:Darwin's fox (
6746:
6736:
6734:
6726:
6725:
6723:
6722:
6713:
6703:
6700:
6699:
6697:
6696:
6686:
6684:
6676:
6675:
6673:
6672:
6662:
6656:
6641:
6629:
6628:
6625:
6624:
6622:
6621:
6612:
6603:
6594:
6585:
6576:
6567:
6558:
6549:
6540:
6531:
6522:
6518:V. bengalensis
6512:
6510:
6492:
6491:
6489:
6488:
6479:
6469:
6467:
6459:
6458:
6456:
6455:
6445:
6439:
6433:
6432:
6430:
6429:
6420:
6410:
6408:
6397:
6387:
6380:
6379:
6371:
6364:
6363:
6360:
6359:
6356:
6355:
6352:
6351:
6348:
6347:
6344:
6343:
6341:
6340:
6331:
6322:
6313:
6304:
6294:
6292:
6284:
6283:
6281:
6280:
6271:
6262:
6258:B. crassicauda
6252:
6250:
6242:
6241:
6239:
6238:
6229:
6220:
6211:
6201:
6198:
6197:
6195:
6194:
6185:
6176:
6167:
6158:
6149:
6140:
6131:
6122:
6112:
6110:
6102:
6101:
6099:
6098:
6089:
6079:
6073:
6067:
6066:
6063:
6062:
6060:
6059:
6050:
6041:
6032:
6022:
6020:
6012:
6011:
6009:
6008:
5999:
5989:
5987:
5979:
5978:
5976:
5975:
5965:
5962:
5961:
5959:
5958:
5949:
5939:
5937:
5929:
5928:
5926:
5925:
5915:
5912:
5911:
5909:
5908:
5898:
5896:
5885:
5876:
5858:
5857:
5854:
5853:
5850:
5849:
5847:
5846:
5837:
5827:
5825:
5817:
5816:
5814:
5813:
5803:
5800:
5799:
5797:
5796:
5792:G. grandidieri
5787:
5777:
5775:
5767:
5766:
5764:
5763:
5753:
5747:
5741:
5740:
5737:
5736:
5734:
5733:
5724:
5714:
5712:
5704:
5703:
5701:
5700:
5691:
5681:
5675:
5666:
5649:
5639:
5636:
5635:
5632:
5631:
5628:
5627:
5624:
5623:
5621:
5620:
5610:
5608:
5600:
5599:
5597:
5596:
5587:
5577:
5571:
5559:
5558:
5556:
5555:
5545:
5543:
5532:
5512:
5511:
5506:
5499:
5498:
5495:
5494:
5491:
5490:
5487:
5486:
5484:
5483:
5482:
5481:
5475:
5473:
5463:
5462:
5459:
5458:
5455:
5454:
5451:
5450:
5447:
5446:
5444:
5443:
5434:
5425:
5416:
5407:
5398:
5389:
5380:
5371:
5362:
5356:Letaba genet (
5353:
5344:
5338:Common genet (
5335:
5326:
5317:
5308:
5298:
5296:
5282:
5281:
5279:
5278:
5269:
5259:
5257:
5240:
5234:
5233:
5230:
5229:
5227:
5226:
5217:
5208:
5199:
5189:
5187:
5179:
5178:
5176:
5175:
5166:
5156:
5150:
5135:
5123:
5116:
5115:
5112:
5111:
5109:
5108:
5104:P. zeylonensis
5099:
5090:
5080:
5078:
5070:
5069:
5067:
5066:
5062:Paguma larvata
5057:
5048:
5039:
5029:
5023:
5017:
5016:
5014:
5013:
5004:
4995:
4986:
4976:
4974:
4965:
4960:
4953:
4952:
4947:
4940:
4939:
4936:
4935:
4932:
4931:
4928:
4927:
4925:
4924:
4923:
4922:
4916:
4914:
4908:
4907:
4904:
4903:
4900:
4899:
4896:
4895:
4892:
4891:
4888:
4887:
4885:
4884:
4875:
4866:
4857:
4848:
4839:
4833:Domestic cat (
4830:
4820:
4818:
4810:
4809:
4806:
4805:
4803:
4802:
4793:
4789:P. rubiginosus
4784:
4775:
4766:
4762:P. bengalensis
4756:
4754:
4746:
4745:
4743:
4742:
4736:Pallas's cat (
4732:
4726:
4720:
4719:
4716:
4715:
4713:
4712:
4702:
4700:
4692:
4691:
4689:
4688:
4678:
4675:
4674:
4672:
4671:
4661:
4659:
4648:
4642:
4641:
4639:
4638:
4629:
4623:Iberian lynx (
4620:
4611:
4601:
4599:
4591:
4590:
4588:
4587:
4578:
4569:
4560:
4551:
4542:
4533:
4524:
4514:
4512:
4504:
4503:
4500:
4499:
4497:
4496:
4487:
4477:
4475:
4467:
4466:
4464:
4463:
4453:
4447:
4441:
4440:
4437:
4436:
4434:
4433:
4424:
4414:
4412:
4404:
4403:
4401:
4400:
4390:
4388:
4377:
4368:
4358:
4355:
4354:
4351:
4350:
4347:
4346:
4344:
4343:
4334:
4325:
4316:
4307:
4301:Snow leopard (
4297:
4295:
4287:
4286:
4284:
4283:
4274:
4264:
4262:
4251:
4238:
4228:
4221:
4220:
4218:
4217:
4208:
4198:
4196:
4180:
4175:
4168:
4167:
4165:
4164:
4154:
4148:
4143:
4136:
4135:
4127:
4120:
4119:
4117:
4116:
4114:Laurasiatheria
4110:
4104:
4098:
4092:
4085:
4082:
4081:
4074:
4072:
4071:
4064:
4057:
4049:
4043:
4042:
4029:
4013:
4000:
3986:
3979:
3978:External links
3976:
3975:
3974:
3963:
3950:
3947:
3946:
3945:
3924:
3899:
3885:
3870:
3856:
3826:
3823:
3820:
3819:
3796:
3772:
3743:
3731:
3719:
3707:
3695:
3683:
3671:
3659:
3647:
3624:
3609:
3596:(1): 129–137.
3576:
3563:
3551:
3549:, pp. 3–4
3539:
3512:(11): 783–96.
3496:
3463:
3413:
3411:, pp. 4–5
3398:
3387:(3): 351–361.
3364:
3349:
3332:
3274:
3261:
3211:
3178:
3166:
3164:, pp. 2–3
3154:
3145:
3093:
3035:
3020:
3002:
2987:
2972:
2960:
2946:
2931:
2919:
2906:978-2831707860
2905:
2877:
2852:
2850:, pp. 7–8
2835:
2832:on 2013-05-16.
2778:Canis simensis
2763:
2686:
2679:
2652:
2632:
2598:Canis simensis
2571:
2505:
2504:
2502:
2499:
2496:
2495:
2483:
2462:
2448:
2447:
2445:
2442:
2441:
2440:
2433:
2430:
2371:
2368:
2364:Canis lupaster
2355:
2352:
2341:
2338:
2312:
2309:
2288:
2285:
2270:
2267:
2253:
2250:
2235:
2232:
2218:
2215:
2201:
2198:
2195:
2194:
2191:
2185:
2182:
2179:
2176:
2175:1,141 km
2173:
2170:Bale Mountains
2166:
2165:
2162:
2159:
2156:
2153:
2150:
2147:
2141:Arsi Mountains
2137:
2136:
2133:
2126:
2123:
2120:
2113:
2110:
2100:
2099:
2091:
2085:
2082:
2079:
2076:
2073:
2069:
2068:
2057:
2054:
2052:
2049:
2046:
2043:
2035:
2034:
2023:
2016:
2014:
2011:
2008:
2005:
1998:
1997:
1990:
1981:
1978:
1975:
1972:
1969:
1959:
1958:
1955:
1952:
1949:
1946:
1943:
1940:
1933:Canis simensis
1925:Bale Mountains
1901:southern Wollo
1884:
1881:
1845:mountain nyala
1825:highland hares
1793:
1790:
1782:giant lobelias
1695:
1692:
1690:
1687:
1659:mountain nyala
1645:
1642:
1600:
1597:
1583:
1580:
1576:Bale Mountains
1567:
1564:
1493:
1490:
1487:
1486:
1484:
1482:Bale Mountains
1474:
1467:
1464:
1456:C. s. citernii
1448:
1447:
1433:C. s. simensis
1412:
1389:
1387:
1381:
1367:C. s. simensis
1359:
1358:
1355:
1352:
1349:
1346:
1329:
1326:
1292:
1285:
1191:
1190:
1152:
1151:
1147:
1146:
1143:
1142:
1138:
1137:
1134:
1133:
1129:
1128:
1125:
1124:
1120:
1119:
1116:
1115:
1103:
1100:
1099:
1096:
1095:
1083:
1078:
1076:
1070:
1068:
1065:
1064:
1061:
1060:
1056:
1055:
1052:
1051:
1039:
1036:
1035:
1032:
1031:
1027:
1026:
1023:
1022:
1010:
1007:
1006:
1003:
1002:
998:
997:
994:
993:
983:
980:
979:
976:
975:
971:
970:
967:
966:
954:
951:
950:
947:
946:
942:
941:
938:
937:
925:
922:
921:
918:
917:
913:
912:
909:
908:
896:
893:
892:
889:
888:
884:
883:
880:
879:
875:
874:
871:
870:
858:
855:
854:
851:
850:
836:
831:
829:
824:
822:
817:
815:
809:
807:
801:
799:
793:
791:
785:
783:
777:
775:
769:
767:
755:
752:
751:
737:
734:
717:Arsi Mountains
655:Charles Darwin
647:British Museum
643:Eduard Rüppell
623:Canis simensis
615:
612:
609:
608:
605:Horse's jackal
603:
597:Jeedala fardaa
594:
588:
587:
582:
569:
563:
562:
559:
556:
549:Canis simensis
537:, you may see
525:
518:
517:
516:
514:
511:
486:
483:
428:Bale Mountains
388:native to the
369:Canis simensis
364:Ethiopian wolf
358:
357:
356:
355:
353:de Beaux, 1922
347:
339:
331:
323:
312:
311:
305:
304:
300:
299:
291:
290:
284:
279:Canis simensis
273:
272:
266:
265:
258:
256:
252:
251:
244:
240:
239:
234:
230:
229:
224:
220:
219:
214:
210:
209:
204:
200:
199:
194:
190:
189:
184:
180:
179:
166:
165:
147:
144:
143:
138:
135:
134:
127:
126:
118:
117:
109:
108:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
46:
37:
35:Ethiopian wolf
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
9918:
9907:
9904:
9902:
9899:
9897:
9894:
9892:
9889:
9887:
9884:
9882:
9879:
9877:
9874:
9872:
9869:
9867:
9864:
9862:
9859:
9858:
9856:
9839:
9834:
9830:
9826:
9821:
9817:
9813:
9808:
9804:
9800:
9795:
9791:
9787:
9782:
9778:
9774:
9769:
9765:
9761:
9756:
9752:
9748:
9743:
9739:
9735:
9730:
9726:
9722:
9717:
9713:
9709:
9704:
9700:
9696:
9691:
9687:
9683:
9678:
9674:
9670:
9665:
9661:
9657:
9652:
9648:
9644:
9638:
9634:
9629:
9625:
9621:
9616:
9612:
9608:
9603:
9599:
9594:
9588:
9584:
9579:
9573:
9569:
9568:
9566:
9564:
9560:
9556:
9551:
9523:
9521:
9516:
9514:
9512:
9507:
9505:
9503:
9498:
9496:
9494:
9489:
9487:
9485:
9480:
9478:
9476:
9471:
9469:
9467:
9462:
9460:
9458:
9457:M. lutreolina
9453:
9451:
9449:
9444:
9442:
9440:
9435:
9433:
9431:
9426:
9424:
9422:
9417:
9415:
9413:
9408:
9406:
9404:
9399:
9397:
9395:
9390:
9388:
9386:
9381:
9379:
9377:
9372:
9371:
9369:
9359:
9358:
9353:
9347:
9345:
9340:
9338:
9336:
9331:
9329:
9327:
9322:
9320:
9318:
9313:
9312:
9310:
9308:
9307:
9302:
9299:
9297:
9293:
9283:
9281:
9276:
9274:
9272:
9267:
9265:
9263:
9258:
9257:
9255:
9253:
9252:
9247:
9241:
9239:
9234:
9233:
9231:
9229:
9228:
9223:
9217:
9215:
9210:
9208:
9206:
9201:
9200:
9198:
9196:
9195:
9190:
9184:
9182:
9177:
9176:
9173:
9167:
9165:
9160:
9159:
9157:
9155:
9154:
9149:
9143:
9141:
9136:
9134:
9132:
9127:
9125:
9123:
9118:
9116:
9114:
9113:L. canadensis
9109:
9108:
9106:
9104:
9103:
9098:
9092:
9090:
9087:Giant otter (
9085:
9084:
9081:
9078:
9068:
9064:
9050:
9048:
9043:
9041:
9039:
9034:
9033:
9031:
9029:
9028:
9023:
9017:
9015:
9010:
9009:
9006:
9000:
8998:
8993:
8992:
8990:
8988:
8987:
8982:
8979:
8969:
8965:
8955:
8953:
8948:
8946:
8944:
8939:
8938:
8936:
8926:
8925:
8920:
8914:
8912:
8907:
8906:
8903:
8900:
8898:
8894:
8891:
8889:
8885:
8875:
8873:
8868:
8866:
8864:
8859:
8857:
8855:
8850:
8848:
8846:
8845:M. gwatkinsii
8841:
8839:
8837:
8832:
8830:
8828:
8823:
8821:
8819:
8814:
8812:
8810:
8805:
8804:
8802:
8792:
8791:
8786:
8780:
8778:
8773:
8772:
8770:
8768:
8767:
8762:
8756:
8754:
8749:
8748:
8746:
8744:
8743:
8738:
8732:
8730:
8725:
8724:
8721:
8718:
8716:
8712:
8706:
8704:
8699:
8697:
8695:
8690:
8688:
8686:
8685:M. orientalis
8681:
8679:
8677:
8672:
8670:
8668:
8663:
8661:
8659:
8654:
8653:
8651:
8641:
8640:
8635:
8625:
8623:
8618:
8616:
8614:
8609:
8607:
8605:
8600:
8598:
8596:
8591:
8590:
8588:
8586:
8585:
8580:
8574:
8572:
8567:
8565:
8563:
8558:
8556:
8554:
8549:
8548:
8546:
8544:
8543:
8538:
8535:
8525:
8521:
8515:
8513:
8508:
8507:
8505:
8503:
8502:
8497:
8491:
8489:
8488:Taxidea taxus
8484:
8483:
8480:
8477:
8474:
8469:
8465:
8461:
8456:
8452:
8435:
8434:
8433:
8432:
8430:
8428:
8424:
8410:
8408:
8403:
8401:
8399:
8398:N. meridensis
8394:
8393:
8391:
8389:
8388:
8383:
8377:
8375:
8370:
8368:
8366:
8361:
8360:
8358:
8356:
8355:
8350:
8347:
8337:
8333:
8327:
8325:
8320:
8318:
8316:
8311:
8309:
8307:
8302:
8300:
8298:
8293:
8292:
8290:
8280:
8279:
8274:
8268:
8266:
8261:
8259:
8257:
8252:
8250:
8248:
8243:
8242:
8240:
8230:
8229:
8224:
8218:
8216:
8211:
8209:
8207:
8202:
8201:
8199:
8197:
8196:
8191:
8185:
8183:
8178:
8177:
8174:
8171:
8169:
8165:
8155:
8153:
8148:
8146:
8144:
8139:
8137:
8135:
8130:
8128:
8126:
8121:
8120:
8118:
8108:
8107:
8102:
8096:
8094:
8089:
8087:
8085:
8084:M. javanensis
8080:
8079:
8077:
8067:
8066:
8061:
8055:
8053:
8048:
8046:
8044:
8039:
8038:
8036:
8034:
8033:
8028:
8022:
8020:
8015:
8013:
8011:
8010:C. leuconotus
8006:
8004:
8002:
8001:C. humboldtii
7997:
7995:
7993:
7988:
7987:
7985:
7975:
7974:
7969:
7966:
7960:
7952:
7948:
7942:
7940:
7935:
7934:
7932:
7930:
7926:
7922:
7918:
7913:
7909:
7883:
7881:
7876:
7874:
7872:
7867:
7865:
7863:
7858:
7856:
7854:
7849:
7848:
7845:
7842:
7832:
7828:
7822:
7820:
7815:
7813:
7811:
7806:
7805:
7803:
7793:
7792:
7787:
7777:
7775:
7770:
7769:
7767:
7765:
7764:
7759:
7753:
7751:
7746:
7745:
7742:
7739:
7729:
7725:
7722:
7712:
7708:
7694:
7692:
7689:Baikal seal (
7687:
7685:
7683:
7680:Ringed seal (
7678:
7676:
7674:
7669:
7668:
7666:
7664:
7663:
7658:
7652:
7650:
7647:Harbor seal (
7645:
7643:
7641:
7636:
7635:
7633:
7631:
7630:
7625:
7619:
7617:
7612:
7610:
7608:
7605:Ribbon seal (
7603:
7601:
7599:
7594:
7593:
7590:
7587:
7585:
7581:
7575:
7573:
7570:Hooded seal (
7568:
7566:
7564:
7559:
7558:
7555:
7552:
7542:
7538:
7535:
7523:
7519:
7509:
7507:
7506:A. tropicalis
7502:
7500:
7498:
7493:
7491:
7489:
7484:
7482:
7480:
7475:
7473:
7471:
7466:
7464:
7462:
7457:
7455:
7453:
7448:
7446:
7444:
7439:
7438:
7436:
7434:
7433:
7432:Arctocephalus
7428:
7422:
7420:
7415:
7414:
7411:
7405:
7403:
7398:
7397:
7395:
7393:
7392:
7387:
7381:
7379:
7374:
7373:
7370:
7360:
7358:
7357:Z. wollebaeki
7353:
7351:
7349:
7344:
7343:
7341:
7339:
7338:
7333:
7327:
7325:
7320:
7319:
7316:
7313:
7311:
7307:
7301:
7299:
7294:
7293:
7291:
7289:
7288:
7283:
7280:
7274:
7269:
7265:(eared seals:
7260:
7255:
7249:
7247:
7242:
7241:
7239:
7237:
7233:
7229:
7225:
7217:
7213:
7188:
7187:
7186:
7185:
7183:
7181:
7177:
7168:
7167:
7166:
7165:
7163:
7154:
7150:
7147:
7145:
7141:
7127:
7125:
7120:
7118:
7116:
7111:
7109:
7107:
7106:U. thibetanus
7102:
7100:
7098:
7097:U. americanus
7093:
7092:
7090:
7088:
7087:
7082:
7076:
7074:
7069:
7067:
7065:
7060:
7059:
7056:
7053:
7051:
7047:
7041:
7039:
7034:
7033:
7031:
7029:
7028:
7023:
7017:
7015:
7012:Giant panda (
7010:
7009:
7007:
7005:
7004:
6999:
6996:
6988:
6984:
6981:
6978:
6973:
6955:
6953:
6948:
6946:
6944:
6939:
6937:
6935:
6930:
6928:
6926:
6921:
6919:
6917:
6912:
6910:
6908:
6903:
6901:
6899:
6898:C. familiaris
6894:
6892:
6890:
6885:
6884:
6882:
6880:
6879:
6874:
6868:
6866:
6861:
6859:
6857:
6856:Lycaon pictus
6852:
6851:
6848:
6842:
6840:
6835:
6833:
6831:
6826:
6825:
6823:
6821:
6820:
6815:
6812:
6804:
6800:
6790:
6788:
6783:
6781:
6779:
6774:
6772:
6770:
6765:
6763:
6761:
6756:
6754:
6752:
6747:
6745:
6743:
6738:
6737:
6735:
6733:
6732:
6727:
6721:
6719:
6714:
6712:
6710:
6705:
6704:
6701:
6695:
6693:
6688:
6687:
6685:
6683:
6682:
6677:
6671:
6669:
6664:
6663:
6660:
6657:
6649:
6645:
6642:
6634:
6630:
6620:
6618:
6613:
6611:
6609:
6604:
6602:
6600:
6595:
6593:
6591:
6586:
6584:
6582:
6577:
6575:
6573:
6568:
6566:
6564:
6559:
6557:
6555:
6552:Tibetan fox (
6550:
6548:
6546:
6541:
6539:
6537:
6532:
6530:
6528:
6523:
6521:
6519:
6514:
6513:
6511:
6506:
6499:
6498:
6493:
6487:
6485:
6484:N. viverrinus
6480:
6478:
6476:
6471:
6470:
6468:
6466:
6465:
6460:
6454:
6452:
6447:
6446:
6443:
6440:
6438:
6434:
6428:
6426:
6425:U. littoralis
6421:
6419:
6417:
6412:
6411:
6409:
6407:
6406:
6401:
6398:
6390:
6385:
6381:
6377:
6369:
6365:
6339:
6337:
6332:
6330:
6328:
6323:
6321:
6319:
6314:
6312:
6310:
6305:
6303:
6301:
6300:H. flavescens
6296:
6295:
6293:
6291:
6290:
6285:
6279:
6277:
6272:
6270:
6268:
6263:
6261:
6259:
6254:
6253:
6251:
6249:
6248:
6243:
6237:
6235:
6230:
6228:
6226:
6221:
6219:
6217:
6212:
6210:
6208:
6203:
6202:
6199:
6193:
6191:
6190:U. vitticolla
6186:
6184:
6182:
6177:
6175:
6173:
6168:
6166:
6164:
6159:
6157:
6155:
6150:
6148:
6146:
6141:
6139:
6137:
6132:
6130:
6128:
6123:
6121:
6119:
6114:
6113:
6111:
6109:
6108:
6103:
6097:
6095:
6094:Xenogale naso
6090:
6088:
6086:
6081:
6080:
6077:
6074:
6072:
6068:
6058:
6056:
6051:
6049:
6047:
6042:
6040:
6038:
6033:
6031:
6029:
6024:
6023:
6021:
6019:
6018:
6013:
6007:
6005:
6000:
5998:
5996:
5991:
5990:
5988:
5986:
5985:
5980:
5974:
5972:
5967:
5966:
5963:
5957:
5955:
5950:
5948:
5946:
5941:
5940:
5938:
5936:
5935:
5930:
5924:
5922:
5917:
5916:
5913:
5907:
5905:
5900:
5899:
5897:
5895:
5894:
5889:
5886:
5884:
5880:
5877:
5868:
5867:sensu stricto
5863:
5859:
5845:
5843:
5838:
5836:
5834:
5829:
5828:
5826:
5824:
5823:
5818:
5812:
5810:
5805:
5804:
5801:
5795:
5793:
5788:
5786:
5784:
5779:
5778:
5776:
5774:
5773:
5768:
5762:
5760:
5755:
5754:
5751:
5748:
5746:
5742:
5732:
5730:
5725:
5723:
5721:
5716:
5715:
5713:
5711:
5710:
5705:
5699:
5697:
5696:Fossa fossana
5692:
5690:
5688:
5683:
5682:
5679:
5676:
5674:
5670:
5667:
5657:
5653:
5650:
5646:
5642:
5637:
5619:
5617:
5612:
5611:
5609:
5607:
5606:
5601:
5595:
5593:
5590:Brown hyena (
5588:
5586:
5584:
5583:Hyaena hyaena
5579:
5578:
5575:
5572:
5564:
5560:
5554:
5552:
5547:
5546:
5544:
5542:
5541:
5536:
5533:
5525:
5521:
5517:
5513:
5509:
5504:
5500:
5479:
5478:
5477:
5476:
5474:
5472:
5468:
5464:
5442:
5440:
5435:
5433:
5431:
5426:
5424:
5422:
5417:
5415:
5413:
5410:Hausa genet (
5408:
5406:
5404:
5399:
5397:
5395:
5390:
5388:
5386:
5381:
5379:
5377:
5372:
5370:
5368:
5363:
5361:
5359:
5354:
5352:
5350:
5345:
5343:
5341:
5336:
5334:
5332:
5327:
5325:
5323:
5318:
5316:
5314:
5313:G. angolensis
5309:
5307:
5305:
5304:G. abyssinica
5300:
5299:
5297:
5289:
5288:
5283:
5277:
5275:
5270:
5268:
5266:
5261:
5260:
5258:
5250:
5249:
5244:
5241:
5239:
5235:
5225:
5223:
5218:
5216:
5214:
5213:V. tangalunga
5209:
5207:
5205:
5200:
5198:
5196:
5191:
5190:
5188:
5186:
5185:
5180:
5174:
5172:
5167:
5165:
5163:
5158:
5157:
5154:
5151:
5148:
5147:sensu stricto
5143:
5139:
5136:
5132:
5127:
5121:
5117:
5107:
5105:
5100:
5098:
5096:
5091:
5089:
5087:
5082:
5081:
5079:
5077:
5076:
5071:
5065:
5063:
5058:
5056:
5054:
5049:
5047:
5045:
5040:
5038:
5036:
5031:
5030:
5027:
5024:
5022:
5021:Paradoxurinae
5018:
5012:
5010:
5005:
5003:
5001:
4996:
4994:
4992:
4989:Otter civet (
4987:
4985:
4983:
4978:
4977:
4975:
4973:
4969:
4966:
4963:
4958:
4954:
4950:
4945:
4941:
4920:
4919:
4918:
4917:
4915:
4913:
4909:
4883:
4881:
4880:F. silvestris
4876:
4874:
4872:
4867:
4865:
4863:
4858:
4856:
4854:
4849:
4847:
4845:
4840:
4838:
4836:
4831:
4829:
4827:
4822:
4821:
4819:
4817:
4816:
4811:
4801:
4799:
4798:P. viverrinus
4796:Fishing cat (
4794:
4792:
4790:
4785:
4783:
4781:
4776:
4774:
4772:
4771:P. javanensis
4767:
4765:
4763:
4760:Leopard cat (
4758:
4757:
4755:
4753:
4752:
4747:
4741:
4739:
4734:
4733:
4730:
4727:
4721:
4711:
4709:
4704:
4703:
4701:
4699:
4698:
4693:
4687:
4685:
4680:
4679:
4676:
4670:
4668:
4663:
4662:
4660:
4658:
4657:
4652:
4649:
4643:
4637:
4635:
4630:
4628:
4626:
4621:
4619:
4617:
4612:
4610:
4608:
4607:L. canadensis
4605:Canada lynx (
4603:
4602:
4600:
4598:
4597:
4592:
4586:
4584:
4579:
4577:
4575:
4570:
4568:
4566:
4561:
4559:
4557:
4552:
4550:
4548:
4543:
4541:
4539:
4534:
4532:
4530:
4525:
4523:
4521:
4516:
4515:
4513:
4511:
4510:
4505:
4495:
4493:
4488:
4486:
4484:
4479:
4478:
4476:
4474:
4473:
4468:
4462:
4460:
4455:
4454:
4451:
4448:
4442:
4432:
4430:
4429:C. temminckii
4425:
4423:
4421:
4416:
4415:
4413:
4411:
4410:
4405:
4399:
4397:
4394:Marbled cat (
4392:
4391:
4389:
4387:
4386:
4381:
4378:
4372:
4369:
4365:
4364:sensu stricto
4361:
4356:
4342:
4340:
4335:
4333:
4331:
4326:
4324:
4322:
4317:
4315:
4313:
4308:
4306:
4304:
4299:
4298:
4296:
4294:
4293:
4288:
4282:
4280:
4275:
4273:
4271:
4266:
4265:
4263:
4261:
4260:
4255:
4252:
4250:
4246:
4242:
4239:
4231:
4226:
4222:
4216:
4214:
4213:P. pardicolor
4209:
4207:
4205:
4200:
4199:
4197:
4190:
4189:
4184:
4181:
4178:
4173:
4169:
4163:
4161:
4156:
4155:
4152:
4149:
4146:
4141:
4137:
4133:
4125:
4121:
4115:
4111:
4109:
4105:
4103:
4099:
4097:
4093:
4091:
4087:
4086:
4083:
4078:
4070:
4065:
4063:
4058:
4056:
4051:
4050:
4047:
4040:
4036:
4035:
4030:
4028:
4024:
4020:
4017:
4014:
4012:
4008:
4004:
4001:
3998:
3994:
3990:
3987:
3985:
3982:
3981:
3977:
3972:
3968:
3964:
3962:, March 2006.
3961:
3957:
3953:
3952:
3948:
3931:
3927:
3921:
3914:
3913:
3908:
3904:
3900:
3896:
3895:
3890:
3886:
3882:
3881:
3875:
3871:
3867:
3866:
3861:
3857:
3847:on 2014-09-08
3843:
3836:
3835:
3829:
3828:
3824:
3807:
3800:
3797:
3793:
3789:
3785:
3782:
3776:
3773:
3761:
3754:
3747:
3744:
3740:
3735:
3732:
3728:
3723:
3720:
3716:
3711:
3708:
3704:
3699:
3696:
3692:
3687:
3684:
3680:
3675:
3672:
3668:
3663:
3660:
3656:
3651:
3648:
3643:
3642:
3637:
3631:
3629:
3625:
3621:
3616:
3614:
3610:
3604:
3599:
3595:
3591:
3587:
3580:
3577:
3573:
3567:
3564:
3561:, p. 207
3560:
3555:
3552:
3548:
3543:
3540:
3535:
3531:
3527:
3523:
3519:
3515:
3511:
3507:
3500:
3497:
3491:
3486:
3483:(3): 579–89.
3482:
3478:
3474:
3467:
3464:
3459:
3455:
3451:
3447:
3443:
3439:
3435:
3431:
3424:
3417:
3414:
3410:
3405:
3403:
3399:
3394:
3390:
3386:
3382:
3375:
3368:
3365:
3361:
3356:
3354:
3350:
3346:
3341:
3339:
3337:
3333:
3328:
3324:
3319:
3314:
3310:
3306:
3302:
3298:
3294:
3290:
3286:
3278:
3275:
3271:
3265:
3262:
3257:
3253:
3248:
3243:
3239:
3235:
3231:
3227:
3223:
3215:
3212:
3206:
3201:
3197:
3193:
3189:
3182:
3179:
3176:, p. 459
3175:
3170:
3167:
3163:
3158:
3155:
3149:
3146:
3141:
3137:
3133:
3129:
3125:
3121:
3117:
3113:
3110:(4): 301–12.
3109:
3105:
3097:
3094:
3089:
3085:
3080:
3075:
3071:
3067:
3063:
3059:
3055:
3051:
3047:
3039:
3036:
3032:
3031:
3024:
3021:
3017:
3016:
3011:
3006:
3003:
2999:
2994:
2992:
2988:
2985:, p. 462
2984:
2983:Lydekker 1908
2979:
2977:
2973:
2969:
2964:
2961:
2956:
2950:
2947:
2943:
2938:
2936:
2932:
2928:
2923:
2920:
2908:
2902:
2898:
2891:
2884:
2882:
2878:
2866:
2862:
2856:
2853:
2849:
2844:
2842:
2840:
2836:
2828:
2824:
2820:
2816:
2812:
2808:
2804:
2800:
2796:
2792:
2788:
2781:
2779:
2770:
2768:
2764:
2753:on 2015-09-24
2749:
2745:
2741:
2737:
2733:
2729:
2725:
2721:
2717:
2710:
2703:
2701:
2699:
2697:
2695:
2693:
2691:
2687:
2682:
2676:
2672:
2668:
2661:
2659:
2657:
2653:
2649:
2645:
2641:
2636:
2633:
2620:
2615:
2611:
2607:
2606:
2601:
2599:
2590:
2588:
2586:
2584:
2582:
2580:
2578:
2576:
2572:
2567:
2563:
2558:
2553:
2549:
2545:
2541:
2537:
2533:
2529:
2525:
2517:
2515:
2513:
2511:
2507:
2500:
2493:
2487:
2484:
2480:
2476:
2472:
2466:
2463:
2459:
2453:
2450:
2443:
2439:
2436:
2435:
2431:
2429:
2426:
2422:
2420:
2416:
2412:
2407:
2402:
2398:
2396:
2392:
2388:
2384:
2379:
2377:
2369:
2367:
2365:
2361:
2353:
2351:
2348:
2339:
2337:
2335:
2331:
2327:
2322:
2318:
2310:
2308:
2306:
2298:
2293:
2286:
2284:
2282:
2277:
2268:
2266:
2263:
2259:
2258:fragmentation
2251:
2249:
2247:
2242:
2233:
2231:
2229:
2225:
2216:
2214:
2212:
2208:
2199:
2192:
2190:
2186:
2183:
2180:
2177:
2174:
2171:
2168:
2167:
2163:
2160:
2157:
2154:
2151:
2148:
2146:
2142:
2139:
2138:
2134:
2131:
2127:
2124:
2121:
2118:
2114:
2111:
2109:
2105:
2102:
2101:
2097:
2092:
2089:
2086:
2083:
2080:
2077:
2074:
2071:
2070:
2066:
2062:
2058:
2055:
2053:
2050:
2047:
2044:
2041:
2037:
2036:
2032:
2028:
2024:
2021:
2017:
2015:
2012:
2009:
2006:
2003:
2000:
1999:
1995:
1991:
1988:
1987:
1982:
1979:
1976:
1973:
1970:
1968:
1964:
1961:
1960:
1957:Conservation
1956:
1953:
1950:
1947:
1944:
1941:
1938:
1937:
1934:
1928:
1926:
1922:
1918:
1914:
1910:
1906:
1902:
1898:
1894:
1890:
1882:
1880:
1878:
1874:
1870:
1866:
1862:
1859:. Undigested
1858:
1854:
1850:
1846:
1842:
1838:
1834:
1830:
1826:
1822:
1818:
1814:
1806:
1802:
1798:
1791:
1789:
1787:
1783:
1779:
1778:
1773:
1772:
1767:
1766:
1761:
1757:
1753:
1750:
1749:
1744:
1740:
1734:
1732:
1727:
1723:
1719:
1715:
1712:
1705:
1700:
1693:
1688:
1686:
1684:
1679:
1674:
1672:
1666:
1662:
1660:
1650:
1643:
1641:
1639:
1635:
1631:
1625:
1623:
1619:
1614:
1610:
1606:
1605:mating season
1598:
1596:
1592:
1589:
1581:
1577:
1572:
1565:
1563:
1560:
1557:
1553:
1548:
1543:
1541:
1540:
1535:
1531:
1527:
1523:
1518:
1514:
1510:
1503:
1498:
1491:
1485:
1483:
1479:
1475:
1472:
1468:
1465:
1462:
1457:
1453:
1450:
1449:
1442:
1434:
1425:
1416:
1413:
1410:
1406:
1402:
1398:
1394:
1390:
1388:
1385:
1382:
1379:
1373:
1368:
1364:
1361:
1360:
1356:
1353:
1350:
1347:
1344:
1340:
1338:
1336:
1327:
1325:
1323:
1319:
1314:
1310:
1309:golden jackal
1306:
1302:
1298:
1290:
1286:
1284:
1282:
1278:
1277:RAD sequences
1274:
1268:
1266:
1262:
1258:
1254:
1253:Oscar Neumann
1250:
1246:
1242:
1238:
1234:
1230:
1229:
1223:
1221:
1217:
1208:
1204:
1200:
1195:
1189:
1188:
1185:See further:
1183:
1182:
1181:
1179:
1175:
1170:
1167:
1163:
1159:
1145:
1144:
1136:
1135:
1127:
1126:
1118:
1117:
1114:
1112:
1108:
1102:
1101:
1098:
1097:
1094:
1092:
1088:
1082:
1081:
1074:
1073:
1067:
1066:
1063:
1062:
1054:
1053:
1050:
1048:
1044:
1038:
1037:
1034:
1033:
1025:
1024:
1021:
1019:
1015:
1009:
1008:
1005:
1004:
996:
995:
992:
990:
982:
981:
978:
977:
969:
968:
965:
963:
959:
958:Golden jackal
953:
952:
949:
948:
940:
939:
936:
934:
930:
924:
923:
920:
919:
911:
910:
907:
905:
901:
895:
894:
891:
890:
882:
881:
873:
872:
869:
867:
863:
857:
856:
853:
852:
849:
846:
841:
835:
834:
828:
827:
821:
820:
813:
812:
805:
804:
797:
796:
789:
788:
781:
780:
773:
772:
766:
762:
759:
758:
754:
753:
749:
745:
742:
741:
735:
733:
731:
727:
723:
718:
714:
710:
706:
701:
699:
695:
694:domestic dogs
691:
687:
683:
679:
675:
669:
667:
663:
659:
656:
652:
648:
644:
635:
628:
624:
620:
613:
604:
602:
598:
595:
593:
590:
589:
583:
580:
574:
570:
568:
565:
564:
560:
557:
554:
553:
550:
540:
536:
532:
530:
529:Ethiopic text
512:
510:
508:
504:
500:
499:Simien jackal
496:
492:
484:
482:
480:
476:
472:
468:
464:
460:
459:interbreeding
456:
452:
448:
444:
440:
436:
431:
429:
425:
421:
416:
414:
410:
407:
403:
399:
395:
391:
387:
383:
379:
378:Simien jackal
375:
371:
370:
365:
351:
348:
345:Heuglin, 1862
343:
340:
335:
332:
329:Heuglin, 1862
327:
324:
319:
316:
315:
313:
310:
306:
301:
297:
292:
287:
282:
280:
274:
271:
270:Binomial name
267:
263:
262:
257:
254:
253:
250:
249:
245:
242:
241:
238:
235:
232:
231:
228:
225:
222:
221:
218:
215:
212:
211:
208:
205:
202:
201:
198:
195:
192:
191:
188:
185:
182:
181:
176:
171:
167:
161:
156:
145:
141:
136:
133:
128:
124:
119:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
44:
33:
30:
19:
9562:
9519:
9510:
9501:
9492:
9483:
9474:
9465:
9456:
9447:
9438:
9429:
9420:
9411:
9402:
9393:
9384:
9375:
9355:
9343:
9334:
9325:
9316:
9304:
9279:
9270:
9261:
9249:
9237:
9225:
9214:L. sumatrana
9213:
9204:
9192:
9180:
9163:
9151:
9139:
9130:
9121:
9112:
9100:
9088:
9046:
9037:
9025:
9013:
8997:V. peregusna
8996:
8984:
8951:
8942:
8922:
8910:
8897:Lyncodontini
8888:Ictonychinae
8872:M. zibellina
8871:
8862:
8853:
8844:
8835:
8827:M. flavigula
8826:
8817:
8809:M. americana
8808:
8788:
8776:
8764:
8752:
8740:
8729:Eira barbara
8728:
8702:
8694:M. personata
8693:
8684:
8675:
8666:
8657:
8637:
8621:
8612:
8604:M. canescens
8603:
8594:
8582:
8570:
8561:
8552:
8540:
8511:
8499:
8487:
8406:
8397:
8385:
8373:
8364:
8352:
8323:
8314:
8305:
8296:
8276:
8264:
8255:
8246:
8226:
8214:
8213:Cacomistle (
8205:
8193:
8182:Potos flavus
8181:
8151:
8142:
8133:
8124:
8104:
8092:
8083:
8063:
8051:
8042:
8030:
8018:
8009:
8000:
7991:
7971:
7938:
7879:
7870:
7861:
7852:
7818:
7809:
7789:
7773:
7761:
7749:
7734:(monk seals)
7690:
7681:
7672:
7660:
7648:
7639:
7627:
7615:
7606:
7597:
7571:
7562:
7505:
7497:A. townsendi
7496:
7487:
7479:A. philippii
7478:
7469:
7460:
7451:
7443:A. australis
7442:
7430:
7418:
7401:
7389:
7377:
7356:
7347:
7335:
7323:
7297:
7285:
7245:
7124:U. maritimus
7123:
7122:Polar bear (
7114:
7113:Brown bear (
7105:
7096:
7084:
7072:
7063:
7062:Sloth bear (
7037:
7025:
7013:
7001:
6951:
6949:
6942:
6933:
6924:
6915:
6906:
6897:
6888:
6876:
6865:Cuon alpinus
6864:
6855:
6839:L. mesomelas
6838:
6829:
6817:
6786:
6777:
6768:
6767:Pampas fox (
6759:
6750:
6741:
6729:
6717:
6708:
6692:S. venaticus
6691:
6679:
6667:
6666:Maned wolf (
6648:Cerdocyonina
6616:
6615:Fennec fox (
6607:
6598:
6590:V. rueppelli
6589:
6580:
6571:
6562:
6561:Arctic fox (
6554:V. ferrilata
6553:
6544:
6543:Corsac fox (
6535:
6526:
6517:
6516:Bengal fox (
6495:
6483:
6474:
6462:
6450:
6424:
6423:Island fox (
6415:
6403:
6336:H. sanguinea
6335:
6326:
6317:
6309:H. ichneumon
6308:
6299:
6287:
6275:
6266:
6257:
6245:
6233:
6224:
6215:
6206:
6189:
6180:
6171:
6162:
6153:
6144:
6136:U. edwardsii
6135:
6127:U. brachyura
6126:
6117:
6105:
6093:
6084:
6054:
6045:
6036:
6028:C. alexandri
6027:
6015:
6003:
5994:
5982:
5970:
5953:
5945:M. gambianus
5944:
5932:
5920:
5904:S. suricatta
5903:
5891:
5866:
5841:
5832:
5820:
5808:
5791:
5782:
5770:
5758:
5728:
5719:
5707:
5695:
5686:
5663:carnivorans)
5644:
5615:
5603:
5591:
5582:
5550:
5538:
5508:Herpestoidea
5471:Herpestoidea
5438:
5430:G. victoriae
5429:
5420:
5419:Cape genet (
5411:
5403:G. servalina
5402:
5393:
5392:King genet (
5385:G. piscivora
5384:
5375:
5366:
5357:
5349:G. johnstoni
5348:
5339:
5330:
5321:
5312:
5303:
5285:
5274:P. leightoni
5273:
5264:
5246:
5221:
5212:
5204:V. megaspila
5203:
5195:V. civettina
5194:
5182:
5170:
5161:
5146:
5130:
5103:
5094:
5085:
5073:
5061:
5052:
5043:
5034:
5008:
4999:
4990:
4981:
4879:
4870:
4862:F. margarita
4861:
4852:
4843:
4842:Jungle cat (
4834:
4825:
4813:
4797:
4788:
4780:P. planiceps
4779:
4770:
4761:
4751:Prionailurus
4749:
4737:
4707:
4695:
4683:
4682:Jaguarundi (
4666:
4654:
4633:
4624:
4615:
4606:
4594:
4582:
4573:
4564:
4555:
4546:
4537:
4529:L. geoffroyi
4528:
4519:
4518:Pampas cat (
4507:
4491:
4482:
4470:
4458:
4428:
4419:
4407:
4396:P. marmorata
4395:
4383:
4363:
4338:
4329:
4320:
4311:
4302:
4290:
4278:
4269:
4257:
4212:
4203:
4186:
4159:
4112:Superorder:
4106:Infraclass:
4033:
3973:, July 2004.
3970:
3959:
3937:. Retrieved
3930:the original
3911:
3893:
3878:
3864:
3849:. Retrieved
3842:the original
3833:
3810:. Retrieved
3799:
3791:
3775:
3763:. Retrieved
3759:
3746:
3741:, p. 30
3734:
3729:, p. 29
3722:
3710:
3698:
3686:
3681:, p. 22
3674:
3662:
3657:, p. 10
3650:
3639:
3593:
3589:
3579:
3566:
3554:
3542:
3509:
3505:
3499:
3480:
3476:
3466:
3433:
3429:
3416:
3384:
3380:
3367:
3347:, p. 32
3292:
3288:
3277:
3269:
3264:
3229:
3225:
3214:
3195:
3191:
3181:
3169:
3157:
3148:
3107:
3103:
3096:
3053:
3049:
3038:
3028:
3023:
3013:
3005:
2970:, p. 39
2968:Rüppell 1835
2963:
2949:
2922:
2910:. Retrieved
2896:
2868:. Retrieved
2864:
2855:
2827:the original
2790:
2786:
2777:
2755:. Retrieved
2748:the original
2722:(485): 1–6.
2719:
2715:
2670:
2640:Rüppell 1835
2635:
2623:. Retrieved
2609:
2603:
2597:
2531:
2527:
2486:
2465:
2452:
2427:
2423:
2403:
2399:
2380:
2373:
2370:Conservation
2363:
2357:
2343:
2314:
2305:soil erosion
2301:
2272:
2255:
2237:
2234:Habitat loss
2224:vaccinations
2220:
2203:
2149:870 km
2129:
2020:habitat loss
1984:
1932:
1886:
1841:rock hyraxes
1810:
1804:
1780:bushes, and
1775:
1769:
1763:
1746:
1735:
1708:
1675:
1668:
1664:
1655:
1626:
1602:
1593:
1585:
1561:
1544:
1537:
1526:canine teeth
1517:African wolf
1506:
1455:
1451:
1440:
1437:(Gray, 1869)
1432:
1423:
1414:
1366:
1362:
1351:Description
1337:Volume Three
1333:
1331:
1300:
1294:
1288:
1273:Canis aureus
1272:
1269:
1260:
1248:
1226:
1224:
1220:interglacial
1212:
1184:
1177:
1171:
1155:
1105:
1085:
1041:
1012:
985:
956:
929:African wolf
927:
898:
860:
840:Domestic dog
838:
702:
670:
658:hypothesised
640:
626:
622:
600:
596:
578:
572:
548:
526:
506:
502:
498:
495:Simenian fox
494:
490:
488:
432:
417:
394:horse jackal
393:
381:
377:
373:
368:
367:
363:
361:
349:
341:
333:
326:C. semiensis
325:
318:C. crinensis
317:
278:
276:
260:
259:
247:
29:
9677:iNaturalist
9587:Wikispecies
9511:M. sibirica
9493:M. putorius
9466:M. nigripes
9448:M. lutreola
9421:M. haidarum
9317:N. africana
9280:A. congicus
9271:A. cinereus
9262:A. capensis
9162:Sea otter (
9140:L. provocax
9047:I. striatus
8968:Ictonychini
8863:M. melampus
8775:Wolverine (
8753:P. pennanti
8676:M. moschata
8667:M. everetti
8613:M. leucurus
8571:A. hoevenii
8562:A. collaris
8512:M. capensis
8407:N. olivacea
8322:Olinguito (
8278:Bassaricyon
8265:P. pygmaeus
8195:Bassariscus
8168:Procyonidae
8143:S. putorius
8134:S. gracilis
8052:M. mephitis
8043:M. macroura
7937:Red panda (
7917:Musteloidea
7878:Ross seal (
7831:Lobodontini
7763:Neomonachus
7691:P. sibirica
7649:P. vitulina
7614:Grey seal (
7596:Harp seal (
7488:A. pusillus
7452:A. forsteri
7287:Callorhinus
7221:Pinnipedia
7180:Musteloidea
6952:C. simensis
6916:C. lupaster
6785:Hoary fox (
6778:L. sechurae
6751:L. fulvipes
6742:L. culpaeus
6638:(true dogs)
6597:Swift fox (
6572:V. macrotis
6464:Nyctereutes
6373:Caniformia
6318:H. ochracea
6276:B. nigripes
6267:B. jacksoni
6154:U. javanica
6071:Herpestinae
6046:C. obscurus
6037:C. ansorgei
6017:Crossarchus
5873:(mongooses)
5862:Herpestidae
5842:S. durrelli
5833:S. concolor
5783:G. fasciata
5720:E. goudotii
5641:Herpestidae
5551:P. cristata
5412:G. thierryi
5367:G. maculata
5331:G. cristata
5322:G. bourloni
5075:Paradoxurus
5033:Binturong (
4972:Hemigalinae
4949:Viverroidea
4912:Viverroidea
4871:F. nigripes
4723:Leopard cat
4708:P. concolor
4625:L. pardinus
4574:L. tigrinus
4565:L. pardalis
4556:L. jacobita
4547:L. guttulus
4520:L. colocola
4279:N. nebulosa
4249:Pantherinae
4129:Feliformia
3939:15 December
3825:Cited works
3812:23 February
3362:, p. 4
3198:: 119–199.
2944:, p. 2
2625:19 November
2334:saddle pads
2287:Overgrazing
2281:brush fires
2211:overgrazing
2130:Helichrysum
2104:Guassa Menz
2088:Overgrazing
1986:Helichrysum
1951:Importance
1917:Mount Choqa
1905:Guassa Menz
1889:Rift Valley
1765:Helichrysum
1530:carnassials
1492:Description
1471:nasal bones
1441:C. s. walgi
1405:Guassa Menz
1393:Rift Valley
1345:Subspecies
1178:C. simensis
1172:In 1994, a
726:Mount Choqa
475:vaccination
451:overgrazing
350:C. citernii
334:C. simensis
41:1.6–0
9855:Categories
9484:M. nudipes
9475:M. nivalis
9439:M. kathiah
9394:M. erminea
9385:M. altaica
9335:N. frenata
9326:N. felipei
9296:Mustelinae
9038:I. libycus
8952:G. vittata
8818:M. caurina
8595:M. anakuma
8473:Mustelidae
8460:Mustelidae
8436:see below↓
8427:Mustelidae
8324:B. neblina
8235:(raccoons)
8206:B. astutus
8204:Ringtail (
8180:Kinkajou (
8152:S. pygmaea
8093:M. marchei
7951:Mephitidae
7819:M. leonina
7711:Monachinae
7682:P. hispida
7673:P. caspica
7470:A. gazella
7402:N. cinerea
7298:C. ursinus
7236:Odobenidae
7189:see below↓
7169:see below↓
7153:Pinnipedia
7071:Sun bear (
7038:T. ornatus
7027:Tremarctos
7003:Ailuropoda
6941:Red wolf (
6907:C. latrans
6830:L. adustus
6787:L. vetulus
6760:L. griseus
6690:Bush dog (
6581:V. pallida
6579:Pale fox (
6563:V. lagopus
6534:Cape fox (
6414:Gray fox (
6172:U. smithii
6004:H. parvula
5995:H. hirtula
5883:Mungotinae
5772:Galidictis
5745:Galidiinae
5673:Euplerinae
5656:Eupleridae
5645:sensu lato
5616:C. crocuta
5549:Aardwolf (
5480:see below↓
5421:G. tigrina
5394:G. poensis
5376:G. pardina
5358:G. letabae
5340:G. genetta
5238:Genettinae
5222:V. zibetha
5142:Viverrinae
5131:sensu lato
5126:Viverrinae
5095:P. jerdoni
4962:Viverridae
4921:see below↓
4860:Sand cat (
4667:A. jubatus
4492:C. caracal
4385:Pardofelis
4204:P. linsang
4145:Feliformia
3851:2012-12-15
3760:Canids.org
2757:2011-09-22
2534:(1): 530.
2501:References
2458:afterbirth
2417:, and the
2042:Highlands
1913:Mount Guna
1817:grass rats
1786:ericaceous
1774:tussocks,
1748:Alchemilla
1718:heathlands
1714:grasslands
1711:Afroalpine
1638:homozygous
1630:inbreeding
1411:highlands
1401:Mount Guna
1391:Northwest
1328:Subspecies
1207:maned wolf
662:greyhounds
586:Trickster
584:Red jackal
491:red jackal
481:programs.
435:endangered
406:Afroalpine
402:generalist
382:Simien fox
374:red jackal
337:Gray, 1869
155:Endangered
18:Red Jackal
9430:M. itatsi
9364:(weasels)
9227:Lutrogale
9164:E. lutris
9122:L. felina
8931:(grisons)
8854:M. martes
8797:(martens)
8715:Guloninae
8501:Mellivora
8374:N. narica
8315:B. medius
8306:B. gabbii
8297:B. alleni
8285:(olingos)
8254:Raccoon (
8106:Spilogale
7992:C. chinga
7973:Conepatus
7929:Ailuridae
7728:Monachini
7640:P. largha
7310:Otariinae
7273:sea lions
7268:fur seals
7259:Otariidae
7144:Mustelida
7115:U. arctos
6977:Arctoidea
6934:C. lycaon
6889:C. aureus
6819:Lupulella
6731:Lycalopex
6608:V. vulpes
6606:Red fox (
6570:Kit fox (
6545:V. corsac
6289:Herpestes
5902:Meerkat (
5661:(Malagasy
5563:Hyaeninae
5524:Hyaenidae
5439:G. felina
4853:F. lybica
4665:Cheetah (
4583:L. wiedii
4572:Oncilla (
4538:L. guigna
4509:Leopardus
4490:Caracal (
4483:C. aurata
4418:Bay cat (
4339:P. pardus
4337:Leopard (
4312:P. tigris
4270:N. diardi
4188:Prionodon
4088:Kingdom:
4077:Carnivora
4005:from the
3436:: 39–45.
2744:253988540
2548:2399-3642
2479:grey wolf
1907:in north
1877:livestock
1829:vlei rats
1741:and deep
1726:tree line
1609:Courtship
1566:Behaviour
1522:premolars
1357:Synonyms
1295:In 2018,
1199:grey wolf
862:Gray wolf
730:carnivore
705:Highlands
651:Abyssinia
573:Ky kebero
413:carnivore
342:C. walgié
255:Species:
227:Carnivora
193:Kingdom:
187:Eukaryota
9820:Species+
9760:14000779
9695:10762843
9572:Wikidata
9410:Ferret (
9344:N. vison
9205:L. lutra
9073:(otters)
9067:Lutrinae
8924:Galictis
8836:M. foina
8751:Fisher (
8639:Melogale
8622:M. meles
8542:Arctonyx
8387:Nasuella
8365:N. nasua
8342:(coatis)
8256:P. lotor
8032:Mephitis
7791:Mirounga
7541:Phocinae
7522:Phocidae
7391:Neophoca
7337:Zalophus
7244:Walrus (
6943:C. rufus
6925:C. lupus
6905:Coyote (
6740:Culpeo (
6681:Speothos
6617:V. zerda
6599:V. velox
6536:V. chama
6247:Bdeogale
6145:U. fusca
5984:Helogale
5954:M. mungo
5893:Suricata
5822:Salanoia
5729:E. major
5709:Eupleres
5540:Proteles
5529:(hyenas)
5293:(genets)
4844:F. chaus
4835:F. catus
4826:F. bieti
4706:Cougar (
4656:Acinonyx
4634:L. rufus
4632:Bobcat (
4581:Margay (
4563:Ocelot (
4536:Kodkod (
4457:Serval (
4420:C. badia
4409:Catopuma
4319:Jaguar (
4303:P. uncia
4292:Panthera
4259:Neofelis
4177:Feloidea
4108:Eutheria
4102:Mammalia
4096:Chordata
4094:Phylum:
4090:Animalia
4019:Archived
3909:(1997).
3891:(1835).
3876:(1902).
3862:(1908).
3784:Archived
3765:19 March
3526:19834483
3458:46671897
3450:23994497
3327:30344120
3256:26234211
3140:25976335
3088:20237475
3012:(1853).
2870:19 March
2823:40027094
2815:15245401
2566:37193884
2557:10187515
2432:See also
2321:firearms
2106:, North
2081:Stable?
2013:Extinct
2002:Mt. Guna
1977:Stable?
1965:, North
1954:Threats
1942:Habitat
1865:roughage
1849:reedbuck
1837:goslings
1762:rich in
1739:drainage
1339:(MSW3).
1162:mandible
607:Reddish
571:ቀይ ቀበሮ (
449:through
309:Synonyms
233:Family:
217:Mammalia
207:Chordata
203:Phylum:
197:Animalia
183:Domain:
160:IUCN 3.1
9833:ZooBank
9747:1005946
9669:5219143
9578:Q188957
9412:M. furo
9357:Mustela
9306:Neogale
9153:Enhydra
9027:Ictonyx
8986:Vormela
8943:G. cuja
8870:Sable (
8777:G. gulo
8742:Pekania
8727:Tayra (
8524:Melinae
8336:Nasuina
8228:Procyon
7584:Phocini
7223:(seals)
7158:(seals)
7050:Ursinae
6992:(bears)
6987:Ursidae
6863:Dhole (
6653:(zorro)
6527:V. cana
6437:Vulpini
6405:Urocyon
6389:Canidae
6181:U. urva
5685:Fossa (
5605:Crocuta
5287:Genetta
5184:Viverra
4725:lineage
4647:lineage
4616:L. lynx
4472:Caracal
4446:lineage
4444:Caracal
4376:lineage
4374:Bay cat
4360:Felinae
4321:P. onca
4310:Tiger (
4230:Felidae
4100:Class:
4079:species
4075:Extant
4025:at the
4009:of the
3995:of the
3993:WildCRU
3880:natives
3318:6224481
3297:Bibcode
3234:Bibcode
3132:7921357
3112:Bibcode
3079:3494089
3058:Bibcode
2865:Awf.org
2795:Bibcode
2736:3504136
2319:, when
2246:Delanta
2217:Disease
2200:Threats
2172:, Bale
2096:Denkoro
1948:Status
1777:Euryops
1771:Festuca
1756:tussock
1729:times,
1722:habitat
1694:Habitat
1689:Ecology
1683:geladas
1534:dewclaw
1386:, 1840
1384:Rüppell
1318:admixed
1291:species
1249:Simenia
1233:jackals
761:Caninae
746:of the
601:Arouayé
567:Amharic
437:by the
409:rodents
384:, is a
286:Ruppell
243:Genus:
237:Canidae
223:Order:
213:Class:
158: (
110:↓
9871:Wolves
9812:433177
9799:752755
9721:621851
9656:327993
9640:ECOS:
9615:ARKive
9102:Lontra
8790:Martes
8065:Mydaus
7959:skunks
6923:Wolf (
6803:Canina
6633:Canini
6497:Vulpes
6393:(dogs)
5934:Mungos
5248:Poiana
4330:P. leo
4328:Lion (
4234:(cats)
4039:ARKive
3922:
3534:771357
3532:
3524:
3456:
3448:
3325:
3315:
3254:
3138:
3130:
3086:
3076:
3050:Nature
2912:4 July
2903:
2821:
2813:
2742:
2734:
2677:
2648:pl. 14
2564:
2554:
2546:
2276:cereal
2241:barley
2098:area.
2038:North
2031:kebele
2027:woreda
1967:Gondar
1897:Gondar
1843:, and
1823:, and
1760:scrubs
1678:cattle
1556:moults
1554:. It
1354:Range
1251:, and
900:Coyote
713:Gojjam
682:rabies
579:Walgie
505:, and
497:, the
493:, the
485:Naming
453:, and
398:coyote
386:canine
376:, the
288:, 1840
9866:Canis
9825:10871
9786:82242
9773:32534
9708:90294
9690:IRMNG
9682:42045
9633:5WXTW
9251:Aonyx
9194:Lutra
8584:Meles
8354:Nasua
7629:Phoca
7086:Ursus
6878:Canis
6505:foxes
4815:Felis
3933:(PDF)
3916:(PDF)
3845:(PDF)
3838:(PDF)
3756:(PDF)
3530:S2CID
3454:S2CID
3426:(PDF)
3377:(PDF)
3136:S2CID
2893:(PDF)
2830:(PDF)
2819:S2CID
2783:(PDF)
2751:(PDF)
2740:S2CID
2732:JSTOR
2712:(PDF)
2644:p. 39
2444:Notes
2376:CITES
2330:goats
2326:sheep
2040:Wollo
1939:Area
1861:sedge
1752:herbs
1743:soils
1588:packs
1547:ochre
1539:Canis
1409:Wollo
1313:basal
1301:Canis
1289:Canis
1261:Canis
1245:genus
1228:Canis
1014:Dhole
814:0.38
798:0.96
592:Oromo
577:ዋልጌ (
461:from
420:range
248:Canis
9768:NCBI
9734:3748
9729:IUCN
9716:ITIS
9664:GBIF
9643:1640
8766:Gulo
7662:Pusa
6107:Urva
4697:Puma
4645:Puma
4596:Lynx
3941:2012
3920:ISBN
3814:2016
3767:2022
3522:PMID
3446:PMID
3323:PMID
3252:PMID
3128:PMID
3084:PMID
2914:2019
2901:ISBN
2872:2022
2811:PMID
2675:ISBN
2627:2021
2610:2011
2562:PMID
2544:ISSN
2328:and
2297:zebu
2145:Bale
2108:Shoa
2059:The
2029:and
1923:and
1921:Arsi
1909:Shoa
1871:and
1792:Diet
1716:and
1620:and
1603:The
1511:and
1480:and
1478:Arsi
1322:wolf
1239:and
1075:2.6
806:0.6
790:2.0
782:2.5
774:3.0
477:and
457:and
439:IUCN
362:The
49:PreꞒ
9755:MSW
9742:MDD
9703:ISC
9651:EoL
9628:CoL
9602:ADW
4037:at
3991:at
3598:doi
3514:doi
3485:doi
3438:doi
3434:122
3389:doi
3385:245
3313:PMC
3305:doi
3242:doi
3200:doi
3120:doi
3074:PMC
3066:doi
3054:464
2803:doi
2724:doi
2720:385
2614:doi
2552:PMC
2536:doi
2475:dog
2471:SNP
1895:in
724:in
501:,
380:or
9857::
9835::
9822::
9809::
9796::
9783::
9770::
9757::
9744::
9731::
9718::
9705::
9692::
9679::
9666::
9653::
9630::
9617::
9604::
9589::
9574::
3969:.
3958:.
3905:;
3790:,
3758:.
3638:.
3627:^
3612:^
3594:96
3592:.
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2990:^
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2863:.
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2817:.
2809:.
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2789:.
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2780:)"
2766:^
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2646:,
2608:.
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2560:.
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2421:.
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765:Ma
732:.
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509:.
430:.
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