1321:
1085:
36:
406:
188:
824:
1406:
763:
629:
2386:
518:) fight to dominate sap sites. Males use their horns to pry rival males off the area, which also may give them the chance to mate with a female. In this and other species that defend mating sites, larger males with larger horns mate more frequently, as they win more contests. Small males often avoid larger males and exhibit alternative strategies to gain access to females.
175:
449:
grew up to 7 cm (3 in). Common names of the
Dynastinae refer to the characteristic horns borne only by the males of most species in the group. Each has a horn on the head and another horn pointing forward from the center of the thorax. The horns are used in fighting other males during
416:
1320:
435:
The
Dynastinae are among the largest of beetles, reaching more than 15 centimetres (6 inches) in length, but are completely harmless to humans because they cannot bite or sting. Some species have been anecdotally claimed to lift up to 850 times their own weight. An extinct
1548:
567:. Since males naturally have the tendency to fight each other for the attention of females, they are the ones used for battle. To get the two male beetles to lock in combat, a female beetle is used, or a small noisemaker duplicating the female's
469:
themselves from the few predators big enough to eat them. If rhinoceros beetles are disturbed, some can release very loud, hissing squeaks created by rubbing their abdomens against the ends of their wing covers.
540:
are not as high as in some other pest insects, and they typically prefer food trees which are already sick or dying from some other cause. Some species' larvae, however, will attack healthy trees or even
1630:
Goczał, Jakub; Rossa, Robert; Tofilski, Adam (17 April 2019). "Intersexual and intrasexual patterns of horn size and shape variation in the
European rhinoceros beetle: quantifying the shape of weapons".
457:. A pair of thick wings lie atop another set of membranous wings underneath, allowing the rhinoceros beetle to fly, although not very efficiently owing to its large size. Their best protection from
1549:
Oryctoantiquus borealis, New Genus and
Species from the Eocene of Oregon, U.S.A., the World's Oldest Fossil Dynastine and Largest Fossil Scarabaeid (Coleoptera: Scarabaeidae: Dynastinae)
1561:
2553:
1084:
1830:
1459:
Bouchard, P., Y. Bousquet, A. Davies, M. Alonso-Zarazaga, J. Lawrence, C. Lyal, A. Newton, et al. (2011). "Family-group names in
Coleoptera (Insecta)".
2619:
2540:
2114:
1682:
1507:
1490:
57:
1764:
79:
1787:
1823:
1839:
2229:
2545:
1581:"Variation in an Extreme Weapon: Horn Performance Differences across Rhinoceros Beetle (Trypoxylus dichotomus) Populations"
2312:
1905:
329:
Many rhinoceros beetles are well known for their unique shapes and large sizes. Some famous species are, for example, the
2614:
2452:
2362:
1816:
598:
50:
44:
2506:
187:
2342:
2322:
1784:
1148:
863:
511:
409:
370:
354:
2558:
2519:
61:
2609:
2307:
2302:
1775:
1475:
Coleoptera, Beetles, Volume 1, Morphology and
Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga partim)
563:, due to being relatively clean, easy to maintain, and safe to handle. Also in Asia, male beetles are used for
338:
2209:
2186:
2099:
946:
890:
828:
390:
304:(Scarabaeidae). Other common names – some for particular groups of rhinoceros beetles – include
450:
mating season, and for digging. The size of the horn is a good indicator of nutrition and physical health.
2405:
2274:
1126:
1089:
692:
2576:
2367:
2259:
2049:
2044:
1895:
1657:
767:
723:
264:
1579:
Buchalski, Benjamin; Gutierrez, Eric; Emlen, Douglas; Lavine, Laura; Swanson, Brook (15 October 2019).
575:
2568:
2214:
2008:
1995:
1325:
1233:
962:
603:
537:
1686:
1303:
1106:
615:
in South Korea. Rhinoceros beetles could play a big part in the next generation of aircraft design.
2604:
2372:
2317:
2084:
2069:
1953:
1283:
1213:
954:
898:
673:
405:
1223:
1202:
735:
714:
704:
415:
1890:
1722:
1342:
1293:
1253:
1164:
1154:
1047:
1026:
528:
182:
1752:
1273:
1067:
2581:
1551:". 2009. The Coleopterists Bulletin 59(Mar 2005):127-135 DOI:10.1649/0010-065X(2005)0592.0.CO;2
1427:
1351:
1263:
1116:
2488:
2094:
2064:
2013:
2003:
1978:
1912:
1760:
1612:
1527:
1486:
1368:
1018:
926:
918:
823:
643:
612:
608:
547:
545:, and when they occur in large numbers, can cause economically significant damage. The fungus
1243:
1057:
683:
2264:
2146:
2039:
2034:
1900:
1640:
1602:
1592:
1519:
1478:
1410:
1405:
1174:
998:
990:
762:
445:
was the largest fossil scarabeid, with a length of 5 centimetres (2.0 in). Some modern
301:
2443:
2199:
564:
533:
362:
346:
306:
482:
stages can be several years long. The larvae feed on rotten wood and the adults feed on
2514:
2501:
2390:
2281:
2254:
2180:
2054:
1938:
1871:
1607:
1580:
772:
756:
542:
441:
1798:
2598:
2337:
1882:
1184:
982:
132:
2493:
2269:
2234:
2158:
1928:
1707:
1330:
1314:
884:
794:
330:
298:
249:
2457:
1808:
628:
17:
2527:
2437:
2332:
2163:
2134:
2059:
2029:
1983:
1933:
1803:
1644:
1036:
653:
591:
568:
454:
107:
2109:
2104:
1961:
1876:
1415:
1399:
1378:
1194:
1008:
871:
845:
805:
784:
466:
152:
117:
2428:
2327:
1943:
1597:
1437:
936:
833:
817:
745:
487:
462:
458:
382:
294:
219:
199:
157:
1616:
1531:
1523:
1482:
2475:
2422:
2224:
2129:
1971:
1966:
1094:
1078:
972:
908:
446:
147:
142:
127:
122:
112:
2467:
2385:
2480:
2219:
2175:
2141:
2119:
2074:
1388:
1358:
1135:
663:
583:
579:
319:
174:
162:
137:
2532:
2153:
1854:
1843:
483:
437:
419:
European rhinoceros beetle – three stages from larva to adult:
323:
239:
229:
209:
2399:
2124:
1404:
1319:
1083:
822:
761:
627:
499:
495:
491:
479:
428:
420:
414:
404:
2168:
587:
560:
465:. When the sun is out, they hide under logs or in vegetation to
424:
2403:
1812:
611:'s Mechanical Engineering Department and in collaboration with
551:
is a proven biocontrol agent for beetle infestation in crops.
29:
1759:. Series Entomologica. Vol. 28. Dr. W. Junk Publishers.
453:
The body of an adult rhinoceros beetle is covered by a thick
601:-funded research, examined the function and aerodynamics of
1473:
Beutel, Rolf G.; Leschen, Richard A.B., eds. (2016-03-21).
494:. First, the larvae hatch from eggs and later develop into
461:
is their size and stature, also avoiding many due to being
559:
Rhinoceros beetles have become popular pets in parts of
526:
Some species, such as the coconut rhinoceros beetle (
1547:
Brett
Ratcliffe, Dena M. Smith, Diane Marie Erwin. "
1042:
Auth. Lacordaire, 1856; all genera from
Madagascar:
536:, e.g., in tree plantations. Usually though, beetle
2412:
2353:
2295:
2247:
2198:
2083:
2022:
1994:
1952:
1921:
1864:
1850:
1508:"Inexpensive load carrying by rhinoceros beetles"
597:Dr. MinJun Kim, leading a team of engineers in
1737:— Rhinoceros beetles could play a big part....
1824:
1658:Featured Creatures: Coconut rhinoceros beetle
590:(approximately 18%), and they could become a
8:
1723:"Engineers Unlock Secrets of Beetle Flight"
1710:- Demain nos enfants mangeront des criquets
1562:"Why horn size matters when picking a mate"
1543:
1541:
592:protein source for a large human population
2400:
1861:
1831:
1817:
1809:
173:
93:
1633:Biological Journal of the Linnean Society
1606:
1596:
1422:Auth: Burmeister, 1847; selected genera:
80:Learn how and when to remove this message
1683:"Rhinoceros beetle gambling in Thailand"
43:This article includes a list of general
1452:
658:Lepeletier & Audinet-Serville, 1828
619:Tribes with selected genera and species
2115:List of crop plants pollinated by bees
1773:Dechambre (R.-P.) & Lachaume (G.)
1337:Auth: Mulsant, 1842. Selected genera:
1101:Auth: Mulsant, 1842. Selected genera:
840:Auth: MacLeay, 1819. Selected genera:
779:Auth: Laporte, 1840. Selected genera:
578:suggests the larvae contain much more
7:
2569:b353af6b-5c28-4965-b432-3b1cb2b0ddf7
2520:9aa5cb31-5738-4393-ab61-abf9d2b0af1f
638:Auth: Burmeister, 1847. all genera:
49:it lacks sufficient corresponding
25:
1208:Auth. Endrödi, 1966; all genera:
1158:– coconut rhinoceros beetle
326:of rhinoceros beetles are known.
2384:
607:with the help of researchers in
186:
34:
2620:Extant Eocene first appearances
1512:Journal of Experimental Biology
950:– Five-horned rhinoceros beetle
2230:Home-stored product entomology
1:
2313:Decline in insect populations
1906:List of insect-inspired songs
1506:Kram, Rodger (1 March 1996).
1735:ScienceDaily (Apr. 10, 2012)
1757:The Dynastinae of the World
1022:– Siamese rhinoceros beetle
599:National Science Foundation
512:Japanese rhinoceros beetles
2636:
2343:Pesticide toxicity to bees
2323:List of endangered insects
1785:Hillside Books, Canterbury
1668:"WHO? KNEW" (May 6, 2005)
1149:European rhinoceros beetle
864:Japanese rhinoceros beetle
410:European rhinoceros beetle
371:Japanese rhinoceros beetle
355:European rhinoceros beetle
2381:
1712:(2010 French documentary)
1645:10.1093/biolinnean/blz026
277:
272:
183:Scientific classification
181:
172:
96:
2308:Colony collapse disorder
2303:Bees and toxic chemicals
1776:The Beetles of the World
339:common rhinoceros beetle
2210:Insect bites and stings
2187:Drosophila melanogaster
2100:Biological pest control
1779:, volume 27, The genus
1598:10.3390/insects10100346
947:Eupatorus gracilicornis
891:Chalcosoma moellenkampi
829:Eupatorus gracilicornis
442:Oryctoantiquus borealis
391:Eastern Hercules beetle
64:more precise citations.
2275:Rats, Lice and History
1419:
1334:
1127:Megaceras briansaltini
1098:
837:
776:
635:
432:
412:
2577:Paleobiology Database
2260:Alfred Russel Wallace
2045:Entomological warfare
1896:Insects in literature
1524:10.1242/jeb.199.3.609
1483:10.1515/9783110373929
1408:
1323:
1087:
894:– Moellenkampi beetle
826:
768:Cyclocephala borealis
765:
631:
418:
408:
2515:Fauna Europaea (new)
2215:Insect sting allergy
2009:Cicadas in mythology
1804:Subfamily Dynastinae
1090:Trichogomphus mongol
963:Eupatorus birmanicus
860:Allomyrina dichotoma
604:Allomyrina dichotoma
538:population densities
532:), can become major
516:Allomyrina dichotoma
379:Allomyrina dichotoma
27:Subfamily of beetles
2318:Habitat destruction
2070:Insects in religion
1799:Family SCARABAEIDAE
1352:Diloboderus abderus
955:Eupatorus siamensis
899:Chalcosoma caucasus
2615:Insects in culture
2391:Insects portal
2363:Insects and humans
1891:Arthropods in film
1840:Human interactions
1732:. April 11, 2012.
1420:
1335:
1155:Oryctes rhinoceros
1145:Oryctes nasicornis
1099:
1027:Xylotrupes ulysses
838:
777:
636:
529:Oryctes rhinoceros
498:before they reach
433:
413:
359:Oryctes nasicornis
343:Xylotrupes ulysses
291:rhinoceros beetles
2592:
2591:
2406:Taxon identifiers
2397:
2396:
2286:Insect Literature
2243:
2242:
2142:Carmine/Cochineal
2095:Beneficial insect
2065:Insects in ethics
2014:Scarab (artifact)
2004:Bees in mythology
1913:Insects on stamps
1492:978-3-11-037392-9
1443:
1433:
1394:
1384:
1374:
1348:
1309:
1299:
1289:
1279:
1269:
1259:
1249:
1239:
1229:
1219:
1190:
1178:– ox beetle
1170:
1141:
1122:
1112:
1073:
1063:
1053:
1019:Xylotrupes gideon
1014:
994:- Elephant beetle
978:
942:
927:Dynastes neptunus
922:– Hercules beetle
919:Dynastes hercules
914:
902:– Caucasus beetle
877:
851:
811:
800:
790:
751:
741:
731:
720:
710:
700:
689:
679:
669:
659:
649:
613:Konkuk University
609:Drexel University
548:Metarhizium majus
367:Dynastes hercules
284:
283:
268:
90:
89:
82:
18:Rhinoceros Beetle
16:(Redirected from
2627:
2585:
2584:
2572:
2571:
2562:
2561:
2549:
2548:
2536:
2535:
2523:
2522:
2510:
2509:
2497:
2496:
2484:
2483:
2471:
2470:
2461:
2460:
2448:
2447:
2446:
2433:
2432:
2431:
2401:
2389:
2388:
2265:Jean-Henri Fabre
2040:Cricket fighting
2035:Cockroach racing
1901:Insects in music
1862:
1833:
1826:
1819:
1810:
1770:
1740:
1739:
1727:
1719:
1713:
1704:
1698:
1697:
1695:
1694:
1685:. Archived from
1679:
1673:
1666:
1660:
1655:
1649:
1648:
1627:
1621:
1620:
1610:
1600:
1576:
1570:
1569:
1558:
1552:
1545:
1536:
1535:
1503:
1497:
1496:
1470:
1464:
1461:ZooKeys, vol. 88
1457:
1441:
1431:
1411:Phileurus valgus
1392:
1383:Burmeister, 1847
1382:
1372:
1364:
1363:Burmeister, 1847
1346:
1307:
1297:
1287:
1277:
1267:
1257:
1247:
1237:
1227:
1217:
1189:Burmeister, 1847
1188:
1175:Strategus aloeus
1168:
1139:
1120:
1110:
1071:
1061:
1051:
1012:
1002:- Actaeon beetle
999:Megasoma actaeon
991:Megasoma elephas
976:
958:– Siamese beetle
941:Burmeister, 1847
940:
930:– Neptune beetle
912:
881:Chalcosoma atlas
875:
849:
809:
801:(masked chafers)
798:
788:
750:Burmeister, 1847
749:
739:
729:
718:
708:
699:Burmeister, 1847
698:
687:
678:Waterhouse, 1881
677:
667:
657:
648:Burmeister, 1847
647:
576:Séverin Tchibozo
387:Strategus aloeus
351:Megasoma elephas
335:Chalcosoma atlas
307:Hercules beetles
263:
191:
190:
177:
167:
104:
100:Temporal range:
94:
85:
78:
74:
71:
65:
60:this article by
51:inline citations
38:
37:
30:
21:
2635:
2634:
2630:
2629:
2628:
2626:
2625:
2624:
2610:Insects as food
2595:
2594:
2593:
2588:
2580:
2575:
2567:
2565:
2557:
2552:
2544:
2539:
2531:
2526:
2518:
2513:
2505:
2500:
2492:
2487:
2479:
2474:
2466:
2464:
2456:
2451:
2442:
2441:
2436:
2427:
2426:
2421:
2408:
2398:
2393:
2383:
2377:
2355:
2349:
2291:
2239:
2201:
2194:
2086:
2079:
2018:
1990:
1948:
1917:
1857:
1852:
1846:
1837:
1795:
1767:
1751:
1748:
1746:Further reading
1743:
1725:
1721:
1720:
1716:
1705:
1701:
1692:
1690:
1681:
1680:
1676:
1670:Current Science
1667:
1663:
1656:
1652:
1629:
1628:
1624:
1578:
1577:
1573:
1560:
1559:
1555:
1546:
1539:
1505:
1504:
1500:
1493:
1472:
1471:
1467:
1458:
1454:
1450:
1442:Latreille, 1807
1403:
1362:
1326:Pentodon idiota
1318:
1308:Dechambre, 1994
1298:Boisduval, 1835
1258:Fairmaire, 1881
1234:Coenoryctoderus
1206:
1082:
1072:Péringuey, 1901
1040:
966:– Rabbit beetle
821:
760:
626:
621:
565:gambling fights
557:
543:root vegetables
524:
508:
478:These beetles'
476:
403:
395:Dynastes tityus
363:Hercules beetle
347:elephant beetle
312:unicorn beetles
262:
185:
168:
166:
165:
160:
155:
150:
145:
140:
135:
130:
125:
120:
115:
110:
102:
101:
98:
86:
75:
69:
66:
56:Please help to
55:
39:
35:
28:
23:
22:
15:
12:
11:
5:
2633:
2631:
2623:
2622:
2617:
2612:
2607:
2597:
2596:
2590:
2589:
2587:
2586:
2573:
2563:
2550:
2537:
2524:
2511:
2502:Fauna Europaea
2498:
2485:
2472:
2462:
2449:
2434:
2418:
2416:
2410:
2409:
2404:
2395:
2394:
2382:
2379:
2378:
2376:
2375:
2370:
2365:
2359:
2357:
2351:
2350:
2348:
2347:
2346:
2345:
2340:
2335:
2325:
2320:
2315:
2310:
2305:
2299:
2297:
2293:
2292:
2290:
2289:
2282:Lafcadio Hearn
2279:
2267:
2262:
2257:
2255:Jan Swammerdam
2251:
2249:
2245:
2244:
2241:
2240:
2238:
2237:
2232:
2227:
2222:
2217:
2212:
2206:
2204:
2196:
2195:
2193:
2192:
2191:
2190:
2181:Model organism
2178:
2173:
2172:
2171:
2161:
2156:
2151:
2150:
2149:
2139:
2138:
2137:
2132:
2127:
2122:
2117:
2112:
2102:
2097:
2091:
2089:
2081:
2080:
2078:
2077:
2072:
2067:
2062:
2057:
2055:Insect farming
2052:
2047:
2042:
2037:
2032:
2026:
2024:
2020:
2019:
2017:
2016:
2011:
2006:
2000:
1998:
1992:
1991:
1989:
1988:
1987:
1986:
1976:
1975:
1974:
1969:
1958:
1956:
1950:
1949:
1947:
1946:
1941:
1939:Artificial fly
1936:
1931:
1925:
1923:
1919:
1918:
1916:
1915:
1910:
1909:
1908:
1898:
1893:
1888:
1887:
1886:
1879:
1872:Insects in art
1868:
1866:
1859:
1848:
1847:
1838:
1836:
1835:
1828:
1821:
1813:
1807:
1806:
1801:
1794:
1793:External links
1791:
1790:
1789:
1783:(Dynastidae),
1771:
1766:978-9061931386
1765:
1747:
1744:
1742:
1741:
1714:
1699:
1674:
1661:
1650:
1622:
1571:
1553:
1537:
1518:(3): 609–612.
1498:
1491:
1477:. De Gruyter.
1465:
1451:
1449:
1446:
1445:
1444:
1434:
1402:
1397:
1396:
1395:
1393:Erichson, 1847
1385:
1375:
1365:
1355:
1317:
1312:
1311:
1310:
1304:Paroryctoderus
1300:
1290:
1280:
1270:
1260:
1250:
1240:
1230:
1220:
1205:
1200:
1199:
1198:
1191:
1181:
1180:
1179:
1161:
1160:
1159:
1151:
1132:
1131:
1130:
1113:
1081:
1076:
1075:
1074:
1064:
1054:
1039:
1034:
1033:
1032:
1031:
1030:
1023:
1005:
1004:
1003:
995:
987:
969:
968:
967:
959:
951:
933:
932:
931:
923:
905:
904:
903:
895:
887:
868:
867:
866:
820:
815:
814:
813:
812:(rice beetles)
802:
791:
789:Erichson, 1847
773:Cyclocephalini
759:
757:Cyclocephalini
754:
753:
752:
742:
732:
721:
711:
701:
690:
680:
670:
660:
650:
625:
622:
620:
617:
556:
553:
523:
520:
507:
504:
475:
472:
427:(center), and
402:
399:
282:
281:
275:
274:
270:
269:
257:
253:
252:
247:
243:
242:
237:
233:
232:
227:
223:
222:
217:
213:
212:
207:
203:
202:
197:
193:
192:
179:
178:
170:
169:
161:
156:
151:
146:
141:
136:
131:
126:
121:
116:
111:
106:
105:
99:
88:
87:
42:
40:
33:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
2632:
2621:
2618:
2616:
2613:
2611:
2608:
2606:
2603:
2602:
2600:
2583:
2578:
2574:
2570:
2564:
2560:
2555:
2551:
2547:
2542:
2538:
2534:
2529:
2525:
2521:
2516:
2512:
2508:
2503:
2499:
2495:
2490:
2486:
2482:
2477:
2473:
2469:
2463:
2459:
2454:
2450:
2445:
2439:
2435:
2430:
2424:
2420:
2419:
2417:
2415:
2411:
2407:
2402:
2392:
2387:
2380:
2374:
2371:
2369:
2366:
2364:
2361:
2360:
2358:
2352:
2344:
2341:
2339:
2338:Neonicotinoid
2336:
2334:
2331:
2330:
2329:
2326:
2324:
2321:
2319:
2316:
2314:
2311:
2309:
2306:
2304:
2301:
2300:
2298:
2294:
2287:
2283:
2280:
2277:
2276:
2271:
2268:
2266:
2263:
2261:
2258:
2256:
2253:
2252:
2250:
2246:
2236:
2233:
2231:
2228:
2226:
2223:
2221:
2218:
2216:
2213:
2211:
2208:
2207:
2205:
2203:
2197:
2189:
2188:
2184:
2183:
2182:
2179:
2177:
2174:
2170:
2167:
2166:
2165:
2162:
2160:
2157:
2155:
2152:
2148:
2145:
2144:
2143:
2140:
2136:
2133:
2131:
2128:
2126:
2123:
2121:
2118:
2116:
2113:
2111:
2108:
2107:
2106:
2103:
2101:
2098:
2096:
2093:
2092:
2090:
2088:
2082:
2076:
2073:
2071:
2068:
2066:
2063:
2061:
2058:
2056:
2053:
2051:
2048:
2046:
2043:
2041:
2038:
2036:
2033:
2031:
2028:
2027:
2025:
2023:Other aspects
2021:
2015:
2012:
2010:
2007:
2005:
2002:
2001:
1999:
1997:
1993:
1985:
1982:
1981:
1980:
1977:
1973:
1970:
1968:
1965:
1964:
1963:
1960:
1959:
1957:
1955:
1951:
1945:
1942:
1940:
1937:
1935:
1932:
1930:
1927:
1926:
1924:
1920:
1914:
1911:
1907:
1904:
1903:
1902:
1899:
1897:
1894:
1892:
1889:
1885:
1884:
1883:Musca depicta
1880:
1878:
1875:
1874:
1873:
1870:
1869:
1867:
1863:
1860:
1856:
1849:
1845:
1841:
1834:
1829:
1827:
1822:
1820:
1815:
1814:
1811:
1805:
1802:
1800:
1797:
1796:
1792:
1788:
1786:
1782:
1778:
1777:
1772:
1768:
1762:
1758:
1754:
1750:
1749:
1745:
1738:
1736:
1731:
1724:
1718:
1715:
1711:
1709:
1703:
1700:
1689:on 2012-04-12
1688:
1684:
1678:
1675:
1671:
1665:
1662:
1659:
1654:
1651:
1646:
1642:
1638:
1634:
1626:
1623:
1618:
1614:
1609:
1604:
1599:
1594:
1590:
1586:
1582:
1575:
1572:
1567:
1566:New Scientist
1563:
1557:
1554:
1550:
1544:
1542:
1538:
1533:
1529:
1525:
1521:
1517:
1513:
1509:
1502:
1499:
1494:
1488:
1484:
1480:
1476:
1469:
1466:
1462:
1456:
1453:
1447:
1440:
1439:
1435:
1432:MacLeay, 1819
1430:
1429:
1425:
1424:
1423:
1417:
1413:
1412:
1407:
1401:
1398:
1391:
1390:
1386:
1381:
1380:
1376:
1371:
1370:
1366:
1361:
1360:
1356:
1354:
1353:
1345:
1344:
1340:
1339:
1338:
1332:
1328:
1327:
1322:
1316:
1313:
1306:
1305:
1301:
1296:
1295:
1291:
1288:Endrödi, 1978
1286:
1285:
1284:Oryctoderinus
1281:
1276:
1275:
1271:
1266:
1265:
1261:
1256:
1255:
1251:
1246:
1245:
1241:
1236:
1235:
1231:
1226:
1225:
1221:
1216:
1215:
1214:Chalcasthenes
1211:
1210:
1209:
1204:
1201:
1197:
1196:
1192:
1187:
1186:
1185:Trichogomphus
1182:
1177:
1176:
1172:
1171:
1167:
1166:
1162:
1157:
1156:
1152:
1150:
1146:
1143:
1142:
1140:Hellwig, 1798
1138:
1137:
1133:
1129:
1128:
1124:
1123:
1119:
1118:
1114:
1109:
1108:
1104:
1103:
1102:
1096:
1092:
1091:
1086:
1080:
1077:
1070:
1069:
1065:
1062:Olivier, 1789
1060:
1059:
1055:
1050:
1049:
1045:
1044:
1043:
1038:
1035:
1029:
1028:
1024:
1021:
1020:
1016:
1015:
1011:
1010:
1006:
1001:
1000:
996:
993:
992:
988:
986:- Mars beetle
985:
984:
983:Megasoma mars
980:
979:
975:
974:
970:
965:
964:
960:
957:
956:
952:
949:
948:
944:
943:
939:
938:
934:
929:
928:
924:
921:
920:
916:
915:
913:MacLeay, 1819
911:
910:
906:
901:
900:
896:
893:
892:
888:
886:
882:
879:
878:
874:
873:
869:
865:
861:
858:
857:
855:
848:
847:
843:
842:
841:
835:
831:
830:
825:
819:
816:
808:
807:
803:
797:
796:
792:
787:
786:
782:
781:
780:
774:
770:
769:
764:
758:
755:
748:
747:
743:
740:Thomson, 1859
738:
737:
733:
728:
726:
722:
719:Endrödi, 1974
717:
716:
712:
707:
706:
702:
697:
695:
691:
686:
685:
681:
676:
675:
674:Brachysiderus
671:
666:
665:
661:
656:
655:
651:
646:
645:
641:
640:
639:
634:
633:Antodon goryi
630:
624:Agaocephalini
623:
618:
616:
614:
610:
606:
605:
600:
595:
593:
589:
585:
581:
577:
574:Entomologist
572:
570:
566:
562:
554:
552:
550:
549:
544:
539:
535:
531:
530:
521:
519:
517:
513:
505:
503:
501:
497:
493:
489:
485:
481:
473:
471:
468:
464:
460:
456:
451:
448:
444:
443:
439:
430:
426:
422:
417:
411:
407:
400:
398:
396:
392:
388:
384:
380:
376:
372:
368:
364:
360:
356:
352:
348:
344:
340:
336:
332:
327:
325:
321:
318:. Over 1,500
317:
313:
309:
308:
303:
300:
299:scarab beetle
296:
292:
288:
280:
276:
271:
266:
261:
258:
255:
254:
251:
248:
245:
244:
241:
238:
235:
234:
231:
228:
225:
224:
221:
218:
215:
214:
211:
208:
205:
204:
201:
198:
195:
194:
189:
184:
180:
176:
171:
164:
159:
154:
149:
144:
139:
134:
129:
124:
119:
114:
109:
103:Eocene–Recent
95:
92:
84:
81:
73:
70:November 2015
63:
59:
53:
52:
46:
41:
32:
31:
19:
2413:
2368:Insecticides
2285:
2273:
2270:Hans Zinsser
2235:Clothes moth
2185:
1996:In mythology
1929:Fishing bait
1881:
1780:
1774:
1756:
1734:
1733:
1730:ScienceDaily
1729:
1726:(news story)
1717:
1708:Global Steak
1706:
1702:
1691:. Retrieved
1687:the original
1677:
1672:Vol.90 No.16
1669:
1664:
1653:
1639:(1): 34–43.
1636:
1632:
1625:
1588:
1584:
1574:
1565:
1556:
1515:
1511:
1501:
1474:
1468:
1460:
1455:
1436:
1426:
1421:
1409:
1387:
1377:
1367:
1357:
1350:
1341:
1336:
1331:Pentodontini
1324:
1315:Pentodontini
1302:
1292:
1282:
1272:
1268:Heller, 1896
1262:
1252:
1242:
1232:
1228:Heller, 1903
1224:Chalcocrates
1222:
1212:
1207:
1203:Oryctoderini
1193:
1183:
1173:
1163:
1153:
1144:
1134:
1125:
1115:
1105:
1100:
1088:
1066:
1056:
1052:Reiche, 1860
1046:
1041:
1025:
1017:
1007:
997:
989:
981:
971:
961:
953:
945:
935:
925:
917:
907:
897:
889:
885:Atlas beetle
880:
870:
859:
853:
844:
839:
827:
810:Harold, 1869
804:
799:Dejean, 1821
795:Cyclocephala
793:
783:
778:
766:
744:
736:Mitracephala
734:
724:
715:Horridocalia
713:
705:Gnathogolofa
703:
693:
682:
672:
662:
652:
642:
637:
632:
602:
596:
573:
558:
546:
527:
525:
515:
509:
477:
452:
440:
434:
394:
386:
378:
374:
366:
358:
350:
342:
334:
331:Atlas beetle
328:
316:horn beetles
315:
311:
305:
290:
286:
285:
278:
259:
250:Scarabaeidae
91:
76:
67:
48:
2528:iNaturalist
2438:Wikispecies
2354:Categories,
2333:Insecticide
2164:Sericulture
2135:Royal jelly
2060:Flea circus
2050:Entomophagy
1984:Cantharidin
1979:Spanish fly
1954:In medicine
1934:Fly fishing
1865:In the arts
1753:Endrödi, S.
1591:(10): 346.
1347:Sturm, 1826
1343:Diloboderus
1294:Oryctoderus
1278:Arrow, 1914
1254:Melanhyphus
1248:Sharp, 1877
1238:Prell, 1933
1218:Arrow, 1937
1169:Kirby, 1828
1048:Hemicyrthus
1037:Hexodontini
977:Kirby, 1825
852:(including
850:Arrow, 1911
730:Breme, 1844
709:Arrow, 1914
688:Ohaus, 1910
668:Brême, 1845
654:Agaocephala
582:(40%) than
569:mating call
455:exoskeleton
401:Description
375:kabutomushi
256:Subfamily:
62:introducing
2605:Dynastinae
2599:Categories
2458:Dynastinae
2444:Dynastinae
2414:Dynastinae
2373:Pesticides
2110:Bee pollen
2105:Beekeeping
2087:entomology
2030:Biomimicry
1962:Apitherapy
1922:In fishing
1877:Beetlewing
1858:in culture
1693:2006-11-03
1416:Phileurini
1400:Phileurini
1379:Pericoptus
1373:Hope, 1837
1349:monotypic
1274:Onychionyx
1195:Xyloryctes
1121:Hope, 1837
1111:Hope, 1837
1068:Hyboschema
1013:Hope, 1837
1009:Xylotrupes
876:Hope, 1837
872:Chalcosoma
854:Trypoxylus
846:Allomyrina
806:Dyscinetus
785:Ancognatha
694:Democrates
586:(20%) and
467:camouflage
389:) and the
287:Dynastinae
260:Dynastinae
240:Coleoptera
220:Arthropoda
97:Dynastinae
45:references
2356:templates
2328:Pesticide
1944:Fly tying
1438:Phileurus
1428:Cryptodus
1264:Neohyphus
1165:Strategus
1117:Megaceras
937:Eupatorus
834:Dynastini
818:Dynastini
746:Spodistes
725:Lycomedes
500:adulthood
488:plant sap
463:nocturnal
459:predators
383:ox beetle
295:subfamily
279:See text
206:Kingdom:
200:Eukaryota
2476:BugGuide
2465:BioLib:
2423:Wikidata
2296:Concerns
2248:Pioneers
2225:Woodworm
2130:Propolis
2085:Economic
1972:Melittin
1967:Apitoxin
1755:(1985).
1617:31618906
1463:, 1-972.
1369:Pentodon
1147:–
1095:Oryctini
1079:Oryctini
973:Megasoma
909:Dynastes
883:–
862:–
727:(beetle)
696:(beetle)
644:Aegopsis
522:As pests
447:Oryctini
423:(back),
322:and 225
246:Family:
216:Phylum:
210:Animalia
196:Domain:
2429:Q641142
2220:Bed bug
2202:insects
2200:Harmful
2176:Shellac
2120:Beeswax
2075:Jingzhe
1855:insects
1851:Aspects
1844:insects
1781:Oryctes
1608:6835817
1585:Insects
1532:9318326
1389:Tomarus
1359:Ligyrus
1244:Hatamus
1136:Oryctes
1058:Hexodon
684:Colacus
664:Antodon
584:chicken
580:protein
474:Ecology
431:(front)
320:species
297:of the
273:Tribes
265:MacLeay
236:Order:
230:Insecta
226:Class:
58:improve
2566:NZOR:
2546:678510
2533:136361
2507:246911
2159:Kermes
2154:Chitin
2147:Polish
1763:
1615:
1605:
1530:
1489:
506:Mating
490:, and
484:nectar
480:larval
438:Eocene
324:genera
302:family
293:are a
267:, 1819
47:, but
2582:70053
2559:50519
2494:7NLQQ
2481:12431
2125:Honey
1842:with
1448:Notes
1107:Enema
534:pests
510:Male
496:pupae
492:fruit
429:imago
421:larva
2554:NCBI
2541:ITIS
2468:8133
2169:Silk
1761:ISBN
1613:PMID
1528:PMID
1487:ISBN
588:beef
561:Asia
555:Uses
425:pupa
108:PreꞒ
2489:CoL
2453:AFD
1853:of
1641:doi
1637:127
1603:PMC
1593:doi
1520:doi
1516:199
1479:doi
397:).
381:),
373:or
369:),
361:),
353:),
345:),
337:),
314:or
289:or
2601::
2579::
2556::
2543::
2530::
2517::
2504::
2491::
2478::
2455::
2440::
2425::
1728:.
1635:.
1611:.
1601:.
1589:10
1587:.
1583:.
1564:.
1540:^
1526:.
1514:.
1510:.
1485:.
856:)
594:.
571:.
502:.
486:,
310:,
158:Pg
2288:)
2284:(
2278:)
2272:(
1832:e
1825:t
1818:v
1769:.
1696:.
1647:.
1643::
1619:.
1595::
1568:.
1534:.
1522::
1495:.
1481::
1418:)
1414:(
1333:)
1329:(
1097:)
1093:(
836:)
832:(
775:)
771:(
514:(
393:(
385:(
377:(
365:(
357:(
349:(
341:(
333:(
163:N
153:K
148:J
143:T
138:P
133:C
128:D
123:S
118:O
113:Ꞓ
83:)
77:(
72:)
68:(
54:.
20:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
