Knowledge (XXG)

Plecoptera

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Nymphs can acquire oxygen via diffusing through the exoskeleton, or through gills located on behind the head, on the thorax, or around the anus. Due to their nymph's requirement for well oxygenated water, the species is very sensitive to water pollution. This makes them important indicators for water quality. Most species are herbivorous as nymphs, feeding on submerged leaves and benthic algae, but many are hunters of other aquatic arthropods.
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The female can lay up to one thousand eggs. It will fly over the water and drop the eggs in the water. It also may hang on a rock or branch. Eggs are covered in a sticky coating which allows them to adhere to rocks without being swept away by swift currents. The eggs typically take two to three weeks
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of well-oxygenated lakes and streams. A few species found in New Zealand and nearby islands have terrestrial nymphs, but even these inhabit only very moist environments. The nymphs physically resemble wingless adults, but often have external gills, which may be present on almost any part of the body.
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The adults generally only survive for a few weeks, and emerge only during specific times of the year when resources are optimal. Some do not feed at all, but those that do are herbivorous. Adults are not strong fliers and generally stay near the stream or lake they hatched from.
293:. Some 3,500 species are described worldwide, with new species still being discovered. Stoneflies are found worldwide, except Antarctica. Stoneflies are believed to be one of the most primitive groups of 797:
Romolo Fochetti & José Manuel Tierno de Figueroa (2008) . "Global diversity of stoneflies (Plecoptera; Insecta) in freshwater". In E. V. Balian; C. Lévêque; H. Segers & K. Martens (eds.).
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In addition, not adopting the clades Antarctoperlaria and Holognatha allows for a systematic layout of the Plecoptera that adequately reproduces phylogeny, while retaining the traditional
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superfamilies of stoneflies, which do not seem to be each other's closest relatives. Thus, the "Antarctoperlaria" are not considered a natural group (despite some claims to the contrary).
556:"Holognatha" together with the Euholognatha (meaning roughly "advanced Holognatha"), but the Scopuridae do not appear significantly closer to the Euholognatha than to the Systellognatha. 2780: 2819: 1314: 552:
must be considered a basal family in the Arctoperlaria, not assignable to any of the infraorders. Alternatively, the Scopuridae were placed in an unranked
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before emerging and becoming terrestrial as adults. Before becoming adults, nymphs will leave the water, attach to a fixed surface and molt one last time.
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geological periods, while true stoneflies are known from fossils only a bit younger. Their modern diversity, however, apparently is of
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and fold flat over their backs. Stoneflies are generally not strong fliers, and some species are entirely wingless.
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Stoneflies have a generalized anatomy, with few specialized features compared to other insects. They have simple
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The insects remain in the nymphal form for one to four years, depending on species, and undergo from 12 to 36
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Ding, Shuangmei; Li, Weihai; Wang, Ying; Cameron, Stephen L.; Murányi, Dávid; Yang, Ding (June 2019).
2883: 2834: 2702: 2640: 2137: 2117: 1201: 389: 2405: 2210: 626: 377: 2906: 2382: 2291: 606: 439: 317: 313: 2225: 471:(Nepomorpha) may also be fully aquatic for their entire lives, but can leave the water to travel. 457: 2235: 2157: 2132: 1233: 363: 249: 147: 2460: 656: 549: 2152: 2842: 2679: 2185: 1225: 1217: 957: 898: 871: 818: 566: 526: 393: 385: 31: 2847: 2785: 2507: 2367: 2127: 1978: 1664: 1209: 863: 810: 802: 695: 571: 2631: 1034: 2857: 2418: 2205: 2122: 2112: 2089: 1656: 638: 451: 409: 408:
is relatively soft, and may include remnants of the nymphal gills even in the adult. Both
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in a stream or still water is usually an indicator of good or excellent water quality.
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Peter Zwick (2000). "Phylogenetic system and zoogeography of the Plecoptera".
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on a number of occasions before once again becoming geographically isolated.
2715: 2478: 2346: 2334: 2142: 1986: 1946: 1921: 1774: 1685: 1644: 1631: 1462: 1449: 1410: 1344: 668: 545: 463: 434:(πτέρυξ, "wing"). This refers to the complex venation of their two pairs of 321: 214: 184: 164: 138: 108: 52: 1229: 875: 1263: 129: 17: 2666: 2610: 2536: 2352: 2190: 2099: 1888: 1758: 1550: 1356: 717: 587: 518: 495: 306: 294: 224: 98: 93: 78: 73: 63: 2259: 1529: 502: 405: 325: 302: 119: 113: 88: 2772: 2340: 2328: 1866: 1766: 1693: 1599: 1508: 1500: 1338: 1323: 920:"Clean water has bugs in it, says BYU Biology Professor Riley Nelson" 814: 769: 413: 401: 286: 194: 174: 2671: 2587: 2759: 2720: 1136: 2865: 581: 565: 553: 544:(also called Setipalpia or Subulipalpia). This corresponds to the 420: 367: 357: 346: 801:. Developments in Hydrobiology. Vol. 198. pp. 365–377. 1954: 1292: 2591: 2439: 2365: 2287: 2074: 1369: 1296: 474:
The nymphs (technically, "naiads") are aquatic and live in the
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English-Greek Dictionary - a Vocabulary of the Attic Language
461:, are the only known insects, perhaps with the exception of 956:(2nd ed.). Oxford University Press. pp. 383–386. 404:. The legs are robust, with each ending in two claws. The 467:, that are exclusively aquatic from birth to death. Some 320:, and the populations are quite distinct, although the 1907:(net-winged insects: lacewings, mantidflies, antlions) 532:
The Arctoperlaria, though, have been divided into two
525:. However, the former simply consists of the two most 521:, the "Antarctoperlaria" (or "Archiperlaria") and the 952:
Hoell, H.V.; Doyen, J.T. & Purcell, A.H. (1998).
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Traditionally, the stoneflies were divided into two
2600: 2519: 2469: 2453: 2404: 2381: 1977: 1935: 1920: 1879: 1848: 1839: 1790: 1744: 1713: 1676: 1643: 1576: 1549: 1520: 1499: 1490: 1461: 1418: 1384: 777:and this species is suspected to belong elsewhere. 748:(about 12 species) – salmonflies, giant stoneflies 324:evidence suggests species may have crossed the 1137:"Order Plecoptera - Stoneflies - BugGuide.Net" 2299: 1308: 671:(about 300 species) – small winter stoneflies 8: 954:Introduction to Insect Biology and Diversity 331:All species of Plecoptera are intolerant of 416:projecting from the tip of their abdomens. 297:, with close relatives identified from the 2588: 2450: 2436: 2378: 2362: 2306: 2292: 2284: 2082: 2071: 1932: 1845: 1818: 1741: 1737: 1584: 1573: 1569: 1496: 1487: 1483: 1442: 1381: 1377: 1366: 1315: 1301: 1293: 1086:"Plecoptera - Stoneflies -- Discover Life" 354:of a golden stonefly, Plecoptera, Perlidae 128: 41: 2497:(spring stoneflies or tiny winter blacks) 734:(about 68 species) – roachlike stoneflies 678:(390+ species) – rolled-winged stoneflies 2036:Four most speciose orders are marked in 1626:(crickets, wetas, grasshoppers, locusts) 1048:United States Department of Agriculture 891:Adults and Nymphs of British Stoneflies 847: 845: 843: 789: 760: 720:(about 400 species) – common stoneflies 698:(about 110 species) – winter stoneflies 419:The name "Plecoptera" literally means " 1033:. In D. D. Murhy; C. M. Knopp (eds.). 983:. London: George Routledge & Sons. 947: 945: 943: 941: 799:Freshwater Animal Diversity Assessment 685:(over 700 species) – spring stoneflies 1194:Molecular Phylogenetics and Evolution 1107: 1105: 713:(over 180 species) – green stoneflies 7: 2871:DA2287BC-FFAC-FFBC-FF54-AA82FCF67D5F 2835:2ffd4e0a-5167-421c-9beb-709b7488bd14 2747:65e60a58-541f-416c-9dfd-8ffe9294b6ed 1899:(alderflies, dobsonflies, fishflies) 445:A few wingless species, such as the 1160:C. Riley Nelson (January 1, 1996). 994:C. Riley Nelson (January 1, 1996). 494:to hatch, but some species undergo 2510:(willowflies or winter stoneflies) 1288:order Plecoptera (Version 5.0/5.0) 25: 2568:(giant stoneflies or salmonflies) 895:Freshwater Biological Association 312:Plecoptera are found in both the 2922:Taxa named by Hermann Burmeister 2917:Extant Permian first appearances 2258: 1409: 1274: 1262: 269: 151: 1036:Lake Tahoe Watershed Assessment 2692:Cockroach Species File (new): 1024:"Lake Tahoe benthic stonefly ( 996:"Capniidae. Winter Stoneflies" 1: 1829:(sawflies, wasps, ants, bees) 868:10.1146/annurev.ento.45.1.709 412:and adults have long, paired 2547:(stripetails or springflies) 1777:(cicadas, aphids, true bugs) 807:10.1007/978-1-4020-8259-7_39 335:, and the presence of their 30:For the genus of moths, see 1214:10.1016/j.ympev.2019.03.005 855:Annual Review of Entomology 447:Lake Tahoe benthic stonefly 392:, long, multiple-segmented 2953: 2489:(rolled-winged stoneflies) 1967:(gnats, mosquitoes, flies) 1860:(twisted-winged parasites) 1532:(dragonflies, damselflies) 481: 430:(πλέκειν, "to braid") and 29: 2855:Plecoptera Species File: 2481:(small winter stoneflies) 2449: 2435: 2377: 2361: 2323: 2252: 2085: 2081: 2070: 2032: 1821: 1740: 1587: 1572: 1486: 1445: 1407: 1380: 1376: 1365: 1333: 1286:Plecoptera Species File: 1050:. pp. O–118 – O–120. 262: 257: 148:Scientific classification 146: 136: 127: 44: 1618:(stick and leaf insects) 1167:Tree of Life Web Project 1162:"Plecoptera. Stoneflies" 1001:Tree of Life Web Project 977:S. C. Woodhouse (1910). 548:with one exception: the 540:(or Filipalpia) and the 2045:are paraphyletic groups 1688:(cockroaches, termites) 1452:(silverfish, firebrats) 889:H. B. N. Hynes (1993). 343:Description and ecology 2555:(roachlike stoneflies) 1396:(jumping bristletails) 595: 579: 381: 355: 601:("Antarctoperlaria") 585: 569: 482:Further information: 361: 350: 2742:Fauna Europaea (new) 1998:(moths, butterflies) 1271:at Wikimedia Commons 1090:www.discoverlife.org 1022:E. M. Holst (2000). 318:Northern Hemispheres 289:, commonly known as 2927:Symbols of Delaware 2539:(common stoneflies) 1206:2019MolPE.135..123D 926:on 6 September 2015 400:, and two or three 27:Order of stoneflies 2531:(green stoneflies) 2265:Insects portal 2236:Triadophlebioptera 741:(about 10 species) 596: 580: 423:-wings", from the 382: 356: 2894: 2893: 2843:Open Tree of Life 2594:Taxon identifiers 2585: 2584: 2581: 2580: 2577: 2576: 2569: 2556: 2548: 2540: 2532: 2511: 2508:Taeniopterygidae 2498: 2490: 2482: 2431: 2430: 2427: 2426: 2368:Antarctoperlaria 2281: 2280: 2248: 2247: 2244: 2243: 2186:Palaeodictyoptera 2171: 2103: 2066: 2065: 2062: 2061: 2028: 2027: 2024: 2023: 2020: 2019: 2016: 2015: 2012: 2011: 2008: 2007: 1999: 1990: 1968: 1958: 1950: 1928: 1916: 1915: 1908: 1900: 1892: 1870: 1861: 1830: 1786: 1785: 1778: 1770: 1762: 1709: 1708: 1705: 1704: 1697: 1689: 1668: 1665:Mantophasmatodea 1660: 1649: 1635: 1627: 1619: 1611: 1603: 1595: 1545: 1544: 1541: 1540: 1533: 1512: 1501:Ephemeropteroidea 1453: 1405: 1404: 1397: 1267:Media related to 1117:www.cals.ncsu.edu 1066:www.cals.ncsu.edu 963:978-0-19-510033-4 904:978-0-900386-28-2 824:978-1-4020-8258-0 452:"Capnia" lacustra 276: 275: 253: 16:(Redirected from 2944: 2887: 2886: 2874: 2873: 2861: 2860: 2851: 2850: 2838: 2837: 2828: 2827: 2815: 2814: 2812:NHMSYS0000841058 2802: 2801: 2789: 2788: 2776: 2775: 2763: 2762: 2750: 2749: 2737: 2736: 2724: 2723: 2711: 2710: 2698: 2697: 2688: 2687: 2675: 2674: 2662: 2661: 2649: 2648: 2636: 2635: 2634: 2621: 2620: 2619: 2589: 2567: 2554: 2546: 2538: 2530: 2509: 2496: 2488: 2480: 2451: 2437: 2379: 2371: 2370:(basal lineages) 2363: 2308: 2301: 2294: 2285: 2263: 2262: 2196:Permoplectoptera 2169: 2128:Diaphanopterodea 2101: 2083: 2072: 1997: 1988: 1979:Amphiesmenoptera 1966: 1960: 1956: 1948: 1938: 1933: 1927: 1924: 1906: 1898: 1890: 1868: 1859: 1846: 1842: 1828: 1819: 1776: 1768: 1761:(barklice, lice) 1760: 1752: 1742: 1738: 1695: 1687: 1666: 1658: 1657:Grylloblattodea 1647: 1633: 1625: 1617: 1609: 1601: 1593: 1585: 1581: 1574: 1570: 1531: 1510: 1497: 1488: 1484: 1451: 1443: 1413: 1395: 1382: 1378: 1367: 1317: 1310: 1303: 1294: 1279:Data related to 1278: 1266: 1251: 1248: 1242: 1241: 1185: 1179: 1178: 1176: 1174: 1157: 1151: 1150: 1148: 1147: 1133: 1127: 1126: 1124: 1123: 1109: 1100: 1099: 1097: 1096: 1082: 1076: 1075: 1073: 1072: 1058: 1052: 1051: 1045: 1032: 1019: 1013: 1012: 1010: 1008: 991: 985: 984: 974: 968: 967: 949: 936: 935: 933: 931: 922:. Archived from 915: 909: 908: 886: 880: 879: 849: 838: 835: 829: 828: 794: 778: 765: 696:Taeniopterygidae 586:Adult of family 572:Taeniopterygidae 570:Adult of family 248: 156: 155: 132: 118: 55: 48:Temporal range: 42: 21: 2952: 2951: 2947: 2946: 2945: 2943: 2942: 2941: 2932:Aquatic insects 2897: 2896: 2895: 2890: 2882: 2877: 2869: 2864: 2856: 2854: 2846: 2841: 2833: 2831: 2823: 2818: 2810: 2805: 2797: 2792: 2784: 2779: 2771: 2766: 2758: 2753: 2745: 2740: 2732: 2727: 2719: 2714: 2706: 2701: 2693: 2691: 2683: 2678: 2670: 2665: 2657: 2652: 2644: 2639: 2630: 2629: 2624: 2615: 2614: 2609: 2596: 2586: 2573: 2566:Pteronarcyidae 2529:Chloroperlidae 2515: 2465: 2445: 2423: 2419:Gripopterygidae 2400: 2373: 2369: 2357: 2319: 2312: 2282: 2277: 2257: 2240: 2206:Protelytroptera 2138:Eudiaphanoptera 2123:Coxoplectoptera 2118:Carbotriplurida 2113:Campylopteridae 2090:Aethiocarenodea 2077: 2058: 2004: 1973: 1949:(scorpionflies) 1936: 1925: 1923: 1912: 1875: 1840: 1835: 1813: 1811: 1809: 1807: 1805: 1803: 1801: 1799: 1797: 1795: 1793: 1782: 1732: 1730: 1728: 1726: 1724: 1722: 1720: 1718: 1716: 1701: 1672: 1646: 1639: 1634:(angel insects) 1577: 1564: 1562: 1560: 1558: 1556: 1554: 1552: 1537: 1516: 1478: 1476: 1474: 1472: 1470: 1468: 1466: 1464: 1457: 1437: 1435: 1433: 1431: 1429: 1427: 1425: 1423: 1421: 1414: 1401: 1372: 1361: 1329: 1321: 1259: 1254: 1249: 1245: 1187: 1186: 1182: 1172: 1170: 1159: 1158: 1154: 1145: 1143: 1135: 1134: 1130: 1121: 1119: 1111: 1110: 1103: 1094: 1092: 1084: 1083: 1079: 1070: 1068: 1060: 1059: 1055: 1039: 1030: 1026:Capnia lacustra 1021: 1020: 1016: 1006: 1004: 993: 992: 988: 976: 975: 971: 964: 951: 950: 939: 929: 927: 918:Nelson, Riley. 917: 916: 912: 905: 888: 887: 883: 851: 850: 841: 836: 832: 825: 796: 795: 791: 787: 782: 781: 766: 762: 757: 639:Gripopterygidae 515: 491: 486: 469:true water bugs 374:Gripopterygidae 371: 345: 333:water pollution 247: 150: 123: 117: 116: 111: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 50: 49: 46: 38: 28: 23: 22: 15: 12: 11: 5: 2950: 2948: 2940: 2939: 2934: 2929: 2924: 2919: 2914: 2909: 2899: 2898: 2892: 2891: 2889: 2888: 2875: 2862: 2852: 2839: 2829: 2816: 2803: 2790: 2777: 2764: 2751: 2738: 2729:Fauna Europaea 2725: 2712: 2699: 2689: 2676: 2663: 2650: 2637: 2622: 2606: 2604: 2598: 2597: 2592: 2583: 2582: 2579: 2578: 2575: 2574: 2572: 2571: 2563: 2558: 2553:Peltoperlidae 2550: 2542: 2534: 2525: 2523: 2521:Systellognatha 2517: 2516: 2514: 2513: 2505: 2503:Notonemouridae 2500: 2492: 2484: 2475: 2473: 2467: 2466: 2464: 2463: 2457: 2455: 2447: 2446: 2440: 2433: 2432: 2429: 2428: 2425: 2424: 2422: 2421: 2416: 2414:Austroperlidae 2410: 2408: 2406:Leptoperloidea 2402: 2401: 2399: 2398: 2393: 2391:Diamphipnoidae 2387: 2385: 2375: 2374: 2366: 2359: 2358: 2356: 2355: 2349: 2343: 2337: 2331: 2324: 2321: 2320: 2313: 2311: 2310: 2303: 2296: 2288: 2279: 2278: 2276: 2275: 2268: 2253: 2250: 2249: 2246: 2245: 2242: 2241: 2239: 2238: 2233: 2228: 2223: 2221:Protozygoptera 2218: 2216:Protorthoptera 2213: 2211:Protephemerida 2208: 2203: 2201:Protanisoptera 2198: 2193: 2188: 2183: 2178: 2173: 2170:(griffinflies) 2168:Meganisoptera 2165: 2160: 2155: 2150: 2148:Glosselytrodea 2145: 2140: 2135: 2130: 2125: 2120: 2115: 2110: 2105: 2097: 2092: 2086: 2079: 2078: 2075: 2068: 2067: 2064: 2063: 2060: 2059: 2057: 2056: 2046: 2040: 2033: 2030: 2029: 2026: 2025: 2022: 2021: 2018: 2017: 2014: 2013: 2010: 2009: 2006: 2005: 2003: 2002: 1992: 1983: 1981: 1975: 1974: 1972: 1971: 1961: 1941: 1939: 1930: 1918: 1917: 1914: 1913: 1911: 1910: 1902: 1894: 1889:Raphidioptera 1885: 1883: 1877: 1876: 1874: 1873: 1863: 1854: 1852: 1843: 1841:Neuropteroidea 1837: 1836: 1834: 1833: 1822: 1816: 1788: 1787: 1784: 1783: 1781: 1780: 1772: 1764: 1755: 1753: 1735: 1711: 1710: 1707: 1706: 1703: 1702: 1700: 1699: 1691: 1682: 1680: 1674: 1673: 1671: 1670: 1662: 1659:(ice-crawlers) 1653: 1651: 1641: 1640: 1638: 1637: 1629: 1621: 1613: 1605: 1597: 1588: 1582: 1567: 1547: 1546: 1543: 1542: 1539: 1538: 1536: 1535: 1526: 1524: 1518: 1517: 1515: 1514: 1509:Ephemeroptera 1505: 1503: 1494: 1481: 1459: 1458: 1456: 1455: 1446: 1440: 1416: 1415: 1408: 1406: 1403: 1402: 1400: 1399: 1394:Archaeognatha 1390: 1388: 1374: 1373: 1370: 1363: 1362: 1360: 1359: 1353: 1347: 1341: 1334: 1331: 1330: 1322: 1320: 1319: 1312: 1305: 1297: 1291: 1290: 1284: 1283:at Wikispecies 1272: 1258: 1257:External links 1255: 1253: 1252: 1250:Nelson (1996b) 1243: 1180: 1152: 1128: 1101: 1077: 1053: 1014: 986: 969: 962: 937: 910: 903: 881: 839: 837:Brittain, 1987 830: 823: 788: 786: 783: 780: 779: 759: 758: 756: 753: 752: 751: 750: 749: 746:Pteronarcyidae 742: 735: 728: 727:(350+ species) 721: 714: 711:Chloroperlidae 705:Systellognatha 701: 700: 699: 692: 690:Notonemouridae 686: 679: 672: 659: 644: 643: 642: 641: 635: 633:Austroperlidae 627:Leptoperloidea 623: 622: 621: 615: 613:Diamphipnoidae 599:Basal lineages 592:Systellognatha 542:Systellognatha 514: 511: 490: 487: 378:Dinotoperlinae 344: 341: 274: 273: 260: 259: 255: 254: 242: 238: 237: 232: 228: 227: 222: 218: 217: 212: 208: 207: 202: 198: 197: 192: 188: 187: 182: 178: 177: 172: 168: 167: 162: 158: 157: 144: 143: 134: 133: 125: 124: 112: 107: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 56: 47: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 2949: 2938: 2935: 2933: 2930: 2928: 2925: 2923: 2920: 2918: 2915: 2913: 2912:Insect orders 2910: 2908: 2905: 2904: 2902: 2885: 2880: 2876: 2872: 2867: 2863: 2859: 2853: 2849: 2844: 2840: 2836: 2830: 2826: 2821: 2817: 2813: 2808: 2804: 2800: 2795: 2791: 2787: 2782: 2778: 2774: 2769: 2765: 2761: 2756: 2752: 2748: 2743: 2739: 2735: 2730: 2726: 2722: 2717: 2713: 2709: 2704: 2700: 2696: 2690: 2686: 2681: 2677: 2673: 2668: 2664: 2660: 2655: 2651: 2647: 2642: 2638: 2633: 2627: 2623: 2618: 2612: 2608: 2607: 2605: 2603: 2599: 2595: 2590: 2570: 2564: 2562: 2561:Styloperlidae 2559: 2557: 2551: 2549: 2543: 2541: 2535: 2533: 2527: 2526: 2524: 2522: 2518: 2512: 2506: 2504: 2501: 2499: 2493: 2491: 2485: 2483: 2477: 2476: 2474: 2472: 2468: 2462: 2459: 2458: 2456: 2452: 2448: 2444: 2443:Arctoperlaria 2438: 2434: 2420: 2417: 2415: 2412: 2411: 2409: 2407: 2403: 2397: 2394: 2392: 2389: 2388: 2386: 2384: 2383:Eusthenioidea 2380: 2376: 2372: 2364: 2360: 2354: 2350: 2348: 2344: 2342: 2338: 2336: 2332: 2330: 2326: 2325: 2322: 2317: 2309: 2304: 2302: 2297: 2295: 2290: 2289: 2286: 2274: 2273: 2269: 2267: 2266: 2261: 2255: 2254: 2251: 2237: 2234: 2232: 2229: 2227: 2224: 2222: 2219: 2217: 2214: 2212: 2209: 2207: 2204: 2202: 2199: 2197: 2194: 2192: 2189: 2187: 2184: 2182: 2179: 2177: 2176:Megasecoptera 2174: 2172: 2166: 2164: 2161: 2159: 2156: 2154: 2151: 2149: 2146: 2144: 2141: 2139: 2136: 2134: 2131: 2129: 2126: 2124: 2121: 2119: 2116: 2114: 2111: 2109: 2106: 2104: 2098: 2096: 2093: 2091: 2088: 2087: 2084: 2080: 2073: 2069: 2055: 2053: 2047: 2044: 2041: 2039: 2035: 2034: 2031: 2001: 2000: 1993: 1991: 1989:(caddisflies) 1985: 1984: 1982: 1980: 1976: 1970: 1969: 1962: 1959: 1955:Siphonaptera 1952: 1951: 1943: 1942: 1940: 1934: 1931: 1929: 1926:(Mecopterida) 1919: 1909: 1903: 1901: 1895: 1893: 1887: 1886: 1884: 1882: 1878: 1872: 1871: 1864: 1862: 1858:Strepsiptera 1856: 1855: 1853: 1851: 1847: 1844: 1838: 1832: 1831: 1824: 1823: 1820: 1817: 1815: 1789: 1779: 1773: 1771: 1767:Thysanoptera 1765: 1763: 1757: 1756: 1754: 1749: 1748: 1743: 1739: 1736: 1734: 1712: 1698: 1692: 1690: 1684: 1683: 1681: 1679: 1675: 1669: 1663: 1661: 1655: 1654: 1652: 1650: 1642: 1636: 1630: 1628: 1622: 1620: 1614: 1612: 1610:(webspinners) 1606: 1604: 1598: 1596: 1590: 1589: 1586: 1583: 1580: 1575: 1571: 1568: 1566: 1548: 1534: 1528: 1527: 1525: 1523: 1519: 1513: 1507: 1506: 1504: 1502: 1498: 1495: 1493: 1489: 1485: 1482: 1480: 1460: 1454: 1448: 1447: 1444: 1441: 1439: 1417: 1412: 1398: 1392: 1391: 1389: 1387: 1383: 1379: 1375: 1368: 1364: 1358: 1354: 1352: 1348: 1346: 1342: 1340: 1336: 1335: 1332: 1328: 1325: 1318: 1313: 1311: 1306: 1304: 1299: 1298: 1295: 1289: 1285: 1282: 1277: 1273: 1270: 1265: 1261: 1260: 1256: 1247: 1244: 1239: 1235: 1231: 1227: 1223: 1219: 1215: 1211: 1207: 1203: 1199: 1195: 1191: 1184: 1181: 1169: 1168: 1163: 1156: 1153: 1142: 1138: 1132: 1129: 1118: 1114: 1108: 1106: 1102: 1091: 1087: 1081: 1078: 1067: 1063: 1057: 1054: 1049: 1043: 1038: 1037: 1029: 1027: 1018: 1015: 1003: 1002: 997: 990: 987: 982: 981: 973: 970: 965: 959: 955: 948: 946: 944: 942: 938: 925: 921: 914: 911: 906: 900: 896: 892: 885: 882: 877: 873: 869: 865: 861: 857: 856: 848: 846: 844: 840: 834: 831: 826: 820: 816: 812: 808: 804: 800: 793: 790: 784: 776: 772: 771: 764: 761: 754: 747: 743: 740: 739:Styloperlidae 736: 733: 732:Peltoperlidae 729: 726: 722: 719: 715: 712: 708: 707: 706: 702: 697: 693: 691: 687: 684: 680: 677: 673: 670: 666: 665: 664: 660: 658: 655:Basal family 654: 653: 652: 651: 650: 649:Arctoperlaria 640: 636: 634: 630: 629: 628: 624: 620: 616: 614: 610: 609: 608: 607:Eusthenioidea 604: 603: 602: 600: 593: 589: 584: 577: 573: 568: 564: 562: 557: 555: 551: 547: 543: 539: 535: 530: 528: 524: 523:Arctoperlaria 520: 512: 510: 506: 504: 499: 497: 488: 485: 480: 477: 472: 470: 466: 465: 460: 459: 454: 453: 448: 443: 441: 437: 433: 429: 426: 425:Ancient Greek 422: 417: 415: 411: 407: 403: 399: 398:compound eyes 395: 391: 388:with chewing 387: 379: 375: 369: 366: 365: 360: 353: 349: 342: 340: 338: 334: 329: 327: 323: 319: 315: 310: 308: 304: 300: 299:Carboniferous 296: 292: 288: 284: 280: 272: 271: 267: 266:Arctoperlaria 261: 256: 251: 246: 243: 240: 239: 236: 233: 230: 229: 226: 223: 220: 219: 216: 213: 210: 209: 206: 203: 200: 199: 196: 193: 190: 189: 186: 183: 180: 179: 176: 173: 170: 169: 166: 163: 160: 159: 154: 149: 145: 141: 140: 135: 131: 126: 121: 115: 110: 105: 100: 95: 90: 85: 80: 75: 70: 65: 60: 54: 43: 40: 36: 34: 19: 2937:Polyneoptera 2601: 2471:Euholognatha 2396:Eustheniidae 2351:Infraclass: 2315: 2270: 2256: 2226:Syntonoptera 2108:Caloneurodea 2100:Blattoptera 2051: 2042: 2037: 1996:Lepidoptera 1994: 1987:Trichoptera 1963: 1945: 1897:Megaloptera 1891:(snakeflies) 1881:Neuropterida 1865: 1850:Coleopterida 1827:Hymenoptera 1825: 1747:Paraneoptera 1745: 1667:(gladiators) 1616:Phasmatodea 1594:(stoneflies) 1591: 1579:Polyneoptera 1522:Odonatoptera 1386:Monocondylia 1351:Pancrustacea 1349:(unranked): 1246: 1197: 1193: 1183: 1171:. Retrieved 1165: 1155: 1144:. Retrieved 1141:bugguide.net 1140: 1131: 1120:. Retrieved 1116: 1093:. Retrieved 1089: 1080: 1069:. Retrieved 1065: 1056: 1035: 1025: 1017: 1005:. Retrieved 999: 989: 979: 972: 953: 928:. Retrieved 924:the original 913: 890: 884: 859: 853: 833: 798: 792: 775:monophyletic 768: 763: 663:Euholognatha 646: 645: 625:Superfamily 619:Eustheniidae 605:Superfamily 598: 597: 576:Euholognatha 558: 538:Euholognatha 531: 516: 507: 500: 492: 476:benthic zone 473: 462: 458:Baikaloperla 456: 450: 444: 438:, which are 431: 427: 418: 383: 362: 330: 322:evolutionary 311: 290: 278: 277: 263: 244: 235:Polyneoptera 221:Infraclass: 201:(unranked): 137: 39: 32: 2768:iNaturalist 2626:Wikispecies 2545:Perlodidae 2495:Nemouridae 2487:Leuctridae 2272:Wikispecies 2231:Titanoptera 2163:Lapeyriidae 2158:Hypoperlida 2133:Eoblattodea 2102:(roachoids) 2095:Archodonata 1937:Antliophora 1905:Neuroptera 1867:Coleoptera 1678:Dictyoptera 1648:(Xenonomia) 1624:Orthoptera 1608:Embioptera 1600:Dermaptera 1592:Plecoptera 1492:Palaeoptera 1355:Subphylum: 1200:: 123–135. 862:: 709–746. 703:Infraorder 661:Infraorder 561:ranked taxa 534:infraorders 513:Systematics 484:Mesoleuctra 364:Dinotoperla 51:299–0  2907:Plecoptera 2901:Categories 2646:Plecoptera 2632:Plecoptera 2602:Plecoptera 2479:Capniidae 2461:Scopuridae 2345:Subclass: 2335:Arthropoda 2316:Plecoptera 2181:Miomoptera 1947:Mecoptera 1775:Hemiptera 1696:(mantises) 1686:Blattodea 1632:Zoraptera 1511:(mayflies) 1450:Zygentoma 1345:Arthropoda 1281:Plecoptera 1269:Plecoptera 1146:2016-04-12 1122:2016-04-12 1095:2016-02-23 1071:2016-02-23 785:References 767:The genus 725:Perlodidae 683:Nemouridae 676:Leuctridae 657:Scopuridae 550:Scopuridae 489:Life cycle 440:membranous 386:mouthparts 301:and Lower 291:stoneflies 279:Plecoptera 258:Suborders 250:Burmeister 245:Plecoptera 211:Subclass: 205:Dicondylia 185:Arthropoda 45:Plecoptera 33:Plecoptera 18:Stoneflies 2537:Perlidae 2441:Suborder 2347:Pterygota 2327:Kingdom: 2153:Heraridea 2143:Geroptera 2048:Based on 1922:Panorpida 1869:(beetles) 1759:Psocodea 1694:Mantodea 1645:Notoptera 1602:(earwigs) 1337:Kingdom: 1222:1055-7903 815:2067/1437 669:Capniidae 647:Suborder 546:phylogeny 519:suborders 464:Halobates 390:mandibles 215:Pterygota 171:Kingdom: 165:Eukaryota 139:Eusthenia 2667:BugGuide 2611:Wikidata 2353:Neoptera 2333:Phylum: 2329:Animalia 2318:families 2191:Paoliida 1965:Diptera 1769:(thrips) 1530:Odonata 1357:Hexapoda 1343:Phylum: 1339:Animalia 1238:80621609 1230:30876966 1173:July 31, 1007:July 31, 876:10761594 718:Perlidae 588:Perlidae 496:diapause 396:, large 394:antennae 314:Southern 309:origin. 307:Mesozoic 295:Neoptera 231:Cohort: 225:Neoptera 181:Phylum: 175:Animalia 161:Domain: 2617:Q203547 2341:Insecta 2339:Class: 2314:Extant 2076:Extinct 2050:Sasaki 1957:(fleas) 1751: * 1202:Bibcode 773:is not 744:Family 737:Family 730:Family 723:Family 716:Family 709:Family 694:Family 688:Family 681:Family 674:Family 667:Family 637:Family 631:Family 617:Family 611:Family 428:plekein 421:braided 406:abdomen 370:(adult) 326:equator 303:Permian 287:insects 264:mostly 241:Order: 195:Insecta 191:Class: 122:–Recent 120:Permian 2884:593173 2858:890815 2848:368358 2832:NZOR: 2799:102467 2721:1PLECO 2695:856749 2054:(2013) 2052:et al. 2043:Italic 1371:Extant 1327:orders 1324:Insect 1236:  1228:  1220:  960:  930:16 May 901:  874:  821:  770:Capnia 536:, the 432:pteryx 410:nymphs 402:ocelli 337:nymphs 281:is an 268:: see 252:, 1839 35:(moth) 2879:WoRMS 2866:Plazi 2825:50622 2786:10508 2781:IRMNG 2773:47504 2734:12088 2685:8MP8Q 2454:basal 1234:S2CID 1031:(PDF) 755:Notes 554:clade 527:basal 503:molts 455:) or 436:wings 414:cerci 368:imago 352:Nymph 283:order 2820:NCBI 2794:ITIS 2755:GBIF 2716:EPPO 2708:1018 2654:BOLD 2038:bold 1226:PMID 1218:ISSN 1175:2008 1009:2008 958:ISBN 932:2013 899:ISBN 872:PMID 819:ISBN 316:and 270:text 142:sp. 59:PreꞒ 2807:NBN 2760:787 2703:EoL 2680:CoL 2659:135 2641:ADW 1210:doi 1198:135 1042:PDF 864:doi 811:hdl 803:doi 285:of 2903:: 2881:: 2868:: 2845:: 2822:: 2809:: 2796:: 2783:: 2770:: 2757:: 2744:: 2731:: 2718:: 2705:: 2682:: 2672:76 2669:: 2656:: 2643:: 2628:: 2613:: 1953:+ 1232:. 1224:. 1216:. 1208:. 1196:. 1192:. 1164:. 1139:. 1115:. 1104:^ 1088:. 1064:. 1046:. 1028:)" 998:. 940:^ 897:. 893:. 870:. 860:45 858:. 842:^ 817:. 809:. 563:. 376:: 109:Pg 53:Ma 2307:e 2300:t 2293:v 1814:a 1812:l 1810:o 1808:b 1806:a 1804:t 1802:e 1800:m 1798:o 1796:l 1794:o 1792:H 1733:a 1731:l 1729:o 1727:b 1725:a 1723:t 1721:e 1719:m 1717:u 1715:E 1565:a 1563:r 1561:e 1559:t 1557:p 1555:o 1553:e 1551:N 1479:a 1477:t 1475:o 1473:g 1471:y 1469:r 1467:e 1465:t 1463:P 1438:a 1436:i 1434:l 1432:y 1430:d 1428:n 1426:o 1424:c 1422:i 1420:D 1316:e 1309:t 1302:v 1240:. 1212:: 1204:: 1177:. 1149:. 1125:. 1098:. 1074:. 1044:) 1040:( 1011:. 966:. 934:. 907:. 878:. 866:: 827:. 813:: 805:: 594:) 590:( 578:) 574:( 449:( 380:) 372:( 114:N 104:K 99:J 94:T 89:P 84:C 79:D 74:S 69:O 64:Ꞓ 37:. 20:)

Index

Stoneflies
Plecoptera (moth)
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N
Permian

Eusthenia
Scientific classification
Edit this classification
Eukaryota
Animalia
Arthropoda
Insecta
Dicondylia
Pterygota
Neoptera
Polyneoptera
Plecoptera
Burmeister

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