1423:. The Newark Supergroup footprints show digits II, III and IV, but no trace of the shorter digits I and V which would likely have been present in a dinosaur of this stage. The outer two digits would have been stubby and ineffective, not touching the ground during walking or running. Despite losing most of their effectiveness, dinosaur evolution had not yet removed these digits to fully streamline the foot. This is known because rare specimens are found with traces of these outer digits. Digits II, III and IV have 3, 4 and 5 phalanges respectively, giving
437:
1531:
145:
1517:
127:
1456:
1401:
409:
footprints are characteristically three-toed (tridactyl) and range from 10 to 20 centimeters (or 4 to 8 inches) long. Though the tracks show only three toes, the trackmakers likely had between four and five toes on their feet. While it is usually impossible to match these prints with the exact
1500:
The scientists called the girl's discovery "the finest impression of a 215 million-year-old dinosaur print found in
Britain in a decade". Karl-James Langford of Archaeology Cymru considered the find to be "internationally important". Cindy Howells, a palaeontologist at
390:
translates into "stilt walker", although the actual length and form of the trackmaking legs varied by species, usually unidentified. The related term "Grallae" is an ancient name for the presumed group of long-legged wading birds, such as
1416:
type are those found on the East Coast of North
America, specifically from the Late Triassic to Early Jurassic Newark Supergroup. These footprints were likely made by an unidentified, primitive dinosaur similar to
1744:
1497:
Experts believe that the footprint was most likely left by a dinosaur (grallator) that stood about 75 centimeters (29.5 inches) tall and 2.5 meters (about 8 feet) long and walked on its two hind feet.
1505:, described it as "one of the best-preserved examples from anywhere in the UK" and said that it "will really aid palaeontologists to get a better idea about how these early dinosaurs walked".
1697:
Molnar, R. E., 2001, Theropod paleopathology: a literature survey: In: Mesozoic
Vertebrate Life, edited by Tanke, D. H., and Carpenter, K., Indiana University Press, p. 337-363.
1329:. The distal end of the digit was consistently flexed. However, this apparent pathology could be caused by the animal rotating the tip of that digit when lifting the foot.
1618:
Zhi-jun. (2009). "Theropod (Dinosauria: Saurischia) tracks from Lower
Cretaceous Yixian Formation at Sihetun Village, Liaoning Province, China and possible track makers."
1969:
1482:
In
January 2021, while walking with her father Richard Wilder, a four-year-old girl called Lily Wilder found a 215- to 220-million year-old dinosaur footprint at
1502:
1907:"Huellas de dinosaurios en la costa oeste del embalse Ezequiel Ramos Mexía y alrededores (Cretácico Superior, Provincia de Neuquén, República Argentina)"
1659:"The first theropod tracks from the Middle Jurassic of Gansu, Northwest China: new and rare evidence of quadrupedal progression in theropod dinosaurs"
2015:
1850:
1718:
1448:. The reason for this similarity is a lack of divergence in the foot evolution of the two distinct groups of dinosaurs: ornithischians and
1823:
410:
dinosaur species that left them, it is sometimes possible to narrow down potential trackmakers by comparing the proportions in individual
468:
1305:
Fossil tracks can be informative about theropod pathologies but apparently pathological traits may be due to unusual behaviors.
2010:
461:
1434:
tracks were made by a bipedal saurischian dinosaur, they can easily be mistaken for those of the late
Triassic ichnogenus
1544:
1876:
1770:
139:
1798:
323:(form taxon based on footprints) which covers a common type of small, three-toed print made by a variety of bipedal
494:
454:
144:
1206:
1083:
938:
1175:
562:
1566:
1056:
534:
1189:
866:
745:
1593:
Weishampel, D. B. & L. Young. 1996. Dinosaurs of the East Coast. The Johns
Hopkins University Press
1129:
634:
1143:
1111:
979:
965:
804:
776:
620:
509:
487:
1937:
1282:
1157:
852:
790:
1038:
1024:
993:
713:
699:
648:
1982:
1373:
1268:
1220:
1069:
1010:
818:
686:
673:
594:
548:
367:
1287:
1097:
998:
952:
896:
884:
879:
839:
607:
436:
1255:
1234:
731:
1745:"Paleontologists stunned by a perfectly preserved dinosaur footprint discovered by a 4-year-old girl"
1670:
924:
910:
762:
518:
1226:
1194:
1149:
1135:
1103:
1089:
1075:
1044:
1030:
985:
971:
957:
944:
916:
902:
871:
858:
810:
796:
737:
723:
705:
640:
626:
612:
567:
554:
540:
522:
474:
1707:
1536:
1167:
1121:
577:
499:
228:
1274:
1117:
295:
1657:
Da-Qing Li; Li-Da Xing; Martin G. Lockley; Anthony
Romilio; Jing-Tao Yang; Long-Feng Li (2019).
1636:
1247:
1061:
930:
844:
768:
751:
1987:
1163:
1048:
1016:
831:
526:
514:
1974:
1487:
1470:
1389:
1291:
1212:
1002:
888:
383:
1243:
1239:
1181:
823:
782:
665:
653:
1918:
1678:
1522:
1345:
1260:
678:
661:
599:
582:
419:
400:
339:
272:
248:
45:
1198:
827:
657:
1722:
1369:
363:
1674:
1385:
1341:
691:
379:
335:
2004:
1483:
1445:
1349:
1325:
343:
84:
586:
257:
1960:
1572:
1449:
1419:
424:
59:
1851:"Girl Discovers 220-Million-Year-Old Dinosaur Footprint On Beach Walk With Dad"
1923:
1906:
1683:
1658:
1512:
1321:
1320:
preserves tracks of an individual with a deformed digit III attributed to the
320:
202:
104:
69:
1824:"Lily's Dinosaur: 4yo girl discovers 220 million year old dinosaur footprint"
1771:"Girl, 4 finds 215 million-year-old dinosaur footprint along the Welsh coast"
1473:
at the St. George
Dinosaur Discovery Site at Johnson Farm, southwestern Utah.
1361:
1306:
355:
215:
156:
109:
53:
1380:, but are most abundant on the east coast of North America, especially the
374:, but are most abundant on the east coast of North America, especially the
17:
126:
1954:
1467:
1436:
1381:
375:
327:
324:
189:
176:
99:
94:
79:
74:
64:
41:
1877:"Perfect 215-million-year-old dinosaur print found by girl, 4, on beach"
1455:
1716:: where are the latest Triassic Ornithischia in the Newark Supergroup?"
1309:
114:
89:
1400:
414:
ichnospecies with known dinosaurs of the same formation. For example,
1365:
1357:
1353:
1313:
719:
359:
351:
347:
166:
1931:
1491:
1454:
1399:
1377:
1317:
435:
396:
392:
371:
1935:
1340:-type footprints have been found in formations dating from the
334:-type footprints have been found in formations dating from the
399:. These footprints were given this name by their discoverer,
1412:
The most famous, and archetypal tracks that conform to the
1725:
Abstracts with
Programs, Geological Society of America,
1708:"Distinguishing the tridactyl dinosaurian ichnogenus
264:
1944:
1769:Wakefield, Owen; Shaw, Amelia (30 January 2021).
1905:Calvo, Jorge Orlando; Rivera, Cynthia (2018).
1603:
1601:
1599:
1706:Safran, J. and Rainforth, E. C. (2004).
1562:
1560:
8:
1589:
1587:
1585:
1583:
1581:
1932:
125:
31:
1922:
1911:Boletín de la Sociedad Geológica Mexicana
1682:
1556:
1503:Amgueddfa Cymru – National Museum Wales
1388:formations of the northern part of the
382:formations of the northern part of the
1793:
1791:
7:
1739:
1737:
1735:
1875:Wakefield, Owen (29 January 2021).
1849:Papenfuss, Mary (30 January 2021).
1799:"Britain's biggest turn-offs named"
1427:a ?-3-4-5-? digital formula.
25:
27:Ichnogenus of dinosaur footprints
1529:
1515:
143:
1430:Although the Newark Supergroup
1348:periods. They are found in the
342:periods. They are found in the
1:
2016:Fossil taxa described in 1858
1620:Geological Bulletin of China
1545:List of dinosaur ichnogenera
287:
1610:Li-da, Harris, J. D.,
418:tracks identified from the
140:Trace fossil classification
2032:
1663:Journal of Palaeogeography
1924:10.18268/BSGM2018v70n2a11
1684:10.1186/s42501-019-0028-4
1444:prints was a quadrupedal
443:from Monte Pisano (Italy)
301:
294:
285:
280:
263:
256:
138:
133:
124:
34:
1408:showing skin impressions
1396:Newark Supergroup tracks
134:Typical footprint form
2011:Theropod trace fossils
1474:
1409:
1404:Negative footprint of
1374:Santa Maria Formations
455:G. (Coelurosaurichnus)
444:
422:may have been left by
368:Santa Maria Formations
1983:Paleobiology Database
1458:
1403:
1207:G. schlauersbachensis
1084:G. protocrassidigitus
939:G. mokanametsongensis
439:
1721:7 March 2016 at the
1440:. The trackmaker of
1828:The Science Insight
1675:2019JPalG...8...10L
1537:Paleontology portal
268:Grallator cursorius
1475:
1410:
1176:G. sassendorfensis
469:G. (C.) p. exiguus
445:
441:Grallator toscanus
1998:
1997:
1938:Taxon identifiers
1669:(1): Article 10.
1488:Vale of Glamorgan
1471:Moenave Formation
1390:Newark Supergroup
1295:
1278:
1264:
1251:
1230:
1216:
1202:
1185:
1171:
1153:
1139:
1125:
1107:
1093:
1079:
1065:
1057:G. pingchuanensis
1052:
1034:
1020:
1006:
989:
975:
961:
948:
934:
920:
906:
892:
875:
862:
848:
835:
814:
800:
786:
772:
758:
741:
727:
709:
695:
682:
669:
644:
630:
616:
603:
590:
571:
558:
544:
535:G. angustidigitus
530:
503:
478:
384:Newark Supergroup
312:
311:
258:Type ichnospecies
252:
16:(Redirected from
2023:
1991:
1990:
1978:
1977:
1965:
1964:
1963:
1933:
1928:
1926:
1917:(2): 449 ‒ 497.
1892:
1891:
1889:
1887:
1872:
1866:
1865:
1863:
1861:
1846:
1840:
1839:
1837:
1835:
1820:
1814:
1813:
1811:
1809:
1795:
1786:
1785:
1783:
1781:
1775:North Wales Live
1766:
1760:
1759:
1757:
1755:
1749:Business Insider
1741:
1730:
1704:
1698:
1695:
1689:
1688:
1686:
1654:
1648:
1647:
1645:
1643:
1633:
1627:
1617:
1614:Xiang-yang, and
1613:
1609:
1605:
1594:
1591:
1576:
1564:
1539:
1534:
1533:
1532:
1525:
1523:Dinosaurs portal
1520:
1519:
1518:
1346:early Cretaceous
1286:
1272:
1259:
1238:
1224:
1210:
1193:
1190:G. sauclierensis
1179:
1161:
1147:
1133:
1115:
1101:
1087:
1073:
1060:
1042:
1028:
1014:
997:
983:
969:
956:
942:
928:
914:
900:
883:
870:
867:G. matsiengensis
856:
843:
822:
808:
794:
780:
766:
749:
746:G. jiuquwanensis
735:
717:
703:
690:
677:
652:
638:
624:
611:
598:
581:
566:
552:
538:
513:
498:
472:
462:G. (C.) palmipes
420:Yixian Formation
401:Edward Hitchcock
340:early Cretaceous
266:
247:
240:
227:
148:
147:
129:
119:
56:
46:Early Cretaceous
40:Temporal range:
32:
21:
2031:
2030:
2026:
2025:
2024:
2022:
2021:
2020:
2001:
2000:
1999:
1994:
1986:
1981:
1973:
1968:
1959:
1958:
1953:
1940:
1904:
1901:
1899:Further reading
1896:
1895:
1885:
1883:
1874:
1873:
1869:
1859:
1857:
1848:
1847:
1843:
1833:
1831:
1830:. February 2021
1822:
1821:
1817:
1807:
1805:
1797:
1796:
1789:
1779:
1777:
1768:
1767:
1763:
1753:
1751:
1743:
1742:
1733:
1723:Wayback Machine
1705:
1701:
1696:
1692:
1656:
1655:
1651:
1641:
1639:
1635:
1634:
1630:
1615:
1611:
1607:
1606:
1597:
1592:
1579:
1565:
1558:
1553:
1535:
1530:
1528:
1521:
1516:
1514:
1511:
1480:
1466:) in the Lower
1398:
1344:through to the
1335:
1303:
1298:
1130:G. quthingensis
635:G. digitigradus
495:G. (G.) zvierzi
434:
338:through to the
276:
270:
246:
238:
225:
142:
120:
118:
117:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
51:
50:
48:
38:
28:
23:
22:
15:
12:
11:
5:
2029:
2027:
2019:
2018:
2013:
2003:
2002:
1996:
1995:
1993:
1992:
1979:
1966:
1950:
1948:
1942:
1941:
1936:
1930:
1929:
1900:
1897:
1894:
1893:
1867:
1841:
1815:
1787:
1761:
1731:
1699:
1690:
1649:
1628:
1595:
1577:
1555:
1554:
1552:
1549:
1548:
1547:
1541:
1540:
1526:
1510:
1507:
1479:
1476:
1397:
1394:
1386:Early Jurassic
1342:Early Triassic
1334:
1331:
1312:dating to the
1302:
1301:Paleopathology
1299:
1297:
1296:
1279:
1265:
1252:
1231:
1217:
1203:
1186:
1172:
1154:
1144:G. rectilineus
1140:
1126:
1112:G. romanovskyi
1108:
1094:
1080:
1066:
1053:
1035:
1021:
1007:
990:
980:G. moshoeshoei
976:
966:G. morijiensis
962:
949:
935:
921:
907:
893:
876:
863:
849:
836:
815:
805:G. leribeensis
801:
787:
777:G. kronbergeri
773:
759:
742:
728:
710:
696:
683:
670:
645:
631:
621:G. deambulator
617:
604:
591:
574:
573:
572:
545:
531:
510:G. andeolensis
506:
505:
504:
488:G. (Grallator)
483:
482:
481:
480:
479:
449:
433:
430:
380:Early Jurassic
336:Early Triassic
310:
309:
308:
307:
299:
298:
292:
291:
288:Many, see text
283:
282:
278:
277:
271:
261:
260:
254:
253:
236:
232:
231:
223:
219:
218:
213:
206:
205:
200:
193:
192:
187:
180:
179:
174:
170:
169:
164:
160:
159:
154:
150:
149:
136:
135:
131:
130:
122:
121:
113:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
57:
42:Early Triassic
39:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
2028:
2017:
2014:
2012:
2009:
2008:
2006:
1989:
1984:
1980:
1976:
1971:
1967:
1962:
1956:
1952:
1951:
1949:
1947:
1943:
1939:
1934:
1925:
1920:
1916:
1912:
1908:
1903:
1902:
1898:
1882:
1878:
1871:
1868:
1856:
1852:
1845:
1842:
1829:
1825:
1819:
1816:
1804:
1800:
1794:
1792:
1788:
1776:
1772:
1765:
1762:
1750:
1746:
1740:
1738:
1736:
1732:
1728:
1724:
1720:
1717:
1715:
1711:
1703:
1700:
1694:
1691:
1685:
1680:
1676:
1672:
1668:
1664:
1660:
1653:
1650:
1638:
1632:
1629:
1626:(6): 705–712.
1625:
1621:
1604:
1602:
1600:
1596:
1590:
1588:
1586:
1584:
1582:
1578:
1574:
1570:
1569:
1563:
1561:
1557:
1550:
1546:
1543:
1542:
1538:
1527:
1524:
1513:
1508:
1506:
1504:
1498:
1495:
1493:
1489:
1485:
1484:Bendricks Bay
1477:
1472:
1469:
1465:
1461:
1457:
1453:
1451:
1447:
1446:ornithischian
1443:
1439:
1438:
1433:
1428:
1426:
1422:
1421:
1415:
1407:
1402:
1395:
1393:
1391:
1387:
1383:
1379:
1375:
1371:
1367:
1363:
1359:
1355:
1351:
1350:United States
1347:
1343:
1339:
1332:
1330:
1328:
1327:
1326:Anchisauripus
1323:
1319:
1315:
1311:
1308:
1300:
1293:
1289:
1285:
1284:
1283:G. variabilis
1280:
1276:
1271:
1270:
1266:
1262:
1258:
1257:
1253:
1249:
1245:
1241:
1237:
1236:
1232:
1228:
1223:
1222:
1218:
1214:
1209:
1208:
1204:
1200:
1196:
1192:
1191:
1187:
1183:
1178:
1177:
1173:
1169:
1165:
1160:
1159:
1158:G. sabinensis
1155:
1151:
1146:
1145:
1141:
1137:
1132:
1131:
1127:
1123:
1119:
1114:
1113:
1109:
1105:
1100:
1099:
1095:
1091:
1086:
1085:
1081:
1077:
1072:
1071:
1067:
1064:et al., 2019)
1063:
1059:
1058:
1054:
1050:
1046:
1041:
1040:
1036:
1032:
1027:
1026:
1022:
1018:
1013:
1012:
1008:
1004:
1000:
996:
995:
991:
987:
982:
981:
977:
973:
968:
967:
963:
959:
955:
954:
950:
946:
941:
940:
936:
932:
927:
926:
922:
918:
913:
912:
908:
904:
899:
898:
894:
890:
886:
882:
881:
877:
873:
869:
868:
864:
860:
855:
854:
853:G. magnificus
850:
846:
842:
841:
837:
833:
829:
825:
821:
820:
816:
812:
807:
806:
802:
798:
793:
792:
791:G. lacunensis
788:
784:
779:
778:
774:
770:
765:
764:
760:
757:
753:
748:
747:
743:
739:
734:
733:
729:
725:
721:
716:
715:
711:
707:
702:
701:
697:
693:
689:
688:
684:
680:
676:
675:
671:
667:
663:
659:
655:
651:
650:
646:
642:
637:
636:
632:
628:
623:
622:
618:
614:
610:
609:
605:
601:
597:
596:
592:
588:
584:
580:
579:
575:
569:
565:
564:
560:
559:
556:
551:
550:
546:
542:
537:
536:
532:
528:
524:
520:
516:
512:
511:
507:
501:
497:
496:
492:
491:
490:
489:
484:
476:
471:
470:
466:
465:
464:
463:
459:
458:
457:
456:
451:
450:
448:
442:
438:
431:
429:
427:
426:
421:
417:
413:
408:
404:
402:
398:
394:
389:
385:
381:
377:
373:
369:
365:
361:
357:
353:
349:
345:
344:United States
341:
337:
333:
329:
326:
322:
318:
317:
306:
303:
302:
300:
297:
293:
290:
289:
284:
281:Ichnospecies
279:
274:
269:
262:
259:
255:
250:
245:
244:
237:
234:
233:
230:
229:Grallatoridae
224:
222:Ichnofamily:
221:
220:
217:
214:
211:
208:
207:
204:
201:
198:
195:
194:
191:
188:
185:
182:
181:
178:
175:
172:
171:
168:
165:
162:
161:
158:
155:
152:
151:
146:
141:
137:
132:
128:
123:
116:
111:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
55:
52:250–100
47:
43:
37:
33:
30:
19:
1945:
1914:
1910:
1884:. Retrieved
1880:
1870:
1858:. Retrieved
1854:
1844:
1832:. Retrieved
1827:
1818:
1806:. Retrieved
1802:
1778:. Retrieved
1774:
1764:
1752:. Retrieved
1748:
1726:
1713:
1709:
1702:
1693:
1666:
1662:
1652:
1640:. Retrieved
1631:
1623:
1619:
1567:
1499:
1496:
1481:
1463:
1459:
1450:saurischians
1441:
1435:
1431:
1429:
1424:
1418:
1413:
1411:
1405:
1337:
1336:
1324:
1316:in southern
1304:
1281:
1267:
1254:
1233:
1219:
1205:
1188:
1174:
1156:
1142:
1128:
1110:
1096:
1082:
1068:
1055:
1039:G. perriauxi
1037:
1025:G. paulstris
1023:
1009:
994:G. olonensis
992:
978:
964:
951:
937:
923:
909:
895:
878:
865:
851:
838:
817:
803:
789:
775:
761:
755:
744:
730:
712:
700:G. graciosus
698:
685:
672:
649:G. emeiensis
647:
633:
619:
606:
593:
578:G. cursorius
576:
563:G. a. cursor
561:
547:
533:
519:Vianey-Liaud
508:
493:
486:
467:
460:
453:
446:
440:
423:
415:
411:
406:
405:
387:
331:
315:
314:
313:
304:
286:
267:
242:
241:
235:Ichnogenus:
209:
196:
183:
35:
29:
1881:WalesOnline
1803:The Week UK
1637:"Paleofile"
1573:Fossilworks
1420:Coelophysis
1406:G. cuneatus
1333:Occurrences
1269:G. toscanus
1227:Ellenberger
1221:G. socialis
1150:Ellenberger
1136:Ellenberger
1104:Ellenberger
1090:Ellenberger
1076:Ellenberger
1070:G. princeps
1031:Ellenberger
1011:G. palissyi
986:Ellenberger
972:Ellenberger
958:Ellenberger
945:Ellenberger
917:Ellenberger
903:Ellenberger
872:Ellenberger
859:Ellenberger
819:G. limnosus
811:Ellenberger
797:Ellenberger
738:Ellenberger
714:G. grancier
706:Ellenberger
687:G. gracilis
674:G. formosus
641:Ellenberger
627:Ellenberger
613:Ellenberger
595:G. cuneatus
568:Ellenberger
555:Ellenberger
549:G. angustus
541:Ellenberger
475:Ellenberger
425:Caudipteryx
403:, in 1858.
386:. The name
18:Saltopoides
2005:Categories
1886:1 February
1860:1 February
1834:1 February
1808:1 February
1780:1 February
1754:1 February
1551:References
1322:ichnogenus
1098:G. rapidus
953:G. molapoi
897:G. minimus
880:G. maximus
840:G. madseni
608:G. damanei
500:Gierlinski
321:ichnogenus
203:Saurischia
190:Dinosauria
1961:Q23691966
1946:Grallator
1714:Grallator
1568:Grallator
1464:Grallator
1460:Eubrontes
1432:Grallator
1425:Grallator
1414:Grallator
1362:Australia
1338:Grallator
1307:Sandstone
1288:Lapparent
1261:Hitchcock
1256:G. tenuis
1235:G. ssatoi
1195:Demathieu
1045:Demathieu
999:Lapparent
885:Lapparent
754:, 1982) =
732:G. ingens
724:Demathieu
692:Hitchcock
679:Hitchcock
600:Hitchcock
587:ichnotype
583:Hitchcock
523:Demathieu
485:Subgenus
452:Subgenus
416:Grallator
412:Grallator
407:Grallator
388:Grallator
356:Australia
332:Grallator
328:dinosaurs
316:Grallator
273:Hitchcock
249:Hitchcock
243:Grallator
216:Theropoda
163:Kingdom:
157:Eukaryota
36:Grallator
1955:Wikidata
1855:HuffPost
1729:(2): 96.
1719:Archived
1710:Atreipus
1509:See also
1468:Jurassic
1442:Atreipus
1437:Atreipus
1382:Triassic
1292:Monetnat
1246:, &
1122:Kurbatov
1003:Monetnat
925:G. moeni
911:G. minor
889:Monetnat
830:, &
763:G. kehli
756:Hunanpus
585:, 1858 (
525:, &
447:Source:
376:Triassic
325:theropod
305:Hunanpus
296:Synonyms
177:Chordata
173:Phylum:
167:Animalia
153:Domain:
1975:1124803
1671:Bibcode
1642:5 April
1486:in the
1310:stratum
1277:, 1941)
1248:Shikama
1229:, 1970)
1215:, 1934)
1184:, 1958)
1170:, 1976)
1168:Pellier
1152:, 1970)
1138:, 1974)
1118:Gabunia
1106:, 1974)
1092:, 1970)
1078:, 1970)
1051:, 1972)
1033:, 1970)
1019:, 1976)
988:, 1970)
974:, 1970)
947:, 1974)
933:, 1950)
931:Beurlen
919:, 1970)
905:, 1970)
861:, 1970)
813:, 1970)
799:, 1970)
785:, 1959)
783:Rehnelt
771:, 1950)
769:Beurlen
740:, 1970)
726:, 2000)
708:, 1970)
643:, 1974)
629:, 1970)
557:, 1974)
543:, 1970)
477:, 1970)
432:Species
1376:) and
1366:Brazil
1358:Europe
1354:Canada
1314:Norian
1294:, 1967
1290:&
1263:, 1858
1250:, 1940
1201:, 1992
1197:&
1166:&
1120:&
1047:&
1005:, 1967
1001:&
960:, 1974
891:, 1967
887:&
874:, 1970
847:, 1995
834:, 1985
722:&
720:Courel
694:, 1865
681:, 1858
668:, 1995
664:&
615:, 1970
602:, 1858
570:, 1974
529:, 2000
527:Garric
502:, 1991
397:herons
393:storks
370:) and
360:Brazil
352:Europe
348:Canada
319:is an
275:, 1858
251:, 1858
1988:64079
1970:IRMNG
1616:Zhang
1492:Wales
1478:Wales
1378:China
1370:Sousa
1318:Wales
1275:Huene
1213:Weiss
1199:Sciau
372:China
364:Sousa
210:Clade
197:Clade
184:Clade
1888:2021
1862:2021
1836:2021
1810:2021
1782:2021
1756:2021
1712:and
1644:2011
1612:Feng
1608:Xing
1575:.org
1384:and
1372:and
1244:Inai
1240:Yabe
1182:Kuhn
1164:Gand
1049:Gand
1017:Gand
845:Irby
824:Zhen
752:Zeng
666:Yang
654:Zhen
515:Gand
395:and
378:and
366:and
60:PreꞒ
1919:doi
1679:doi
1571:at
1494:.
1462:(=
832:Rao
662:Han
2007::
1985::
1972::
1957::
1915:70
1913:.
1909:.
1879:.
1853:.
1826:.
1801:.
1790:^
1773:.
1747:.
1734:^
1727:36
1677:.
1665:.
1661:.
1624:28
1622:,
1598:^
1580:^
1559:^
1490:,
1452:.
1392:.
1364:,
1360:,
1356:,
1352:,
1242:,
1062:Li
828:Li
826:,
660:,
658:Li
656:,
521:,
517:,
428:.
358:,
354:,
350:,
346:,
330:.
212::
199::
186::
110:Pg
54:Ma
1927:.
1921::
1890:.
1864:.
1838:.
1812:.
1784:.
1758:.
1687:.
1681::
1673::
1667:8
1646:.
1368:(
1273:(
1225:(
1211:(
1180:(
1162:(
1148:(
1134:(
1124:)
1116:(
1102:(
1088:(
1074:(
1043:(
1029:(
1015:(
984:(
970:(
943:(
929:(
915:(
901:(
857:(
809:(
795:(
781:(
767:(
750:(
736:(
718:(
704:(
639:(
625:(
589:)
553:(
539:(
473:(
362:(
265:†
239:†
226:†
115:N
105:K
100:J
95:T
90:P
85:C
80:D
75:S
70:O
65:Ꞓ
49:~
44:-
20:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
