1218:
992:
536:
1128:
359:
544:
777:
745:
1412:
713:
1153:
680:
878:
1439:
1193:
121:
1268:
1243:
335:
1385:
1094:
810:
1307:
582:
The extent of the Crato unit and its relationship to the
Romualdo Formation had long been ill-defined. It was not until a 2007 volume on the unit by Martill, Bechly and Loveridge that the Crato Formation was given a formal type locality, and was formally made a distinct formation separate from the
2193:
Custódio, M.A.; Guaglio, F.; Warren, L.V.; Simões, M.G.; Fürsich, F.T.; Perinotto, J.A.; Assine, M.L. (2017). "The transgressive-regressive cycle of the
Romualdo Formation (Araripe Basin): Sedimentary archive of the Early Cretaceous marine ingression in the interior of Northeast Brazil".
1896:
Aranciaga
Rolando, Alexis M.; Brissón Egli, Federico; Sales, Marcos A.F.; Martinelli, Agustín G.; Canale, Juan I.; Ezcurra, Martín D. (2018). "A supposed Gondwanan oviraptorosaur from the Albian of Brazil represents the oldest South American megaraptoran".
503:
collected fossils has sprung up from the decade of 1970, driven by the remarkable state of preservation and beauty of these fossils and amounting to a considerable local industry. An urgent preservation program is being called for by paleontologists.
2407:"Novos Sítios Fossilíferos da Formação Romualdo, Cretáceo Inferior, Bacia do Araripe, Exu, Pernambuco, Nordeste do Brasil - New Fossiliferous Sites of the Romualdo Formation, Lower Cretaceous, Araripe Basin, Exu, Pernambuco, Northeast of Brazil"
566:
reflect changeable conditions in the opening sea. The age of the
Romualdo Formation, formerly known as the Romualdo Member of the Santana Formation, has been controversial, though most workers have agreed that it lies on or near the
1959:
Juliana Manso Sayão; Antônio Álamo
Feitosa Saraiva; Arthur Souza Brum; Renan Alfredo Machado Bantim; Rafael Cesar Lima Pedroso de Andrade; Xin Cheng; Flaviana Jorge de Lima; Helder de Paula Silva; Alexander W. A. Kellner (2020).
561:
was previously considered the lowest member of the then
Santana Formation, but has been elevated to a formal formation. The Crato Formation is the product of a single phase, where complicated sequence of sediment
2541:"Controlled excavations in the Romualdo Member of the Santana Formation (Early Cretaceous, Araripe Basin, northeastern Brazil): stratigraphic, palaeoenvironmental and palaeoecological implications"
2721:
Neumann, V.H.; Borrego, A.G.; Cabrera, L.; Dino, R. (2003). "Organic matter composition and distribution through the Aptian–Albian lacustrine sequences of the
Araripe Basin, northeastern Brazil".
2865:
2637:"The age of the Cretaceous Santana Formation fossil Konservat Lagerstätte of north-east Brazil: a historical review and an appraisal of the biochronostratigraphic utility of its palaeobiota"
492:
that formed nodules around dead organisms, preserving even soft parts of their anatomy. In preservation, the nodules are etched away with acid, and the fossils often prepared by the
2490:
Fabin, Carlos E.; Correia Filho, Osvaldo J.; Alencar, Márcio L.; Barbosa, José A.; de
Miranda, Tiago S.; Neumann, Virgínio H.; Gomes, Igor F.; de Santana, Felipe R. (2018).
2548:
991:
2765:
2499:
523:) is a very rare bird that was discovered only in the late 20th century; it is not known from anywhere outside the characteristic forest that grows on the
1217:
2761:"Shrimps from the Santana Group (Cretaceous: Albian): new species (Crustacea: Decapoda: Dendrobranchiata) and new record (Crustacea: Decapoda: Caridea)"
2441:"A new ornithocheirid, Barbosania gracilirostris gen. et sp. nov. (Pterosauria, Pterodactyloidea) from the Santana Formation (Cretaceous) of NE Brazil"
1740:
2855:
2819:
2540:
2618:
1962:"The first theropod dinosaur (Coelurosauria, Theropoda) from the base of the Romualdo Formation (Albian), Araripe Basin, Northeast Brazil"
2890:
2824:
870:
2915:
2715:
2690:
2905:
2895:
2885:
2870:
2593:
869:
Type specimen 423-R is a single skull articulating with part of a lower jaw. A more complete specimen, AMNH 24450, is at the
2910:
2900:
2723:
2384:
2318:
2238:
2128:
2112:
2276:
2176:
2147:
2875:
535:
233:
2850:
2535:
Fara, Emmanuel; Saraiva, Antônio Á.F.; Campos, Díogenes de
Almeida; Moreira, João K.R.; de Carvalho Siebra, Daniele;
1127:
358:
2860:
2610:
279:
240:
2880:
2094:
776:
543:
2292:
441:
The
Romualdo Formation earns the designation of Lagerstätte due to an exceedingly well preserved and diverse
2305:
1770:
1750:
412:
2492:"Stratigraphic Relations of the Ipubi Formation: Siliciclastic-Evaporitic Succession of the Araripe Basin"
996:
946:
127:
2491:
170:
2339:
2732:
2650:
2557:
2452:
2351:
2340:"Continental break up and the distribution of fishes of Western Gondwana during the Early Cretaceous"
2203:
2035:
1973:
1906:
1813:
1760:
1732:
1203:
146:
1598:
2641:
1765:
1689:
1675:
489:
2478:
2059:
612:
have been found in Ceara state, Brazil. The oviraptorosaurian remains have been re-identified as
524:
493:
408:
2813:
1469:
744:
1449:
2809:
2802:
2784:
2711:
2686:
2614:
2604:
2589:
2586:
Laboratory Preparation Techniques. In Maisey, j:G. (ed): Santana Fossils: An Illustrated Atlas
2536:
2516:
2470:
2367:
2051:
2001:
1932:
1839:
1831:
404:
2021:"A new crested maniraptoran dinosaur from the Santana Formation (Lower Cretaceous) of Brazil"
2019:
Martill, D. M.; Cruickshank, A. R. I.; Frey, E.; Small, P. G.; Clarke, M. (1 February 1996).
1626:
397:
come together. The geological formation, previously designated as the Romualdo Member of the
2793:
2774:
2749:
2740:
2658:
2565:
2508:
2460:
2418:
2359:
2219:
2211:
2043:
1991:
1981:
1922:
1914:
1821:
1755:
1583:
1411:
1152:
1061:
802:
712:
609:
474:
424:
679:
1228:
1163:
1007:
769:
558:
516:
478:
466:
184:
180:
133:
2736:
2654:
2561:
2456:
2355:
2207:
2039:
1977:
1910:
1817:
1996:
1961:
1633:
1590:
877:
829:
374:
272:
2753:
2744:
2020:
2844:
2406:
2063:
1745:
1696:
1612:
1368:
1253:
1138:
1079:
1025:
903:
855:
787:
740:. Partial skull (posterior half); one of the most complete spinosaurid skulls known.
737:
668:
552:
470:
435:
399:
382:
350:
166:
156:
72:
29:
2482:
1438:
1192:
888:
605:
500:
462:
449:
are often found with stomach contents preserved, enabling paleontologists to study
120:
2798:
2779:
2760:
2662:
2512:
1918:
2683:
Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria
2569:
2215:
2636:
2388:
2325:
2280:
2242:
2180:
2151:
2135:
2116:
2100:
1619:
1545:
1395:
1267:
1113:
705:
690:
613:
446:
438:. At that time, the South Atlantic was opening up in a long narrow shallow sea.
47:
1986:
2465:
2440:
1682:
1661:
1605:
1531:
1422:
1242:
1043:
654:
576:
508:
431:
428:
386:
314:
92:
57:
2474:
2371:
2055:
2047:
1936:
1835:
363:
Extent of the Santana Group, to which the Romualdo Formation belongs, in blue
294:
281:
255:
242:
1717:
1703:
1640:
1278:
923:
755:
723:
597:
486:
482:
454:
450:
217:
97:
41:
2806:
2788:
2520:
2363:
2005:
1843:
1384:
1093:
809:
2423:
1710:
1654:
1647:
1317:
628:
394:
390:
322:
318:
207:
87:
82:
67:
62:
52:
2224:
2405:
Cavalcanti Duque, Rudah Ruano; Franca Barreto, Alcina Magnólia (2018).
1927:
1668:
1538:
601:
563:
458:
416:
102:
77:
1306:
499:
Local mining activities for cement and construction damage the sites.
2759:
Pinheiro, Allysson P.; Saraiva, Antônio Á.; Santana, William (2014).
572:
568:
442:
420:
378:
340:
33:
1826:
1801:
1504:
2710:(Field Guide to Fossils no. 5) (The Palaeontological Association)
1509:
671:. Partial skull (rostralmost portion). Possible junior synonym of
542:
534:
221:
805:. Some caudal vertebrae, partial pelvis, most of both hindlimbs.
453:
relationships in this ecosystem. There are also fine examples of
2603:
Martill, David M.; Bechly, Günter; Loveridge, Robert F. (2007).
512:
2685:. Berkeley: University of California Press. pp. 563–570.
2606:
The Crato Fossil Beds of Brazil: Window into an Ancient World
1437:
1410:
1383:
1305:
1266:
1241:
1216:
1191:
1151:
1126:
1092:
990:
876:
808:
775:
743:
711:
678:
2679:
Dinosaur distribution (Early Cretaceous, South America)
1882:
1880:
1878:
1876:
1874:
2584:
Maisey, J.G.; Rutzky, I.; Blum, S.; Elvers, W. (1991).
1339:
918:
824:
623:
575:
boundary, about 112 million years ago. Nevertheless, a
527:
soils formed ultimately from Romualdo Formation rocks.
2820:"Pterosaurs Diorama Depicts Ancient Brazilian Coast"
2076:"Table 5.1," in Weishampel, et al. (2004). Page 114.
539:
Outcrop and interpretation of the Romualdo Formation
2708:
Fossils of the Santana and Crato Formations, Brazil
1949:"Table 4.1," in Weishampel, et al. (2004). Page 73.
346:
328:
310:
271:
232:
227:
213:
203:
198:
190:
176:
162:
152:
142:
23:
583:Romualdo Formation, which is about 10 Ma younger.
2866:Cretaceous paleontological sites of South America
2549:Palaeogeography, Palaeoclimatology, Palaeoecology
1855:
1853:
2439:Elgin, Ross A.; Frey, Eberhard (20 April 2011).
2263:
2261:
2259:
2257:
2255:
2253:
2251:
515:conditions unlike elsewhere in the region. The
8:
1802:"Brazil clamps down on illegal fossil trade"
547:Outcrop of the Romualdo and Ipubi Formations
511:of Romualdo Formation rocks has contributed
2677:Weishampel, David B.; et al. (2004).
2172:
2170:
2168:
2166:
2164:
2162:
2160:
357:
119:
20:
2797:
2778:
2588:. Tfh Publications Inc. pp. 99–103.
2464:
2422:
2411:Anuário do Instituto de Geociências, UFRJ
2223:
1995:
1985:
1926:
1825:
2766:Anais da Academia Brasileira de Ciências
2500:Anais da Academia Brasileira de Ciências
1741:List of dinosaur-bearing rock formations
1342:
921:
827:
626:
1783:
445:faunal assemblage. Some 25 species of
434:with shallow marine incursions of the
2724:International Journal of Coal Geology
7:
772:. Pelvis and partial left hindlimb.
137:(bottom) from the Romualdo Formation
1561:Uncertain or tentative taxa are in
596:Indeterminate remains of non-avian
2825:American Museum of Natural History
1800:Gibney, Elizabeth (6 March 2014).
871:American Museum of Natural History
553:Santana Group § Basin history
469:. Even dinosaurs are represented (
14:
2028:Journal of the Geological Society
1521:Taxon falsely reported as present
2338:Maisey, John G. (1 April 2000).
333:
1790:Maisey et al., 1991, pp. 99–103
1526:Dubious taxon or junior synonym
2445:Swiss Journal of Palaeontology
194:2–10 m (6.6–32.8 ft)
1:
2856:Geologic formations of Brazil
2799:10.1590/0001-3765201420130338
2780:10.1590/0001-3765201420130338
2754:10.1016/S0166-5162(03)00018-1
2745:10.1016/S0166-5162(03)00018-1
2663:10.1016/j.cretres.2007.01.002
2513:10.1590/0001-3765201820170526
1919:10.1016/j.cretres.2017.10.019
1886:Weishampel, 2004, pp. 563–570
403:, named after the village of
2570:10.1016/j.palaeo.2004.12.012
2216:10.1016/j.sedgeo.2017.07.010
1599:Beurlenichthys ouricuriensis
1481:
1461:
1434:
1407:
1380:
273:Approximate paleocoordinates
16:Brazilian geologic formation
1972:(1): Article number 10892.
1488:
2932:
2611:Cambridge University Press
1987:10.1038/s41598-020-67822-9
1697:Santanaclupea silvasantosi
1591:Axelrodichthys araripensis
708:. Partial right hindlimb.
550:
419:were deposited during the
407:, lies at the base of the
2891:Environment of Pernambuco
2466:10.1007/s13358-011-0017-4
2277:Buxéxé, Santana do Cariri
1495:
1491:
1091:
1077:
579:age cannot be ruled out.
356:
118:
28:
2916:Northeast Region, Brazil
2706:David M. Martill, 1993.
2507:(2 suppl 1): 2049–2071.
2267:Fara et al., 2005, p.152
2048:10.1144/gsjgs.153.1.0005
1676:Placidichthys bidorsalis
1493:
485:of the site resulted in
2906:Landforms of Pernambuco
2537:Kellner, Alexander W.A.
1771:Mata Amarilla Formation
1751:Cerro Barcino Formation
413:Johann Baptist von Spix
411:. It was discovered by
2871:Paleontology in Brazil
2364:10.1006/cres.1999.0195
2096:Santanachelys gaffneyi
2085:Elgin & Frey, 2011
1569:taxa are discredited.
1442:
1415:
1388:
1310:
1271:
1246:
1221:
1196:
1156:
1131:
1097:
1000:
881:
813:
780:
748:
716:
683:
548:
540:
2635:Martill, D.M (2007).
2424:10.11137/2018_1_05_14
1736:(with link directory)
1441:
1414:
1387:
1309:
1270:
1245:
1220:
1195:
1155:
1130:
1096:
994:
880:
812:
779:
747:
715:
682:
673:Irritator challengeri
546:
538:
521:Antilophia bokermanni
375:Konservat-Lagerstätte
2896:Environment of Piauí
2886:Environment of Ceará
1859:Martill et al., 2007
1733:List of fossil sites
385:where the states of
147:Geological formation
2876:Mudstone formations
2737:2003IJCG...54...21N
2655:2007CrRes..28..895M
2642:Cretaceous Research
2562:2005PPP...218..145F
2457:2011SwJP..130..259E
2356:2000CrRes..21..281M
2344:Cretaceous Research
2208:2017SedG..359....1C
2196:Sedimentary Geology
2130:Araripemys barretoi
2040:1996JGSoc.153....5M
1978:2020NatSR..1010892S
1911:2018CrRes..84..107A
1899:Cretaceous Research
1818:2014Natur.507...20G
1766:Itapecuru Formation
899:Romualdo Formation
866:Romualdo Formation
291: /
252: /
30:Stratigraphic range
2911:Landforms of Piauí
2901:Landforms of Ceará
2851:Romualdo Formation
2177:Chapada do Araripe
1966:Scientific Reports
1516:Reclassified taxon
1443:
1416:
1389:
1311:
1272:
1247:
1222:
1197:
1157:
1132:
1098:
1001:
882:
814:
781:
749:
717:
684:
549:
541:
531:Geology and dating
525:Chapada do Araripe
501:Trade in illegally
494:transfer technique
371:Romualdo Formation
171:Arajara Formations
24:Romualdo Formation
2861:Cretaceous Brazil
2620:978-1-139-46776-6
2113:Juazeiro do Norte
1761:Guarujá Formation
1648:Microdon penalvai
1573:
1572:
1564:
1552:
1551:
1548:
1541:
1534:
1527:
1522:
1517:
1512:
1487:
1486:
1333:
1332:
1297:T. longicristatus
1050:B. gracilirostris
955:A. blittersdorffi
912:
911:
818:
817:
697:A. museunacionali
507:In addition, the
405:Santana do Cariri
400:Santana Formation
367:
366:
2923:
2881:Deltaic deposits
2837:
2835:
2833:
2801:
2792:
2782:
2748:
2696:
2673:
2671:
2669:
2631:
2629:
2627:
2599:
2580:
2578:
2576:
2556:(1–2): 145–160.
2545:
2531:
2529:
2527:
2496:
2486:
2468:
2435:
2433:
2431:
2426:
2392:
2382:
2376:
2375:
2335:
2329:
2316:
2310:
2303:
2297:
2290:
2284:
2274:
2268:
2265:
2246:
2236:
2230:
2229:
2227:
2190:
2184:
2174:
2155:
2145:
2139:
2126:
2120:
2110:
2104:
2092:
2086:
2083:
2077:
2074:
2068:
2067:
2025:
2016:
2010:
2009:
1999:
1989:
1956:
1950:
1947:
1941:
1940:
1930:
1893:
1887:
1884:
1869:
1866:
1860:
1857:
1848:
1847:
1829:
1797:
1791:
1788:
1756:Elrhaz Formation
1641:Lepidotes wenzai
1584:Araripelepidotes
1562:
1544:
1537:
1530:
1525:
1520:
1515:
1508:
1505:
1489:
1340:
1062:Brasileodactylus
919:
825:
624:
475:Tyrannosauroidea
425:Early Cretaceous
377:in northeastern
361:
339:
337:
336:
306:
305:
303:
302:
301:
296:
292:
289:
288:
287:
284:
267:
266:
264:
263:
262:
257:
253:
250:
249:
248:
245:
185:Ipubi Formations
123:
114:
44:
21:
2931:
2930:
2926:
2925:
2924:
2922:
2921:
2920:
2841:
2840:
2831:
2829:
2828:. 14 April 2014
2818:
2758:
2720:
2703:
2701:Further reading
2693:
2676:
2667:
2665:
2634:
2625:
2623:
2621:
2613:. p. 236.
2602:
2596:
2583:
2574:
2572:
2543:
2534:
2525:
2523:
2494:
2489:
2438:
2429:
2427:
2404:
2401:
2396:
2395:
2383:
2379:
2337:
2336:
2332:
2317:
2313:
2304:
2300:
2291:
2287:
2275:
2271:
2266:
2249:
2237:
2233:
2192:
2191:
2187:
2175:
2158:
2146:
2142:
2127:
2123:
2111:
2107:
2093:
2089:
2084:
2080:
2075:
2071:
2023:
2018:
2017:
2013:
1958:
1957:
1953:
1948:
1944:
1895:
1894:
1890:
1885:
1872:
1867:
1863:
1858:
1851:
1827:10.1038/507020a
1799:
1798:
1794:
1789:
1785:
1780:
1775:
1728:
1723:
1690:Iansan beurleni
1578:
1560:
1503:
1338:
1229:Thalassodromeus
1164:Santanadactylus
1008:Araripedactylus
917:
906:crocodyliform.
830:Crocodylomorphs
823:
821:Crocodylomorphs
770:Compsognathidae
622:
610:oviraptorosaurs
608:, and possibly
594:
589:
559:Crato Formation
555:
533:
517:Araripe manakin
481:). The unusual
479:Compsognathidae
467:crocodylomorphs
409:Araripe Plateau
362:
334:
332:
299:
297:
293:
290:
285:
282:
280:
278:
277:
260:
258:
254:
251:
246:
243:
241:
239:
238:
138:
134:Santanadactylus
113:
112:
111:
110:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
39:
38:
36:
17:
12:
11:
5:
2929:
2927:
2919:
2918:
2913:
2908:
2903:
2898:
2893:
2888:
2883:
2878:
2873:
2868:
2863:
2858:
2853:
2843:
2842:
2839:
2838:
2816:
2773:(2): 663–670.
2756:
2731:(1–2): 21–40.
2718:
2702:
2699:
2698:
2697:
2691:
2674:
2649:(6): 895–920.
2632:
2619:
2600:
2594:
2581:
2532:
2487:
2436:
2400:
2397:
2394:
2393:
2377:
2350:(2): 281–314.
2330:
2311:
2298:
2285:
2269:
2247:
2231:
2185:
2156:
2140:
2121:
2105:
2087:
2078:
2069:
2011:
1951:
1942:
1888:
1870:
1861:
1849:
1792:
1782:
1781:
1779:
1776:
1774:
1773:
1768:
1763:
1758:
1753:
1748:
1743:
1738:
1729:
1727:
1724:
1722:
1721:
1714:
1711:Tribodus limae
1707:
1700:
1693:
1686:
1679:
1672:
1669:Oshunia brevis
1665:
1658:
1651:
1644:
1637:
1630:
1623:
1616:
1609:
1602:
1595:
1587:
1579:
1577:
1574:
1571:
1570:
1555:
1553:
1550:
1549:
1542:
1535:
1528:
1523:
1518:
1513:
1497:
1496:
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1492:
1485:
1484:
1482:
1480:
1478:
1473:
1465:
1464:
1462:
1460:
1458:
1453:
1445:
1444:
1435:
1433:
1431:
1426:
1418:
1417:
1408:
1406:
1404:
1399:
1391:
1390:
1381:
1379:
1377:
1372:
1364:
1363:
1360:
1357:
1354:
1351:
1347:
1346:
1337:
1334:
1331:
1330:
1328:
1326:
1321:
1313:
1312:
1303:
1301:
1300:
1299:
1294:
1289:
1287:T. deliradamus
1282:
1274:
1273:
1264:
1262:
1260:T. mesembrinus
1257:
1249:
1248:
1239:
1237:
1232:
1224:
1223:
1214:
1212:
1210:T. wellnhoferi
1207:
1199:
1198:
1189:
1187:
1186:
1185:
1180:
1174:
1172:S. brasilensis
1167:
1159:
1158:
1149:
1147:
1142:
1134:
1133:
1124:
1122:
1117:
1109:
1108:
1106:
1100:
1099:
1090:
1088:
1083:
1075:
1074:
1072:
1070:
1068:B. araripensis
1065:
1057:
1056:
1054:
1052:
1047:
1039:
1038:
1036:
1034:
1029:
1021:
1020:
1018:
1016:
1011:
1003:
1002:
988:
986:
985:
984:
979:
974:
969:
967:A. araripensis
964:
958:
957:
950:
942:
941:
938:
935:
932:
928:
927:
916:
913:
910:
909:
907:
900:
897:
892:
884:
883:
874:
867:
864:
859:
851:
850:
847:
844:
841:
838:
834:
833:
822:
819:
816:
815:
806:
803:tyrannosauroid
799:
796:
791:
783:
782:
773:
767:
764:
762:M. asymmetrica
759:
751:
750:
741:
735:
732:
730:I. challengeri
727:
719:
718:
709:
702:
699:
694:
686:
685:
676:
666:
663:
658:
650:
649:
646:
643:
640:
637:
633:
632:
621:
618:
606:ornithischians
593:
590:
588:
587:Fossil content
585:
551:Main article:
532:
529:
436:proto-Atlantic
373:is a geologic
365:
364:
354:
353:
348:
344:
343:
330:
326:
325:
312:
308:
307:
275:
269:
268:
236:
230:
229:
225:
224:
215:
211:
210:
205:
201:
200:
196:
195:
192:
188:
187:
178:
174:
173:
164:
160:
159:
154:
150:
149:
144:
140:
139:
124:
116:
115:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
45:
26:
25:
15:
13:
10:
9:
6:
4:
3:
2:
2928:
2917:
2914:
2912:
2909:
2907:
2904:
2902:
2899:
2897:
2894:
2892:
2889:
2887:
2884:
2882:
2879:
2877:
2874:
2872:
2869:
2867:
2864:
2862:
2859:
2857:
2854:
2852:
2849:
2848:
2846:
2827:
2826:
2821:
2817:
2815:
2811:
2808:
2804:
2800:
2795:
2790:
2786:
2781:
2776:
2772:
2768:
2767:
2762:
2757:
2755:
2751:
2746:
2742:
2738:
2734:
2730:
2726:
2725:
2719:
2717:
2716:0-901702-46-3
2713:
2709:
2705:
2704:
2700:
2694:
2692:0-520-24209-2
2688:
2684:
2680:
2675:
2664:
2660:
2656:
2652:
2648:
2644:
2643:
2638:
2633:
2622:
2616:
2612:
2608:
2607:
2601:
2597:
2591:
2587:
2582:
2571:
2567:
2563:
2559:
2555:
2551:
2550:
2542:
2538:
2533:
2522:
2518:
2514:
2510:
2506:
2502:
2501:
2493:
2488:
2484:
2480:
2476:
2472:
2467:
2462:
2458:
2454:
2450:
2446:
2442:
2437:
2425:
2420:
2416:
2412:
2408:
2403:
2402:
2398:
2390:
2386:
2381:
2378:
2373:
2369:
2365:
2361:
2357:
2353:
2349:
2345:
2341:
2334:
2331:
2327:
2323:
2322:type locality
2321:
2320:Placidichthys
2315:
2312:
2309:
2308:
2307:Iemanja palma
2302:
2299:
2296:
2295:
2294:Enneles audax
2289:
2286:
2282:
2278:
2273:
2270:
2264:
2262:
2260:
2258:
2256:
2254:
2252:
2248:
2244:
2240:
2239:Casa de Pedra
2235:
2232:
2226:
2221:
2217:
2213:
2209:
2205:
2202:(359): 1–15.
2201:
2197:
2189:
2186:
2182:
2178:
2173:
2171:
2169:
2167:
2165:
2163:
2161:
2157:
2153:
2149:
2144:
2141:
2137:
2133:
2132:type locality
2131:
2125:
2122:
2118:
2114:
2109:
2106:
2102:
2098:
2097:
2091:
2088:
2082:
2079:
2073:
2070:
2065:
2061:
2057:
2053:
2049:
2045:
2041:
2037:
2033:
2029:
2022:
2015:
2012:
2007:
2003:
1998:
1993:
1988:
1983:
1979:
1975:
1971:
1967:
1963:
1955:
1952:
1946:
1943:
1938:
1934:
1929:
1924:
1920:
1916:
1912:
1908:
1904:
1900:
1892:
1889:
1883:
1881:
1879:
1877:
1875:
1871:
1868:Martill, 2007
1865:
1862:
1856:
1854:
1850:
1845:
1841:
1837:
1833:
1828:
1823:
1819:
1815:
1811:
1807:
1803:
1796:
1793:
1787:
1784:
1777:
1772:
1769:
1767:
1764:
1762:
1759:
1757:
1754:
1752:
1749:
1747:
1746:Araripe Basin
1744:
1742:
1739:
1737:
1734:
1731:
1730:
1725:
1720:
1719:
1715:
1713:
1712:
1708:
1706:
1705:
1701:
1699:
1698:
1694:
1692:
1691:
1687:
1685:
1684:
1680:
1678:
1677:
1673:
1671:
1670:
1666:
1664:
1663:
1659:
1657:
1656:
1652:
1650:
1649:
1645:
1643:
1642:
1638:
1636:
1635:
1634:Iemanja palma
1631:
1629:
1628:
1627:Enneles audax
1624:
1622:
1621:
1617:
1615:
1614:
1613:Calamopleurus
1610:
1608:
1607:
1603:
1601:
1600:
1596:
1593:
1592:
1588:
1586:
1585:
1581:
1580:
1575:
1568:
1559:
1556:
1554:
1547:
1543:
1540:
1536:
1533:
1529:
1524:
1519:
1514:
1511:
1507:
1506:
1502:
1499:
1498:
1490:
1483:
1479:
1477:
1474:
1472:
1471:
1467:
1466:
1463:
1459:
1457:
1454:
1452:
1451:
1447:
1446:
1440:
1436:
1432:
1430:
1427:
1425:
1424:
1420:
1419:
1413:
1409:
1405:
1403:
1400:
1398:
1397:
1393:
1392:
1386:
1382:
1378:
1376:
1373:
1371:
1370:
1369:Santanachelys
1366:
1365:
1361:
1358:
1355:
1352:
1349:
1348:
1345:
1341:
1335:
1329:
1327:
1325:
1322:
1320:
1319:
1315:
1314:
1308:
1304:
1302:
1298:
1295:
1293:
1290:
1288:
1285:
1284:
1283:
1281:
1280:
1276:
1275:
1269:
1265:
1263:
1261:
1258:
1256:
1255:
1254:Tropeognathus
1251:
1250:
1244:
1240:
1238:
1236:
1233:
1231:
1230:
1226:
1225:
1219:
1215:
1213:
1211:
1208:
1206:
1205:
1201:
1200:
1194:
1190:
1188:
1184:
1181:
1179:
1175:
1173:
1170:
1169:
1168:
1166:
1165:
1161:
1160:
1154:
1150:
1148:
1146:
1143:
1141:
1140:
1139:Maaradactylus
1136:
1135:
1129:
1125:
1123:
1121:
1118:
1116:
1115:
1111:
1110:
1107:
1105:
1102:
1101:
1095:
1089:
1087:
1084:
1082:
1081:
1080:Cearadactylus
1076:
1073:
1071:
1069:
1066:
1064:
1063:
1059:
1058:
1055:
1053:
1051:
1048:
1046:
1045:
1041:
1040:
1037:
1035:
1033:
1030:
1028:
1027:
1026:Araripesaurus
1023:
1022:
1019:
1017:
1015:
1012:
1010:
1009:
1005:
1004:
999:
998:
993:
989:
987:
983:
982:A. spielbergi
980:
978:
975:
973:
970:
968:
965:
963:
960:
959:
956:
953:
952:
951:
949:
948:
944:
943:
939:
936:
933:
930:
929:
926:
925:
920:
914:
908:
905:
901:
898:
896:
893:
891:
890:
886:
885:
879:
875:
872:
868:
865:
863:
860:
858:
857:
856:Araripesuchus
853:
852:
848:
845:
842:
839:
836:
835:
832:
831:
826:
820:
811:
807:
804:
800:
797:
795:
792:
790:
789:
788:Santanaraptor
785:
784:
778:
774:
771:
768:
765:
763:
760:
758:
757:
753:
752:
746:
742:
739:
738:Spinosauridae
736:
733:
731:
728:
726:
725:
721:
720:
714:
710:
707:
703:
700:
698:
695:
693:
692:
688:
687:
681:
677:
674:
670:
669:Spinosauridae
667:
664:
662:
659:
657:
656:
652:
651:
647:
644:
641:
638:
635:
634:
631:
630:
625:
619:
617:
615:
611:
607:
603:
599:
591:
586:
584:
580:
578:
574:
570:
565:
560:
554:
545:
537:
530:
528:
526:
522:
518:
514:
510:
505:
502:
497:
495:
491:
488:
484:
480:
476:
472:
471:Spinosauridae
468:
464:
463:invertebrates
460:
456:
452:
451:predator–prey
448:
447:fossil fishes
444:
439:
437:
433:
430:
426:
423:stage of the
422:
418:
415:in 1819. The
414:
410:
406:
402:
401:
396:
392:
388:
384:
383:Araripe Basin
380:
376:
372:
360:
355:
352:
351:Araripe Basin
349:
345:
342:
331:
327:
324:
320:
316:
313:
309:
304:
295:12.2°S 10.7°W
276:
274:
270:
265:
237:
235:
231:
226:
223:
219:
216:
212:
209:
206:
202:
197:
193:
189:
186:
182:
179:
175:
172:
168:
165:
161:
158:
157:Santana Group
155:
151:
148:
145:
141:
136:
135:
130:
129:
122:
117:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
43:
40:111–108
35:
31:
27:
22:
19:
2830:. Retrieved
2823:
2770:
2764:
2728:
2722:
2707:
2682:
2678:
2666:. Retrieved
2646:
2640:
2624:. Retrieved
2605:
2585:
2573:. Retrieved
2553:
2547:
2524:. Retrieved
2504:
2498:
2448:
2444:
2428:. Retrieved
2414:
2410:
2399:Bibliography
2380:
2347:
2343:
2333:
2319:
2314:
2306:
2301:
2293:
2288:
2272:
2234:
2225:11449/175030
2199:
2195:
2188:
2143:
2129:
2124:
2108:
2095:
2090:
2081:
2072:
2031:
2027:
2014:
1969:
1965:
1954:
1945:
1902:
1898:
1891:
1864:
1812:(7490): 20.
1809:
1805:
1795:
1786:
1735:
1716:
1709:
1702:
1695:
1688:
1681:
1674:
1667:
1660:
1653:
1646:
1639:
1632:
1625:
1618:
1611:
1604:
1597:
1589:
1582:
1566:
1557:
1500:
1475:
1468:
1455:
1448:
1428:
1421:
1401:
1394:
1374:
1367:
1343:
1323:
1316:
1296:
1292:T. leonardii
1291:
1286:
1277:
1259:
1252:
1234:
1227:
1209:
1202:
1182:
1177:
1171:
1162:
1144:
1137:
1119:
1112:
1103:
1085:
1078:
1067:
1060:
1049:
1042:
1032:A. castilhoi
1031:
1024:
1013:
1006:
995:
981:
976:
971:
966:
961:
954:
945:
922:
894:
889:Caririsuchus
887:
861:
854:
828:
793:
786:
761:
754:
729:
722:
696:
689:
672:
660:
653:
627:
614:megaraptoran
595:
581:
556:
520:
506:
498:
440:
398:
370:
368:
300:-12.2; -10.7
256:7.2°S 39.3°W
132:
126:
34:Early Albian
18:
2832:21 December
2389:Fossilworks
2326:Fossilworks
2281:Fossilworks
2243:Fossilworks
2181:Fossilworks
2152:Fossilworks
2136:Fossilworks
2117:Fossilworks
2101:Fossilworks
1928:11336/93821
1905:: 107–119.
1620:Cladocyclus
1567:crossed out
1546:Morphotaxon
1456:E. essweini
1429:A. barretoi
1402:C. placidoi
1396:Cearachelys
1375:S. gaffneyi
1324:U. trigonus
1145:M. kellneri
1114:Kariridraco
1104:C. ligabuei
977:A. piscator
972:A. robustus
962:A. santanae
904:peirosaurid
801:A possible
794:S. placidus
766:Pernambuco
706:coelurosaur
691:Aratasaurus
298: /
261:-7.2; -39.3
259: /
234:Coordinates
125:Fossils of
2845:Categories
2595:0866225498
2451:(2): 259.
2034:(1): 5–8.
1778:References
1683:Rhacolepis
1662:Obaichthys
1606:Brannerion
1563:small text
1532:Ichnotaxon
1470:Brasilemys
1423:Araripemys
1044:Barbosania
997:Anhanguera
947:Anhanguera
924:Pterosaurs
915:Pterosaurs
895:C. camposi
862:A. gomesii
655:Angaturama
592:Archosaurs
577:Cenomanian
509:weathering
490:accretions
455:pterosaurs
432:rift basin
429:lacustrine
387:Pernambuco
315:Pernambuco
131:(top) and
128:Anhanguera
2814:0001-3765
2668:6 October
2626:6 October
2575:6 October
2526:5 October
2475:1664-2384
2430:6 October
2372:0195-6671
2064:131339386
2056:0016-7649
1937:0195-6671
1836:0028-0836
1718:Vinctifer
1704:Tharrhias
1501:Color key
1450:Euraxemys
1279:Tupuxuara
1178:S. pricei
1120:K. dianae
756:Mirischia
724:Irritator
629:Dinosaurs
620:Dinosaurs
616:fossils.
598:theropods
487:limestone
483:taphonomy
218:Limestone
199:Lithology
191:Thickness
163:Underlies
2807:24789213
2789:24789213
2539:(2005).
2521:29947671
2483:89178816
2417:: 5–14.
2385:Ze Gomes
2006:32651406
1844:24598620
1726:See also
1655:Notelops
1476:B. josai
1356:Presence
1318:Unwindia
1235:T. sethi
1204:Tapejara
1183:S. spixi
1086:C. atrox
1014:A. dehmi
937:Presence
843:Presence
661:A. limai
642:Presence
602:avialans
459:reptiles
228:Location
208:Mudstone
177:Overlies
2733:Bibcode
2651:Bibcode
2558:Bibcode
2453:Bibcode
2352:Bibcode
2204:Bibcode
2036:Bibcode
1997:7351750
1974:Bibcode
1907:Bibcode
1814:Bibcode
1539:Ootaxon
1362:Images
1353:Species
1344:Turtles
1336:Turtles
940:Images
934:Species
849:Images
840:Species
648:Images
639:Species
329:Country
286:10°42′W
283:12°12′S
247:39°18′W
204:Primary
153:Unit of
109:↓
2812:
2805:
2787:
2714:
2689:
2617:
2592:
2519:
2481:
2473:
2370:
2062:
2054:
2004:
1994:
1935:
1842:
1834:
1806:Nature
798:Ceará
734:Ceará
701:Ceará
665:Ceará
573:Albian
569:Aptian
564:strata
465:, and
443:fossil
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